343 lines
70 KiB
XML
343 lines
70 KiB
XML
<document id="2132950B6B800C3A843D0A6BAF525012" ID-CLB-Dataset="32293" ID-DOI="10.1206/0003-0090-415.1.1" ID-GBIF-Dataset="22b02e67-634f-4e71-a8fa-3392aff63985" ID-Handle="2246/6723" ID-Zenodo-Dep="851382" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1503785870183" checkinUser="admin" docAuthor="Lauren A. Esposito, Humberto Y. Yamaguti, Cláudio A. Souza, Ricardo Pinto-Da-Rocha & Lorenzo Prendini" docDate="2017" docId="8F65ED57FF97B10A38C7CCC8B2696FF4" docLanguage="en" docName="BulAmeMusNatHis.2017.415.1-63.pdf.imf" docOrigin="Bulletin of the American Museum of Natural History 415" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Physoctonus debilis C.L. Koch 1840" docType="treatment" docVersion="19" lastPageNumber="93" masterDocId="735C952FFFCAB1543860CC2CB35C6E75" masterDocTitle="Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="93" updateTime="1698467378382" updateUser="ExternalLinkService">
|
||
<mods:mods id="01121FC44BAB57E1ADBB0981E59050DF" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="9D1AC515C186FC05B34AD38643315704">
|
||
<mods:title id="6E661EF8B464B51AED039DA349F64470">Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="399AD7361BB26CB9052E4B1CB0FEA824" type="personal">
|
||
<mods:role id="F328B78D2026135E2323A9E3C3F0E99B">
|
||
<mods:roleTerm id="F8694FBB2814676E7171A93993C5E7AD">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="30E4666A001A5272A5257FC2E8BF5412">Lauren A. Esposito</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="47B80AFB3E5F0AAF8342ADBB75FB813D" type="personal">
|
||
<mods:role id="795C1579C4F3106831C3F32FB92D9590">
|
||
<mods:roleTerm id="06218146960AEF6BB9B1A566E375BC1A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="36315641031466C2802049ED226EF85E">Humberto Y. Yamaguti</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="FFF4C3779DF639A1A42A00E2A2D3B7DE" type="personal">
|
||
<mods:role id="CE13A5A839E575E820391371B0621961">
|
||
<mods:roleTerm id="9688F32BB7AA6DDF4527EEAD68D82732">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="70706DF04F45FBF3A61FEE2C9237EEC8">Cláudio A. Souza</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="1A8D8BBE873DE89DD42F9D0C469E75A3" type="personal">
|
||
<mods:role id="70ACBBE3101F73CA58FC7F5552740843">
|
||
<mods:roleTerm id="A26310EE48DE145D381708E053B04584">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="19D8DC37472D799ADECD221CE36795D3">Ricardo Pinto-Da-Rocha</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="657AF83BDD03BBAFD2857E7D550C72EC" type="personal">
|
||
<mods:role id="23F256233AD271979DF11357E0F30231">
|
||
<mods:roleTerm id="E5AAE1F27C39140175BC316F620608BF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="33A6F82BA320AE18CD6D1E9F28C82DC5">Lorenzo Prendini</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="6F9E76F8439D5E3D9DA59030C80BD4D5">text</mods:typeOfResource>
|
||
<mods:relatedItem id="0D91EC1D6902920B6CFD11E629FA972D" type="host">
|
||
<mods:titleInfo id="BB36803D0AF53A4C451F38D6AE52882B">
|
||
<mods:title id="660D0742BF836CB09DE77C7D8CFFFE65">Bulletin of the American Museum of Natural History</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B4F50CEB356E0A5BF09AB3B93A659030">
|
||
<mods:date id="7680CBC968450BB515927E9215114F74">2017</mods:date>
|
||
<mods:detail id="B973BC456DFC0300C6E55902369A1623" type="pubDate">
|
||
<mods:number id="C12AD05C340F40A0A2DEAE19DBF1E1B7">2017-06-26</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CBCB803C8065C52BD15606377B65D182" type="volume">
|
||
<mods:number id="AC7E82F01089D68DFF475636491258BC">415</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="90186DB8BFE59EF4BC2288BB9C969A92" unit="page">
|
||
<mods:start id="DEF67BCB3415B54164C196116525B3B0">1</mods:start>
|
||
<mods:end id="12EB6F94AC82D033B004E23A9B76E9F9">134</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="8479D5B438D32E90A16724216B095869">journal article</mods:classification>
|
||
<mods:identifier id="A6D0F14CC7F86FF31E29CC139ECDB269" type="CLB-Dataset">32293</mods:identifier>
|
||
<mods:identifier id="E636406137C0B319A1DCBAEDB72426F6" type="DOI">10.1206/0003-0090-415.1.1</mods:identifier>
|
||
<mods:identifier id="60D0D663CB54CFB997581EADF1C74167" type="GBIF-Dataset">22b02e67-634f-4e71-a8fa-3392aff63985</mods:identifier>
|
||
<mods:identifier id="FE0AC21735036B57AB8274407616E2CE" type="Handle">2246/6723</mods:identifier>
|
||
<mods:identifier id="FCF6537EAE2D7997076401CACABE141C" type="Zenodo-Dep">851382</mods:identifier>
|
||
<mods:identifier id="EE6D5D786E5471FDBF6997CBB3DC7E94" type="ZooBank">146A0539-0A2C-44CD-986C-8F8A8EB4598C</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="8F65ED57FF97B10A38C7CCC8B2696FF4" ID-DOI="http://doi.org/10.5281/zenodo.4610684" ID-GBIF-Taxon="132953594" ID-Zenodo-Dep="4610684" LSID="urn:lsid:plazi:treatment:8F65ED57FF97B10A38C7CCC8B2696FF4" httpUri="http://treatment.plazi.org/id/8F65ED57FF97B10A38C7CCC8B2696FF4" lastPageId="94" lastPageNumber="93" pageId="93" pageNumber="93">
|
||
<subSubSection id="4FD60FCAFF97B10938C7CCC8B1636E89" box="[167,575,228,252]" pageId="93" pageNumber="93" type="nomenclature">
|
||
<paragraph id="07735C41FF97B10938C7CCC8B1636E89" blockId="93.[167,575,228,252]" box="[167,575,228,252]" pageId="93" pageNumber="93">
|
||
<taxonomicName id="C0CC27C2FF97B10938C7CCC8B1636E89" ID-CoL="4HHR8" authority="(C. L. Koch, 1840)" authorityName="C.L. Koch" authorityYear="1840" baseAuthorityName="C. L. Koch" baseAuthorityYear="1840" box="[167,575,228,252]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B10938C7CCC8B2296E89" box="[167,373,228,252]" italics="true" pageId="93" pageNumber="93">Physoctonus debilis</emphasis>
|
||
(C.L.
|
||
<bibRefCitation id="635D21B0FF97B10939DACCC8B1646E89" author="Koch" box="[442,568,228,252]" firstAuthor="Koch" journalOrPublisher="Nurnberg: C. H. Zeh'sche Buchhandlung" pageId="93" pageNumber="93" pagination="1 - 114" part="8" refId="ref66739" refString="Koch, C. L. 1840. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung 8: 1 - 114 (pts. 1, 2, 1840; pts. 2, 3, 1841)." title="Die Arachniden" type="journal article" year="1840">Koch, 1840</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFF97B109381BCD3AB10D6F25" pageId="93" pageNumber="93" type="description">
|
||
<paragraph id="07735C41FF97B109381BCD3AB10D6F25" blockId="93.[123,619,278,336]" pageId="93" pageNumber="93">
|
||
<figureCitation id="9FF740C4FF97B109381BCD3AB3AD6F5B" box="[123,241,278,302]" captionStart="FIG" captionStartId="16.[107,147,1517,1538]" captionTargetBox="[188,1135,224,1487]" captionTargetId="figure@16.[142,1179,224,1487]" captionTargetPageId="16" captionText="FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil." httpUri="https://zenodo.org/record/851384/files/figure.png" pageId="93" pageNumber="93">
|
||
<figureCitation id="9FF740C4FF97B109381BCD3AB3816F5B" box="[123,221,278,302]" captionStart="FIG. 1" captionStartId="16.[107,147,1517,1538]" captionTargetBox="[188,1135,224,1487]" captionTargetId="figure@16.[142,1179,224,1487]" captionTargetPageId="16" captionText="FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil." httpUri="https://zenodo.org/record/851384/files/figure.png" pageId="93" pageNumber="93">Figures 1</figureCitation>
|
||
E
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FF97B109389DCD3AB2436F5B" box="[253,287,278,302]" captionStart="FIG" captionStartId="17.[111,153,1486,1507]" captionTargetBox="[155,1166,224,1455]" captionTargetId="figure@17.[111,1214,224,1455]" captionTargetPageId="17" captionText="FIG. 2. Representative habitats of species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971. A. Isla Mona, Puerto Rico, habitat of Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov. B. Sierra de Cubitas, Camagüey, Cuba, habitat of Heteroctenus junceus (Herbst, 1800), comb. nov. C. Guanambí, Bahía, Brazil, habitat of Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., and Jaguajir rochae (Borelli, 1910), comb. nov. D. Exu, Pernambuco, Brazil, habitat of J. rochae and Physoctonus debilis (C.L. Koch, 1840). E. Santarém, Pará, Brazil, habitat of Rhopalurus laticauda Thorell, 1876. F. Normandia, Roraima, Brazil, habitat of R. laticauda. G. Lapa do Bode, Itaeté, Bahía, Brazil, habitat of Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov. H. Gruta do Lapão, Lençóis, Bahía, Brazil, habitat of Troglorhopalurus translucidus Lourenço et al., 2004." httpUri="https://zenodo.org/record/851386/files/figure.png" pageId="93" pageNumber="93">2D</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FF97B109394BCD3AB2156F5B" box="[299,329,278,302]" captionStart="FIG" captionStartId="24.[107,149,1416,1437]" captionTargetBox="[138,1207,234,1374]" captionTargetId="figure@24.[107,1211,224,1383]" captionTargetPageId="24" captionText="FIG. 9. Map of northeastern South America, plotting known locality records of six species of Rhopalurusinae Bücherl, 1971: A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov. (star), Ischnotelson peruassu, sp. nov. (triangle), Physoctonus striatus, sp. nov. (circle), Troglorhopalurus lacrau (Lourenço and Pinto-da- Rocha, 1997), comb. nov (diamonds). B. Physoctonus debilis (C.L. Koch, 1840) (squares), Troglorhopalurus translucidus Lourenço et al., 2004 (crosses)." httpUri="https://zenodo.org/record/851400/files/figure.png" pageId="93" pageNumber="93">9B</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FF97B1093935CD3AB2DF6F5B" box="[341,387,278,302]" captionStart="FIG" captionStartId="26.[109,151,1396,1417]" captionTargetBox="[109,1213,224,1368]" captionTargetId="figure@26.[109,1213,224,1368]" captionTargetPageId="26" captionText="FIG. 10. Rhopalurusinae Bücherl, 1971, sternites III (A–C, E, F), V and VI (D), ventral aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). B. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (MZSP). C, D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E. Physoctonus debilis (C.L. Koch, 1840), ♂ (MZSP). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 1 mm (A, E, F), 2 mm (B, C, D)." httpUri="https://zenodo.org/record/851402/files/figure.png" pageId="93" pageNumber="93">10D</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FF97B10939EFCD3AB2E36F5B" box="[399,447,278,302]" captionStart="FIG" captionStartId="28.[108,150,1167,1188]" captionTargetBox="[108,1213,224,1136]" captionTargetId="figure@28.[108,1213,224,1136]" captionTargetPageId="28" captionText="FIG. 11. Rhopalurusinae Bücherl, 1971, dextral pecten, ventral aspect (A, C, E, G) and pectinal teeth, dorsal aspect (B, D, F). A. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♀ (AMNH). C. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP) D. Jaguajir rochae (Borelli, 1910), comb. nov., (AMNH). E. Rhopalurus laticauda Thorell, 1876, ♂ (AMCC [LP 2845]). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (AMCC [LP 3260]). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP)." httpUri="https://zenodo.org/record/851404/files/figure.png" pageId="93" pageNumber="93">11G</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FF97B10939ABCD3AB2AE6F5B" box="[459,498,278,302]" captionStart="FIG" captionStartId="29.[107,149,984,1005]" captionTargetBox="[107,1211,224,956]" captionTargetId="figure@29.[107,1211,224,956]" captionTargetPageId="29" captionText="FIG. 12. Rhopalurusinae Bücherl, 1971, pectinal teeth, dorsal aspect, fine structure. A. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., (MZSP). B. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). C. Rhopalurus laticauda Thorell, 1876, (MZSP). D. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (MZSP). E. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). F. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP)." httpUri="https://zenodo.org/record/851406/files/figure.png" pageId="93" pageNumber="93">12F</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FF97B109399ECD3AB1716F5B" box="[510,557,278,302]" captionStart="FIG" captionStartId="34.[109,151,1628,1649]" captionTargetBox="[208,1116,224,1599]" captionTargetId="figure@34.[109,1212,224,1599]" captionTargetPageId="34" captionText="FIG. 16. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP). B. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). C. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). D. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. E. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Guatire, Venezuela. F. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). Scale bars = 1 mm (A, B), 2 mm (C–F)." httpUri="https://zenodo.org/record/851414/files/figure.png" pageId="93" pageNumber="93">16A</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FF97B1093A58CD3AB13B6F5B" box="[568,615,278,302]" captionStart="FIG" captionStartId="40.[108,151,1490,1511]" captionTargetBox="[108,1212,224,1461]" captionTargetId="figure@40.[108,1212,224,1462]" captionTargetPageId="40" captionText="FIG. 20. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Physoctonus debilis (C.L. Koch, 1840), ♂ (MZSP). B. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). C. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). D. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. E. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Puerto Colombia, Venezuela. F. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851422/files/figure.png" pageId="93" pageNumber="93">20A</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FF97B10938F6CD14B3996F25" box="[150,197,312,336]" captionStart="FIG" captionStartId="44.[107,149,1352,1373]" captionTargetBox="[108,1212,224,1323]" captionTargetId="figure@44.[108,1212,224,1323]" captionTargetPageId="44" captionText="FIG. 21. Rhopalurusinae Bücherl, 1971, telson, lateral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851424/files/figure.png" pageId="93" pageNumber="93">21G</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FF97B10938B1CD14B25D6F25" box="[209,257,312,336]" captionStart="FIG" captionStartId="45.[108,147,1513,1534]" captionTargetBox="[176,1145,224,1484]" captionTargetId="figure@45.[108,1212,224,1484]" captionTargetPageId="45" captionText="FIG. 22. Rhopalurusinae Bücherl, 1971, telson, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH). J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851426/files/figure.png" pageId="93" pageNumber="93">22G</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FF97B109396CCD14B20A6F25" box="[268,342,312,336]" captionStart="FIG" captionStartId="47.[107,149,1521,1542]" captionTargetBox="[107,1211,228,1487]" captionTargetId="figure@47.[107,1211,228,1493]" captionTargetPageId="47" captionText="FIG. 24. Rhopalurusinae Bücherl, 1971, hemispermatophores, dorsal aspect (E), dorsal aspect, detail (B, H, M, P), ectal aspect (F, I, N, S), ectal aspect, detail (D, J, R, T, U), ental aspect (A, G, K, O), and ental aspect, detail (C, L, Q). A–D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E–H. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). I–M. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP). N–R. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (MZSP). S–U. Physoctonus debilis (C.L. Koch, 1840), ♂ (MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/851430/files/figure.png" pageId="93" pageNumber="93">24S–U</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FF97B1093901CD14B2F16F3A" box="[353,429,312,336]" captionStart="FIG" captionStartId="88.[108,150,1167,1188]" captionTargetBox="[108,1212,224,1137]" captionTargetId="figure@88.[108,1212,224,1137]" captionTargetPageId="88" captionText="FIG. 48. Physoctonus Mello-Leitão, 1934, pedipalp chela, retrolateral aspect. A. P. debilis (C.L. Koch, 1840), ♂ (MZSP). B. P. debilis (C.L. Koch, 1840), ♀ (MZSP). C. P. striatus, sp. nov., ♂ (MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/851478/files/figure.png" pageId="93" pageNumber="93">48A, B</figureCitation>
|
||
,
|
||
<figureCitation id="9FF740C4FF97B10939D8CD1BB2A16F3A" box="[440,509,311,335]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="90.[107,149,1293,1314]" captionStartId-1="91.[107,149,1276,1297]" captionStartId-2="92.[107,149,1467,1488]" captionTargetBox-0="[120,1209,224,1263]" captionTargetBox-1="[115,1204,224,1246]" captionTargetBox-2="[201,1115,224,1429]" captionTargetId-0="figure@90.[120,1209,224,1263]" captionTargetId-1="figure@91.[115,1204,224,1246]" captionTargetId-2="figure@92.[201,1115,224,1429]" captionTargetPageId-0="90" captionTargetPageId-1="91" captionTargetPageId-2="92" captionText-0="FIG. 49. Physoctonus debilis (C.L. Koch, 1840), metasoma and telson, dorsal (A–C) and ventral (D–F) aspects. A, C, D, F. 2 ♂ (MZSP). B, E. ♀ (AMNH). Scale bars = 2.5 mm." captionText-1="FIG. 50. Physoctonus debilis (C.L. Koch, 1840), metasoma and telson, lateral aspect. A, C. 2♂ (MZSP). B. ♀ (AMNH). Scale bars = 2.5 mm." captionText-2="FIG. 51. Physoctonus debilis (C.L. Koch, 1840), 2 ♀ (AMNH), habitus, dorsal (A, C) and ventral (B, D) aspects. Scale bars = 1 mm." httpUri-0="https://zenodo.org/record/851480/files/figure.png" httpUri-1="https://zenodo.org/record/851482/files/figure.png" httpUri-2="https://zenodo.org/record/851484/files/figure.png" pageId="93" pageNumber="93">49–51</figureCitation>
|
||
;
|
||
<tableCitation id="4A4E69FAFF97B1093A68CD14B10D6F25" box="[520,593,312,336]" captionStart="TABLE 3" captionStartId="69.[610,690,228,252]" captionTargetBox="[131,1171,439,1728]" captionText="TABLE 3 Meristic data for type material of Ischnotelson peruassu, sp. n., Physoctonus striatus, sp. n., and Physoctonus debilis (C. L. Koch, 1840). Measurements follow Prendini (2000, 2003, 2004). 1 Sum of carapace, tergites I – VII, metasomal segments I – V, and telson; 2 sum of metasomal segments I – V and telson; 3 measured from base of condyle to tip of fixed finger." httpUri="http://table.plazi.org/id/53B30CC9FF8FB1113A02CCC8B0CF6FEA" pageId="93" pageNumber="93" tableUuid="53B30CC9FF8FB1113A02CCC8B0CF6FEA">table 3</tableCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFF97B109380BCD43B2356AD7" pageId="93" pageNumber="93" type="reference_group">
|
||
<paragraph id="07735C41FF97B109380BCD43B3E76FBE" blockId="93.[107,636,366,1186]" pageId="93" pageNumber="93">
|
||
<taxonomicName id="C0CC27C2FF97B109380BCD43B2BB6FF3" authority="C. L. Koch, 1840: 21" authorityName="C. L. Koch" authorityPageNumber="21" authorityYear="1840" box="[107,487,366,390]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B109380BCD43B2576FF3" box="[107,267,366,390]" italics="true" pageId="93" pageNumber="93">Vaejovis debilis</emphasis>
|
||
C.L.
|
||
<bibRefCitation id="635D21B0FF97B1093926CD42B2BB6FF3" author="Koch" box="[326,487,366,390]" firstAuthor="Koch" journalOrPublisher="Nurnberg: C. H. Zeh'sche Buchhandlung" pageId="93" pageNumber="93" pagination="1 - 114" part="8" refId="ref66739" refString="Koch, C. L. 1840. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung 8: 1 - 114 (pts. 1, 2, 1840; pts. 2, 3, 1841)." title="Die Arachniden" type="journal article" year="1840">Koch, 1840: 21</bibRefCitation>
|
||
</taxonomicName>
|
||
, 22, pl. CCLIX, fig. 605; 1850: 89;
|
||
<bibRefCitation id="635D21B0FF97B10939D8CDBCB3EB6FBE" author="Kraepelin" bookContentInfo="265 pp." editor="F. Dahl" firstAuthor="Kraepelin" journalOrPublisher="Friedlander and Sohn Verlag" pageId="93" pageNumber="93" refId="ref66921" refString="Kraepelin, K. 1899. Scorpiones und Pedipalpi. In F. Dahl (editor), Das Tierreich. Berlin: R. Friedlander and Sohn Verlag. 265 pp." title="Scorpiones und Pedipalpi" type="book" volumeTitle="Das Tierreich. Berlin: R" year="1899">Kraepelin, 1899: 96</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF97B109380BCDFAB2A66F9B" blockId="93.[107,636,366,1186]" box="[107,506,470,494]" pageId="93" pageNumber="93">
|
||
<taxonomicName id="C0CC27C2FF97B109380BCDFAB24E6F9B" authority=": Gervais, 1844 b: 458" authorityName=": Gervais" authorityPageNumber="458" authorityYear="1844" box="[107,274,470,494]" class="Arachnida" family="Scorpionidae" genus="Waejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B109380BCDFAB24E6F9B" box="[107,274,470,494]" italics="true" pageId="93" pageNumber="93">Waejovis debilis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<bibRefCitation id="635D21B0FF97B1093940CDFAB2E96F9B" author="Gervais" box="[288,437,470,494]" firstAuthor="Gervais" journalOrPublisher="Archives du Museum d'Histoire Naturelle, Paris" pageId="93" pageNumber="93" pagination="201 - 240" part="4" refId="ref66060" refString="Gervais, P. M. 1844 a. Remarques sur la famille des scorpions et description des plusieurs especes nouvelles de la collection du Museum. Archives du Museum d'Histoire Naturelle, Paris 4: 201 - 240." title="Remarques sur la famille des scorpions et description des plusieurs especes nouvelles de la collection du Museum" type="journal article" year="1844">Gervais, 1844</bibRefCitation>
|
||
b: 458.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF97B109380BCDD4B2F96D1F" blockId="93.[107,636,366,1186]" pageId="93" pageNumber="93">
|
||
<taxonomicName id="C0CC27C2FF97B109380BCDD4B2A46C65" authority=": Borelli, 1910: 5" authorityName=": Borelli" authorityPageNumber="5" authorityYear="1910" box="[107,504,504,528]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B109380BCDD4B2716C65" box="[107,301,504,528]" italics="true" pageId="93" pageNumber="93">Rhopalurus debilis</emphasis>
|
||
:
|
||
<bibRefCitation id="635D21B0FF97B109395ACDD4B2A46C65" author="Borelli" box="[314,504,504,528]" firstAuthor="Borelli" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Realle Universita di Torino" pageId="93" pageNumber="93" pagination="1 - 8" part="25" refId="ref64441" refString="Borelli, A. 1910. Scorpioni nuovi e poco noti del Brasile. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Realle Universita di Torino 25 (629): 1 - 8." title="Scorpioni nuovi e poco noti del Brasile" type="journal article" year="1910">Borelli, 1910: 5–8</bibRefCitation>
|
||
</taxonomicName>
|
||
, fig. 1;
|
||
<bibRefCitation id="635D21B0FF97B10938FBCE37B2216C46" author="Mello-Campos" box="[155,381,539,563]" firstAuthor="Mello-Campos" journalOrPublisher="Memorias do Instituto Oswaldo Cruz" pageId="93" pageNumber="93" pagination="237 - 369" part="17" refId="ref69164" refString="Mello-Campos, O. de. 1924 a. Os escorpioes brasileiros. Memorias do Instituto Oswaldo Cruz 17 (2): 237 - 369." title="Os escorpioes brasileiros" type="journal article" year="1924">Mello-Campos, 1924</bibRefCitation>
|
||
a: 252, 275, 276, 277; 1924b: 318, 341, 342, 343;
|
||
<bibRefCitation id="635D21B0FF97B10939D8CE11B3AA6C0D" author="Mello-Leitao" firstAuthor="Mello-Leitao" journalOrPublisher="Arquivos do Museu Nacional" pageId="93" pageNumber="93" pagination="9 - 46" part="34" refId="ref69227" refString="Mello-Leitao, C. F. de. 1932. Notas sobre escorpioes sulamericanos. Arquivos do Museu Nacional 34: 9 - 46." title="Notas sobre escorpioes sulamericanos" type="journal article" year="1932">Mello-Leitão, 1932: 14</bibRefCitation>
|
||
, 30;
|
||
<bibRefCitation id="635D21B0FF97B1093949CE4CB28E6C0D" author="Meise" box="[297,466,608,632]" firstAuthor="Meise" journalOrPublisher="Nyt Magazin for Naturvidenskaberne" pageId="93" pageNumber="93" pagination="25 - 43" part="72" refId="ref69146" refString="Meise, W. 1934. Scorpiones. Nyt Magazin for Naturvidenskaberne 72: 25 - 43." title="Scorpiones" type="journal article" year="1934">Meise, 1934: 42</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FF97B10939BDCE4CB3EB6CEE" author="Prado" firstAuthor="Prado" journalOrPublisher="Sinopse das especies de Rhopalurus. Memorias do Instituto Butantan" pageId="93" pageNumber="93" pagination="25 - 36" part="13" refId="ref70234" refString="Prado, A. 1940. Contribuicao ao conhecimento dos escorpioes sul-americanos. Sinopse das especies de Rhopalurus. Memorias do Instituto Butantan 13: 25 - 36." title="Contribuicao ao conhecimento dos escorpioes sul-americanos" type="journal article" year="1940">Prado, 1940: 26</bibRefCitation>
|
||
, 29–30;
|
||
<bibRefCitation id="635D21B0FF97B1093970CEAFB1536CEE" author="Mello-Leitao" box="[272,527,642,667]" firstAuthor="Mello-Leitao" journalOrPublisher="Arquivos do Museu Nacional" pageId="93" pageNumber="93" pagination="1 - 468" part="40" refId="ref69384" refString="Mello-Leitao, C. F. de. 1945. Escorpioes sul americanos. Arquivos do Museu Nacional 40: 1 - 468." title="Escorpioes sul americanos" type="journal article" year="1945">Mello-Leitão, 1945: 266</bibRefCitation>
|
||
, 272, 273;
|
||
<bibRefCitation id="635D21B0FF97B10938FBCE89B23F6CC8" author="Bucherl" box="[155,355,677,701]" firstAuthor="Bucherl" journalOrPublisher="Catalogo da colecao escorpionica do Instituto Butantan. Memorias do Instituto Butantan" pageId="93" pageNumber="93" pagination="255 - 275" part="29" refId="ref64652" refString="Bucherl, W. 1959. Escorpioes e escorpionismo no Brasil. X. Catalogo da colecao escorpionica do Instituto Butantan. Memorias do Instituto Butantan 29: 255 - 275." title="Escorpioes e escorpionismo no Brasil. X" type="journal article" year="1959">Bücherl, 1959: 268</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FF97B109390FCE89B28B6CC8" author="Bucherl" box="[367,471,677,701]" editor="W. Bucherl & E. R. Buckley" firstAuthor="Bucherl" journalOrPublisher="New York: Academic Press" pageId="93" pageNumber="93" pagination="317 - 348" part="3" refId="ref64808" refString="Bucherl, W. 1971. Classification, biology and venom extraction of scorpions. In W. Bucherl and E. R. Buckley (editors), Venomous animals and their venoms 3: 317 - 348. New York: Academic Press." title="Classification, biology and venom extraction of scorpions" type="journal article" volumeTitle="Venomous animals and their venoms" year="1971">1971: 327</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FF97B1093983CE89B38D6C95" author="Francke" firstAuthor="Francke" journalOrPublisher="Journal of Arachnology" pageId="93" pageNumber="93" pagination="125 - 135" part="4" refId="ref65579" refString="Francke, O. F. 1977 a. Two emendations to Stahnke's (1974) Vaejovidae revision (Scorpionida, Vaejovidae). Journal of Arachnology 4 (2): 125 - 135." title="Two emendations to Stahnke's (1974) Vaejovidae revision (Scorpionida, Vaejovidae)" type="journal article" year="1977">Francke, 1977</bibRefCitation>
|
||
a: 127–134, figs. 1–15;
|
||
<bibRefCitation id="635D21B0FF97B10939A2CEE4B1376C95" author="Lourenco" box="[450,619,712,736]" firstAuthor="Lourenco" journalOrPublisher="Revue Arachnologique" pageId="93" pageNumber="93" pagination="107 - 141" part="4" refId="ref67639" refString="Lourenco, W. R. 1982 a. Revision du genre Rhopalurus Thorell, 1876 (Scorpiones, Buthidae). Revue Arachnologique 4: 107 - 141." title="Revision du genre Rhopalurus Thorell, 1876 (Scorpiones, Buthidae)" type="journal article" year="1982">Lourenço, 1982</bibRefCitation>
|
||
a: 133, 136, fig. 78; 1986a: 133, figs. 12, 16; 1986b: 165, fig. 7; 1990: 161; 1992: 55;
|
||
<bibRefCitation id="635D21B0FF97B10938FBCF03B2396D3D" author="Kovarik" bookContentInfo="175 pp." box="[155,357,815,840]" firstAuthor="Kovarik" journalOrPublisher="Madagaskar: Jihlava" pageId="93" pageNumber="93" refId="ref66846" refString="Kovarik, F. 1998. Stiri [Scorpions]. Madagaskar: Jihlava. 175 pp. [in Czech]" title="Stiri [Scorpions]" type="book" year="1998">Kovařík, 1998: 118</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FF97B1093911CF1CB1256D3D" author="Fet and Lowe" box="[369,633,815,840]" editor="V. Fet & W. D. Sissom" firstAuthor="Fet" journalOrPublisher="New York: New York Entomological Society" pageId="93" pageNumber="93" pagination="54 - 286" refId="ref65304" refString="Fet, V., and G. Lowe. 2000. Family Buthidae C. L. Koch, 1837. In V. Fet, W. D. Sissom, G., Lowe, and M. E. Braunwalder, Catalog of the scorpions of the world (1758 - 1998): 54 - 286. New York: New York Entomological Society." title="Family Buthidae C. L. Koch, 1837" type="book chapter" volumeTitle="G., Lowe, and M. E" year="2000">Fet and Lowe, 2000: 219</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FF97B10938FBCF7EB2FD6D1F" author="Ubinski" box="[155,417,850,874]" etAl="et al." firstAuthor="Ubinski" journalOrPublisher="Cytogenetic and Genome Research" pageId="93" pageNumber="93" pagination="83 - 155" part="148" refId="ref72292" refString="Ubinski, C. V., L. S. Carvalho, and M. C. Schneider. 2016. Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil. Cytogenetic and Genome Research 148: 83 - 155. [doi: 10.1159 / 000446523]" title="Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil" type="journal article" year="2016">Ubinski et al., 2016: 122</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF97B109380BCF59B2376A62" blockId="93.[107,636,366,1186]" pageId="93" pageNumber="93">
|
||
<taxonomicName id="C0CC27C2FF97B109380BCF59B10B6DF8" authority="Mello-Leitao, 1934 b: 76" authorityName="Mello-Leitao" authorityPageNumber="76" authorityYear="1934" box="[107,599,885,909]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="physurus">
|
||
<emphasis id="35B88053FF97B109380BCF59B2136DF8" box="[107,335,885,909]" italics="true" pageId="93" pageNumber="93">Physoctonus physurus</emphasis>
|
||
<bibRefCitation id="635D21B0FF97B1093937CF59B17F6DF8" author="Mello-Leitao" box="[343,547,885,909]" firstAuthor="Mello-Leitao" journalOrPublisher="Octava Reunion de la Sociedad Argentina, Santiago del Estero" pageId="93" pageNumber="93" pagination="1 - 97" part="1933" refId="ref69252" refString="Mello-Leitao, C. de. 1934 a. (" 1933 "). Estudo monografico dos escorpioes da Republica Argentina. Octava Reunion de la Sociedad Argentina, Santiago del Estero 1933: 1 - 97." title="Estudo monografico dos escorpioes da Republica Argentina" type="journal article" year="1934">Mello-Leitão, 1934</bibRefCitation>
|
||
b: 76
|
||
</taxonomicName>
|
||
, 77, figs. 1–7 (synonymized by
|
||
<bibRefCitation id="635D21B0FF97B1093981CFBBB38D6DA7" author="Francke" firstAuthor="Francke" journalOrPublisher="Journal of Arachnology" pageId="93" pageNumber="93" pagination="125 - 135" part="4" refId="ref65579" refString="Francke, O. F. 1977 a. Two emendations to Stahnke's (1974) Vaejovidae revision (Scorpionida, Vaejovidae). Journal of Arachnology 4 (2): 125 - 135." title="Two emendations to Stahnke's (1974) Vaejovidae revision (Scorpionida, Vaejovidae)" type="journal article" year="1977">Francke, 1977</bibRefCitation>
|
||
a: 127); 1942: 129; 1945: 129–132, figs. 40, 41;
|
||
<bibRefCitation id="635D21B0FF97B1093889CFF0B2ED6D81" author="Bucherl" box="[233,433,988,1012]" editor="H. Lent" firstAuthor="Bucherl" journalOrPublisher="Zoologia" pageId="93" pageNumber="93" pagination="111 - 125" part="5" refId="ref64708" refString="Bucherl, W. 1967. Escorpioes, aranhas e escolopendromorfos da Amazonia. In H. Lent (editor), Atas do Simposio sobre a Biota Amazonica 5 (Zoologia): 111 - 125." title="Escorpioes, aranhas e escolopendromorfos da Amazonia" type="journal article" volumeTitle="Atas do Simposio sobre a Biota Amazonica" year="1967">Bücherl, 1967: 115</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FF97B10939DDCFF0B17A6D81" author="Bucherl" box="[445,550,988,1012]" editor="E. J. Fittkau & J. Illies & H. King & G. H. Schwabe & H. Sioli" firstAuthor="Bucherl" journalOrPublisher="2" pageId="93" pageNumber="93" pagination="764 - 793" part="19" refId="ref64746" refString="Bucherl, W. 1969. Giftige Arthropoden. In E. J. Fittkau, J. Illies, H. King, G. H. Schwabe, and H. Sioli (editors), Biogeography and ecology in South America. Monographiae Biologicae 19 (2): 764 - 793. Dordrecht: W. Junk." title="Giftige Arthropoden" type="journal article" volumeTitle="Biogeography and ecology in South America" year="1969">1969: 768</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FF97B10938FBCFD3B23B6A62" author="Stahnke" box="[155,359,1023,1047]" firstAuthor="Stahnke" journalOrPublisher="Journal of Arachnology" pageId="93" pageNumber="93" pagination="107 - 141" part="1" refId="ref71310" refString="Stahnke, H. L. 1974. Revision and keys to the higher categories of Vejovidae (Scorpionida). Journal of Arachnology 1 (2): 107 - 141." title="Revision and keys to the higher categories of Vejovidae (Scorpionida)" type="journal article" year="1974">Stahnke, 1974: 129</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF97B109380BC80EB2356AD7" blockId="93.[107,636,366,1186]" pageId="93" pageNumber="93">
|
||
<taxonomicName id="C0CC27C2FF97B109380BC80EB1426A4F" ID-CoL="4HHR8" authority=": Lourenco, 2007: 360" authorityName=": Lourenco" authorityPageNumber="360" authorityYear="2007" box="[107,542,1058,1082]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B109380BC80EB2686A4F" box="[107,308,1058,1082]" italics="true" pageId="93" pageNumber="93">Physoctonus debilis</emphasis>
|
||
:
|
||
<bibRefCitation id="635D21B0FF97B1093922C80EB1426A4F" author="Lourenco" box="[322,542,1058,1082]" firstAuthor="Lourenco" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="93" pageNumber="93" pagination="359 - 365" part="40" refId="ref68211" refString="Lourenco, W. R. 2007. New considerations on the taxonomic status of the genus Physoctonus Mello-Leitao, 1934 (Scorpiones, Buthidae). Boletin de la Sociedad Entomologica Aragonesa 40: 359 - 365." title="New considerations on the taxonomic status of the genus Physoctonus Mello-Leitao, 1934 (Scorpiones, Buthidae)" type="journal article" year="2007">Lourenço, 2007: 360</bibRefCitation>
|
||
</taxonomicName>
|
||
, figs. 4–5,
|
||
<figureCitation id="9FF740C4FF97B10938AFC868B24F6A29" box="[207,275,1092,1116]" captionStart-0="FIG. 14" captionStart-1="FIG. 15" captionStart-10="FIG. 24" captionStart-11="FIG. 25" captionStart-2="FIG. 16" captionStart-3="FIG. 17" captionStart-4="FIG. 18" captionStart-5="FIG. 19" captionStart-6="FIG. 20" captionStart-7="FIG. 21" captionStart-8="FIG. 22" captionStart-9="FIG. 23" captionStartId-0="32.[107,149,1598,1619]" captionStartId-1="33.[108,150,1627,1648]" captionStartId-10="47.[107,149,1521,1542]" captionStartId-11="48.[108,150,1256,1277]" captionStartId-2="34.[109,151,1628,1649]" captionStartId-3="36.[106,149,1153,1174]" captionStartId-4="38.[107,149,1554,1575]" captionStartId-5="39.[108,151,1503,1524]" captionStartId-6="40.[108,151,1490,1511]" captionStartId-7="44.[107,149,1352,1373]" captionStartId-8="45.[108,147,1513,1534]" captionStartId-9="46.[109,151,1497,1518]" captionTargetBox-0="[220,1108,224,1568]" captionTargetBox-1="[235,1082,224,1599]" captionTargetBox-10="[107,1211,228,1487]" captionTargetBox-11="[108,1212,224,1227]" captionTargetBox-2="[208,1116,224,1599]" captionTargetBox-3="[106,634,224,1125]" captionTargetBox-4="[108,1212,224,1525]" captionTargetBox-5="[108,1212,227,1475]" captionTargetBox-6="[108,1212,224,1461]" captionTargetBox-7="[108,1212,224,1323]" captionTargetBox-8="[176,1145,224,1484]" captionTargetBox-9="[117,1187,233,1462]" captionTargetId-0="figure@32.[118,1207,224,1568]" captionTargetId-1="figure@33.[108,1209,224,1599]" captionTargetId-10="figure@47.[107,1211,228,1493]" captionTargetId-11="figure@48.[108,1212,224,1227]" captionTargetId-2="figure@34.[109,1212,224,1599]" captionTargetId-3="figure@36.[106,634,224,1125]" captionTargetId-4="figure@38.[108,1212,224,1526]" captionTargetId-5="figure@39.[108,1212,227,1475]" captionTargetId-6="figure@40.[108,1212,224,1462]" captionTargetId-7="figure@44.[108,1212,224,1323]" captionTargetId-8="figure@45.[108,1212,224,1484]" captionTargetId-9="figure@46.[109,1213,228,1467]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionTargetPageId-10="47" captionTargetPageId-11="48" captionTargetPageId-2="34" captionTargetPageId-3="36" captionTargetPageId-4="38" captionTargetPageId-5="39" captionTargetPageId-6="40" captionTargetPageId-7="44" captionTargetPageId-8="45" captionTargetPageId-9="46" captionText-0="FIG. 14. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. Heteroctenus garridoi (Armas, 1974), comb. nov., ♂ (AMNH). D. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). E. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." captionText-1="FIG. 15. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). B. Ischnotelson peruassu, sp. nov., paratype ♂ (MZSP). C. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♂ (MZSP). D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♀ (MZSP). E. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (AMNH). Scale bars = 2 mm." captionText-10="FIG. 24. Rhopalurusinae Bücherl, 1971, hemispermatophores, dorsal aspect (E), dorsal aspect, detail (B, H, M, P), ectal aspect (F, I, N, S), ectal aspect, detail (D, J, R, T, U), ental aspect (A, G, K, O), and ental aspect, detail (C, L, Q). A – D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E – H. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). I – M. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP). N – R. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (MZSP). S – U. Physoctonus debilis (C. L. Koch, 1840), ♂ (MZSP). Scale bars = 1 mm." captionText-11="FIG. 25. Heteroctenus Pocock, 1893, pedipalp chela, retrolateral aspect. A. H. abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. H. bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. H. garridoi (Armas, 1974), comb. nov., ♂ (AMNH). Scale bars = 2 mm." captionText-2="FIG. 16. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Physoctonus debilis (C. L. Koch, 1840), ♀ (MZSP). B. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). C. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). D. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. E. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Guatire, Venezuela. F. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). Scale bars = 1 mm (A, B), 2 mm (C – F)." captionText-3="FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." captionText-4="FIG. 18. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. Heteroctenus junceus (Herbst, 1800), comb. nov., ♂ (AMNH). D. Heteroctenus junceus (Herbst, 1800), ♀ (AMNH). E. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." captionText-5="FIG. 19. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♀ (MZSP). B. Ischnotelson peruassu, sp. nov., paratype ♂ (MZSP). C. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♂ (MZSP). D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). Scale bars = 2 mm." captionText-6="FIG. 20. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Physoctonus debilis (C. L. Koch, 1840), ♂ (MZSP). B. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). C. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). D. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. E. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Puerto Colombia, Venezuela. F. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). Scale bars = 2 mm." captionText-7="FIG. 21. Rhopalurusinae Bücherl, 1971, telson, lateral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C. L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." captionText-8="FIG. 22. Rhopalurusinae Bücherl, 1971, telson, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C. L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH). J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." captionText-9="FIG. 23. Rhopalurusinae Bücherl, 1971, hemispermatophores, dorsal aspect (E), dorsal aspect, detail (D, L, R), ectal aspect (A, F, H, O, T), ectal aspect, detail (B, K, M, V), ental aspect (G, I, P, S) and ental aspect, detail (C, J, N, Q, U). A – D. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). E – G. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). H – K. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). L – N. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). O – R. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). S – V. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♂ (MZSP). Scale bars = 1 mm." httpUri-0="https://zenodo.org/record/851410/files/figure.png" httpUri-1="https://zenodo.org/record/851412/files/figure.png" httpUri-10="https://zenodo.org/record/851430/files/figure.png" httpUri-11="https://zenodo.org/record/851432/files/figure.png" httpUri-2="https://zenodo.org/record/851414/files/figure.png" httpUri-3="https://zenodo.org/record/851416/files/figure.png" httpUri-4="https://zenodo.org/record/851418/files/figure.png" httpUri-5="https://zenodo.org/record/851420/files/figure.png" httpUri-6="https://zenodo.org/record/851422/files/figure.png" httpUri-7="https://zenodo.org/record/851424/files/figure.png" httpUri-8="https://zenodo.org/record/851426/files/figure.png" httpUri-9="https://zenodo.org/record/851428/files/figure.png" pageId="93" pageNumber="93">14–25</figureCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FF97B109397EC868B1706A29" author="Prendini" box="[286,556,1092,1116]" etAl="et al." firstAuthor="Prendini" journalOrPublisher="Journal of Arachnology" pageId="93" pageNumber="93" pagination="206 - 224" part="37" refId="ref70522" refString="Prendini, L., L. A. Esposito, J. Huff, and E. S. Volschenk. 2009. Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola. Journal of Arachnology 37: 206 - 224." title="Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola" type="journal article" year="2009">Prendini et al., 2009: 222</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FF97B1093A57C868B2326A0A" author="Brazil and Porto" bookContentInfo="84 pp." firstAuthor="Brazil" journalOrPublisher="Salvador, BA, Brazil: EDUFBA" pageId="93" pageNumber="93" refId="ref64569" refString="Brazil, T. K., and Porto, T. J. 2010. Os escorpioes. Salvador, BA, Brazil: EDUFBA. 84 pp." title="Os escorpioes" type="book" year="2010">Brazil and Porto, 2010: 50</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="635D21B0FF97B1093919C84BB1396A0A" author="Porto" box="[377,613,1127,1151]" etAl="et al." firstAuthor="Porto" journalOrPublisher="Check List" pageId="93" pageNumber="93" pagination="292 - 297" part="6" refId="ref70138" refString="Porto, T. J., T. K. Brazil, and R. M. Lira-da-Silva. 2010. Scorpions, state of Bahia, northeastern Brazil. Check List 6 (2): 292 - 297." title="Scorpions, state of Bahia, northeastern Brazil" type="journal article" year="2010">Porto et al., 2010: 293</bibRefCitation>
|
||
, 295, fig. 1F,
|
||
<tableCitation id="4A4E69FAFF97B109397CC8A5B2386AD7" box="[284,356,1161,1186]" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="93" pageNumber="93" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFF97B10938EBC892B16B6B6C" pageId="93" pageNumber="93" type="materials_examined">
|
||
<paragraph id="07735C41FF97B10938EBC892B16B6B6C" blockId="93.[107,637,1214,1741]" pageId="93" pageNumber="93">
|
||
TYPE MATERIAL:
|
||
<emphasis id="35B88053FF97B1093936C892B2E26AA3" bold="true" box="[342,446,1214,1238]" pageId="93" pageNumber="93">BRAZIL:</emphasis>
|
||
<taxonomicName id="C0CC27C2FF97B10939ABC892B12A6AA3" authorityName="C. L. Koch" authorityYear="1840" box="[459,630,1214,1238]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B10939ABC892B12A6AA3" box="[459,630,1214,1238]" italics="true" pageId="93" pageNumber="93">Vaejovis debilis</emphasis>
|
||
</taxonomicName>
|
||
: Holotype?♀ (ZSM) [lost].
|
||
<taxonomicName id="C0CC27C2FF97B10939F1C8F3B12A6A82" authorityName="Mello-Leitao" authorityYear="1934" box="[401,630,1247,1271]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="physurus">
|
||
<emphasis id="35B88053FF97B10939F1C8F3B12A6A82" box="[401,630,1247,1271]" italics="true" pageId="93" pageNumber="93">Physoctonus physurus</emphasis>
|
||
</taxonomicName>
|
||
: Holotype ♂ (MNRJ),
|
||
<emphasis id="35B88053FF97B1093937C92DB2F46B6C" box="[343,424,1281,1305]" italics="true" pageId="93" pageNumber="93">Paraíba</emphasis>
|
||
: Santa Luzia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFF97B10938EBC90EB0006C33" pageId="93" pageNumber="93" type="diagnosis">
|
||
<paragraph id="07735C41FF97B10938EBC90EB0006C33" blockId="93.[107,637,1214,1741]" lastBlockId="93.[684,1214,228,1740]" pageId="93" pageNumber="93">
|
||
DIAGNOSIS:
|
||
<taxonomicName id="C0CC27C2FF97B1093976C90EB2B16B4F" authorityName=": Lourenco" authorityYear="2007" box="[278,493,1314,1338]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B1093976C90EB2B16B4F" box="[278,493,1314,1338]" italics="true" pageId="93" pageNumber="93">Physoctonus debilis</emphasis>
|
||
</taxonomicName>
|
||
differs from its sister species,
|
||
<taxonomicName id="C0CC27C2FF97B109394CC969B2C46B29" authority="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[300,408,1349,1372]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
|
||
<emphasis id="35B88053FF97B109394CC969B2C46B29" box="[300,408,1349,1372]" italics="true" pageId="93" pageNumber="93">P. striatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="2E8B3D28FF97B10939C6C968B2A16B2E" box="[422,509,1348,1371]" pageId="93" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, as follows. The mesosomal tergites of
|
||
<taxonomicName id="C0CC27C2FF97B10939CEC94AB14D6B08" authorityName=": Lourenco" authorityYear="2007" box="[430,529,1381,1405]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B10939CEC94AB14D6B08" box="[430,529,1381,1405]" italics="true" pageId="93" pageNumber="93">P. debilis</emphasis>
|
||
</taxonomicName>
|
||
are uniformly pale, except for the infuscate dorsomedian carinae, which form a thin stripe longitudinally, whereas the tergites of
|
||
<emphasis id="35B88053FF97B1093A45C9E7B3D16876" italics="true" pageId="93" pageNumber="93">
|
||
<taxonomicName id="C0CC27C2FF97B1093A45C9E7B1336B97" box="[549,623,1483,1506]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="stria">P. stria</taxonomicName>
|
||
- tus
|
||
</emphasis>
|
||
are darker, with transverse bands of infuscation. The intercarinal surfaces of the carapace and metasomal segments II–IV are more coarsely and densely granular in
|
||
<taxonomicName id="C0CC27C2FF97B1093A7DCA7DB125681D" authorityName=": Lourenco" authorityYear="2007" box="[541,633,1616,1640]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B1093A7DCA7DB125681D" box="[541,633,1616,1640]" italics="true" pageId="93" pageNumber="93">P. debilis</emphasis>
|
||
</taxonomicName>
|
||
but weakly granular to smooth in
|
||
<taxonomicName id="C0CC27C2FF97B1093A64CA5EB12968FC" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[516,629,1650,1673]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="35B88053FF97B1093A64CA5EB12968FC" box="[516,629,1650,1673]" italics="true" pageId="93" pageNumber="93">P. striatus</emphasis>
|
||
</taxonomicName>
|
||
. The carinae of the carapace, pedipalps, and metasoma are more pronounced in
|
||
<taxonomicName id="C0CC27C2FF97B1093A77CA99B12568B8" authorityName=": Lourenco" authorityYear="2007" box="[535,633,1717,1741]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B1093A77CA99B12568B8" box="[535,633,1717,1741]" italics="true" pageId="93" pageNumber="93">P. debilis</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="C0CC27C2FF97B1093B73CCC9B0D66E89" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[787,906,229,252]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="35B88053FF97B1093B73CCC9B0D66E89" box="[787,906,229,252]" italics="true" pageId="93" pageNumber="93">P. striatus</emphasis>
|
||
</taxonomicName>
|
||
: for example, the lateral inframedian carinae of metasomal segments II and III are complete in
|
||
<taxonomicName id="C0CC27C2FF97B1093BDBCD0BB7466F4B" authorityName=": Lourenco" authorityYear="2007" box="[955,1050,294,318]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B1093BDBCD0BB7466F4B" box="[955,1050,294,318]" italics="true" pageId="93" pageNumber="93">P. debilis</emphasis>
|
||
</taxonomicName>
|
||
but restricted to the posterior third of the segment in
|
||
<emphasis id="35B88053FF97B1093C09CD64B1916FF5" italics="true" pageId="93" pageNumber="93">
|
||
<taxonomicName id="C0CC27C2FF97B1093C09CD64B7EC6F2A" box="[1129,1200,328,351]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="stria">P. stria</taxonomicName>
|
||
- tus
|
||
</emphasis>
|
||
; and the retromedian and secondary accessory carinae of the pedipalp chela manus are complete and granular in
|
||
<taxonomicName id="C0CC27C2FF97B1093BAFCD87B7776FB7" authorityName=": Lourenco" authorityYear="2007" box="[975,1067,426,450]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B1093BAFCD87B7776FB7" box="[975,1067,426,450]" italics="true" pageId="93" pageNumber="93">P. debilis</emphasis>
|
||
</taxonomicName>
|
||
but vestigial or absent in
|
||
<taxonomicName id="C0CC27C2FF97B1093B28CDE0B0E06F96" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[840,956,460,483]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="35B88053FF97B1093B28CDE0B0E06F96" box="[840,956,460,483]" italics="true" pageId="93" pageNumber="93">P. striatus</emphasis>
|
||
</taxonomicName>
|
||
. Pedipalp chela fixed finger trichobothrium
|
||
<emphasis id="35B88053FF97B1093BD7CDC1B08F6C70" box="[951,979,493,517]" italics="true" pageId="93" pageNumber="93">db</emphasis>
|
||
is situated distal to trichobothrium
|
||
<emphasis id="35B88053FF97B1093B04CE23B0246C53" box="[868,888,527,550]" italics="true" pageId="93" pageNumber="93">et</emphasis>
|
||
in
|
||
<taxonomicName id="C0CC27C2FF97B1093BFDCE23B0A66C53" authorityName=": Lourenco" authorityYear="2007" box="[925,1018,526,550]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B1093BFDCE23B0A66C53" box="[925,1018,526,550]" italics="true" pageId="93" pageNumber="93">P. debilis</emphasis>
|
||
</taxonomicName>
|
||
but aligned with
|
||
<emphasis id="35B88053FF97B1093ACCCE1CB19C6C32" box="[684,704,560,583]" italics="true" pageId="93" pageNumber="93">et</emphasis>
|
||
in
|
||
<taxonomicName id="C0CC27C2FF97B1093A87CE1CB0096C32" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[743,853,560,583]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="35B88053FF97B1093A87CE1CB0096C32" box="[743,853,560,583]" italics="true" pageId="93" pageNumber="93">P. striatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFF97B1093AACCE7CB7F96CDF" pageId="93" pageNumber="93" type="distribution">
|
||
<paragraph id="07735C41FF97B1093AACCE7CB7F96CDF" blockId="93.[684,1214,228,1740]" pageId="93" pageNumber="93">
|
||
DISTRIBUTION:
|
||
<taxonomicName id="C0CC27C2FF97B1093B0ECE7CB7736C1D" authorityName=": Lourenco" authorityYear="2007" box="[878,1071,592,616]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B1093B0ECE7CB7736C1D" box="[878,1071,592,616]" italics="true" pageId="93" pageNumber="93">Physoctonus debilis</emphasis>
|
||
</taxonomicName>
|
||
is endemic to Brazil, where it has been recorded in the states of Ceará, Paraíba, Pernambuco, and Piauí (fig. 9B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFF97B1093AACCE9FB7436D05" pageId="93" pageNumber="93" type="biology_ecology">
|
||
<paragraph id="07735C41FF97B1093AACCE9FB7436D05" blockId="93.[684,1214,228,1740]" pageId="93" pageNumber="93">
|
||
ECOLOGY: The known locality records are situated in the Brazilian caatinga (fig. 2D). Specimens collected by the authors were found under stones during the day or with UV light detection at night. The habitus is consistent with the lapidicolous ecomorphotype (
|
||
<bibRefCitation id="635D21B0FF97B1093B03CF74B7596D05" author="Prendini" box="[867,1029,856,880]" firstAuthor="Prendini" journalOrPublisher="Journal of Arachnology" pageId="93" pageNumber="93" pagination="173 - 188" part="29" refId="ref70328" refString="Prendini, L. 2001 a. Further additions to the scorpion fauna of Trinidad and Tobago. Journal of Arachnology 29 (2): 173 - 188." title="Further additions to the scorpion fauna of Trinidad and Tobago" type="journal article" year="2001">Prendini, 2001</bibRefCitation>
|
||
b).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFF97B1093AACCF56B7346ACE" pageId="93" pageNumber="93" type="discussion">
|
||
<paragraph id="07735C41FF97B1093AACCF56B7346ACE" blockId="93.[684,1214,228,1740]" pageId="93" pageNumber="93">
|
||
REMARKS:
|
||
<taxonomicName id="C0CC27C2FF97B1093B27CF55B74A6DE4" authorityName=": Lourenco" authorityYear="2007" box="[839,1046,889,913]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B1093B27CF55B74A6DE4" box="[839,1046,889,913]" italics="true" pageId="93" pageNumber="93">Physoctonus debilis</emphasis>
|
||
</taxonomicName>
|
||
was originally placed in the nonbuthid genus
|
||
<taxonomicName id="C0CC27C2FF97B1093B92CFB7B1BC6DA1" authority="C. L. Koch, 1836" authorityName="C. L. Koch" authorityYear="1836" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FF97B1093B92CFB7B71A6DC7" box="[1010,1094,923,946]" italics="true" pageId="93" pageNumber="93">Vaejovis</emphasis>
|
||
C.L. Koch, 1836
|
||
</taxonomicName>
|
||
, but was transferred to
|
||
<taxonomicName id="C0CC27C2FF97B1093BBACF90B70C6DA1" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[986,1104,956,980]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FF97B1093BBACF90B70C6DA1" box="[986,1104,956,980]" italics="true" pageId="93" pageNumber="93">Rhopalurus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="635D21B0FF97B1093C18CF90B1AF6D80" author="Borelli" firstAuthor="Borelli" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Realle Universita di Torino" pageId="93" pageNumber="93" pagination="1 - 8" part="25" refId="ref64441" refString="Borelli, A. 1910. Scorpioni nuovi e poco noti del Brasile. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Realle Universita di Torino 25 (629): 1 - 8." title="Scorpioni nuovi e poco noti del Brasile" type="journal article" year="1910">Borelli (1910)</bibRefCitation>
|
||
, where it remained until
|
||
<bibRefCitation id="635D21B0FF97B1093C6ACFF1B7E16D80" author="Lourenco" bookContentInfo="306 pp." box="[1034,1213,989,1013]" firstAuthor="Lourenco" journalOrPublisher="Paris: Les Editions de l'If" pageId="93" pageNumber="93" refId="ref68189" refString="Lourenco, W. R. 2002. Scorpions of Brazil. Paris: Les Editions de l'If. 306 pp." title="Scorpions of Brazil" type="book" year="2002">Lourenço (2002)</bibRefCitation>
|
||
revalidated the genus
|
||
<taxonomicName id="C0CC27C2FF97B1093BEFCFD2B7506A63" authorityName="Mello-Leitao" authorityYear="1934" box="[911,1036,1022,1046]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35B88053FF97B1093BEFCFD2B7506A63" box="[911,1036,1022,1046]" italics="true" pageId="93" pageNumber="93">Physoctonus</emphasis>
|
||
</taxonomicName>
|
||
. As noted by
|
||
<bibRefCitation id="635D21B0FF97B1093CFDCFD2B03A6A42" author="Fet and Lowe" editor="V. Fet & W. D. Sissom" firstAuthor="Fet" journalOrPublisher="New York: New York Entomological Society" pageId="93" pageNumber="93" pagination="54 - 286" refId="ref65304" refString="Fet, V., and G. Lowe. 2000. Family Buthidae C. L. Koch, 1837. In V. Fet, W. D. Sissom, G., Lowe, and M. E. Braunwalder, Catalog of the scorpions of the world (1758 - 1998): 54 - 286. New York: New York Entomological Society." title="Family Buthidae C. L. Koch, 1837" type="book chapter" volumeTitle="G., Lowe, and M. E" year="2000">Fet and Lowe (2000)</bibRefCitation>
|
||
, the true date of publication of C.L. Koch’s name is 1840 (see
|
||
<bibRefCitation id="635D21B0FF97B1093B8FC86CB7DA6A2D" author="Brignoli" box="[1007,1158,1088,1112]" firstAuthor="Brignoli" journalOrPublisher="Bulletin of the British Arachnological Society" pageId="93" pageNumber="93" pagination="414 - 416" part="6" refId="ref64599" refString="Brignoli, P. M. 1985. On the correct dates of publication of the arachnid taxa described in some works by C. W. Hahn and C. L. Koch (Arachnida). Bulletin of the British Arachnological Society 6 (9): 414 - 416." title="On the correct dates of publication of the arachnid taxa described in some works by C. W. Hahn and C. L. Koch (Arachnida)" type="journal article" year="1985">Brignoli, 1985</bibRefCitation>
|
||
), not 1841 as commonly quoted. This species has a junior homonym,
|
||
<taxonomicName id="C0CC27C2FF97B1093B4CC8AFB7256AEF" authority="L. Koch, 1865" authorityName="L. Koch" authorityYear="1865" box="[812,1145,1154,1178]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="93" pageNumber="93" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF97B1093B4CC8AFB08C6AEF" box="[812,976,1154,1178]" italics="true" pageId="93" pageNumber="93">Vaejovis debilis</emphasis>
|
||
L. Koch, 1865
|
||
</taxonomicName>
|
||
, from Mexico, the identity of which is unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFF97B10A3AACC8E8B2696FF4" lastPageId="94" lastPageNumber="94" pageId="93" pageNumber="93" type="materials_examined">
|
||
<paragraph id="07735C41FF97B10A3AACC8E8B2696FF4" blockId="93.[684,1214,228,1740]" lastBlockId="94.[107,636,225,385]" lastPageId="94" lastPageNumber="94" pageId="93" pageNumber="93">
|
||
MATERIAL EXAMINED:
|
||
<emphasis id="35B88053FF97B1093BA7C8E8B7776AA9" bold="true" box="[967,1067,1220,1244]" pageId="93" pageNumber="93">BRAZIL:</emphasis>
|
||
<emphasis id="35B88053FF97B1093C55C8E8B7EB6AA9" box="[1077,1207,1220,1244]" italics="true" pageId="93" pageNumber="93">Pernambuco</emphasis>
|
||
: Exu, 18 km N, 5.iii.1977, L.J. Vitt, under leaf of granite on boulder, caatinga habitat, 1 ♀ (AMNH); Exu, 5 km N, 4.x.1977, L.J. Vitt, 1 ♀ (AMNH), 18.i.1978, L.J. Vitt and K.E. Streilein, 1 ♀ (AMNH); Fazenda Batente, 13 km E Exu, 10.xi.1977, L.J. Vitt and K.E. Streilein, 1 ♀ (AMNH); Fazenda Caterino, 10 km NE Exu, 9.vii.1977, L.J. Vitt, 1 ♀ (AMNH), 25.ix.1977, L.J. Vitt, 1 ♀ (AMNH).
|
||
<emphasis id="35B88053FF97B1093CE1C9E1B7EA6B91" box="[1153,1206,1485,1508]" italics="true" pageId="93" pageNumber="93">Piauí</emphasis>
|
||
: Castelo do Piauí, 05°13′43″S 41°41′57″W, 13.viii.2008, R. Pinto-da-Rocha et al., 2 ♀ (MZSP 30868/AMCC [LP 9680]), MZSP 31158/AMCC [LP 9931]); Oeiras, 06°58′28″S 42°06′31″W, 2–3. vi.2008, H.Y. Yamaguti et al., 1 ♀ (MZSP 31162/ AMCC [LP 9940]), 1 subad. ♀ (MZSP 31164/ AMCC [LP 9962]), 3.vi.2008, H.Y. Yamaguti et al., 1 ♀ (MZSP 30866/AMCC [LP 9678]); near Parque Nacional Sete Cidades, Brasileira e Piracuruca, 04°10′0 2″S 41°41′56.7″W, 16.viii.2008, R. Pinto-da- Rocha and L.S. Carvalho, 1 ♀ (MZSP 30867/ AMCC [LP 9679]).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |