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<document id="C76A6068C3760C1BE2D0D4249D014A19" ID-CLB-Dataset="55750" ID-DOI="10.5281/zenodo.5385198" ID-GBIF-Dataset="c7f95b94-53f8-420d-97c5-4e27d7c211bf" ID-ISSN="2345-7600" ID-Zenodo-Dep="5385198" ID-ZooBank="AC7F1491-CD89-4F10-9D61-55354B55E368" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630300948061" checkinUser="valdenar" docAuthor="Hájek, Jiří &amp; Brancucci, Michel" docDate="2015" docId="038A87C4FFA9FF90F3529344FB2FBDB3" docLanguage="en" docName="RafflesBZool.63.309-326.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Laccophilus mahakamensis Balke, Mazzoldi &amp; Hendrich 1998" docType="treatment" docVersion="9" lastPageNumber="321" masterDocId="FFB3FFBCFFA5FF9CF02A9279FFD5BB5C" masterDocTitle="A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)" masterLastPageNumber="326" masterPageNumber="309" pageNumber="321" updateTime="1699240741915" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="044BCDC20EF0378FE609489AC637AB11" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="599F1CEF4510F0E44A2A384AF2BC40FC">
<mods:title id="617803297CC642FE49CD8D475FC85EAD">A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name id="F43EF8F1B4296EA19234DC0DB081A3B9" type="personal">
<mods:role id="ED890C9A5F7A255500488301B68A53A9">
<mods:roleTerm id="8193B09CFDF1D22F1AC838DA8505673F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CFB594728CEBEA0CD843D39B9CE9E8F">Hájek, Jiří</mods:namePart>
<mods:affiliation id="FEDFD6B3D193BCBBC3C8CBD6ECD629F8">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 – Horní Počernice, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="BB9159046C6E0397A257480E81D17749" type="email">jiri_hajek@nm.cz</mods:nameIdentifier>
</mods:name>
<mods:name id="1C62F65D942615B84B9DD3C7ACA84F9E" type="personal">
<mods:role id="7F4F5B214D22DCAD4AEFA9DD5568B9B5">
<mods:roleTerm id="2F88064B6D5BA0467E0FA19AAC9903CE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E191515D7432312163EB5F0AD2FEF66">Brancucci, Michel</mods:namePart>
<mods:affiliation id="48372A2925FBF90D148C48707D893841">Naturhistorisches Museum, Basel, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource id="56F06E03216F20A6774E1D379D3897A5">text</mods:typeOfResource>
<mods:relatedItem id="7A43FF6CD11A018C68C6BA4B1168F3C4" type="host">
<mods:titleInfo id="DFDF6C094C8E37E5E17AEA1F4E42A29B">
<mods:title id="AC0AF2747919FDDE5AC29E446F8F86E0">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="A97E606CBC0898C1608754716F7454B3">
<mods:date id="7B2958F84969E9EF0B416D636E1B7B35">2015</mods:date>
<mods:detail id="8927AF961F532777ADA49A54C9950734" type="pubDate">
<mods:number id="08644ED3EE12CB294B94A2FBB94E528E">2015-08-06</mods:number>
</mods:detail>
<mods:detail id="2AC2D1E325B8FA53C24C4B535CD77B08" type="volume">
<mods:number id="BBA3921921FA0392E3CF0DA5B1F26AA1">63</mods:number>
</mods:detail>
<mods:extent id="2B3F6A4BF55B13662A0FB286B212B2B9" unit="page">
<mods:start id="5120DB8DA4F9EB6110658DCFEBBF9A87">309</mods:start>
<mods:end id="FCAE8FD5BE953B38943C4CC2489488AA">326</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="C0AD705D12A63603D81FCED554C85EEE">journal article</mods:classification>
<mods:identifier id="4500A6E904802169666E77E4496F7676" type="CLB-Dataset">55750</mods:identifier>
<mods:identifier id="D02BEA3D52F1E7991B84C7BA4EEDA9F2" type="DOI">10.5281/zenodo.5385198</mods:identifier>
<mods:identifier id="A863676D73D428415104C833E948428E" type="GBIF-Dataset">c7f95b94-53f8-420d-97c5-4e27d7c211bf</mods:identifier>
<mods:identifier id="621872CF9D7DDFE57E9A106E4449ABB9" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="3050ECBC610280422672B0CC21AA8906" type="Zenodo-Dep">5385198</mods:identifier>
<mods:identifier id="D14AEF6548BF3FAF8E1258AB7FAE28B3" type="ZooBank">AC7F1491-CD89-4F10-9D61-55354B55E368</mods:identifier>
</mods:mods>
<treatment id="038A87C4FFA9FF90F3529344FB2FBDB3" ID-DOI="http://doi.org/10.5281/zenodo.5466897" ID-GBIF-Taxon="194955231" ID-Zenodo-Dep="5466897" LSID="urn:lsid:plazi:treatment:038A87C4FFA9FF90F3529344FB2FBDB3" httpUri="http://treatment.plazi.org/id/038A87C4FFA9FF90F3529344FB2FBDB3" lastPageNumber="321" pageId="12" pageNumber="321">
<subSubSection id="C3396559FFA9FF90F3529344FB08BA29" pageId="12" pageNumber="321" type="nomenclature">
<paragraph id="8B9C36D2FFA9FF90F3529344FB08BA29" blockId="12.[888,1423,317,405]" pageId="12" pageNumber="321">
<heading id="D0D481BEFFA9FF90F3529344FB08BA29" bold="true" centered="true" fontSize="10" level="2" pageId="12" pageNumber="321" reason="2">
<taxonomicName id="4C234D51FFA9FF90F3529344FB08BA29" ID-CoL="3RM8S" authority="Balke, Mazzoldi &amp; Hendrich, 1998" authorityName="Balke, Mazzoldi &amp; Hendrich" authorityYear="1998" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="321" phylum="Arthropoda" rank="species" species="mahakamensis">
<emphasis id="B957EAC0FFA9FF90F3529344FB08BA29" bold="true" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F3529344FB79BA09" bold="true" box="[888,1196,317,341]" italics="true" pageId="12" pageNumber="321">Laccophilus mahakamensis</emphasis>
<bibRefCitation id="EFB24B23FFA9FF90F49D9344FB08BA29" author="Balke M &amp; Mazzoldi P &amp; Hendrich L" firstAuthor="Balke" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="12" pageNumber="321" pagination="71 - 77" part="46" refId="ref15090" refString="Balke M, Mazzoldi P &amp; Hendrich L (1998) Two new Laccophilus species (Coleoptera: Dytiscidae) from Southeast Asia, and notes on other species of the genus. The Raffles Bulletin of Zoology, 46: 71 - 77." title="Two new Laccophilus species (Coleoptera: Dytiscidae) from Southeast Asia, and notes on other species of the genus" type="journal article" year="1998">Balke, Mazzoldi &amp; Hendrich, 1998</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF90F4139304FB18BAC9" box="[1081,1229,381,405]" pageId="12" pageNumber="321" type="description">
<paragraph id="8B9C36D2FFA9FF90F4139304FB18BAC9" blockId="12.[888,1423,317,405]" box="[1081,1229,381,405]" pageId="12" pageNumber="321">
(
<figureCitation id="13182A57FFA9FF90F46B9304FB49BAC9" box="[1089,1180,381,405]" captionStart="Figs" captionStartId="4.[135,179,2064,2086]" captionTargetBox="[168,1456,189,2051]" captionTargetId="figure-8@4.[133,1476,180,2059]" captionTargetPageId="4" captionText="Figs. 10–17. Habitus of Laccophilus: 10, L. mahakamensis; 11, L. mazzoldii, new species; 12, L. nusatenggaraensis, new species; 13, L. schillhammeri, new species; 14, L. stastnyi, new species; 15, L. tobaensis; 16, L. tonkinensis; 17, L. wittmeri. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385202" httpUri="https://zenodo.org/record/5385202/files/figure.png" pageId="12" pageNumber="321">Figs. 10</figureCitation>
,
<figureCitation id="13182A57FFA9FF90F4839304FB10BAC9" box="[1193,1221,381,405]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="12" pageNumber="321">28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF90F36993C5FB88BAB3" pageId="12" pageNumber="321" type="reference_group">
<paragraph id="8B9C36D2FFA9FF90F36993C5FB88BAB3" blockId="12.[835,1475,444,495]" pageId="12" pageNumber="321">
<treatmentCitationGroup id="AB3311FCFFA9FF90F36993C5FB88BAB3" pageId="12" pageNumber="321">
<treatmentCitation id="0A8210C3FFA9FF90F36993C5FC54BAB2" author="Balke M &amp; Mazzoldi P &amp; Hendrich L" page="74" pageId="12" pageNumber="321" year="1998">
<taxonomicName id="4C234D51FFA9FF90F36993C5FC54BAB2" ID-CoL="3RM8S" authority="Balke, Mazzoldi &amp; Hendrich, 1998: 74" authorityName="Balke, Mazzoldi &amp; Hendrich" authorityPageNumber="74" authorityYear="1998" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="321" phylum="Arthropoda" rank="species" species="mahakamensis">
<emphasis id="B957EAC0FFA9FF90F36993C5FB84BA8D" box="[835,1105,444,465]" italics="true" pageId="12" pageNumber="321">Laccophilus mahakamensis</emphasis>
<bibRefCitation id="EFB24B23FFA9FF90F47093C5FC54BAB2" author="Balke M &amp; Mazzoldi P &amp; Hendrich L" firstAuthor="Balke" journalOrPublisher="The Raffles Bulletin of Zoology" page="74" pageId="12" pageNumber="321" pagination="71 - 77" part="46" refId="ref15090" refString="Balke M, Mazzoldi P &amp; Hendrich L (1998) Two new Laccophilus species (Coleoptera: Dytiscidae) from Southeast Asia, and notes on other species of the genus. The Raffles Bulletin of Zoology, 46: 71 - 77." title="Two new Laccophilus species (Coleoptera: Dytiscidae) from Southeast Asia, and notes on other species of the genus" type="journal article" year="1998">Balke, Mazzoldi &amp; Hendrich, 1998: 74</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF90F369906EFB7BBF13" pageId="12" pageNumber="321" type="materials_examined">
<paragraph id="8B9C36D2FFA9FF90F369906EFAAAB913" blockId="12.[835,1474,535,591]" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F369906EFC35B973" bold="true" box="[835,992,535,559]" pageId="12" pageNumber="321">
<typeStatus id="54988870FFA9FF90F369906EFCA8B973" box="[835,893,535,559]" pageId="12" pageNumber="321">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3347642FFA9FF90F3C3906EFB86B973" box="[1001,1107,535,559]" name="Indonesia" pageId="12" pageNumber="321">Indonesia</collectingCountry>
,
<collectingRegion id="49E7F830FFA9FF90F44B906EFAC2B973" box="[1121,1303,535,559]" country="Indonesia" name="Kalimantan Timur" pageId="12" pageNumber="321">East Kalimantan</collectingRegion>
, district barong
<collectingCounty id="62FD4E5EFFA9FF90F369904EFC73B913" box="[835,934,567,591]" pageId="12" pageNumber="321">Tongkok</collectingCounty>
,
<collectingMunicipality id="6BF8ACA8FFA9FF90F399904EFBF3B913" box="[947,1062,567,591]" pageId="12" pageNumber="321">Juhan Asa</collectingMunicipality>
,
<location id="8EFC6009FFA9FF90F419904EFBBDB913" LSID="urn:lsid:plazi:treatment:038A87C4FFA9FF90F3529344FB2FBDB3:8EFC6009FFA9FF90F419904EFBBDB913" box="[1075,1128,567,591]" country="Indonesia" county="Tongkok" municipality="Juhan Asa" name="Aput" pageId="12" pageNumber="321" stateProvince="Kalimantan Timur">Aput</location>
stream in primary forest.
</paragraph>
<paragraph id="8B9C36D2FFA9FF90F369900EFB7BBF13" blockId="12.[835,1475,631,1103]" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F369900EFBECB9D3" bold="true" box="[835,1081,631,655]" pageId="12" pageNumber="321">Material examined.</emphasis>
<specimenCount id="9D25FD5BFFA9FF90F46F900EFB3FB9D3" box="[1093,1258,631,655]" pageId="12" pageNumber="321" type="generic">35 specimens</specimenCount>
.
<materialsCitation id="3B4B3C8FFFA9FF90F4D1900EFC71B853" ID-GBIF-Occurrence="3760488354" collectingDate="2011-11-30" collectionCode="NMPC" collectorName="J. Hajek &amp; J. Schneider &amp; P. Votruba" country="Indonesia" elevation="66" latitude="-1.2233334" location="Kalimantan" longLatPrecision="130" longitude="116.37666" pageId="12" pageNumber="321" specimenCount="5" stateProvince="Kalimantan Utara">
<emphasis id="B957EAC0FFA9FF90F4D1900EFC1EB9F3" bold="true" pageId="12" pageNumber="321">
<collectingCountry id="F3347642FFA9FF90F4D1900EFA4EB9D3" box="[1275,1435,631,655]" name="Indonesia" pageId="12" pageNumber="321">INDONESIA</collectingCountry>
: E
<collectingRegion id="49E7F830FFA9FF90F36990EEFC1EB9F3" box="[835,971,663,687]" country="Indonesia" name="Kalimantan Utara" pageId="12" pageNumber="321">Kalimantan</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFA9FF90F3F690EEFBC0B9F2" box="[988,1045,663,687]" pageId="12" pageNumber="321" type="generic">5 ex.</specimenCount>
, ca.
<locationDeviation id="9A2E2FF0FFA9FF90F46390EEFA83B9F3" box="[1097,1366,663,687]" location="Balikpapan" pageId="12" pageNumber="321">
<quantity id="4CDB9B37FFA9FF90F46390EEFB5AB9F3" box="[1097,1167,663,687]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="321" unit="km" value="50.0">50 km</quantity>
W of Balikpapan
</locationDeviation>
, PT
<location id="8EFC6009FFA9FF90F5A190EEFC3CB993" LSID="urn:lsid:plazi:treatment:038A87C4FFA9FF90F3529344FB2FBDB3:8EFC6009FFA9FF90F5A190EEFC3CB993" country="Indonesia" latitude="-1.2233334" longLatPrecision="130" longitude="116.37666" name="Fajar Surya Swadaya" pageId="12" pageNumber="321">Fajar Surya Swadaya</location>
[area],
<geoCoordinate id="EE175015FFA9FF90F41790CEFB72B993" box="[1085,1191,693,719]" degrees="01" direction="south" minutes="13.4" orientation="latitude" pageId="12" pageNumber="321" precision="92" value="-1.2233334">01°13.4′S</geoCoordinate>
,
<geoCoordinate id="EE175015FFA9FF90F49990CEFAFBB992" box="[1203,1326,693,719]" degrees="116" direction="east" minutes="22.6" orientation="longitude" pageId="12" pageNumber="321" precision="92" value="116.37666">116°22.6′E</geoCoordinate>
,
<quantity id="4CDB9B37FFA9FF90F51390CEFAA7B992" box="[1337,1394,695,719]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="12" pageNumber="321" unit="m" value="66.0">
<elevation id="000ED1E1FFA9FF90F51390CEFAA7B992" box="[1337,1394,695,719]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="12" pageNumber="321" unit="m" value="66.0">66 m</elevation>
</quantity>
, 27+
<date id="FF9D1012FFA9FF90F58D90CEFC20B9B3" pageId="12" pageNumber="321" value="2011-11-30">
<collectingDate id="EFD9E9FAFFA9FF90F58D90CEFC20B9B3" pageId="12" pageNumber="321" value="2011-11-30">30 November 2011</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA9FF90F41390AEFB43B9B3" box="[1081,1174,727,751]" pageId="12" pageNumber="321">J. Hájek</collectorName>
,
<collectorName id="26D65304FFA9FF90F48890AEFAF2B9B3" box="[1186,1319,727,751]" pageId="12" pageNumber="321">J. Schneider</collectorName>
&amp;
<collectorName id="26D65304FFA9FF90F56690AEFA17B9B3" box="[1356,1474,727,751]" pageId="12" pageNumber="321">P. Votruba</collectorName>
(
<collectionCode id="ED32AE17FFA9FF90F361908EFC48B853" box="[843,925,759,783]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="12" pageNumber="321">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFA9FF90F39A908EFBD2B833" ID-GBIF-Occurrence="3760488433" collectingDate="2011-12-08" collectingDateMax="2011-12-11" collectingDateMin="2011-12-08" collectionCode="NMPC, ZSMG" collectorName="J. Hajek &amp; J. Schneider &amp; P. Votruba" country="Indonesia" elevation="35" latitude="-1.133333" location="Sungai Wain Protection Forest" longitude="116.833333" pageId="12" pageNumber="321" specimenCount="22">
<specimenCount id="9D25FD5BFFA9FF90F39A908EFC20B852" box="[944,1013,759,783]" pageId="12" pageNumber="321" type="generic">22 ex.</specimenCount>
, ca.
<locationDeviation id="9A2E2FF0FFA9FF90F402908EFAFFB853" box="[1064,1322,759,783]" location="Balikpapan" pageId="12" pageNumber="321">
<quantity id="4CDB9B37FFA9FF90F402908EFBB9B853" box="[1064,1132,759,783]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="321" unit="km" value="15.0">15 km</quantity>
N of Balikpapan
</locationDeviation>
,
<location id="8EFC6009FFA9FF90F51C908EFBDBB872" LSID="urn:lsid:plazi:treatment:038A87C4FFA9FF90F3529344FB2FBDB3:8EFC6009FFA9FF90F51C908EFBDBB872" country="Indonesia" latitude="-1.135" name="Sungai Wain Protection Forest" pageId="12" pageNumber="321">Sungai Wain Protection Forest</location>
, 01°08.1′S, 116°49.9′’E,
<quantity id="4CDB9B37FFA9FF90F511916EFAAFB872" box="[1339,1402,791,815]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="321" unit="m" value="35.0">
<elevation id="000ED1E1FFA9FF90F511916EFAAFB872" box="[1339,1402,791,815]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="321" unit="m" value="35.0">35 m</elevation>
</quantity>
,
<date id="FF9D1012FFA9FF90F5A0916EFC26B813" pageId="12" pageNumber="321" value="2011-12-08" valueMax="2011-12-11" valueMin="2011-12-08">
<collectingDate id="EFD9E9FAFFA9FF90F5A0916EFC26B813" pageId="12" pageNumber="321" value="2011-12-08" valueMax="2011-12-11" valueMin="2011-12-08">8–11 December 2011</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA9FF90F412914EFB40B813" box="[1080,1173,823,847]" pageId="12" pageNumber="321">J. Hájek</collectorName>
,
<collectorName id="26D65304FFA9FF90F48B914EFAF3B813" box="[1185,1318,823,847]" pageId="12" pageNumber="321">J. Schneider</collectorName>
&amp;
<collectorName id="26D65304FFA9FF90F566914EFA17B813" box="[1356,1474,823,847]" pageId="12" pageNumber="321">P. Votruba</collectorName>
(
<collectionCode id="ED32AE17FFA9FF90F361912EFC4BB833" box="[843,926,855,879]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="12" pageNumber="321">NMPC</collectionCode>
,
<collectionCode id="ED32AE17FFA9FF90F387912EFC2AB833" box="[941,1023,855,879]" pageId="12" pageNumber="321">ZSMG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4B3C8FFFA9FF90F43F912EFAEAB8F3" ID-GBIF-Occurrence="3760488349" collectingDate="1993-02-20" collectionCode="NHMW" collectorName="E Bandar Sri Ammen &amp; M. A. Jach" country="Malaysia" location="Batang Ai National Park" pageId="12" pageNumber="321" specimenCount="1" specimenCount-female="1">
<emphasis id="B957EAC0FFA9FF90F43F912EFAFDB833" bold="true" box="[1045,1320,855,879]" pageId="12" pageNumber="321">
<collectingCountry id="F3347642FFA9FF90F43F912EFB7EB833" box="[1045,1195,855,879]" name="Malaysia" pageId="12" pageNumber="321">MALAYSIA</collectingCountry>
: Sarawak
</emphasis>
:
<specimenCount id="9D25FD5BFFA9FF90F511912EFA76B833" box="[1339,1443,855,879]" pageId="12" pageNumber="321" type="female">1 female</specimenCount>
,
<collectorName id="26D65304FFA9FF90F598912EFBCCB8D3" pageId="12" pageNumber="321">E Bandar Sri Ammen</collectorName>
[Sri Aman],
<location id="8EFC6009FFA9FF90F487910EFA6AB8D3" LSID="urn:lsid:plazi:treatment:038A87C4FFA9FF90F3529344FB2FBDB3:8EFC6009FFA9FF90F487910EFA6AB8D3" box="[1197,1471,887,911]" country="Malaysia" name="Batang Ai National Park" pageId="12" pageNumber="321">Batang Ai National Park</location>
,
<date id="FF9D1012FFA9FF90F36991EEFBD9B8F3" box="[835,1036,919,943]" pageId="12" pageNumber="321" value="1993-02-20">
<collectingDate id="EFD9E9FAFFA9FF90F36991EEFBD9B8F3" box="[835,1036,919,943]" pageId="12" pageNumber="321" value="1993-02-20">20 February 1993</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA9FF90F47E91EEFB12B8F3" box="[1108,1223,919,943]" pageId="12" pageNumber="321">M.A. Jäch</collectorName>
(
<collectionCode id="ED32AE17FFA9FF90F4F391EEFAE2B8F3" box="[1241,1335,919,943]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="321" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFA9FF90F56791EEFB94B8B3" ID-GBIF-Occurrence="3760488431" collectingDate="2003-05-30" collectionCode="JSCL" collectorName="J. Stastny" country="Malaysia" location="Gading National Park" pageId="12" pageNumber="321" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="9D25FD5BFFA9FF90F56791EEFA73B8F3" box="[1357,1446,919,943]" pageId="12" pageNumber="321" type="male">2 males</specimenCount>
,
<specimenCount id="9D25FD5BFFA9FF90F59F91EEFC5BB893" pageId="12" pageNumber="321" type="female">1 female</specimenCount>
, G. [Gunung]
<location id="8EFC6009FFA9FF90F40491CEFAC8B893" LSID="urn:lsid:plazi:treatment:038A87C4FFA9FF90F3529344FB2FBDB3:8EFC6009FFA9FF90F40491CEFAC8B893" box="[1070,1309,951,975]" country="Malaysia" name="Gading National Park" pageId="12" pageNumber="321">Gading National Park</location>
,
<date id="FF9D1012FFA9FF90F50091CEFA6BB893" box="[1322,1470,951,975]" pageId="12" pageNumber="321" value="2003-05-30">
<collectingDate id="EFD9E9FAFFA9FF90F50091CEFA6BB893" box="[1322,1470,951,975]" pageId="12" pageNumber="321" value="2003-05-30">30 May 2003</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA9FF90F35191AEFC3DB8AD" box="[891,1000,983,1009]" pageId="12" pageNumber="321">J. Šťastný</collectorName>
(
<collectionCode id="ED32AE17FFA9FF90F3D291AEFBECB8B3" box="[1016,1081,983,1007]" pageId="12" pageNumber="321">JSCL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4B3C8FFFA9FF90F46491AEFB7EBF13" ID-GBIF-Occurrence="3760488367" collectingDate="1996-07-26" collectionCode="PMCB" collectorName="P. Mazzoldi" country="Thailand" county="Khao Sok National Park" location="Bang Leap Nam" municipality="Park Hq" pageId="12" pageNumber="321" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Surat Thani">
<emphasis id="B957EAC0FFA9FF90F46491AEFB0BB8B3" bold="true" box="[1102,1246,983,1007]" pageId="12" pageNumber="321">
<collectingCountry id="F3347642FFA9FF90F46491AEFB0BB8B3" box="[1102,1246,983,1007]" name="Thailand" pageId="12" pageNumber="321">THAILAND</collectingCountry>
</emphasis>
:
<specimenCount id="9D25FD5BFFA9FF90F4C491AEFA90B8B3" box="[1262,1349,983,1007]" pageId="12" pageNumber="321" type="male">2 males</specimenCount>
,
<specimenCount id="9D25FD5BFFA9FF90F57891AEFA6BB8B3" box="[1362,1470,983,1007]" pageId="12" pageNumber="321" type="female">2 females</specimenCount>
,
<collectingRegion id="49E7F830FFA9FF90F369918EFBE6BF53" box="[835,1075,1015,1039]" country="Thailand" name="Surat Thani" pageId="12" pageNumber="321">Surat Thani Province</collectingRegion>
,
<collectingCounty id="62FD4E5EFFA9FF90F46A918EFA81BF53" box="[1088,1364,1015,1039]" pageId="12" pageNumber="321">Khao Sok National Park</collectingCounty>
, pools on road from
<collectingMunicipality id="6BF8ACA8FFA9FF90F391966EFBC2BF72" box="[955,1047,1047,1071]" pageId="12" pageNumber="321">Park Hq</collectingMunicipality>
to
<location id="8EFC6009FFA9FF90F46A966EFB23BF72" LSID="urn:lsid:plazi:treatment:038A87C4FFA9FF90F3529344FB2FBDB3:8EFC6009FFA9FF90F46A966EFB23BF72" box="[1088,1270,1047,1071]" country="Thailand" county="Khao Sok National Park" municipality="Park Hq" name="Bang Leap Nam" pageId="12" pageNumber="321" stateProvince="Surat Thani">Bang Leap Nam</location>
waterfall,
<date id="FF9D1012FFA9FF90F55A966EFCAEBF13" pageId="12" pageNumber="321" value="1996-07-26">
<collectingDate id="EFD9E9FAFFA9FF90F55A966EFCAEBF13" pageId="12" pageNumber="321" value="1996-07-26">26 July 1996</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA9FF90F395964EFB96BF13" box="[959,1091,1079,1103]" pageId="12" pageNumber="321">P. Mazzoldi</collectorName>
(
<collectionCode id="ED32AE17FFA9FF90F478964EFB77BF13" box="[1106,1186,1079,1103]" pageId="12" pageNumber="321">PMCB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF90F369960EFB5FBE53" pageId="12" pageNumber="321" type="diagnosis">
<paragraph id="8B9C36D2FFA9FF90F369960EFB5FBE53" blockId="12.[835,1474,1143,1295]" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F369960EFC62BFD3" bold="true" box="[835,951,1143,1167]" pageId="12" pageNumber="321">Diagnosis.</emphasis>
Small, elongate oval species (body length
<quantity id="4CDB9B37FFA9FF90F558960EFCA4BFF3" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.7" metricValueMin="3.4" pageId="12" pageNumber="321" unit="mm" value="3.55" valueMax="3.7" valueMin="3.4">3.4–3.7 mm</quantity>
). Elytra with subbasal testaceous band sinuous, narrower, usually well separated from elytral base (
<figureCitation id="13182A57FFA9FF90F53A96CEFAB1BF93" box="[1296,1380,1207,1231]" captionStart="Figs" captionStartId="4.[135,179,2064,2086]" captionTargetBox="[168,1456,189,2051]" captionTargetId="figure-8@4.[133,1476,180,2059]" captionTargetPageId="4" captionText="Figs. 10–17. Habitus of Laccophilus: 10, L. mahakamensis; 11, L. mazzoldii, new species; 12, L. nusatenggaraensis, new species; 13, L. schillhammeri, new species; 14, L. stastnyi, new species; 15, L. tobaensis; 16, L. tonkinensis; 17, L. wittmeri. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385202" httpUri="https://zenodo.org/record/5385202/files/figure.png" pageId="12" pageNumber="321">Fig. 10</figureCitation>
). Apical part of median lobe in lateral view distinctly sinuous with elongate pointed tip (
<figureCitation id="13182A57FFA9FF90F406968EFBABBE53" box="[1068,1150,1271,1295]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 18–35. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="12" pageNumber="321">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF90F369974EFC03BEB2" pageId="12" pageNumber="321" type="biology_ecology">
<paragraph id="8B9C36D2FFA9FF90F369974EFC03BEB2" blockId="12.[835,1475,1335,1519]" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F369974EFC76BE13" bold="true" box="[835,931,1335,1359]" pageId="12" pageNumber="321">Habitat.</emphasis>
In
<collectingRegion id="49E7F830FFA9FF90F3E7974EFB52BE13" box="[973,1159,1335,1359]" country="Indonesia" name="Kalimantan Timur" pageId="12" pageNumber="321">East Kalimantan</collectingRegion>
,
<taxonomicName id="4C234D51FFA9FF90F4BF974EFA16BE13" authorityName="Balke, Mazzoldi &amp; Hendrich" authorityYear="1998" box="[1173,1475,1335,1359]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="321" phylum="Arthropoda" rank="species" species="mahakamensis">
<emphasis id="B957EAC0FFA9FF90F4BF974EFA16BE13" box="[1173,1475,1335,1359]" italics="true" pageId="12" pageNumber="321">Laccophilus mahakamensis</emphasis>
</taxonomicName>
was collected in remnant pools of small stream in remains of lowland primary forest; pools were usually with clay bottom and a thick layer of decaying leaves (
<figureCitation id="13182A57FFA9FF90F4CA97EEFAE4BEF3" box="[1248,1329,1431,1455]" captionStart="Figs" captionStartId="8.[135,179,2035,2057]" captionTargetBox="[187,1423,189,2020]" captionTargetId="figure-8@8.[187,1423,189,2020]" captionTargetPageId="8" captionText="Figs. 36–39. Habitats: 36, E Kalimantan, Muara Ritan (type locality of Laccophilus kalimantanensis, new species); 37, China, Guangxi, Shiwandashan (locality of L. komareki, new species); 38, E Kalimantan, Sungai Wain (locality of L. mahakamensis); 39, Laos, Attapeu (locality of L. tonkinensis)." figureDoi="http://doi.org/10.5281/zenodo.5385206" httpUri="https://zenodo.org/record/5385206/files/figure.png" pageId="12" pageNumber="321">Fig. 38</figureCitation>
), some pools were very small (diameter less than
<quantity id="4CDB9B37FFA9FF90F4E797CEFAC4BE93" box="[1229,1297,1463,1487]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="321" unit="cm" value="20.0">20 cm</quantity>
) with almost no visible water.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA9FF90F369946EFB2FBDB3" pageId="12" pageNumber="321" type="distribution">
<paragraph id="8B9C36D2FFA9FF90F369946EFB2FBDB3" blockId="12.[835,1475,1559,1775]" pageId="12" pageNumber="321">
<emphasis id="B957EAC0FFA9FF90F369946EFC03BD73" bold="true" box="[835,982,1559,1583]" pageId="12" pageNumber="321">Distribution.</emphasis>
The species was described and is so far only known from Indonesian
<collectingRegion id="49E7F830FFA9FF90F468944EFB3ABD13" box="[1090,1263,1591,1615]" country="Indonesia" name="Kalimantan Timur" pageId="12" pageNumber="321">East Kalimantan</collectingRegion>
(
<bibRefCitation id="EFB24B23FFA9FF90F4D6944EFA62BD13" author="Balke M &amp; Mazzoldi P &amp; Hendrich L" box="[1276,1463,1591,1615]" etAl="et al." firstAuthor="Balke" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="12" pageNumber="321" pagination="71 - 77" part="46" refId="ref15090" refString="Balke M, Mazzoldi P &amp; Hendrich L (1998) Two new Laccophilus species (Coleoptera: Dytiscidae) from Southeast Asia, and notes on other species of the genus. The Raffles Bulletin of Zoology, 46: 71 - 77." title="Two new Laccophilus species (Coleoptera: Dytiscidae) from Southeast Asia, and notes on other species of the genus" type="journal article" year="1998">Balke et al., 1998</bibRefCitation>
). The Thai specimens correspond well with the
<typeStatus id="54988870FFA9FF90F51F942EFAB6BD33" box="[1333,1379,1623,1647]" pageId="12" pageNumber="321">type</typeStatus>
material stored in NHMW. The present new records markedly extend the distributional area of
<taxonomicName id="4C234D51FFA9FF90F46394EEFAB8BDF3" authorityName="Balke, Mazzoldi &amp; Hendrich" authorityYear="1998" box="[1097,1389,1687,1711]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="321" phylum="Arthropoda" rank="species" species="mahakamensis">
<emphasis id="B957EAC0FFA9FF90F46394EEFAB8BDF3" box="[1097,1389,1687,1711]" italics="true" pageId="12" pageNumber="321">Laccophilus mahakamensis</emphasis>
</taxonomicName>
suggest the likelihood of more records from the Malay Peninsula and the Greater Sunda Islands (
<figureCitation id="13182A57FFA9FF90F4B694AEFB3BBDB3" box="[1180,1262,1751,1775]" captionStart="Figs" captionStartId="9.[113,157,2093,2115]" captionTargetBox="[141,1427,190,2077]" captionTargetPageId="9" captionText="Figs. 40–41. Distribution of the species of the Laccophilus javanicus group" figureDoi="http://doi.org/10.5281/zenodo.5385208" httpUri="https://zenodo.org/record/5385208/files/figure.png" pageId="12" pageNumber="321">Fig. 41</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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