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<document id="3A8C2E832F68E414A11ECC383F0F67F5" ID-CLB-Dataset="55750" ID-DOI="10.5281/zenodo.5385198" ID-GBIF-Dataset="c7f95b94-53f8-420d-97c5-4e27d7c211bf" ID-ISSN="2345-7600" ID-Zenodo-Dep="5385198" ID-ZooBank="AC7F1491-CD89-4F10-9D61-55354B55E368" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630300948061" checkinUser="valdenar" docAuthor="Hájek, Jiří &amp; Brancucci, Michel" docDate="2015" docId="038A87C4FFA0FF99F3B49124FB5DBDEE" docLanguage="en" docName="RafflesBZool.63.309-326.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Laccophilus girardi Brancucci 1983" docType="treatment" docVersion="9" lastPageNumber="314" masterDocId="FFB3FFBCFFA5FF9CF02A9279FFD5BB5C" masterDocTitle="A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)" masterLastPageNumber="326" masterPageNumber="309" pageNumber="314" updateTime="1699240741915" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1C4FD2BD93C1BFE96882C7EB5635490B">A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)</mods:title>
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<mods:namePart id="C4F3E39EBC8D4D85EA9F7AD81E91F270">Hájek, Jiří</mods:namePart>
<mods:affiliation id="6F9336EBBCA60EEA1DFC74709F98F236">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 Horní Počernice, Czech Republic</mods:affiliation>
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<mods:namePart id="BD1677414E005D3293A21D82DC791E92">Brancucci, Michel</mods:namePart>
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<treatment id="038A87C4FFA0FF99F3B49124FB5DBDEE" ID-DOI="http://doi.org/10.5281/zenodo.5466885" ID-GBIF-Taxon="194955236" ID-Zenodo-Dep="5466885" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F3B49124FB5DBDEE" httpUri="http://treatment.plazi.org/id/038A87C4FFA0FF99F3B49124FB5DBDEE" lastPageNumber="314" pageId="5" pageNumber="314">
<subSubSection id="C3396559FFA0FF99F3B49124FAE9B829" box="[926,1340,861,885]" pageId="5" pageNumber="314" type="nomenclature">
<paragraph id="8B9C36D2FFA0FF99F3B49124FAE9B829" blockId="5.[926,1340,861,917]" box="[926,1340,861,885]" pageId="5" pageNumber="314">
<heading id="D0D481BEFFA0FF99F3B49124FAE9B829" bold="true" box="[926,1340,861,885]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="314" reason="2">
<taxonomicName id="4C234D51FFA0FF99F3B49124FAE9B829" ID-CoL="3RM7J" authority="Brancucci, 1983" authorityName="Brancucci" authorityYear="1983" box="[926,1340,861,885]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="314" phylum="Arthropoda" rank="species" species="girardi">
<emphasis id="B957EAC0FFA0FF99F3B49124FAE9B829" bold="true" box="[926,1340,861,885]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F3B49124FBAFB829" bold="true" box="[926,1146,861,885]" italics="true" pageId="5" pageNumber="314">Laccophilus girardi</emphasis>
<bibRefCitation id="EFB24B23FFA0FF99F4AE9124FAE9B829" author="Brancucci M" box="[1156,1340,861,885]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" pageId="5" pageNumber="314" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA0FF99F4039104FB65B8C9" box="[1065,1200,893,917]" pageId="5" pageNumber="314" type="description">
<paragraph id="8B9C36D2FFA0FF99F4039104FB65B8C9" blockId="5.[926,1340,861,917]" box="[1065,1200,893,917]" pageId="5" pageNumber="314">
(
<figureCitation id="13182A57FFA0FF99F41B9104FBABB8C9" box="[1073,1150,893,917]" captionStart="Figs" captionStartId="3.[113,157,2051,2073]" captionTargetBox="[115,1452,190,2035]" captionTargetId="figure-10@3.[97,1463,180,2039]" captionTargetPageId="3" captionText="Figs. 19. Habitus of Laccophilus: 1, L. aurofasciatus; 2, L. auropictus; 3, L. ceylonicus; 4, L. elegans; 5, L. girardi; 6, L. hendrichi, new species; 7, L. jaechi, new species; 8, L. javanicus; 9, L. komareki, new species. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385200" httpUri="https://zenodo.org/record/5385200/files/figure.png" pageId="5" pageNumber="314">Figs. 5</figureCitation>
,
<figureCitation id="13182A57FFA0FF99F4A69104FB7DB8C9" box="[1164,1192,893,917]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 1835. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="5" pageNumber="314">22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA0FF99F30791C5FA76B88E" box="[813,1443,956,978]" pageId="5" pageNumber="314" type="reference_group">
<paragraph id="8B9C36D2FFA0FF99F30791C5FA76B88E" blockId="5.[813,1443,956,978]" box="[813,1443,956,978]" pageId="5" pageNumber="314">
<treatmentCitationGroup id="AB3311FCFFA0FF99F30791C5FA76B88E" box="[813,1443,956,978]" pageId="5" pageNumber="314">
<treatmentCitation id="0A8210C3FFA0FF99F30791C5FB12B88E" author="Brancucci M" box="[813,1223,956,978]" page="256" pageId="5" pageNumber="314" year="1983">
<taxonomicName id="4C234D51FFA0FF99F30791C5FB12B88E" ID-CoL="3RM7J" authority="Brancucci, 1983: 256" authorityName="Brancucci" authorityPageNumber="256" authorityYear="1983" box="[813,1223,956,978]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="314" phylum="Arthropoda" rank="species" species="girardi">
<emphasis id="B957EAC0FFA0FF99F30791C5FC3BB88D" box="[813,1006,956,977]" italics="true" pageId="5" pageNumber="314">Laccophilus girardi</emphasis>
<bibRefCitation id="EFB24B23FFA0FF99F3DC91C5FB12B88E" author="Brancucci M" box="[1014,1223,956,978]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" page="256" pageId="5" pageNumber="314" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983: 256</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA0FF99F3079183FAF4BF8E" pageId="5" pageNumber="314" type="materials_examined">
<paragraph id="8B9C36D2FFA0FF99F3079183FCA8BF6E" blockId="5.[813,1451,1018,1074]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F3079183FC1CBF4E" bold="true" box="[813,969,1018,1042]" pageId="5" pageNumber="314">
<typeStatus id="54988870FFA0FF99F3079183FCB2BF4E" box="[813,871,1018,1042]" pageId="5" pageNumber="314">Type</typeStatus>
locality.
</emphasis>
<collectorName id="26D65304FFA0FF99F3FB9182FBEBBF4E" box="[977,1086,1019,1042]" pageId="5" pageNumber="314">N Borneo</collectorName>
, Kudat [=
<collectingCountry id="F3347642FFA0FF99F49F9183FACCBF4E" box="[1205,1305,1018,1042]" name="Malaysia" pageId="5" pageNumber="314">Malaysia</collectingCountry>
,
<collectingRegion id="49E7F830FFA0FF99F50C9183FA72BF4E" box="[1318,1447,1018,1042]" country="Malaysia" name="Sabah" pageId="5" pageNumber="314">Sabah State</collectingRegion>
,
<location id="8EFC6009FFA0FF99F3079663FCBBBF6E" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F3B49124FB5DBDEE:8EFC6009FFA0FF99F3079663FCBBBF6E" box="[813,878,1050,1074]" country="Malaysia" name="Kudat" pageId="5" pageNumber="314" stateProvince="Sabah">Kudat</location>
].
</paragraph>
<paragraph id="8B9C36D2FFA0FF99F3079623FAF4BF8E" blockId="5.[812,1452,1114,1234]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F3079623FBD3BF2E" bold="true" box="[813,1030,1114,1138]" pageId="5" pageNumber="314">Material examined.</emphasis>
<specimenCount id="9D25FD5BFFA0FF99F4209623FB4FBF2E" box="[1034,1178,1114,1138]" pageId="5" pageNumber="314" type="generic">15 specimens</specimenCount>
.
<materialsCitation id="3B4B3C8FFFA0FF99F4889623FC51BFEE" ID-GBIF-Occurrence="3760488327" collectingDate="1996-06-24" collectingDateMax="1996-07-01" collectingDateMin="1996-06-24" collectionCode="NHMW, NMPC" country="Malaysia" location="Sabah" pageId="5" pageNumber="314" specimenCount="14" stateProvince="Sabah">
<emphasis id="B957EAC0FFA0FF99F4889623FA50BF2E" bold="true" box="[1186,1413,1114,1138]" pageId="5" pageNumber="314">
<collectingCountry id="F3347642FFA0FF99F4889623FAE0BF2E" box="[1186,1333,1114,1138]" name="Malaysia" pageId="5" pageNumber="314">MALAYSIA</collectingCountry>
:
<collectingRegion id="49E7F830FFA0FF99F5159623FA50BF2E" box="[1343,1413,1114,1138]" country="Malaysia" name="Sabah" pageId="5" pageNumber="314">Sabah</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFA0FF99F5BB9623FC9BBFCE" pageId="5" pageNumber="314" type="generic">14 ex.</specimenCount>
,
<location id="8EFC6009FFA0FF99F37D9602FBFABFCE" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F3B49124FB5DBDEE:8EFC6009FFA0FF99F37D9602FBFABFCE" box="[855,1071,1146,1170]" country="Malaysia" name="Batu Punggul Resort" pageId="5" pageNumber="314" stateProvince="Sabah">Batu Punggul Resort</location>
env.,
<date id="FF9D1012FFA0FF99F4469603FA94BFCE" box="[1132,1345,1146,1170]" pageId="5" pageNumber="314" value="1996-06-24" valueMax="1996-07-01" valueMin="1996-06-24">
<collectingDate id="EFD9E9FAFFA0FF99F4469603FA94BFCE" box="[1132,1345,1146,1170]" pageId="5" pageNumber="314" value="1996-06-24" valueMax="1996-07-01" valueMin="1996-06-24">24 June1 July 1996</collectingDate>
</date>
(
<collectionCode id="ED32AE17FFA0FF99F5679603FA7DBFCE" box="[1357,1448,1146,1170]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="314" type="Museum">NHMW</collectionCode>
,
<collectionCode id="ED32AE17FFA0FF99F30696E3FCA8BFEE" box="[812,893,1178,1202]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="314">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFA0FF99F3A596E3FAC8BF8E" ID-GBIF-Occurrence="3760488319" collectingDate="1996-09-27" collectionCode="SMNS" collectorName="D. Grimm" country="Malaysia" elevation="1000" location="Sayap" municipality="Kinabalu National Park" pageId="5" pageNumber="314" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<specimenCount id="9D25FD5BFFA0FF99F3A596E3FC3ABFEE" box="[911,1007,1178,1202]" pageId="5" pageNumber="314" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6BF8ACA8FFA0FF99F3D096E3FB2FBFEE" box="[1018,1274,1178,1202]" pageId="5" pageNumber="314">Kinabalu National Park</collectingMunicipality>
,
<location id="8EFC6009FFA0FF99F52E96E3FA9DBFEE" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F3B49124FB5DBDEE:8EFC6009FFA0FF99F52E96E3FA9DBFEE" box="[1284,1352,1178,1202]" country="Malaysia" municipality="Kinabalu National Park" name="Sayap" pageId="5" pageNumber="314" stateProvince="Sabah">Sayap</location>
,
<quantity id="4CDB9B37FFA0FF99F57996E3FA72BFEE" box="[1363,1447,1178,1202]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="314" unit="m" value="1000.0">
<elevation id="000ED1E1FFA0FF99F57996E3FA72BFEE" box="[1363,1447,1178,1202]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="314" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FF9D1012FFA0FF99F30796C3FBD6BF8E" box="[813,1027,1210,1234]" pageId="5" pageNumber="314" value="1996-09-27">
<collectingDate id="EFD9E9FAFFA0FF99F30796C3FBD6BF8E" box="[813,1027,1210,1234]" pageId="5" pageNumber="314" value="1996-09-27">27 September 1996</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA0FF99F46D96C2FB63BF8E" box="[1095,1206,1210,1234]" pageId="5" pageNumber="314">D. Grimm</collectorName>
(
<collectionCode id="ED32AE17FFA0FF99F4ED96C3FAC0BF8E" box="[1223,1301,1210,1234]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="5" pageNumber="314">SMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA0FF99F3079683FC38BD0E" pageId="5" pageNumber="314" type="diagnosis">
<paragraph id="8B9C36D2FFA0FF99F3079683FBFDBEEE" blockId="5.[812,1452,1274,1458]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F3079683FC75BE4E" bold="true" box="[813,928,1274,1298]" pageId="5" pageNumber="314">Diagnosis.</emphasis>
Small, elongate oval species (body length
<quantity id="4CDB9B37FFA0FF99F5769683FC8EBE6E" metricMagnitude="-3" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="5" pageNumber="314" unit="mm" value="3.6" valueMax="3.7" valueMin="3.5">3.53.7 mm</quantity>
). Elytra with subbasal testaceous band very broad, covering at least basal third of elytra; or elytra almost uniformly testaceous (
<figureCitation id="13182A57FFA0FF99F4099723FBB2BE2E" box="[1059,1127,1370,1394]" captionStart="Figs" captionStartId="3.[113,157,2051,2073]" captionTargetBox="[115,1452,190,2035]" captionTargetId="figure-10@3.[97,1463,180,2039]" captionTargetPageId="3" captionText="Figs. 19. Habitus of Laccophilus: 1, L. aurofasciatus; 2, L. auropictus; 3, L. ceylonicus; 4, L. elegans; 5, L. girardi; 6, L. hendrichi, new species; 7, L. jaechi, new species; 8, L. javanicus; 9, L. komareki, new species. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385200" httpUri="https://zenodo.org/record/5385200/files/figure.png" pageId="5" pageNumber="314">Fig. 5</figureCitation>
). Apical part of median lobe in lateral view almost straight, narrowing to apex which is bent dorsally (
<figureCitation id="13182A57FFA0FF99F3E097E3FBC9BEEE" box="[970,1052,1434,1458]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 1835. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="5" pageNumber="314">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="8B9C36D2FFA0FF99F30797A3FC38BD0E" blockId="5.[813,1452,1498,1618]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F30797A3FC65BEAE" bold="true" box="[813,944,1498,1522]" pageId="5" pageNumber="314">Variability.</emphasis>
A species rather variable in elytral colouration: specimens from Batu Punggul are almost completely testaceous with only indistinct infuscation posteriorly to elytral midlength.
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA0FF99F3079403FB5DBDEE" pageId="5" pageNumber="314" type="distribution">
<paragraph id="8B9C36D2FFA0FF99F3079403FB5DBDEE" blockId="5.[813,1452,1658,1714]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F3079403FC6ABDCE" bold="true" box="[813,959,1658,1682]" pageId="5" pageNumber="314">Distribution.</emphasis>
<taxonomicName id="4C234D51FFA0FF99F3EE9403FB4DBDCE" authorityName="Brancucci" authorityYear="1983" box="[964,1176,1658,1682]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="314" phylum="Arthropoda" rank="species" species="girardi">
<emphasis id="B957EAC0FFA0FF99F3EE9403FB4DBDCE" box="[964,1176,1658,1682]" italics="true" pageId="5" pageNumber="314">Laccophilus girardi</emphasis>
</taxonomicName>
is so far known only from
<collectingRegion id="49E7F830FFA0FF99F30794E3FC7CBDEE" box="[813,937,1690,1714]" country="Malaysia" name="Sabah" pageId="5" pageNumber="314">Sabah state</collectingRegion>
,
<collectingCountry id="F3347642FFA0FF99F39D94E3FBCCBDEE" box="[951,1049,1690,1714]" name="Malaysia" pageId="5" pageNumber="314">Malaysia</collectingCountry>
(
<figureCitation id="13182A57FFA0FF99F40394E3FBA9BDEE" box="[1065,1148,1690,1714]" captionStart="Figs" captionStartId="9.[113,157,2093,2115]" captionTargetBox="[141,1427,190,2077]" captionTargetPageId="9" captionText="Figs. 4041. Distribution of the species of the Laccophilus javanicus group" figureDoi="http://doi.org/10.5281/zenodo.5385208" httpUri="https://zenodo.org/record/5385208/files/figure.png" pageId="5" pageNumber="314">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>