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<document id="24DF5D5C23250533A6090901FA68EAC9" ID-DOI="10.21248/contrib.entomol.67.2.325-352" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742448" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267809870" checkinUser="felipe" docAuthor="Zorn, Carsten, Kobayashi, Hirokazu &amp; Wada, Kaoru" docDate="2017" docId="03826012FFDDFFAB2B730371CED2F90B" docLanguage="en" docName="ContribEntomol.67.2.325-352.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Anomala herbacea Zorn &amp; Kobayashi &amp; Wada 2017, spec. nov." docType="treatment" docUuid="D39067BB-B013-4D67-9E01-BEA126CF5EA8" docUuidSource="ZooBank" docVersion="3" lastPageNumber="337" masterDocId="FFBB186AFFD7FFA7280F027BCF12FFB3" masterDocTitle="Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records" masterLastPageNumber="352" masterPageNumber="325" pageNumber="335" updateTime="1699436963292" updateUser="plazi">
<mods:mods id="3D21011DE60045DBE775B210809C3B05" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DFB96AB1D81E126DF8B91A2696C9D60F">
<mods:title id="91979CD6566DCC69CC1D80CD971DC53B">Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records</mods:title>
</mods:titleInfo>
<mods:name id="A64A66564D9FC493CCF751188B444464" type="personal">
<mods:role id="8CC1C227494F6993F21B81A911C97E0A">
<mods:roleTerm id="982BDFA0C6E3514BA1FC84E69D0CCB3F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="10B5E06DF2E0B4FCEE74D6ACC9792849">Zorn, Carsten</mods:namePart>
</mods:name>
<mods:name id="4E3FB2FB775D13B9C845E1F0CCED1DEC" type="personal">
<mods:role id="1D26B1984AC9F1C2ECBA0E0A8D8096B0">
<mods:roleTerm id="B3CAD78D7E3716BB7A1840A4C227F6B3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0D746ED0E2E13FDE49A106889BDD53BE">Kobayashi, Hirokazu</mods:namePart>
</mods:name>
<mods:name id="95AF122BFE95F6A26874E2CADA48DABA" type="personal">
<mods:role id="AD55C0F22F361F724A82F8F9DB0246CA">
<mods:roleTerm id="B8CA8A9B0D99DF629D5F646C270DCD8D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="152D09E0184FAB13F2BA13C73E0D0B57">Wada, Kaoru</mods:namePart>
</mods:name>
<mods:typeOfResource id="887D80326F32AC2AD4A0ED4BE346496B">text</mods:typeOfResource>
<mods:relatedItem id="7EFDE3AAF0C02E0037F95599FC91EF86" type="host">
<mods:titleInfo id="AE4E9AC25AA66D7FABD2EFEE27B81136">
<mods:title id="8504F28126F5FB4D1664940886E1CED3">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="23BE94DEBFE8DABA4AE6033CE911B125">
<mods:date id="7AF966894C47AB0945A8DB655FDC3DF5">2017</mods:date>
<mods:detail id="8F2E3C6DC6AD740E65FEA3B687D2C323" type="volume">
<mods:number id="D2ECCA774A58224D8FEF949E453EBCFB">67</mods:number>
</mods:detail>
<mods:detail id="FCFC0699804559EFB9BCE93D7B434BE3" type="issue">
<mods:number id="BED0FC05DFAD0B4F0BB41E284F76E49A">2</mods:number>
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<mods:extent id="45B6B40E053E1EF962E17E6AE06EC5A7" unit="page">
<mods:start id="7FA26B3EBC90A253B980C50AF201C3AD">325</mods:start>
<mods:end id="B279C28777C11946B25F6684F616ADBE">352</mods:end>
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<mods:classification id="1339FD97D930FCA746864AF60546E1EE">journal article</mods:classification>
<mods:identifier id="B06EA8BCA3BDF83A5EC95EFECEB9A9B5" type="DOI">10.21248/contrib.entomol.67.2.325-352</mods:identifier>
<mods:identifier id="31B731E4DE122E83F52B667642F69CE1" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="A46F199A35D1A3C4342BE995C1949938" type="Zenodo-Dep">5742448</mods:identifier>
<mods:identifier id="CB9E6219B7B6F4E2EC69EA2CDDF8D3F0" type="ZooBank">F2CE9642-9537-46FF-BA3F-5D4148AC1A0B</mods:identifier>
</mods:mods>
<treatment id="03826012FFDDFFAB2B730371CED2F90B" LSID="urn:lsid:zoobank.org:act:D39067BB-B013-4D67-9E01-BEA126CF5EA8" httpUri="http://treatment.plazi.org/id/03826012FFDDFFAB2B730371CED2F90B" lastPageId="12" lastPageNumber="337" pageId="10" pageNumber="335">
<subSubSection id="C331828FFFDDFFAD2B730371CA95FEF5" pageId="10" pageNumber="335" type="nomenclature">
<paragraph id="8B94D104FFDDFFAD2B730371CBBCFE96" blockId="10.[892,1415,266,358]" box="[892,1198,266,293]" pageId="10" pageNumber="335">
<heading id="D0DC6668FFDDFFAD2B730371CBBCFE96" box="[892,1198,266,293]" fontSize="12" level="2" pageId="10" pageNumber="335" reason="2">
<taxonomicName id="4C2BAA87FFDDFFAD2B730371CB2CFE96" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[892,1086,266,293]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="335" phylum="Arthropoda" rank="species" species="herbacea" status="sp. nov.">
<emphasis id="B95F0D16FFDDFFAD2B730371CB2CFE96" box="[892,1086,266,293]" italics="true" pageId="10" pageNumber="335">Anomala herbacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFDDFFAD2C4A0371CBBCFE96" box="[1093,1198,266,293]" pageId="10" pageNumber="335" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B94D104FFDDFFAD2B730349CA95FEF5" blockId="10.[892,1415,266,358]" box="[892,1415,306,326]" pageId="10" pageNumber="335">
<uri id="FFBADD06FFDDFFAD2B730349CA95FEF5" box="[892,1415,306,326]" pageId="10" pageNumber="335">
urn:lsid:zoobank.org:act:
<uuid id="FF8DEBD1FFDDFFAD2C230349CA95FEF5" box="[1068,1415,306,326]" pageId="10" pageNumber="335">D39067BB-B013-4D67-9E01-BEA126CF5EA8</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFDDFFAD2B730329CCDCFED5" box="[892,974,338,358]" pageId="10" pageNumber="335" type="description">
<paragraph id="8B94D104FFDDFFAD2B730329CCDCFED5" blockId="10.[892,1415,266,358]" box="[892,974,338,358]" pageId="10" pageNumber="335">
(
<figureCitation id="1310CD81FFDDFFAD2B8F0329CCBDFED5" box="[896,943,338,358]" captionStart="Figs 1–9" captionStartId="7.[174,214,2023,2045]" captionTargetBox="[214,1362,260,1985]" captionTargetId="figure-122@7.[203,1392,255,2011]" captionTargetPageId="7" captionText="Figs 1–9: Anomala spp., habitus – 1. A. okushimai spec. nov., holotype. 2. A. immeliorata spec. nov., holotype. 3. A. confrater spec. nov., holotype.4. A. imperspicabilis spec.nov., holotype.5. A. ordinata spec.nov., holotype. 6. A. recordata spec.nov., holotype. 7. A. asaitoae spec. nov., holotype. 8. A. asaitoae spec. nov., paratype (female). 9. A. herbacea spec. nov., holotype. Scale = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5742452" httpUri="https://zenodo.org/record/5742452/files/figure.png" pageId="10" pageNumber="335">Figs 9</figureCitation>
,
<figureCitation id="1310CD81FFDDFFAD2BB80329CCD8FED5" box="[951,970,338,358]" captionStart="Figs 14–17" captionStartId="11.[174,214,2052,2074]" captionTargetBox="[179,1404,258,2008]" captionTargetId="figure-99@11.[174,1421,251,2026]" captionTargetPageId="11" captionText="Figs 14–17: Anomala spp., aedeagus dorsal (a), ventral (b) and lateral (c). – 14. A. ordinata spec. nov., holotype. 15. A. recordata" figureDoi="http://doi.org/10.5281/zenodo.5742456" httpUri="https://zenodo.org/record/5742456/files/figure.png" pageId="10" pageNumber="335">17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFDDFFAD2B5903FBCB4AFDEB" pageId="10" pageNumber="335" type="materials_examined">
<paragraph id="8B94D104FFDDFFAD2B5903FBCB4AFDEB" blockId="10.[854,1459,383,600]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD2B5903FBCCC3FE24" bold="true" box="[854,977,383,408]" pageId="10" pageNumber="335">Type series</emphasis>
:
<materialsCitation id="3B43DB59FFDDFFAD2BD003FBCBF0FE0B" collectingDate="2010-05" collectionCode="NSMT" country="Vietnam" location="Ngoe Thinh" pageId="10" pageNumber="335" specimenCount="1" stateProvince="Kon Tum" typeStatus="holotype">
<emphasis id="B95F0D16FFDDFFAD2BD003FBCB57FE2B" bold="true" box="[991,1093,384,408]" pageId="10" pageNumber="335">
<typeStatus id="54906FA6FFDDFFAD2BD003FBCB57FE2B" box="[991,1093,384,408]" pageId="10" pageNumber="335" type="holotype">Holotype</typeStatus>
</emphasis>
: , C.
<collectingCountry id="F33C9194FFDDFFAD2C9903FBCA03FE2B" box="[1174,1297,384,408]" name="Vietnam" pageId="10" pageNumber="335">VIETNAM</collectingCountry>
:
<collectingRegion id="49EF1FE6FFDDFFAD2D1203FBCA96FE24" box="[1309,1412,384,407]" country="Vietnam" name="Kon Tum" pageId="10" pageNumber="335">Kon Tum</collectingRegion>
dist nr.
<location id="8EF487DFFFDDFFAD2B7603DBCCE9FE0B" LSID="urn:lsid:plazi:treatment:03826012FFDDFFAB2B730371CED2F90B:8EF487DFFFDDFFAD2B7603DBCCE9FE0B" box="[889,1019,416,440]" country="Vietnam" name="Ngoe Thinh" pageId="10" pageNumber="335" stateProvince="Kon Tum">Ngoe Thinh</location>
<date id="FF95F7C4FFDDFFAD2C0D03DBCB68FE0B" box="[1026,1146,416,440]" pageId="10" pageNumber="335" value="2010-05">
<collectingDate id="EFD10E2CFFDDFFAD2C0D03DBCB68FE0B" box="[1026,1146,416,440]" pageId="10" pageNumber="335" value="2010-05">MAY. 2010</collectingDate>
</date>
. (
<collectionCode id="ED3A49C1FFDDFFAD2C8303DBCBC9FE0B" box="[1164,1243,416,440]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="10" pageNumber="335">NSMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B43DB59FFDDFFAD2CE303DBCB46FDEB" collectingDate="2016-05-04" collectingDateMax="2016-05-07" collectingDateMin="2016-05-04" collectionCode="MFPC, MZUF" collectorName="L. Bartolozzi &amp; Bartolozzi, A &amp; Bandinelli, S &amp; Bambi, V" country="Vietnam" elevation="1250" latitude="14.672" location="Kon Plong" longLatPrecision="1" longitude="108.26382" pageId="10" pageNumber="335" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<typeStatus id="54906FA6FFDDFFAD2CE303DBCA44FE0B" box="[1260,1366,416,440]" pageId="10" pageNumber="335" type="paratype">Paratypes</typeStatus>
: 1 , the same data as for the holotype (
<collectionCode id="ED3A49C1FFDDFFAD2CB603BBCA17FE6B" box="[1209,1285,448,472]" pageId="10" pageNumber="335">MFPC</collectionCode>
); 1 , C. VIET- NAM:
<collectingRegion id="49EF1FE6FFDDFFAD2BAC039BCB65FE4B" box="[931,1143,480,504]" country="Vietnam" name="Kon Tum" pageId="10" pageNumber="335">Kon Tum Province</collectingRegion>
, ca.
<quantity id="4CD37CE1FFDDFFAD2CA3039BCBFDFE4B" box="[1196,1263,480,504]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="335" unit="km" value="30.0">30 km</quantity>
from
<location id="8EF487DFFFDDFFAD2D37039BCABDFE4B" LSID="urn:lsid:plazi:treatment:03826012FFDDFFAB2B730371CED2F90B:8EF487DFFFDDFFAD2D37039BCABDFE4B" box="[1336,1455,480,504]" latitude="14.672" longLatPrecision="1" longitude="108.26382" name="Kon Plong" pageId="10" pageNumber="335" stateProvince="Kon Tum">Kon Plong</location>
,
<quantity id="4CD37CE1FFDDFFAD2B59007BCCB7FDA4" box="[854,933,512,536]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="10" pageNumber="335" unit="m" value="1250.0">
<elevation id="00063637FFDDFFAD2B59007BCCB7FDA4" box="[854,933,512,536]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="10" pageNumber="335" unit="m" value="1250.0">1250 m</elevation>
</quantity>
<geoCoordinate id="EE1FB7C3FFDDFFAD2BB9007BCB21FDAB" box="[950,1075,512,536]" degrees="14" direction="north" minutes="40,320" orientation="latitude" pageId="10" pageNumber="335" precision="1" value="14.672">14°40,320N</geoCoordinate>
<geoCoordinate id="EE1FB7C3FFDDFFAD2C4C007BCBDAFDAB" box="[1091,1224,512,536]" degrees="108" direction="east" minutes="15,829" orientation="longitude" pageId="10" pageNumber="335" precision="1" value="108.26382">108°15,829E</geoCoordinate>
| leg.
<collectorName id="26DEB4D2FFDDFFAD2D29007BCABDFDAB" box="[1318,1455,512,536]" pageId="10" pageNumber="335">L. Bartolozzi</collectorName>
, A.
<collectorName id="26DEB4D2FFDDFFAD2B7A005BCB12FD8B" box="[885,1024,544,568]" pageId="10" pageNumber="335">Bandinelli, S</collectorName>
.
<collectorName id="26DEB4D2FFDDFFAD2C02005BCB66FD84" box="[1037,1140,544,568]" pageId="10" pageNumber="335">Bambi, V</collectorName>
. Sbordoni at light
<date id="FF95F7C4FFDDFFAD2D4D005BCAA1FD8B" box="[1346,1459,544,568]" pageId="10" pageNumber="335" value="2016-05-04" valueMax="2016-05-07" valueMin="2016-05-04">
<collectingDate id="EFD10E2CFFDDFFAD2D4D005BCAA1FD8B" box="[1346,1459,544,568]" pageId="10" pageNumber="335" value="2016-05-04" valueMax="2016-05-07" valueMin="2016-05-04">4-7.V.2016</collectingDate>
</date>
(n°Mag. 3078) (
<collectionCode id="ED3A49C1FFDDFFAD2BF2003BCB5FFDEB" box="[1021,1101,576,600]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="10" pageNumber="335">MZUF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFDDFFAB2B590004CC23FC24" lastPageId="12" lastPageNumber="337" pageId="10" pageNumber="335" type="description">
<paragraph id="8B94D104FFDDFFAD2B590004CA5FFD04" blockId="10.[854,1460,639,2104]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD2B590004CCCAFD24" bold="true" box="[854,984,639,663]" pageId="10" pageNumber="335">Description</emphasis>
: Body shape elongate ovoid, weakly convex; body length:
<quantity id="4CD37CE1FFDDFFAD2BED00DBCB6AFD04" box="[994,1144,672,696]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" metricValueMax="1.5699999999999998" metricValueMin="1.47" pageId="10" pageNumber="335" unit="mm" value="15.2" valueMax="15.7" valueMin="14.7">14.7-15.7 mm</quantity>
, width: 8.8-9.0 mm.
</paragraph>
<paragraph id="8B94D104FFDDFFAD2B5900C4CA7EFC2B" blockId="10.[854,1460,639,2104]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD2B5900C4CC87FD64" bold="true" box="[854,917,703,727]" pageId="10" pageNumber="335">Color</emphasis>
. Dorsal surface light green with metallic shine, elytra with brown suffusions; pronotum with narrow yellow band along lateral margins; front of clypeus and antennae yellowish brown; ventral surface and legs yellowish brown with weak metallic reflections except for mesotarsi, metatibia and -tarsi (greenish to reddish copper) and abdominal ventrites (reddish brown).
</paragraph>
<paragraph id="8B94D104FFDDFFAD2B5901DBCA24FB4B" blockId="10.[854,1460,639,2104]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD2B5901DBCC82FC0B" bold="true" box="[854,912,928,952]" pageId="10" pageNumber="335">Head</emphasis>
with erect yellowish brown setae (
<quantity id="4CD37CE1FFDDFFAD2D0F01DBCA85FC04" box="[1280,1431,928,952]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" metricValueMax="3.8" metricValueMin="0.7" pageId="10" pageNumber="335" unit="mm" value="0.225" valueMax="0.38" valueMin="0.07">0.07-0.38 mm</quantity>
in length) becoming sparser and shorter anteriad, longer and denser latero-posteriad; interocular distance equals 0.65 times the maximum transverse head width. Antennal club 0.9 times as long as footstalk. Clypeus semicircular, finely and densely rugoso-punctate; anterior margin distinctly reflexed; fronto-clypeal suture almost straight. Frons densely and coarsely rugoso-punctate anteriorly and in the middle, punctures becoming sparser and smaller in postero-lateral portions. Vertex coarsely punctate, the punctures somewhat sparser laterad.
</paragraph>
<paragraph id="8B94D104FFDDFFAD2B59077BCA95F9AB" blockId="10.[854,1460,639,2104]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD2B59077BCCD5FAAB" bold="true" box="[854,967,1280,1304]" pageId="10" pageNumber="335">Pronotum</emphasis>
1.6 times as wide as long; sides arcuate in the middle, subparallel basad, nearly straight and strongly convergent anteriorly; widest at base; lateral marginal line almost reaching the hind angles; basal marginal line missing; front angles produced and acute, hind angles obtuse, slightly rounded off; surface evenly covered with moderately large, moderately dense, round to weakly transverse punctures; punctures smaller laterad; each puncture bearing an erect yellow seta (
<quantity id="4CD37CE1FFDDFFAD2C87047BCA00F9A4" box="[1160,1298,1536,1560]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="10" pageNumber="335" unit="mm" value="0.275" valueMax="0.4" valueMin="0.15">0.15-0.4 mm</quantity>
in length).
</paragraph>
<paragraph id="8B94D104FFDDFFAD2B590464CBC5F9E4" blockId="10.[854,1460,639,2104]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD2B590464CCD7F984" bold="true" box="[854,965,1567,1591]" pageId="10" pageNumber="335">Scutellum</emphasis>
1.6 times as wide as long; apex rounded; punctures like those of disc of pronotum.
</paragraph>
<paragraph id="8B94D104FFDDFFAD2B59041BCCD1F824" blockId="10.[854,1460,639,2104]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD2B59041BCC8BF9CB" bold="true" box="[854,921,1632,1656]" pageId="10" pageNumber="335">Elytra</emphasis>
. Regularly striate-sulcate; intervals weakly convex; each stria marked by a regular line of almost round, hairless punctures; entire surface covered with transverse, seta-bearing (
<quantity id="4CD37CE1FFDDFFAD2C2004BBCBD4F964" box="[1071,1222,1728,1752]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="10" pageNumber="335" unit="mm" value="0.1" valueMax="0.15" valueMin="0.05">0.05-0.15 mm</quantity>
in length) punctures; this irregular punctation sparser on two inner primary costae as well as on the two secondary costae formed within the subsutural interstice; epipleura disappearing shortly before apical curvature which is fringed by thick erect setae; marginal membrane starting near middle of metacoxa.
</paragraph>
<paragraph id="8B94D104FFDDFFAD2B5905E4CB08F78B" blockId="10.[854,1460,639,2104]" pageId="10" pageNumber="335">
<emphasis id="B95F0D16FFDDFFAD2B5905E4CCD0F804" bold="true" box="[854,962,1951,1975]" pageId="10" pageNumber="335">Pygidium</emphasis>
triangular, widely rounded at apex; weakly tumid; disc with reticulate striolation which is arranged concentrically around the tumidity; with rather dense yellow setosity (
<quantity id="4CD37CE1FFDDFFAD2C0B0A7BCB9CF7A4" box="[1028,1166,2048,2072]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="7.5" metricValueMin="1.0" pageId="10" pageNumber="335" unit="mm" value="0.425" valueMax="0.75" valueMin="0.1">0.1-0.75 mm</quantity>
in length), setae longer in the apical portion.
</paragraph>
<caption id="DF54818CFFDCFFAC28A10A7FCA9FF7AA" ID-DOI="http://doi.org/10.5281/zenodo.5742456" ID-Zenodo-Dep="5742456" box="[174,1421,2052,2074]" httpUri="https://zenodo.org/record/5742456/files/figure.png" pageId="11" pageNumber="336" startId="11.[174,214,2052,2074]" targetBox="[179,1404,258,2008]" targetPageId="11">
<paragraph id="8B94D104FFDCFFAC28A10A7FCA9FF7AA" blockId="11.[174,1421,2052,2103]" box="[174,1421,2052,2074]" pageId="11" pageNumber="336">
<emphasis id="B95F0D16FFDCFFAC28A10A7FCE0BF7AA" bold="true" box="[174,281,2052,2074]" pageId="11" pageNumber="336">Figs 14–17</emphasis>
:
<taxonomicName id="4C2BAA87FFDCFFAC29290A7FCEBEF7AA" box="[294,428,2052,2073]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="336" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95F0D16FFDCFFAC29290A7FCE6FF7AA" box="[294,381,2052,2073]" italics="true" pageId="11" pageNumber="336">Anomala</emphasis>
spp.
</taxonomicName>
, aedeagus dorsal (
<emphasis id="B95F0D16FFDCFFAC2A500A7FCD78F7AA" bold="true" box="[607,618,2052,2073]" pageId="11" pageNumber="336">a</emphasis>
), ventral (
<emphasis id="B95F0D16FFDCFFAC2ADE0A7FCDCCF7AA" bold="true" box="[721,734,2052,2073]" pageId="11" pageNumber="336">b</emphasis>
) and lateral (
<emphasis id="B95F0D16FFDCFFAC2B6C0A7FCC7FF7AA" bold="true" box="[867,877,2052,2073]" pageId="11" pageNumber="336">c</emphasis>
). –
<emphasis id="B95F0D16FFDCFFAC2B9A0A7FCCBFF7AA" bold="true" box="[917,941,2052,2073]" pageId="11" pageNumber="336">14</emphasis>
.
<taxonomicName id="4C2BAA87FFDCFFAC2BB60A7FCB37F7AA" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[953,1061,2052,2073]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="336" phylum="Arthropoda" rank="species" species="ordinata" status="sp. nov.">
<emphasis id="B95F0D16FFDCFFAC2BB60A7FCB37F7AA" box="[953,1061,2052,2073]" italics="true" pageId="11" pageNumber="336">A. ordinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFDCFFAC2C230A7FCB9FF7AA" box="[1068,1165,2052,2073]" pageId="11" pageNumber="336" rank="species">spec. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="B95F0D16FFDCFFAC2CF80A7FCA1DF7AA" bold="true" box="[1271,1295,2052,2073]" pageId="11" pageNumber="336">15</emphasis>
.
<taxonomicName id="4C2BAA87FFDCFFAC2D150A7FCA9FF7AA" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[1306,1421,2052,2073]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="336" phylum="Arthropoda" rank="species" species="recordata" status="sp. nov.">A. recordata</taxonomicName>
</paragraph>
</caption>
<paragraph id="8B94D104FFDCFFAC28A10A59CBA3F784" blockId="11.[174,1421,2052,2103]" box="[174,1201,2081,2103]" pageId="11" pageNumber="336">
spec. nov.,
<typeStatus id="54906FA6FFDCFFAC29190A5ACE7FF784" box="[278,365,2081,2103]" pageId="11" pageNumber="336" type="holotype">holotype</typeStatus>
.
<emphasis id="B95F0D16FFDCFFAC297A0A5ACE9FF785" bold="true" box="[373,397,2081,2102]" pageId="11" pageNumber="336">16</emphasis>
.
<taxonomicName id="4C2BAA87FFDCFFAC29980A59CEEDF784" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[407,511,2082,2103]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="336" phylum="Arthropoda" rank="species" species="asaitoae" status="sp. nov.">
<emphasis id="B95F0D16FFDCFFAC29980A59CEEDF784" box="[407,511,2082,2103]" italics="true" pageId="11" pageNumber="336">A. asaitoae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFDCFFAC2A0A0A59CD76F784" box="[517,612,2082,2103]" pageId="11" pageNumber="336" rank="species">spec. nov.</taxonomicNameLabel>
,
<typeStatus id="54906FA6FFDCFFAC2A630A5ACDD1F784" box="[620,707,2081,2103]" pageId="11" pageNumber="336" type="holotype">holotype</typeStatus>
.
<emphasis id="B95F0D16FFDCFFAC2AC40A5ACDF1F785" bold="true" box="[715,739,2081,2102]" pageId="11" pageNumber="336">17</emphasis>
.
<taxonomicName id="4C2BAA87FFDCFFAC2AE10A59CC4FF785" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[750,861,2081,2103]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="336" phylum="Arthropoda" rank="species" species="herbacea" status="sp. nov.">
<emphasis id="B95F0D16FFDCFFAC2AE10A59CC4FF785" box="[750,861,2081,2103]" italics="true" pageId="11" pageNumber="336">A. herbacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFDCFFAC2B6D0A59CCD3F784" box="[866,961,2082,2103]" pageId="11" pageNumber="336" rank="species">spec. nov.</taxonomicNameLabel>
,
<typeStatus id="54906FA6FFDCFFAC2BC60A5ACB32F784" box="[969,1056,2081,2103]" pageId="11" pageNumber="336" type="holotype">holotype</typeStatus>
. Scale =
<quantity id="4CD37CE1FFDCFFAC2C7D0A5ACBBFF784" box="[1138,1197,2081,2103]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="336" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8B94D104FFDBFFAB28DB0284CD62FE84" blockId="12.[212,817,255,920]" pageId="12" pageNumber="337">
<emphasis id="B95F0D16FFDBFFAB28DB0284CE71FEA4" bold="true" box="[212,355,255,279]" pageId="12" pageNumber="337">Mesoventrite</emphasis>
very narrow and somewhat concave between the approximated mesocoxae.
</paragraph>
<paragraph id="8B94D104FFDBFFAB28DB0344CD87FE6B" blockId="12.[212,817,255,920]" pageId="12" pageNumber="337">
<emphasis id="B95F0D16FFDBFFAB28DB0344CE72FEE4" bold="true" box="[212,352,319,343]" pageId="12" pageNumber="337">Metaventrite</emphasis>
with very dense, partly transversely confluent punctures at the sides, sparsely punctate, almost smooth in the middle; with dense, long, pale yellowish brown setae (0.13-1.0 mm in length) at the sides which become sparser and shorter in the middle.
</paragraph>
<paragraph id="8B94D104FFDBFFAB28DB03A4CE3DFD2B" blockId="12.[212,817,255,920]" pageId="12" pageNumber="337">
<emphasis id="B95F0D16FFDBFFAB28DB03A4CEA6FE44" bold="true" box="[212,436,479,503]" pageId="12" pageNumber="337">Abdominal ventrites</emphasis>
with rather sparse, transverse punctures (somewhat denser at the sides); each ventrite with a transverse row of semierect yellow setae in the apical half (
<quantity id="4CD37CE1FFDBFFAB28D4003BCE63FDE4" box="[219,369,576,600]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" metricValueMax="13.0" metricValueMin="5.0" pageId="12" pageNumber="337" unit="mm" value="0.09" valueMax="0.13" valueMin="0.05">0.05-0.13 mm</quantity>
in length) and denser adpressed setosity laterally; sides of abdominal ventrites 1-4 distinctly carinate.
</paragraph>
<paragraph id="8B94D104FFDBFFAB28DB00DBCC23FC24" blockId="12.[212,817,255,920]" pageId="12" pageNumber="337">
<emphasis id="B95F0D16FFDBFFAB28DB00DBCE14FD0B" bold="true" box="[212,262,672,696]" pageId="12" pageNumber="337">Legs</emphasis>
slender and long; protibia bidentate; terminal tooth rather short, somewhat obtuse; protarsal segment 5 ventrally with a sharp tooth in middle; inner protarsal claw and outer mesotarsal claw apically incised; the upper branch of the inner protarsal claw small and slender, the lower branch broad, somewhat enlarged ventrally with obtuse tooth in the middle; the upper branch of outer mesotarsal claw slightly more slender than the lower one.
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFDBFFAB28DB01C4CEB5F9AB" pageId="12" pageNumber="337" type="diagnosis">
<paragraph id="8B94D104FFDBFFAB28DB01C4CEB5F9AB" blockId="12.[212,817,959,1560]" pageId="12" pageNumber="337">
<emphasis id="B95F0D16FFDBFFAB28DB01C4CE53FC64" bold="true" box="[212,321,959,983]" pageId="12" pageNumber="337">Diagnosis</emphasis>
: This species is easily distinguished from all other known
<taxonomicName id="4C2BAA87FFDBFFAB2963019BCEDEFC4B" authorityName="SAMOUELLE. It" authorityYear="1819" box="[364,460,992,1016]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="B95F0D16FFDBFFAB2963019BCEDEFC4B" box="[364,460,992,1016]" italics="true" pageId="12" pageNumber="337">Anomala</emphasis>
</taxonomicName>
by the combination of a setose dorsal surface, a light metallic green dorsum and sulcate elytra. Furthermore, the shape of the aedeagus with its long, bifid parameres is unique (fig. 17). No described species seems to be closely related. Several species with dorsal setosity occur in
<collectingCountry id="F33C9194FFDBFFAB29DC06FBCD06FB2B" box="[467,532,1152,1176]" name="China" pageId="12" pageNumber="337">China</collectingCountry>
and Indo-China and were subsumed in the “
<taxonomicName id="4C2BAA87FFDBFFAB299A06DBCEE1FB0B" authorityName="Nonfried" authorityYear="1892" box="[405,499,1184,1208]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="hirsutula">
<emphasis id="B95F0D16FFDBFFAB299A06DBCEE1FB0B" box="[405,499,1184,1208]" italics="true" pageId="12" pageNumber="337">hirsutula</emphasis>
</taxonomicName>
-group” (LIN, 1996).
<taxonomicName id="4C2BAA87FFDBFFAB2ADF06DBCE23FB6B" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="herbacea">
<emphasis id="B95F0D16FFDBFFAB2ADF06DBCE23FB6B" italics="true" pageId="12" pageNumber="337">Anomala herbacea</emphasis>
</taxonomicName>
, however, does not fit in this group because the dorsal setosity is rather semi-erect to erect (adpressed to semi-erect in species of the
<taxonomicName id="4C2BAA87FFDBFFAB2A07077BCD74FAAB" authorityName="Nonfried" authorityYear="1892" box="[520,614,1280,1304]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="hirsutula">
<emphasis id="B95F0D16FFDBFFAB2A07077BCD74FAAB" box="[520,614,1280,1304]" italics="true" pageId="12" pageNumber="337">hirsutula</emphasis>
</taxonomicName>
-group), the punctures of pronotum and elytra are less transverse, the propygidium is not exposed in
<taxonomicName id="4C2BAA87FFDBFFAB2A11073BCD8BFAEB" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[542,665,1344,1368]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="herbacea">
<emphasis id="B95F0D16FFDBFFAB2A11073BCD8BFAEB" box="[542,665,1344,1368]" italics="true" pageId="12" pageNumber="337">A. herbacea</emphasis>
</taxonomicName>
, and the body is more depressed and broader than in all species of the
<taxonomicName id="4C2BAA87FFDBFFAB28DB07FBCE20FA2B" authorityName="Nonfried" authorityYear="1892" box="[212,306,1408,1432]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="337" phylum="Arthropoda" rank="species" species="hirsutula">
<emphasis id="B95F0D16FFDBFFAB28DB07FBCE20FA2B" box="[212,306,1408,1432]" italics="true" pageId="12" pageNumber="337">hirsutula</emphasis>
</taxonomicName>
-group. In general, the dorsal setosity is a quantitative rather than a qualitative character in
<taxonomicName id="4C2BAA87FFDBFFAB2AC507DBCC38FA0B" authorityName="SAMOUELLE. It" authorityYear="1819" box="[714,810,1440,1464]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="B95F0D16FFDBFFAB2AC507DBCC38FA0B" box="[714,810,1440,1464]" italics="true" pageId="12" pageNumber="337">Anomala</emphasis>
</taxonomicName>
, because seta-bearing punctures on pronotum and elytra might be observed in many known species depending on magnification used.
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFDBFFAB28DB043BCED2F90B" pageId="12" pageNumber="337" type="etymology">
<paragraph id="8B94D104FFDBFFAB28DB043BCED2F90B" blockId="12.[212,817,1600,1720]" pageId="12" pageNumber="337">
<emphasis id="B95F0D16FFDBFFAB28DB043BCE59F9EB" bold="true" box="[212,331,1600,1624]" pageId="12" pageNumber="337">Etymology</emphasis>
: The species epithet is formed by the Latin adjective “herbaceus, -a, -um”. It translates as “grassy” or “grass-colored”. The name refers to the green, hairy dorsal surface of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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