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<document id="11C0B6D85CE9A274E12BB847888F304D" ID-CLB-Dataset="55750" ID-DOI="10.5281/zenodo.5385198" ID-GBIF-Dataset="c7f95b94-53f8-420d-97c5-4e27d7c211bf" ID-ISSN="2345-7600" ID-Zenodo-Dep="5385198" ID-ZooBank="AC7F1491-CD89-4F10-9D61-55354B55E368" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630300948061" checkinUser="valdenar" docAuthor="Hájek, Jiří &amp; Brancucci, Michel" docDate="2015" docId="038A87C4FFA9FF91F397956FFD4EB349" docLanguage="en" docName="RafflesBZool.63.309-326.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Laccophilus mazzoldii Hájek &amp; Brancucci 2015, new species" docType="treatment" docUuid="D324DBE2-9273-4B55-B432-852A25E46FB0" docUuidSource="ZooBank" docVersion="9" lastPageNumber="322" masterDocId="FFB3FFBCFFA5FF9CF02A9279FFD5BB5C" masterDocTitle="A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)" masterLastPageNumber="326" masterPageNumber="309" pageNumber="321" updateTime="1699240741915" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="80D5430F64D790D3390B371565E5C7D8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="454AB3B71D1AF7DA5E0ABDE0C87D4710">
<mods:title id="4ABA71CCCE8D59B4CC4F233120DF9CDE">A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name id="D9C7304DBEBC41EEBA8480E60BE23063" type="personal">
<mods:role id="7D1B8AF1673EF5BD35AC70A725AFE87B">
<mods:roleTerm id="5407203296470F6DE189E1C00C1AA068">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="17321E0AA5C3FCD5282F5B4900BD5633">Hájek, Jiří</mods:namePart>
<mods:affiliation id="623209F98E80ABF495B622DC778EA692">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 – Horní Počernice, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="AAF6462DFD492C37171ACD5502DA194B" type="email">jiri_hajek@nm.cz</mods:nameIdentifier>
</mods:name>
<mods:name id="AE5F76E5B7F2ADFCD5BF345579E15874" type="personal">
<mods:role id="FF6C747290C115487326A881EB83AC0B">
<mods:roleTerm id="D4DAE764DEA447D77AC77DB3EAA2FAB6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="271E05C0F12279EBFA26BBA5063F1FB5">Brancucci, Michel</mods:namePart>
<mods:affiliation id="AF0C29663F1400945191D2BCCB7D9D48">Naturhistorisches Museum, Basel, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource id="5BAE20E81F629B407A15B1FB83F09227">text</mods:typeOfResource>
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<mods:title id="EF92B80AF25227CDC53E9E9260C2D2E9">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="E710D8AC07AB7679EF4959642AB4DBE4">
<mods:date id="11488AA8B7161FAD2D45149AA8462CA7">2015</mods:date>
<mods:detail id="0900FC32EE505A0E500181F210A9C4D0" type="pubDate">
<mods:number id="D3BEE43BEE5762B57C01FBF3BF2342D7">2015-08-06</mods:number>
</mods:detail>
<mods:detail id="7939BDF2EA625C68BE5B7E2B3A229304" type="volume">
<mods:number id="1DC673303E0CC9B3CDD04F9F73DB8294">63</mods:number>
</mods:detail>
<mods:extent id="ACBCCBA2A4915C2F9339F8DD397FEDF8" unit="page">
<mods:start id="FE2617D88320F6B496B11471B2245896">309</mods:start>
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<mods:classification id="A2F713861D54F19033B86764A9C0BFD0">journal article</mods:classification>
<mods:identifier id="C04FE1C11DA26B4172A7AFB9E597A93D" type="CLB-Dataset">55750</mods:identifier>
<mods:identifier id="2DB081872251547CFF651F118CCBEB3B" type="DOI">10.5281/zenodo.5385198</mods:identifier>
<mods:identifier id="B8EE6348AA4E62BFDD42C8C2099E442D" type="GBIF-Dataset">c7f95b94-53f8-420d-97c5-4e27d7c211bf</mods:identifier>
<mods:identifier id="D4C936EAD113ED8269E1ED32CE983FDA" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="558D79BE5DB1F4498F2E284D7A15B261" type="Zenodo-Dep">5385198</mods:identifier>
<mods:identifier id="04106619DE85F9F1D6182D3E0C1113F1" type="ZooBank">AC7F1491-CD89-4F10-9D61-55354B55E368</mods:identifier>
</mods:mods>
<treatment id="038A87C4FFA9FF91F397956FFD4EB349" ID-DOI="http://doi.org/10.5281/zenodo.6512183" ID-GBIF-Taxon="194955230" ID-Zenodo-Dep="6512183" LSID="urn:lsid:zoobank.org:act:D324DBE2-9273-4B55-B432-852A25E46FB0" httpUri="http://treatment.plazi.org/id/038A87C4FFA9FF91F397956FFD4EB349" lastPageId="13" lastPageNumber="322" pageId="12" pageNumber="321">
<subSubSection id="C3396559FFA9FF90F397956FFA9CBC73" box="[957,1353,1814,1839]" pageId="12" pageNumber="321" type="nomenclature">
<paragraph id="8B9C36D2FFA9FF90F397956FFA9CBC73" blockId="12.[957,1353,1814,1871]" box="[957,1353,1814,1839]" pageId="12" pageNumber="321">
<heading id="D0D481BEFFA9FF90F397956FFA9CBC73" bold="true" box="[957,1353,1814,1839]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="321" reason="2">
<emphasis id="B957EAC0FFA9FF90F397956FFA9CBC73" bold="true" box="[957,1353,1814,1839]" pageId="12" pageNumber="321">
<taxonomicName id="4C234D51FFA9FF90F397956FFB61BC72" ID-CoL="8LXXJ" authority="Hájek &amp; Brancucci, 2015" authorityName="Hájek &amp; Brancucci" authorityYear="2015" box="[957,1204,1814,1838]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="321" phylum="Arthropoda" rank="species" species="mazzoldii" status="sp. nov.">
<emphasis id="B957EAC0FFA9FF90F397956FFB61BC72" bold="true" box="[957,1204,1814,1838]" italics="true" pageId="12" pageNumber="321">Laccophilus mazzoldii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26457BBFFA9FF90F4E9956EFA9CBC73" box="[1219,1353,1815,1839]" pageId="12" pageNumber="321" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF90F413954EFB18BC13" box="[1081,1229,1847,1871]" pageId="12" pageNumber="321" type="description">
<paragraph id="8B9C36D2FFA9FF90F413954EFB18BC13" blockId="12.[957,1353,1814,1871]" box="[1081,1229,1847,1871]" pageId="12" pageNumber="321">
(
<figureCitation id="13182A57FFA9FF90F46B954EFB4EBC13" box="[1089,1179,1847,1871]" captionStart="Figs" captionStartId="4.[135,179,2064,2086]" captionTargetBox="[168,1456,189,2051]" captionTargetId="figure-8@4.[133,1476,180,2059]" captionTargetPageId="4" captionText="Figs. 10–17. Habitus of Laccophilus: 10, L. mahakamensis; 11, L. mazzoldii, new species; 12, L. nusatenggaraensis, new species; 13, L. schillhammeri, new species; 14, L. stastnyi, new species; 15, L. tobaensis; 16, L. tonkinensis; 17, L. wittmeri. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385202" httpUri="https://zenodo.org/record/5385202/files/figure.png" pageId="12" pageNumber="321">Figs. 11</figureCitation>
,
<figureCitation id="13182A57FFA9FF90F483954EFB10BC13" box="[1193,1221,1847,1871]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="12" pageNumber="321">29</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF91F369950EFEDFBAC9" lastPageId="13" lastPageNumber="322" pageId="12" pageNumber="321" type="materials_examined">
<paragraph id="8B9C36D2FFA9FF90F369950EFB37BCF2" blockId="12.[835,1474,1911,1967]" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F369950EFC0FBCD3" bold="true" box="[835,986,1911,1935]" pageId="12" pageNumber="321">
<typeStatus id="54988870FFA9FF90F369950EFCA9BCD3" box="[835,892,1911,1935]" pageId="12" pageNumber="321">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3347642FFA9FF90F3F4950EFBEBBCD3" box="[990,1086,1911,1935]" name="Thailand" pageId="12" pageNumber="321">Thailand</collectingCountry>
,
<collectingRegion id="49E7F830FFA9FF90F46D950EFACCBCD3" box="[1095,1305,1911,1935]" country="Thailand" name="Mukdahan" pageId="12" pageNumber="321">Mukdahan Province</collectingRegion>
,
<location id="8EFC6009FFA9FF90F50E950EFC09BCF3" LSID="urn:lsid:plazi:treatment:038A87C4FFA9FF91F397956FFD4EB349:8EFC6009FFA9FF90F50E950EFC09BCF3" country="Thailand" latitude="16.433332" longLatPrecision="1282" longitude="104.8" name="Phu Pha Thoep National Park" pageId="12" pageNumber="321" stateProvince="Mukdahan">Phu Pha Thoep National Park</location>
, ca.
<geoCoordinate id="EE175015FFA9FF90F42595ECFBBFBCF2" box="[1039,1130,1941,1966]" degrees="16" direction="north" minutes="26" orientation="latitude" pageId="12" pageNumber="321" precision="925" value="16.433332">16°26′N</geoCoordinate>
,
<geoCoordinate id="EE175015FFA9FF90F45D95ECFB0BBCF2" box="[1143,1246,1941,1966]" degrees="104" direction="east" minutes="48" orientation="longitude" pageId="12" pageNumber="321" precision="925" value="104.8">104°48′E</geoCoordinate>
.
</paragraph>
<paragraph id="8B9C36D2FFA9FF91F36995AEFEDFBAC9" blockId="12.[835,1475,2007,2095]" lastBlockId="13.[113,753,189,405]" lastPageId="13" lastPageNumber="322" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F36995AEFBE3BCB3" bold="true" box="[835,1078,2007,2031]" pageId="12" pageNumber="321">Material examined.</emphasis>
<materialsCitation id="3B4B3C8FFFA9FF91F46895AEFE6FBA49" ID-GBIF-Occurrence="3760488337" collectingDate="2011-05-31" collectingDateMax="2011-06-06" collectingDateMin="2011-05-31" collectionCode="NHMB" collectorName="M. Brancucci &amp; M. Geiser &amp; D. Hauck &amp; Z. Kraus &amp; A. Phantala &amp; E. Vongphachan" country="Laos" elevation="325" lastPageId="13" lastPageNumber="322" latitude="17.0" location="Phou Xang He" longLatPrecision="1280" longitude="105.63333" municipality="Phou Xang He" pageId="12" pageNumber="321" specimenCount="1" specimenCount-male="1" stateProvince="Savannakhet Prov." typeStatus="holotype">
<typeStatus id="54988870FFA9FF90F46895AEFB79BCB3" box="[1090,1196,2007,2031]" pageId="12" pageNumber="321" type="holotype">Holotype</typeStatus>
<specimenCount id="9D25FD5BFFA9FF90F49295AEFB27BCB3" box="[1208,1266,2007,2031]" pageId="12" pageNumber="321" type="male">male</specimenCount>
, labelled:
<collectingCountry id="F3347642FFA9FF90F55895AEFA68BCB3" box="[1394,1469,2007,2031]" name="Laos" pageId="12" pageNumber="321">LAOS</collectingCountry>
,
<collectingRegion id="49E7F830FFA9FF90F36E958EFBC4B352" box="[836,1041,2039,2063]" country="Laos" name="Savannakhet" pageId="12" pageNumber="321">Savannakhet Prov.</collectingRegion>
, /
<collectingMunicipality id="6BF8ACA8FFA9FF90F406958EFB19B352" box="[1068,1228,2039,2063]" pageId="12" pageNumber="321">Phou Xang He</collectingMunicipality>
NBCA,
<locationDeviation id="9A2E2FF0FFA9FF90F507958EFA56B353" box="[1325,1411,2039,2063]" pageId="12" pageNumber="321">
ca.
<quantity id="4CDB9B37FFA9FF90F57E958EFA56B353" box="[1364,1411,2039,2063]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="321" unit="km" value="5.0">5km</quantity>
</locationDeviation>
/
<locationDeviation id="9A2E2FF0FFA9FF90F5B1958EFC23B373" pageId="12" pageNumber="321">SW Ban Pa Phaknau</locationDeviation>
,
<quantity id="4CDB9B37FFA9FF90F42B9A6EFBADB372" box="[1025,1144,2071,2095]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="12" pageNumber="321" unit="m" value="325.0" valueMax="400.0" valueMin="250.0">
<elevation id="000ED1E1FFA9FF90F42B9A6EFBADB372" box="[1025,1144,2071,2095]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="12" pageNumber="321" unit="m" value="325.0" valueMax="400.0" valueMin="250.0">250-400 m</elevation>
</quantity>
, /
<geoCoordinate id="EE175015FFA9FF90F4B89A6CFB20B372" box="[1170,1269,2069,2094]" degrees="17" direction="north" minutes="00" orientation="latitude" pageId="12" pageNumber="321" precision="925" value="17.0">17°00′ N</geoCoordinate>
,
<geoCoordinate id="EE175015FFA9FF90F52A9A6CFAB8B372" box="[1280,1389,2069,2094]" degrees="105" direction="east" minutes="38" orientation="longitude" pageId="12" pageNumber="321" precision="925" value="105.63333">105°38′ E</geoCoordinate>
, / 31.V. – 6.VI. 2011 // NHMB Basel / Laos 2011 Expedition, /
<collectorName id="26D65304FFA8FF91F2F892C7FF35BBA9" pageId="13" pageNumber="322">M. Brancucci</collectorName>
,
<collectorName id="26D65304FFA8FF91F0C792A7FE89BBA9" box="[237,348,221,245]" pageId="13" pageNumber="322">M. Geiser</collectorName>
, /
<collectorName id="26D65304FFA8FF91F15092A7FE30BBA9" box="[378,485,221,245]" pageId="13" pageNumber="322">D. Hauck</collectorName>
,
<collectorName id="26D65304FFA8FF91F1D892A7FD81BBA9" box="[498,596,222,245]" pageId="13" pageNumber="322">Z. Kraus</collectorName>
, /
<collectorName id="26D65304FFA8FF91F25B92A4FE93BA49" pageId="13" pageNumber="322">A. Phantala &amp; E. Vongphachan</collectorName>
(
<collectionCode id="ED32AE17FFA8FF91F1729284FE7BBA49" box="[344,430,253,277]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="13" pageNumber="322">NHMB</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B4B3C8FFFA8FF91F1EE9287FDFFBA69" ID-GBIF-Occurrence="3760488368" collectingDate="2011-05-31" collectingDateMax="2011-06-06" collectingDateMin="2011-05-31" collectionCode="NHMB, NMPC" collectorName="M. Brancucci &amp; M. Geiser &amp; D. Hauck &amp; Z. Kraus &amp; A. Phantala &amp; E. Vongphachan" country="Laos" elevation="325" lastPageId="13" lastPageNumber="322" latitude="17.0" location="Phou Xang He" longLatPrecision="1280" longitude="105.63333" municipality="Phou Xang He" pageId="13" pageNumber="322" specimenCount="7" specimenCount-female="7" stateProvince="Savannakhet Prov." typeStatus="paratype">
<typeStatus id="54988870FFA8FF91F1EE9287FDE4BA4A" box="[452,561,254,278]" pageId="13" pageNumber="322" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D25FD5BFFA8FF91F2149287FD7EBA49" box="[574,683,253,277]" pageId="13" pageNumber="322" type="female">7 females</specimenCount>
, same label data as holotype (
<collectionCode id="ED32AE17FFA8FF91F1449364FE17BA69" box="[366,450,285,309]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="13" pageNumber="322">NHMB</collectionCode>
,
<collectionCode id="ED32AE17FFA8FF91F1E49364FDCABA69" box="[462,543,285,309]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="13" pageNumber="322">NMPC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B4B3C8FFFA8FF91F2189364FED3BAC9" ID-GBIF-Occurrence="3760488312" collectionCode="PMCB" collectorName="Phu Pha Thoep N. P. &amp; Mazzoldi P." country="Thailand" location="Mukdahan" pageId="13" pageNumber="322" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mukdahan" typeStatus="paratype">
<specimenCount id="9D25FD5BFFA8FF91F2189364FDABBA69" box="[562,638,285,309]" pageId="13" pageNumber="322" type="male">1 male</specimenCount>
,
<specimenCount id="9D25FD5BFFA8FF91F2A19364FD38BA69" box="[651,749,285,309]" pageId="13" pageNumber="322" type="female">1 female</specimenCount>
:
<collectingCountry id="F3347642FFA8FF91F05B9344FF00BA09" box="[113,213,317,341]" name="Thailand" pageId="13" pageNumber="322">Thailand</collectingCountry>
,
<collectingRegion id="49E7F830FFA8FF91F0CE9344FE8DBA09" box="[228,344,317,341]" country="Thailand" name="Mukdahan" pageId="13" pageNumber="322">Mukdahan</collectingRegion>
[Prov.]
<date id="FF9D1012FFA8FF91F1909344FDE1BA09" box="[442,564,317,341]" pageId="13" pageNumber="322" value="2000-01-01">1.I. / 2000</date>
,
<collectorName id="26D65304FFA8FF91F2699344FF77BA29" pageId="13" pageNumber="322">Phu Pha Thoep N.P.</collectorName>
/ Small pools on dry stream / bed,
<collectorName id="26D65304FFA8FF91F2139324FD6ABA29" box="[569,703,349,373]" pageId="13" pageNumber="322">Mazzoldi P.</collectorName>
leg. (23) (
<collectionCode id="ED32AE17FFA8FF91F0849304FF2BBAC9" box="[174,254,381,405]" pageId="13" pageNumber="322">PMCB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA8FF91F05B93C4FF0AB9C9" pageId="13" pageNumber="322" type="diagnosis">
<paragraph id="8B9C36D2FFA8FF91F05B93C4FF0AB9C9" blockId="13.[113,753,445,661]" pageId="13" pageNumber="322">
<emphasis id="B957EAC0FFA8FF91F05B93C4FF3DBA89" bold="true" box="[113,232,445,469]" pageId="13" pageNumber="322">Diagnosis.</emphasis>
Large, oblong oval species (body length
<quantity id="4CDB9B37FFA8FF91F2EB93C4FF18BAA9" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.7" metricValueMin="4.6" pageId="13" pageNumber="322" unit="mm" value="4.65" valueMax="4.7" valueMin="4.6">4.6– 4.7 mm</quantity>
). The new species is characteristic, with elytral colouration that resembles
<taxonomicName id="4C234D51FFA8FF91F18E9387FD8FB949" authorityName="Vazirani" authorityYear="1972" box="[420,602,509,533]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="322" phylum="Arthropoda" rank="species" species="aurofasciatus">
<emphasis id="B957EAC0FFA8FF91F18E9387FD8FB949" box="[420,602,509,533]" italics="true" pageId="13" pageNumber="322">L. aurofasciatus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C234D51FFA8FF91F2419384FF02B969" authorityName="Hájek &amp; Brancucci" authorityYear="2015" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="322" phylum="Arthropoda" rank="species" species="mazzoldii">
<emphasis id="B957EAC0FFA8FF91F2419384FF02B969" italics="true" pageId="13" pageNumber="322">Laccophilus mazzoldii</emphasis>
</taxonomicName>
differs from that species by narrower body shape, elytral sculpture moderately impressed with meshes complete, and by distinct shape of median lobe, resembling that of
<taxonomicName id="4C234D51FFA8FF91F2F19027FF02B9C9" authorityName="Regimbart" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="322" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="B957EAC0FFA8FF91F2F19027FF02B9C9" italics="true" pageId="13" pageNumber="322">L. javanicus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA8FF91F05B90C4FD8BBC09" pageId="13" pageNumber="322" type="description">
<paragraph id="8B9C36D2FFA8FF91F05B90C4FD25B829" blockId="13.[113,754,701,885]" pageId="13" pageNumber="322">
<emphasis id="B957EAC0FFA8FF91F05B90C4FF29B989" bold="true" box="[113,252,701,725]" pageId="13" pageNumber="322">Description.</emphasis>
Colouration: Head and pronotum testaceous. Elytra blackish with broad testaceous subbasal band not separated from elytral base but interrupted on suture, mediolateral spot very small, preapical spot large (preapical transverse band broadly interrupted in sutural area), and small apical spot (
<figureCitation id="13182A57FFA8FF91F0C49124FEEFB829" box="[238,314,861,885]" captionStart="Figs" captionStartId="4.[135,179,2064,2086]" captionTargetBox="[168,1456,189,2051]" captionTargetId="figure-8@4.[133,1476,180,2059]" captionTargetPageId="4" captionText="Figs. 10–17. Habitus of Laccophilus: 10, L. mahakamensis; 11, L. mazzoldii, new species; 12, L. nusatenggaraensis, new species; 13, L. schillhammeri, new species; 14, L. stastnyi, new species; 15, L. tobaensis; 16, L. tonkinensis; 17, L. wittmeri. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385202" httpUri="https://zenodo.org/record/5385202/files/figure.png" pageId="13" pageNumber="322">Fig. 11</figureCitation>
). Appendages and ventral part testaceous.
</paragraph>
<paragraph id="8B9C36D2FFA8FF91F05B91E4FE75BD49" blockId="13.[113,753,925,1557]" pageId="13" pageNumber="322">Surface structures and sculpture: Head reticulation simple, composed of medium sized irregular polygonal meshes, which are moderately deeply impressed. Frons between eyes, and clypeus with several large punctures. Reticulation of pronotum same as that of head. Pronotum along margins with numerous large setigerous punctures. Reticulation of elytra simple, composed of well impressed medium sized irregular polygonal meshes similar to that of head or pronotum. Meshes mostly complete, their inner space usually with several (1–3) small punctures; in some parts secondary microreticulation indicated. Each elytron with indistinct sutural, discal and lateral rows of large serial setigerous punctures. Ventral side reticulated with fine elongate meshes. Prosternal apophysis lanceolate. Abdominal ventrites with numerous oblique grooves and several large setigerous punctures, lateral margin bordered. Apical ventrite rounded posteriorly, with only few large setigerous punctures and transverse grooves on disc. Male pro- and mesotarsomeres 1–3 weakly dilated with bundle of adhesive setae on ventral side. Claws of pro- and mesotarsi simple and equal.</paragraph>
<paragraph id="8B9C36D2FFA8FF91F05B9444FF36BD29" blockId="13.[113,752,1597,1653]" pageId="13" pageNumber="322">
Male genitalia: Median lobe of aedeagus in lateral view as in
<figureCitation id="13182A57FFA8FF91F0A59424FF35BD29" box="[143,224,1629,1653]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="13" pageNumber="322">Fig. 29</figureCitation>
.
</paragraph>
<paragraph id="8B9C36D2FFA8FF91F05B94E4FF39BD89" blockId="13.[113,752,1693,1749]" pageId="13" pageNumber="322">Female genitalia: Similar to male; pro- and mesotarsomeres not dilated.</paragraph>
<paragraph id="8B9C36D2FFA8FF91F05B9484FD8BBC09" blockId="13.[113,752,1789,1877]" pageId="13" pageNumber="322">
<emphasis id="B957EAC0FFA8FF91F05B9484FF20BC49" bold="true" box="[113,245,1789,1813]" pageId="13" pageNumber="322">Variability.</emphasis>
Measurements (N= 10): Body length
<quantity id="4CDB9B37FFA8FF91F28A9484FF4FBC69" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.7" metricValueMin="4.5" pageId="13" pageNumber="322" unit="mm" value="4.6" valueMax="4.7" valueMin="4.5">4.5–4.7 mm</quantity>
(mean value
<quantity id="4CDB9B37FFA8FF91F1079564FE57BC69" box="[301,386,1821,1845]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="322" unit="mm" value="4.5">4.5 mm</quantity>
,
<typeStatus id="54988870FFA8FF91F1A49564FE3EBC69" box="[398,491,1821,1845]" pageId="13" pageNumber="322" type="holotype">holotype</typeStatus>
<quantity id="4CDB9B37FFA8FF91F1D89564FD9CBC69" box="[498,585,1821,1845]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="322" unit="mm" value="4.5">4.5 mm</quantity>
); width
<quantity id="4CDB9B37FFA8FF91F28A9564FF4FBC09" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="13" pageNumber="322" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.5–2.6 mm</quantity>
(mean value
<quantity id="4CDB9B37FFA8FF91F11B9544FE52BC09" box="[305,391,1853,1877]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="322" unit="mm" value="2.5">2.5 mm</quantity>
,
<typeStatus id="54988870FFA8FF91F1BE9544FE24BC09" box="[404,497,1853,1877]" pageId="13" pageNumber="322" type="holotype">holotype</typeStatus>
<quantity id="4CDB9B37FFA8FF91F1D09544FD87BC09" box="[506,594,1853,1877]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="322" unit="mm" value="2.5">2.5 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA8FF91F05B9504FD24BCE9" pageId="13" pageNumber="322" type="etymology">
<paragraph id="8B9C36D2FFA8FF91F05B9504FD24BCE9" blockId="13.[113,753,1917,1973]" pageId="13" pageNumber="322">
<emphasis id="B957EAC0FFA8FF91F05B9504FF26BCC9" bold="true" box="[113,243,1917,1941]" pageId="13" pageNumber="322">Etymology.</emphasis>
The new species is dedicated to its collector, Paolo Mazzoldi (Brescia,
<collectingCountry id="F3347642FFA8FF91F1A695E4FE14BCE9" box="[396,449,1949,1973]" name="Italy" pageId="13" pageNumber="322">Italy</collectingCountry>
), a specialist on
<taxonomicName id="4C234D51FFA8FF91F25D95E4FD38BCE9" authorityName="Leach" authorityYear="1815" box="[631,749,1949,1973]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="322" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA8FF91F05B95A4FD4EB349" pageId="13" pageNumber="322" type="distribution">
<paragraph id="8B9C36D2FFA8FF91F05B95A4FD4EB349" blockId="13.[113,752,2013,2069]" pageId="13" pageNumber="322">
<emphasis id="B957EAC0FFA8FF91F05B95A4FED1BCA9" bold="true" box="[113,260,2013,2037]" pageId="13" pageNumber="322">Distribution.</emphasis>
So far known only from two close localities on both sides of the Thai-Laotian border (
<figureCitation id="13182A57FFA8FF91F2179584FD5AB349" box="[573,655,2045,2069]" captionStart="Figs" captionStartId="9.[113,157,2093,2115]" captionTargetBox="[141,1427,190,2077]" captionTargetPageId="9" captionText="Figs. 40–41. Distribution of the species of the Laccophilus javanicus group" figureDoi="http://doi.org/10.5281/zenodo.5385208" httpUri="https://zenodo.org/record/5385208/files/figure.png" pageId="13" pageNumber="322">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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