232 lines
28 KiB
XML
232 lines
28 KiB
XML
<document id="7EAD3E29F1550C6947846E94FA233493" ID-DOI="10.1080/00222933.2012.708452" ID-ISSN="1464-5262" ID-Zenodo-Dep="4590828" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615293597861" checkinUser="carolina" docAuthor="Kramer, Bernd, Bills, Roger, Skelton, Paul & Wink, Michael" docDate="2012" docId="038087B4FFDCFFADA382FE4C627F3019" docLanguage="en" docName="JNATHIST.46.35-36.2179-2258.pdf" docOrigin="Journal of Natural History 46 (35 - 36)" docSource="http://dx.doi.org/10.1080/00222933.2012.708452" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Petrocephalus longianalis Kramer & Bills & Skelton & Wink 2012, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2238" masterDocId="FFB9FFCCFFE0FF90A279FFDA6003327D" masterDocTitle="A critical revision of the churchill snoutfish, genus Petrocephalus Marcusen, 1854 (Actinopterygii: Teleostei: Mormyridae), from southern and eastern Africa, with the recognition of Petrocephalus tanensis, and the description of five new species" masterLastPageNumber="2258" masterPageNumber="2179" pageNumber="2237" updateTime="1698882054463" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1B8732E80CD80B8A37B3E90754308014">A critical revision of the churchill snoutfish, genus Petrocephalus Marcusen, 1854 (Actinopterygii: Teleostei: Mormyridae), from southern and eastern Africa, with the recognition of Petrocephalus tanensis, and the description of five new species</mods:title>
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<mods:namePart id="89C9B20BD1C3F30EB81165179AB9B700">Kramer, Bernd</mods:namePart>
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<mods:namePart id="E9D4419814DE1CE25C9CCAC59BB9FD20">Bills, Roger</mods:namePart>
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<mods:roleTerm id="F77346BA0BB110A7D470D96B8F480D73">Author</mods:roleTerm>
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<mods:namePart id="999910EABBED0BB7B513EA62A9D0BC2D">Skelton, Paul</mods:namePart>
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<mods:roleTerm id="719E1223C02FCC0BF1DA19FCF3DF3DB1">Author</mods:roleTerm>
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<mods:namePart id="1C4043D1AA9BB652DC877DD3072E603C">Wink, Michael</mods:namePart>
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<mods:title id="0DB5A05A0A106020DB8AB070CCCDE76D">Journal of Natural History</mods:title>
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<mods:part id="750E7E7B36E1A88261D96EC578272084">
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<mods:date id="D6E9EFB58F0E12ABFE5197A5DEB0DFA0">2012</mods:date>
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<mods:number id="A331458C658D22C667533FE8F6AF329A">2012-09-30</mods:number>
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<mods:detail id="B0D591B811DC046AD8DEB8096104C8EF" type="volume">
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<mods:number id="967F75BDEE0B4258CFF1F4EBD9F88869">46</mods:number>
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<mods:number id="38C4EB2D1C02A8B17DEFB6211513970E">35 - 36</mods:number>
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<mods:start id="560DAA41BB1AF003A1722AE82F7A0B95">2179</mods:start>
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<mods:url id="62C869CDC92BA3EA6A00B0A615BCCD51">http://dx.doi.org/10.1080/00222933.2012.708452</mods:url>
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<mods:identifier id="356FDA23E350F79FB6966FD0FF42EB91" type="DOI">10.1080/00222933.2012.708452</mods:identifier>
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<subSubSection id="C3336529FFDCFFACA382FE4C636E33D3" box="[507,877,406,431]" pageId="60" pageNumber="2237" type="nomenclature">
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<paragraph id="8B9636A2FFDCFFACA382FE4C636E33D3" blockId="60.[507,877,406,462]" box="[507,877,406,431]" pageId="60" pageNumber="2237">
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<heading id="D0DE81CEFFDCFFACA382FE4C636E33D3" box="[507,877,406,431]" centered="true" fontSize="36" level="2" pageId="60" pageNumber="2237" reason="7">
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<taxonomicName id="4C294D21FFDCFFACA382FE4C631133D2" ID-CoL="4FDXH" authority="Kramer & Bills & Skelton & Wink, 2012" authorityName="Kramer & Bills & Skelton & Wink" authorityYear="2012" box="[507,786,406,431]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="60" pageNumber="2237" phylum="Chordata" rank="species" species="longianalis" status="sp. nov.">
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<emphasis id="B95DEAB0FFDCFFACA382FE4C631133D2" bold="true" box="[507,786,406,431]" italics="true" pageId="60" pageNumber="2237">Petrocephalus longianalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26E57CBFFDCFFACA161FE4D636E33D3" box="[792,877,407,430]" pageId="60" pageNumber="2237" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C3336529FFDCFFADA02BFE6C627F3019" lastPageId="61" lastPageNumber="2238" pageId="60" pageNumber="2237" type="description">
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<paragraph id="8B9636A2FFDCFFACA02BFE6C631733B3" blockId="60.[507,877,406,462]" box="[594,788,438,462]" pageId="60" pageNumber="2237">
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(
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<figureCitation id="13122A27FFDCFFACA023FE6C62BF33B3" box="[602,700,438,462]" captionStart="Figure 2" captionStartId="5.[143,210,1254,1276]" captionText="Figure 2(A–D). Photographs of members of southern and eastern African Petrocephalus species studied (numbers refer to localities given in Figures 1, 6 and 9). (1) P. catostoma (Günther, 1866), Lectotype, BMNH 1863.10.12.4, right side, SL 4.7 cm. (2) P. stuhlmanni Boulenger, 1909, holotype, BMNH 1907.12.3.1, SL 7.9 cm. (3) P. wesselsi Kramer and Van der Bank, 2000, right side, SL 9.7 cm, SAIAB 85922 (R1). (7) P. longicapitis sp. nov., SMF 28265 (R1), SL 9 cm. (9) P. okavangensis sp. nov., holotype, SAIAB 030046, right side, SL 6.1 cm. (10) P. catostoma tanensis Whitehead and Greenwood, 1959, holotype, BMNH 1963.11.29.1, right side, SL 6.7 cm. (10a) P. tanensis, field no. Ta05na, SL 8.7 cm, SAIAB 85907. (12) P. longianalis sp. nov., holotype, SL 8.2 cm, right side, SAIAB 76758. (21) P. petersi sp. nov., holotype, SAIAB 060846, right side, SL 6.4 cm. (23) P. magnoculis sp. nov., SL 8.9 cm, ZSM 38659. (24) P. haullevillii Boulenger, 1912, BMNH 1912.4.1.186–188 (R1), right side, SL 5.7 cm. (25) P. steindachneri Fowler 1958, syntype, NMW 55118:3, right side, SL 6.4 cm. (26) P. stuhlmanni congicus David and Poll, 1937, syntype, MRAC 30807-30808, SL 7.8 cm. (27) P. degeni Boulenger, 1906, BMNH 1906.5.30.84, right side, SL 8.12 cm. (28) P. catostoma, SAIAB 73894 (R1), right side, SL 6.4 cm. (31) P. magnitrunci sp. nov., SL 8.5 cm, SAIAB 67069 (R5, right side), see map Figure 6. Scale bar, 1 cm." pageId="60" pageNumber="2237">Figure 2</figureCitation>
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, no. 12)
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</paragraph>
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<paragraph id="8B9636A2FFDCFFACA2AAFE2D61253033" blockId="60.[211,1172,502,590]" pageId="60" pageNumber="2237">
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<emphasis id="B95DEAB0FFDCFFACA2AAFE2D618F3073" box="[211,396,503,526]" italics="true" pageId="60" pageNumber="2237">Type specimens.</emphasis>
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<materialsCitation id="3B413CFFFFDCFFACA3EDFE2C62EF3053" collectionCode="SAIAB, R" country="Zambia" county="Province" location="Luongo River" municipality="System" pageId="60" pageNumber="2237" specimenCode="SAIAB 76758, R9" specimenCount="1" stateProvince="Luapula" typeStatus="holotype">
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<typeStatus id="54928800FFDCFFACA3EDFE2C62003073" box="[404,515,502,526]" pageId="60" pageNumber="2237" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="DB8F9ED9FFDCFFACA06DFE2C62BB3073" box="[532,696,502,526]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="60" pageNumber="2237" type="Other type of research institution/biorepository">SAIAB 76758</specimenCode>
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(specimen
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<specimenCode id="DB8F9ED9FFDCFFACA13FFE2C636B3073" box="[838,872,502,526]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="60" pageNumber="2237">R9</specimenCode>
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) from
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<collectingCountry id="F33E7632FFDCFFACA1BDFE2C641F3073" box="[964,1052,502,526]" name="Zambia" pageId="60" pageNumber="2237">Zambia</collectingCountry>
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<collectingCounty id="62F74E2EFFDCFFACA652FE2C64923073" box="[1067,1169,502,526]" pageId="60" pageNumber="2237">Province</collectingCounty>
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:
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<collectingRegion id="49EDF840FFDCFFACA2AAFDCC61313053" box="[211,306,534,558]" country="Zambia" name="Luapula" pageId="60" pageNumber="2237">Luapula</collectingRegion>
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<collectingMunicipality id="6BF2ACD8FFDCFFACA345FDCC61933053" box="[316,400,534,558]" pageId="60" pageNumber="2237">System</collectingMunicipality>
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:
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<location id="8EF66079FFDCFFACA3E4FDCD62413053" LSID="urn:lsid:plazi:treatment:038087B4FFDCFFADA382FE4C627F3019:8EF66079FFDCFFACA3E4FDCD62413053" box="[413,578,534,559]" country="Zambia" county="Province" municipality="System" name="Luongo River" pageId="60" pageNumber="2237" stateProvince="Luapula">Luongo River</location>
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:
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<location id="8EF66079FFDCFFACA037FDCC62EF3053" LSID="urn:lsid:plazi:treatment:038087B4FFDCFFADA382FE4C627F3019:8EF66079FFDCFFACA037FDCC62EF3053" box="[590,748,534,558]" country="Zambia" county="Province" municipality="System" name="Lufubu River" pageId="60" pageNumber="2237" stateProvince="Luapula">Lufubu River</location>
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</materialsCitation>
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.
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<materialsCitation id="3B413CFFFFDCFFACA080FDCD61213033" collectionCode="SAIAB" country="Zambia" pageId="60" pageNumber="2237" specimenCode="SAIAB 186060" specimenCount="48" stateProvince="Luapula" typeStatus="paratype">
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<typeStatus id="54928800FFDCFFACA080FDCD63683052" box="[761,875,535,559]" pageId="60" pageNumber="2237" type="paratype">Paratypes</typeStatus>
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:
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<specimenCode id="DB8F9ED9FFDCFFACA101FDCC64273053" box="[888,1060,534,558]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="60" pageNumber="2237" type="Other type of research institution/biorepository">SAIAB 186060</specimenCode>
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(
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<specimenCount id="9D2FFD2BFFDCFFACA64CFDCC611A3033" pageId="60" pageNumber="2237" type="generic">48 specimens</specimenCount>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B9636A2FFDCFFACA2AAFDA161DB30AC" blockId="60.[211,1172,634,721]" pageId="60" pageNumber="2237">
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<materialsCitation id="3B413CFFFFDCFFACA2AAFDA161D730AC" country="Zambia" county="Province" location="Luongo River" municipality="System" pageId="60" pageNumber="2237" specimenCount="1" stateProvince="Luapula" typeStatus="holotype">
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<typeStatus id="54928800FFDCFFACA2AAFDA1610A30EF" box="[211,265,635,658]" pageId="60" pageNumber="2237">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F33E7632FFDCFFACA30CFDA061D330EF" box="[373,464,634,658]" name="Zambia" pageId="60" pageNumber="2237">Zambia</collectingCountry>
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,
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<collectingCountry id="F33E7632FFDCFFACA398FDA0623A30EF" box="[481,569,634,658]" name="Zambia" pageId="60" pageNumber="2237">Zambia</collectingCountry>
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<collectingCounty id="62F74E2EFFDCFFACA03CFDA062AB30EF" box="[581,680,634,658]" pageId="60" pageNumber="2237">Province</collectingCounty>
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,
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<collectingRegion id="49EDF840FFDCFFACA0C0FDA0631B30EF" box="[697,792,634,658]" country="Zambia" name="Luapula" pageId="60" pageNumber="2237">Luapula</collectingRegion>
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<collectingMunicipality id="6BF2ACD8FFDCFFACA15CFDA0637A30EF" box="[805,889,634,658]" pageId="60" pageNumber="2237">System</collectingMunicipality>
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,
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<location id="8EF66079FFDCFFACA1F0FDA0642D30EF" LSID="urn:lsid:plazi:treatment:038087B4FFDCFFADA382FE4C627F3019:8EF66079FFDCFFACA1F0FDA0642D30EF" box="[905,1070,634,658]" country="Zambia" county="Province" municipality="System" name="Luongo River" pageId="60" pageNumber="2237" stateProvince="Luapula">Luongo River</location>
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,
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<location id="8EF66079FFDCFFACA646FDA0611130CC" LSID="urn:lsid:plazi:treatment:038087B4FFDCFFADA382FE4C627F3019:8EF66079FFDCFFACA646FDA0611130CC" country="Zambia" county="Province" municipality="System" name="Lufubu River" pageId="60" pageNumber="2237" stateProvince="Luapula">Lufubu River</location>
|
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,
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<location id="8EF66079FFDCFFACA351FD43621730CC" LSID="urn:lsid:plazi:treatment:038087B4FFDCFFADA382FE4C627F3019:8EF66079FFDCFFACA351FD43621730CC" box="[296,532,665,689]" country="Zambia" county="Province" municipality="System" name="Lufubu River Falls" pageId="60" pageNumber="2237" stateProvince="Luapula">Lufubu River Falls</location>
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below bridge at
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||
<location id="8EF66079FFDCFFACA085FD43644E30CF" LSID="urn:lsid:plazi:treatment:038087B4FFDCFFADA382FE4C627F3019:8EF66079FFDCFFACA085FD43644E30CF" box="[764,1101,665,690]" country="Zambia" county="Province" municipality="System" name="Chipili on Mensa-Mununga" pageId="60" pageNumber="2237" stateProvince="Luapula">Chipili on Mensa-Mununga</location>
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road 10.7297
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<superScript id="7C5C9BEAFFDCFFACA353FD6F613130BB" attach="left" box="[298,306,693,710]" fontSize="7" pageId="60" pageNumber="2237">◦</superScript>
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S, 29.0936
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<superScript id="7C5C9BEAFFDCFFACA3D7FD6F61B530BB" attach="left" box="[430,438,693,710]" fontSize="7" pageId="60" pageNumber="2237">◦</superScript>
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E
|
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B9636A2FFDCFFACA2AAFD24640731A9" blockId="60.[211,1172,765,980]" pageId="60" pageNumber="2237">
|
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<emphasis id="B95DEAB0FFDCFFACA2AAFD2461443168" box="[211,327,766,789]" italics="true" pageId="60" pageNumber="2237">Diagnosis.</emphasis>
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||
Number of anal fin rays, nA, median 33 (range 30–35); anal fin length, LA, mean 0.261 (range
|
||
<geoCoordinate id="EE1D5065FFDCFFACA39DFCC762223148" box="[484,545,797,821]" degrees="0.244" direction="north" orientation="latitude" pageId="60" pageNumber="2237" precision="55" value="0.244">0.244</geoCoordinate>
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<geoCoordinate id="EE1D5065FFDCFFACA058FCC762683148" box="[545,619,797,821]" degrees="0.278" direction="west" orientation="longitude" pageId="60" pageNumber="2237" precision="55" value="-0.278">–0.278</geoCoordinate>
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||
) of SL; preanal length, PAL, mean 0.569 (range 0.543–0.59) of SL; depth of caudal peduncle, CPD, mean 0.298 (range
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<geoCoordinate id="EE1D5065FFDCFFACA19EFCE764263128" box="[999,1061,829,853]" degrees="0.269" direction="north" orientation="latitude" pageId="60" pageNumber="2237" precision="55" value="0.269">0.269</geoCoordinate>
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<geoCoordinate id="EE1D5065FFDCFFACA65DFCE7646D3128" box="[1060,1134,829,853]" degrees="0.331" direction="west" orientation="longitude" pageId="60" pageNumber="2237" precision="55" value="-0.331">–0.331</geoCoordinate>
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||
) of CPL, length of caudal peduncle; body depth, BD, mean 0.267 (range
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<geoCoordinate id="EE1D5065FFDCFFACA19CFC8764213108" box="[997,1058,861,885]" degrees="0.225" direction="north" orientation="latitude" pageId="60" pageNumber="2237" precision="55" value="0.225">0.225</geoCoordinate>
|
||
<geoCoordinate id="EE1D5065FFDCFFACA65BFC87646E3108" box="[1058,1133,861,885]" degrees="0.293" direction="west" orientation="longitude" pageId="60" pageNumber="2237" precision="55" value="-0.293">–0.293</geoCoordinate>
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) of SL; number of dorsal fin rays, nD, median 24 (range 22–26); number of scales in lateral line row, SLS, median 40 (range 39–42); distance between anterior base of pectoral fin to anterior base of pelvic fin, PPf, mean 0.152 (range
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<geoCoordinate id="EE1D5065FFDCFFACA151FC66636631A9" box="[808,869,956,980]" degrees="0.128" direction="north" orientation="latitude" pageId="60" pageNumber="2237" precision="55" value="0.128">0.128</geoCoordinate>
|
||
<geoCoordinate id="EE1D5065FFDCFFACA11CFC6663B331A9" box="[869,944,956,980]" degrees="0.169" direction="west" orientation="longitude" pageId="60" pageNumber="2237" precision="55" value="-0.169">–0.169</geoCoordinate>
|
||
) of SL.
|
||
</paragraph>
|
||
<paragraph id="8B9636A2FFDCFFACA2AAFBDB6412344B" blockId="60.[211,1172,1024,1590]" pageId="60" pageNumber="2237">
|
||
<emphasis id="B95DEAB0FFDCFFACA2AAFBDB61593665" box="[211,346,1025,1048]" italics="true" pageId="60" pageNumber="2237">Description.</emphasis>
|
||
Body long-oval shape (
|
||
<figureCitation id="13122A27FFDCFFACA013FBDA62CC3665" box="[618,719,1024,1048]" captionStart="Figure 2" captionStartId="5.[143,210,1254,1276]" captionText="Figure 2(A–D). Photographs of members of southern and eastern African Petrocephalus species studied (numbers refer to localities given in Figures 1, 6 and 9). (1) P. catostoma (Günther, 1866), Lectotype, BMNH 1863.10.12.4, right side, SL 4.7 cm. (2) P. stuhlmanni Boulenger, 1909, holotype, BMNH 1907.12.3.1, SL 7.9 cm. (3) P. wesselsi Kramer and Van der Bank, 2000, right side, SL 9.7 cm, SAIAB 85922 (R1). (7) P. longicapitis sp. nov., SMF 28265 (R1), SL 9 cm. (9) P. okavangensis sp. nov., holotype, SAIAB 030046, right side, SL 6.1 cm. (10) P. catostoma tanensis Whitehead and Greenwood, 1959, holotype, BMNH 1963.11.29.1, right side, SL 6.7 cm. (10a) P. tanensis, field no. Ta05na, SL 8.7 cm, SAIAB 85907. (12) P. longianalis sp. nov., holotype, SL 8.2 cm, right side, SAIAB 76758. (21) P. petersi sp. nov., holotype, SAIAB 060846, right side, SL 6.4 cm. (23) P. magnoculis sp. nov., SL 8.9 cm, ZSM 38659. (24) P. haullevillii Boulenger, 1912, BMNH 1912.4.1.186–188 (R1), right side, SL 5.7 cm. (25) P. steindachneri Fowler 1958, syntype, NMW 55118:3, right side, SL 6.4 cm. (26) P. stuhlmanni congicus David and Poll, 1937, syntype, MRAC 30807-30808, SL 7.8 cm. (27) P. degeni Boulenger, 1906, BMNH 1906.5.30.84, right side, SL 8.12 cm. (28) P. catostoma, SAIAB 73894 (R1), right side, SL 6.4 cm. (31) P. magnitrunci sp. nov., SL 8.5 cm, SAIAB 67069 (R5, right side), see map Figure 6. Scale bar, 1 cm." pageId="60" pageNumber="2237">Figure 2</figureCitation>
|
||
, no. 12). Head broadly rounded with a small ventrally positioned subterminal mouth, situated ventral to the eye; head and body dorsolaterally compressed. Dorsal fin (a) origin situated about two-thirds of standard length from snout, (b) obliquely orientated, anteriorly higher and posteriorly lower, (c) distal margin crescentic with anterior two or three rays longer than posterior rays, and (d) number of rays 22 (
|
||
<emphasis id="B95DEAB0FFDCFFACA03FFB7B625036C5" box="[582,595,1185,1208]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 8), 23 (
|
||
<emphasis id="B95DEAB0FFDCFFACA0BEFB7B62D736C5" box="[711,724,1185,1208]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 20), 24 (
|
||
<emphasis id="B95DEAB0FFDCFFACA12FFB7B636036C5" box="[854,867,1185,1208]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 18), 25 (
|
||
<emphasis id="B95DEAB0FFDCFFACA19DFB7B63F236C5" box="[996,1009,1185,1208]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 3), (e) dark spot below fin origin (lacking in samples from
|
||
<collectingRegion id="49EDF840FFDCFFACA09DFB65638A36AA" box="[740,905,1215,1239]" country="Zambia" name="Luapula" pageId="60" pageNumber="2237">Luapula River</collectingRegion>
|
||
bridge, SAIAB 76582). Anal fin (a) longer than dorsal fin, (b) opposite dorsal fin with slightly more anterior origin, (c) obliquely orientated, anteriorly lower and posteriorly higher, (d) anterior 10 or so rays longer than posterior ones, especially in males where they also appear stronger, (e) margin broadly rounded, (f) rays posterior to first 10 with distal margin straight, (g) number of rays 30 (
|
||
<emphasis id="B95DEAB0FFDCFFACA04EFABA6247370A" box="[567,580,1376,1399]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 2), 31 (
|
||
<emphasis id="B95DEAB0FFDCFFACA0CDFABA62C2370A" box="[692,705,1376,1399]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 6), 32 (
|
||
<emphasis id="B95DEAB0FFDCFFACA14BFABA633C370A" box="[818,831,1376,1399]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 16), 33 (
|
||
<emphasis id="B95DEAB0FFDCFFACA1C5FABA63CA370A" box="[956,969,1376,1399]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 17), 34 (
|
||
<emphasis id="B95DEAB0FFDCFFACA63EFABA6457370A" box="[1095,1108,1376,1399]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 6), 35 (
|
||
<emphasis id="B95DEAB0FFDCFFACA378FA5A610D37EA" box="[257,270,1408,1431]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 2). Forked tail fin with rounded lobes. Scales cycloid with reticulate striae, scales extending anteriorly to operculum and pectoral fins (beyond pelvics). Scales in lateral series, 39 (
|
||
<emphasis id="B95DEAB0FFDCFFACA3EAFA1A61A337AA" box="[403,416,1472,1495]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 9), 40 (
|
||
<emphasis id="B95DEAB0FFDCFFACA077FA1A621837AA" box="[526,539,1472,1495]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 17), 41 (
|
||
<emphasis id="B95DEAB0FFDCFFACA0EFFA1A62A037AA" box="[662,675,1472,1495]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 21), 42 (
|
||
<emphasis id="B95DEAB0FFDCFFACA167FA1A632837AA" box="[798,811,1472,1495]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 2). Scales on caudal peduncle circumference, 11 (
|
||
<emphasis id="B95DEAB0FFDCFFACA3D1FA0561B6378B" box="[424,437,1503,1526]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 2), 12 (
|
||
<emphasis id="B95DEAB0FFDCFFACA050FA056235378B" box="[553,566,1503,1526]" italics="true" pageId="60" pageNumber="2237">n</emphasis>
|
||
= 47). Caudal peduncle slender, subcylindrical entire length, usually 19.3% (18.4–19.9%) of SL (
|
||
<tableCitation id="C6AB0319FFDCFFACA0CCFA24630A346B" box="[693,777,1534,1558]" captionStart="Table 1" captionStartId="18.[58,113,442,464]" captionText="Table 1. Morphometrics of the southern and eastern African Petrocephalus species." pageId="60" pageNumber="2237">Table 1</tableCitation>
|
||
). Males with kink in anal fin base, which is absent in juveniles and females where the anal fin base is straight.
|
||
</paragraph>
|
||
<paragraph id="8B9636A2FFDCFFACA2AAF9B8649034E7" blockId="60.[211,1171,1634,1690]" pageId="60" pageNumber="2237">
|
||
<emphasis id="B95DEAB0FFDCFFACA2AAF9B861D73407" box="[211,468,1634,1658]" italics="true" pageId="60" pageNumber="2237">Colour in preservation.</emphasis>
|
||
Body brown, back only slightly darker, breast and belly fair, fins rather unpigmented and transparent, anal and dorsal fin bases emphasized as dark
|
||
</paragraph>
|
||
<paragraph id="8B9636A2FFDDFFADA2F6FF7D623832A2" blockId="61.[143,1104,167,223]" pageId="61" pageNumber="2238">
|
||
lines. Well-circumscribed dark spot below dorsal fin origin (except for specimens from
|
||
<collectingRegion id="49EDF840FFDDFFADA2F6FF1D613032A2" box="[143,307,199,223]" country="Zambia" name="Luapula" pageId="61" pageNumber="2238">Luapula River</collectingRegion>
|
||
bridge, SAIAB 76582).
|
||
</paragraph>
|
||
<paragraph id="8B9636A2FFDDFFADA2F6FED0624D33BF" blockId="61.[143,1103,266,450]" pageId="61" pageNumber="2238">
|
||
<emphasis id="B95DEAB0FFDDFFADA2F6FED060ED335F" box="[143,238,266,290]" italics="true" pageId="61" pageNumber="2238">Ecology.</emphasis>
|
||
The Lufubu
|
||
<collectingRegion id="49EDF840FFDDFFADA3F1FED061C5335F" box="[392,454,266,290]" country="Gambia" name="Upper River" pageId="61" pageNumber="2238">River</collectingRegion>
|
||
just above a
|
||
<quantity id="4CD19B47FFDDFFADA019FED062A4335F" box="[608,679,266,290]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="61" pageNumber="2238" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">3–4 m</quantity>
|
||
falls is a braided rocky channel with dense riparian forest. Water flow was rapid in section but there were also deeper pools with large rocks providing slacker flow areas. The substrate was rocky but with considerable amounts of leaf litter. In some sections there were mats of aquatic water ferns and mosses and filamentous algae. Fishes were collected with rotenone. Altitude, approximately
|
||
<quantity id="4CD19B47FFDDFFADA34FFE70618833BC" box="[310,395,426,450]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="61" pageNumber="2238" unit="m" value="1200.0">1200 m</quantity>
|
||
. Online
|
||
<figureCitation id="13122A27FFDDFFADA391FE70624933BF" box="[488,586,426,450]" captionStart="Figure 6" captionStartId="38.[211,278,892,914]" captionTargetBox="[211,1171,169,859]" captionTargetId="graphics-5@38.[211,1171,169,859]" captionTargetPageId="38" captionText="Figure 6. As Figure 1, but area of Petrocephalus okavangensis sp. nov. and location from where samples were taken shown at better resolution. (9) Nguma (Guma) Lagoon. (31) Boro River. (32) Gadikwe Lagoon. (33) Xakanixa River. (34) Xakanixa Channel. (35) Popa Rapids. (8) unique specimen from Kwando River." figureDoi="http://doi.org/10.5281/zenodo.4590846" httpUri="https://zenodo.org/record/4590846/files/figure.png" pageId="61" pageNumber="2238">Figure 6</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9636A2FFDDFFADA2F6FE37627F3019" blockId="61.[143,1103,492,612]" pageId="61" pageNumber="2238">
|
||
<emphasis id="B95DEAB0FFDDFFADA2F6FE3760FA3078" box="[143,249,493,517]" italics="true" pageId="61" pageNumber="2238">Remarks.</emphasis>
|
||
A median of nA = 33 is highest and distinctive among the
|
||
<taxonomicName id="4C294D21FFDDFFADA1CFFE37644C3078" authorityName="Kramer and Van der Bank" authorityYear="2000" box="[950,1103,493,517]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="61" pageNumber="2238" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95DEAB0FFDDFFADA1CFFE37644C3078" box="[950,1103,493,517]" italics="true" pageId="61" pageNumber="2238">Petrocephalus</emphasis>
|
||
</taxonomicName>
|
||
species of the present study. Except for samples from
|
||
<collectingRegion id="49EDF840FFDDFFADA08DFDD663993059" box="[756,922,524,548]" country="Zambia" name="Luapula" pageId="61" pageNumber="2238">Luapula River</collectingRegion>
|
||
bridge (SAIAB 76582), all other samples referable to
|
||
<taxonomicName id="4C294D21FFDDFFADA055FDF762BE3038" authority="Kramer & Bills & Skelton & Wink, 2012" authorityName="Kramer & Bills & Skelton & Wink" authorityYear="2012" box="[556,701,557,581]" class="Actinopterygii" family="Mormyridae" genus="Petrocephalus" kingdom="Animalia" order="Osteoglossiformes" pageId="61" pageNumber="2238" phylum="Chordata" rank="species" species="longianalis" status="sp. nov.">
|
||
<emphasis id="B95DEAB0FFDDFFADA055FDF762BE3038" box="[556,701,557,581]" italics="true" pageId="61" pageNumber="2238">P. longianalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26E57CBFFDDFFADA0BBFDF763163039" box="[706,789,557,580]" pageId="61" pageNumber="2238" rank="species">sp. nov.</taxonomicNameLabel>
|
||
with dark spot below dorsal fin origin, usually round, small and distinct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |