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<document id="8BCE9B049118002BAC6D63B74E6678F8" ID-CLB-Dataset="55750" ID-DOI="10.5281/zenodo.5385198" ID-GBIF-Dataset="c7f95b94-53f8-420d-97c5-4e27d7c211bf" ID-ISSN="2345-7600" ID-Zenodo-Dep="5385198" ID-ZooBank="AC7F1491-CD89-4F10-9D61-55354B55E368" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630300948061" checkinUser="valdenar" docAuthor="Hájek, Jiří &amp; Brancucci, Michel" docDate="2015" docId="038A87C4FFAAFF93F3BB9124FA42BEEE" docLanguage="en" docName="RafflesBZool.63.309-326.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Laccophilus tobaensis Brancucci 1983" docType="treatment" docVersion="9" lastPageNumber="324" masterDocId="FFB3FFBCFFA5FF9CF02A9279FFD5BB5C" masterDocTitle="A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)" masterLastPageNumber="326" masterPageNumber="309" pageNumber="324" updateTime="1699240741915" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="88CF7539BAFA0EB657DCDC8AED956438" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EE5699DC173BF960B3EDE8B0217B7612">
<mods:title id="8669C5B668784722F7DC1E50ADF162C7">A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name id="5B647B5AEECDAC6B2F6980542FA539B9" type="personal">
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</mods:role>
<mods:namePart id="30A26483144D8B0DDF37B1C952C4F830">Hájek, Jiří</mods:namePart>
<mods:affiliation id="0D57A0391EF6FD4D019EDCEEB36BD084">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 – Horní Počernice, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="1BE82B485D18F30826EE79A1962E7350" type="email">jiri_hajek@nm.cz</mods:nameIdentifier>
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<mods:name id="1DDFED2C403E8BAB2DBB01E89ED2985E" type="personal">
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<mods:roleTerm id="6CA93BCF22DD8E60F07B7A345E9A0DE5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E95B551B84602F06DE0C8A1BD8F166F5">Brancucci, Michel</mods:namePart>
<mods:affiliation id="09B148018365073BDD8AD47C6CDAE738">Naturhistorisches Museum, Basel, Switzerland</mods:affiliation>
</mods:name>
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<mods:title id="9C900A8AB4175DDFBD3D24ACCFEDBF12">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="8267F5C2973E4BA8AB0025ED6DCFFE67">
<mods:date id="49950E82CF5A5123093250484BBCE9A2">2015</mods:date>
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<mods:number id="6B1592752AE8BF7A13D8D1B091D94ECB">2015-08-06</mods:number>
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<mods:detail id="A3D333D0742615C02ED4A24A5BF7D298" type="volume">
<mods:number id="F1E017B1CBCE54758A168326EAC88ABB">63</mods:number>
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<mods:start id="2D1DB041F8331946CF2231873B22B6AC">309</mods:start>
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<mods:classification id="D8F06171095BBA144DFB37C56757EFFB">journal article</mods:classification>
<mods:identifier id="ABDE96CCF7B18B33CF947408ED0FC1C0" type="CLB-Dataset">55750</mods:identifier>
<mods:identifier id="74D25F1707ED023C68D02C7161EF1174" type="DOI">10.5281/zenodo.5385198</mods:identifier>
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<mods:identifier id="1281FCC12B81348E47474E618D4E93D0" type="ISSN">2345-7600</mods:identifier>
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<treatment id="038A87C4FFAAFF93F3BB9124FA42BEEE" ID-DOI="http://doi.org/10.5281/zenodo.5466907" ID-GBIF-Taxon="194955227" ID-Zenodo-Dep="5466907" LSID="urn:lsid:plazi:treatment:038A87C4FFAAFF93F3BB9124FA42BEEE" httpUri="http://treatment.plazi.org/id/038A87C4FFAAFF93F3BB9124FA42BEEE" lastPageNumber="324" pageId="15" pageNumber="324">
<subSubSection id="C3396559FFAAFF93F3BB9124FA9CB829" box="[913,1353,861,885]" pageId="15" pageNumber="324" type="nomenclature">
<paragraph id="8B9C36D2FFAAFF93F3BB9124FA9CB829" blockId="15.[913,1353,861,917]" box="[913,1353,861,885]" pageId="15" pageNumber="324">
<heading id="D0D481BEFFAAFF93F3BB9124FA9CB829" bold="true" box="[913,1353,861,885]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="324" reason="2">
<taxonomicName id="4C234D51FFAAFF93F3BB9124FA9CB829" ID-CoL="3RMC9" authority="Brancucci, 1983" authorityName="Brancucci" authorityYear="1983" box="[913,1353,861,885]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="324" phylum="Arthropoda" rank="species" species="tobaensis">
<emphasis id="B957EAC0FFAAFF93F3BB9124FA9CB829" bold="true" box="[913,1353,861,885]" pageId="15" pageNumber="324">
<emphasis id="B957EAC0FFAAFF93F3BB9124FB53B829" bold="true" box="[913,1158,861,885]" italics="true" pageId="15" pageNumber="324">Laccophilus tobaensis</emphasis>
<bibRefCitation id="EFB24B23FFAAFF93F4BB9124FA9CB829" author="Brancucci M" box="[1169,1353,861,885]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" pageId="15" pageNumber="324" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAAFF93F4089104FB63B8C9" box="[1058,1206,893,917]" pageId="15" pageNumber="324" type="description">
<paragraph id="8B9C36D2FFAAFF93F4089104FB63B8C9" blockId="15.[913,1353,861,917]" box="[1058,1206,893,917]" pageId="15" pageNumber="324">
(
<figureCitation id="13182A57FFAAFF93F4009104FB50B8C9" box="[1066,1157,893,917]" captionStart="Figs" captionStartId="4.[135,179,2064,2086]" captionTargetBox="[168,1456,189,2051]" captionTargetId="figure-8@4.[133,1476,180,2059]" captionTargetPageId="4" captionText="Figs. 10–17. Habitus of Laccophilus: 10, L. mahakamensis; 11, L. mazzoldii, new species; 12, L. nusatenggaraensis, new species; 13, L. schillhammeri, new species; 14, L. stastnyi, new species; 15, L. tobaensis; 16, L. tonkinensis; 17, L. wittmeri. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385202" httpUri="https://zenodo.org/record/5385202/files/figure.png" pageId="15" pageNumber="324">Figs. 15</figureCitation>
,
<figureCitation id="13182A57FFAAFF93F4B89104FB7BB8C9" box="[1170,1198,893,917]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="15" pageNumber="324">33</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAAFF93F30791C5FA78B88E" box="[813,1453,956,978]" pageId="15" pageNumber="324" type="reference_group">
<paragraph id="8B9C36D2FFAAFF93F30791C5FA78B88E" blockId="15.[813,1453,956,978]" box="[813,1453,956,978]" pageId="15" pageNumber="324">
<treatmentCitationGroup id="AB3311FCFFAAFF93F30791C5FA78B88E" box="[813,1453,956,978]" pageId="15" pageNumber="324">
<treatmentCitation id="0A8210C3FFAAFF93F30791C5FB00B88E" author="Brancucci M" box="[813,1237,956,978]" page="255" pageId="15" pageNumber="324" year="1983">
<taxonomicName id="4C234D51FFAAFF93F30791C5FB00B88E" ID-CoL="3RMC9" authority="Brancucci, 1983: 255" authorityName="Brancucci" authorityPageNumber="255" authorityYear="1983" box="[813,1237,956,978]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="324" phylum="Arthropoda" rank="species" species="tobaensis">
<emphasis id="B957EAC0FFAAFF93F30791C5FBD6B88D" box="[813,1027,956,977]" italics="true" pageId="15" pageNumber="324">Laccophilus tobaensis</emphasis>
<bibRefCitation id="EFB24B23FFAAFF93F42291C5FB00B88E" author="Brancucci M" box="[1032,1237,956,978]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" page="255" pageId="15" pageNumber="324" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983: 255</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAAFF93F3079183FBC3BFCE" pageId="15" pageNumber="324" type="materials_examined">
<paragraph id="8B9C36D2FFAAFF93F3079183FA7EBF4E" blockId="15.[813,1451,1018,1042]" box="[813,1451,1018,1042]" pageId="15" pageNumber="324">
<emphasis id="B957EAC0FFAAFF93F3079183FC10BF4E" bold="true" box="[813,965,1018,1042]" pageId="15" pageNumber="324">
<typeStatus id="54988870FFAAFF93F3079183FCB2BF4E" box="[813,871,1018,1042]" pageId="15" pageNumber="324">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3347642FFAAFF93F3E29183FBE7BF4E" box="[968,1074,1018,1042]" name="Indonesia" pageId="15" pageNumber="324">Indonesia</collectingCountry>
,
<collectingCounty id="62FD4E5EFFAAFF93F4119183FBA7BF4E" box="[1083,1138,1018,1042]" pageId="15" pageNumber="324">Nord</collectingCounty>
<collectingRegion id="49E7F830FFAAFF93F45D9183FB07BF4E" box="[1143,1234,1018,1042]" country="Indonesia" name="Sumatera Utara" pageId="15" pageNumber="324">Sumatra</collectingRegion>
,
<collectingMunicipality id="6BF8ACA8FFAAFF93F4F19183FAE0BF4E" box="[1243,1333,1018,1042]" pageId="15" pageNumber="324">Samosir</collectingMunicipality>
,
<location id="8EFC6009FFAAFF93F5149183FA72BF4E" LSID="urn:lsid:plazi:treatment:038A87C4FFAAFF93F3BB9124FA42BEEE:8EFC6009FFAAFF93F5149183FA72BF4E" box="[1342,1447,1018,1042]" country="Indonesia" county="Nord" municipality="Samosir" name="Toba-See" pageId="15" pageNumber="324" stateProvince="Sumatera Utara">Toba-See</location>
.
</paragraph>
<paragraph id="8B9C36D2FFAAFF93F3079643FBC3BFCE" blockId="15.[813,1452,1082,1170]" pageId="15" pageNumber="324">
<emphasis id="B957EAC0FFAAFF93F3079643FBD9BF0E" bold="true" box="[813,1036,1082,1106]" pageId="15" pageNumber="324">Material examined.</emphasis>
<specimenCount id="9D25FD5BFFAAFF93F4389643FB5ABF0E" box="[1042,1167,1082,1106]" pageId="15" pageNumber="324" type="generic">1 specimen</specimenCount>
.
<materialsCitation id="3B4B3C8FFFAAFF93F4B09643FA71BF0E" ID-GBIF-Occurrence="3760488412" box="[1178,1444,1082,1106]" country="Indonesia" location="Sumatra" pageId="15" pageNumber="324" specimenCount="1" stateProvince="Sumatera Utara">
<emphasis id="B957EAC0FFAAFF93F4B09643FA71BF0E" bold="true" box="[1178,1444,1082,1106]" pageId="15" pageNumber="324">
<collectingCountry id="F3347642FFAAFF93F4B09643FAE4BF0E" box="[1178,1329,1082,1106]" name="Indonesia" pageId="15" pageNumber="324">INDONESIA</collectingCountry>
:
<collectingRegion id="49E7F830FFAAFF93F56B9643FA71BF0E" box="[1345,1444,1082,1106]" country="Indonesia" name="Sumatera Utara" pageId="15" pageNumber="324">Sumatra</collectingRegion>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B4B3C8FFFAAFF93F3079623FBC7BFCE" ID-GBIF-Occurrence="3760488358" collectingDate="1991-02-04" collectionCode="NHMW" collectorName="M. A. Jach" country="Indonesia" pageId="15" pageNumber="324" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara">
<specimenCount id="9D25FD5BFFAAFF93F3079623FCAEBF2E" box="[813,891,1114,1138]" pageId="15" pageNumber="324" type="male">1 male</specimenCount>
,
<collectorName id="26D65304FFAAFF93F3A09622FBC7BF2E" box="[906,1042,1114,1138]" pageId="15" pageNumber="324">NW Sipirok</collectorName>
,
<collectorName id="26D65304FFAAFF93F40B9623FB70BF2E" box="[1057,1189,1114,1138]" pageId="15" pageNumber="324">Hutaimbaru</collectorName>
,
<date id="FF9D1012FFAAFF93F4989623FAB8BF2E" box="[1202,1389,1114,1138]" pageId="15" pageNumber="324" value="1991-02-04">
<collectingDate id="EFD9E9FAFFAAFF93F4989623FAB8BF2E" box="[1202,1389,1114,1138]" pageId="15" pageNumber="324" value="1991-02-04">4 February 1991</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFAAFF93F3079603FC48BFCE" box="[813,925,1146,1170]" pageId="15" pageNumber="324">M.A. Jäch</collectorName>
(
<collectionCode id="ED32AE17FFAAFF93F3879603FBDEBFCE" box="[941,1035,1146,1170]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="15" pageNumber="324" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAAFF93F30796C3FAB7BE0E" pageId="15" pageNumber="324" type="diagnosis">
<paragraph id="8B9C36D2FFAAFF93F30796C3FAB7BE0E" blockId="15.[813,1452,1210,1363]" pageId="15" pageNumber="324">
<emphasis id="B957EAC0FFAAFF93F30796C3FC74BF8E" bold="true" box="[813,929,1210,1234]" pageId="15" pageNumber="324">Diagnosis.</emphasis>
Medium sized, broadly oval species (body length
<quantity id="4CDB9B37FFAAFF93F30796A3FC66BFAE" box="[813,947,1242,1266]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="15" pageNumber="324" unit="mm" value="4.45" valueMax="4.5" valueMin="4.4">4.4–4.5 mm</quantity>
). Elytra with subbasal testaceous band divided longitudinally into separate spots (
<figureCitation id="13182A57FFAAFF93F4839683FB2FBE4E" box="[1193,1274,1274,1298]" captionStart="Figs" captionStartId="4.[135,179,2064,2086]" captionTargetBox="[168,1456,189,2051]" captionTargetId="figure-8@4.[133,1476,180,2059]" captionTargetPageId="4" captionText="Figs. 10–17. Habitus of Laccophilus: 10, L. mahakamensis; 11, L. mazzoldii, new species; 12, L. nusatenggaraensis, new species; 13, L. schillhammeri, new species; 14, L. stastnyi, new species; 15, L. tobaensis; 16, L. tonkinensis; 17, L. wittmeri. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385202" httpUri="https://zenodo.org/record/5385202/files/figure.png" pageId="15" pageNumber="324">Fig. 15</figureCitation>
). Apical part of median lobe rather short, almost subparallel; in lateral view with dorsal side only very slightly curved (
<figureCitation id="13182A57FFAAFF93F52E9743FA83BE0E" box="[1284,1366,1338,1362]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="15" pageNumber="324">Fig. 33</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFAAFF93F3079703FA42BEEE" pageId="15" pageNumber="324" type="distribution">
<paragraph id="8B9C36D2FFAAFF93F3079703FA42BEEE" blockId="15.[813,1452,1402,1458]" pageId="15" pageNumber="324">
<emphasis id="B957EAC0FFAAFF93F3079703FC6ABECE" bold="true" box="[813,959,1402,1426]" pageId="15" pageNumber="324">Distribution.</emphasis>
The species was described and is so far known only from northern
<collectingRegion id="49E7F830FFAAFF93F42E97E3FB88BEEE" box="[1028,1117,1434,1458]" country="Indonesia" name="Sumatera Utara" pageId="15" pageNumber="324">Sumatra</collectingRegion>
(
<bibRefCitation id="EFB24B23FFAAFF93F44797E3FACABEEE" author="Brancucci M" box="[1133,1311,1434,1458]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" pageId="15" pageNumber="324" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
) (
<figureCitation id="13182A57FFAAFF93F51397E3FA5CBEEE" box="[1337,1417,1434,1458]" captionStart="Figs" captionStartId="9.[113,157,2093,2115]" captionTargetBox="[141,1427,190,2077]" captionTargetPageId="9" captionText="Figs. 40–41. Distribution of the species of the Laccophilus javanicus group" figureDoi="http://doi.org/10.5281/zenodo.5385208" httpUri="https://zenodo.org/record/5385208/files/figure.png" pageId="15" pageNumber="324">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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