395 lines
47 KiB
XML
395 lines
47 KiB
XML
<document ID-DOI="10.1038/s41598-021-96221-x" ID-GBIF-Dataset="11088aaa-ac64-4b8f-bb4f-5ab4dd5baa6f" ID-PMC="PMC8421421" ID-PubMed="34489502" ID-Zenodo-Dep="5645747" approvalRequired="62" approvalRequired_for_document="1" approvalRequired_for_taxonomicNames="29" approvalRequired_for_treatments="32" checkinTime="1636046587746" checkinUser="valdenar" docAuthor="Mennecart, Bastien, Aiglstorfer, Manuela, Li, Yikun, Li, Chunxiao & Wang, ShiQi" docDate="2021" docId="E97287E44C457A081F8AFA3DFA72CBF7" docLanguage="en" docName="SciRep.11.17710.pdf" docOrigin="Scientific Reports 1" docTitle="Iberomeryx miaoi Mennecart & Aiglstorfer & Li & Li & Wang 2021, nov.sp." docType="treatment" docVersion="4" lastPageNumber="8" masterDocId="154BFF9C4C437A0F1E4AFFE2FFB5C812" masterDocTitle="Ruminants reveal Eocene Asiatic palaeobiogeographical provinces as the origin of diachronous mammalian Oligocene dispersals into Europe" masterLastPageNumber="12" masterPageNumber="1" pageNumber="7" updateTime="1668144176883" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Ruminants reveal Eocene Asiatic palaeobiogeographical provinces as the origin of diachronous mammalian Oligocene dispersals into Europe</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mennecart, Bastien</mods:namePart>
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<mods:affiliation>Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria & Naturhistorisches Museum Basel, Augustinergasse 2, 4001 Basel, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="email">mennecartbastien@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Aiglstorfer, Manuela</mods:namePart>
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<mods:affiliation>Naturhistorisches Museum Mainz / Landessammlung Für Naturkunde Rheinland-Pfalz, Reichklarastrasse 10, 55116 Mainz, Germany.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Yikun</mods:namePart>
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<mods:affiliation>Center for Research and Education On Biological Evolution and Environment, Nanjing Univeristy, Nanjing 210023, China.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Chunxiao</mods:namePart>
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<mods:affiliation>Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China & CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wang, ShiQi</mods:namePart>
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<mods:affiliation>Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China & CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China</mods:affiliation>
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<mods:nameIdentifier type="email">wangshiqi@ivpp.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Scientific Reports</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-09-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>12</mods:end>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1038/s41598-021-96221-x</mods:identifier>
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<mods:identifier type="GBIF-Dataset">11088aaa-ac64-4b8f-bb4f-5ab4dd5baa6f</mods:identifier>
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<mods:identifier type="PMC">PMC8421421</mods:identifier>
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<mods:identifier type="PubMed">34489502</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5645747</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5645765" ID-GBIF-Taxon="189990180" ID-Zenodo-Dep="5645765" LSID="urn:lsid:plazi:treatment:E97287E44C457A081F8AFA3DFA72CBF7" httpUri="http://treatment.plazi.org/id/E97287E44C457A081F8AFA3DFA72CBF7" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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<subSubSection box="[448,695,1503,1525]" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph blockId="6.[448,926,1503,1579]" box="[448,695,1503,1525]" pageId="6" pageNumber="7">
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<taxonomicName authority="Mennecart & Aiglstorfer & Li & Li & Wang, 2021" authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" box="[448,617,1503,1525]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="miaoi" status="nov.sp.">
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<emphasis box="[448,617,1503,1525]" italics="true" pageId="6" pageNumber="7">Iberomeryx miaoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[622,695,1504,1525]" pageId="6" pageNumber="7" rank="species">nov.sp.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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||
<subSubSection box="[448,689,1530,1552]" pageId="6" pageNumber="7" type="description">
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||
<paragraph blockId="6.[448,926,1503,1579]" box="[448,689,1530,1552]" pageId="6" pageNumber="7">
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<figureCitation box="[448,545,1530,1552]" captionStart="Figure 1" captionStartId="2.[416,470,1525,1544]" captionTargetBox="[422,1363,140,1491]" captionTargetId="figure-31@2.[416,1368,135,1496]" captionTargetPageId="2" captionText="Figure 1. Dentition of Krabimeryx gracilis nov. comb. (Miao, 1982)20 (A, B, G, H), Chiyoumeryx nov. gen. shinaoensis (Miao, 1982)20 (C, D), Chiyoumeryx nov. gen. flavimperatoris nov. sp. (E) and Iberomeryx miaoi nov. sp. (F–I). Krabimeryx gracilis nov. comb.(Miao, 1982)20: (A) IVPP V 6546-1 (holotype), partial skull with right and lef M1–M3; (B) IVPP V 6546-2 (holotype), right fragmented mandible with m2–m3. Chiyoumeryx nov.gen. shinaoensis (Miao, 1982)20: (C) IVPP V 6531 (holotype), right mandible with p2–m3 and tooth socket of p1; (D) IVPP V 6532 (paratype), right fragmented maxillary with P4-M3. Chiyoumeryx nov. gen. flavimperatoris nov. sp.: (E) IVPP V 6547 (holotype), right mandible with p4–m3; Iberomeryx miaoi nov. sp.: (F) IVPP V 6551 (holotype), lef mandible with m1–m3 (mirrored); (G) lower molar Lophiomerycidae dental nomenclature (based on the m3 of IVPP 6546-2): 1 internal postmetacristid, 2 metaconid, 3 external postmetacristid, 4 internal preentocristid, 5 entoconidian groove, 6 external preentocristid, 7 entoconid, 8 posthypoconulidcristid, 9 hypoconulid, 10 prehypoconuldicristid, 11 posthypocristid, 12 hypoconid, 13 prehypocristid, 14 ectostylid, 15 postprotocristid, 16 protoconid, 17 preprotocristid, 18 anterior cingulid; (H) upper molar Lophiomerycidae dental nomenclature (based on the M2 of IVPP 6546-1): 1 postmetacrista, 2 metacone, 3 premetacrista, 4 mesostyle, 5 postparacrista, 6 paracone, 7 paraconid labial groove, 8 preparacrista, 9 parastyle, 10 preprotocrista, 11 anterolingual cingulum, 12 protocone, 13 postprotocrista, 14 entostyle, 15 additional cone, 16 premetaconulecrista, 17 metaconule, 18 postmetaconulecrista; (I) lower molar Tragulidae dental nomenclature (based on the m2 of IVPP V 6551, reversed): 1 metaconid, 2 external postmetacristid, 3 Dorcatherium fold, 4 internal postmetacristid, 5 preentocristid, 6 entoconid, 7 postentocristid, 8 posterior cingulid, 9 posthypocristid, 10 hypoconid, 11 prehypocristid, 12 ectostylid, 13 external postprotocristid, 14 Tragulus fold, 15 internal postprotocristid, 16 protoconid, 17 preprotocristid, 18 paraconid, 19 preparacristid. (J) phylogenetic position and stratigraphie of the Shinao/ Yangjiachong/Xiaerhete ruminants (topology2). a stem Ruminantia, b Archaeomeryx, c Chiyoumeryx nov.gen. and Krabimeryx gracilis, d crown Ruminantia, e Iberomeryx miaoi nov. sp.; 1 lingual view, 2 occlusal view. Scale bare is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5645749" httpUri="https://zenodo.org/record/5645749/files/figure.png" pageId="6" pageNumber="7">Figure 1D</figureCitation>
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and Figure S4.
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</paragraph>
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</subSubSection>
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<subSubSection box="[448,926,1554,1579]" pageId="6" pageNumber="7" type="reference_group">
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<paragraph blockId="6.[448,926,1503,1579]" box="[448,926,1554,1579]" pageId="6" pageNumber="7">
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v 1982
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<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[516,714,1557,1578]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[516,714,1557,1578]" italics="true" pageId="6" pageNumber="7">Lophiomeryx gracilis</emphasis>
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</taxonomicName>
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?—Miao: 536, Fig.
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<superScript attach="none" box="[906,921,1554,1568]" fontSize="6" pageId="6" pageNumber="7">820</superScript>
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.
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</paragraph>
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</subSubSection>
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<subSubSection box="[416,1295,1614,1638]" pageId="6" pageNumber="7" type="description">
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<paragraph blockId="6.[416,1295,1614,1638]" box="[416,1295,1614,1638]" pageId="6" pageNumber="7">
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<heading bold="true" box="[416,588,1614,1638]" fontSize="10" level="2" pageId="6" pageNumber="7" reason="0">
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<emphasis bold="true" box="[416,588,1614,1638]" pageId="6" pageNumber="7">ZooBank LSID.</emphasis>
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</heading>
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urn:lsid:zoobank.org:act:EE3F88E9-0EAF-4EC6-A46F-8623241E614B.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="7" type="diagnosis">
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<paragraph blockId="6.[416,1479,1673,1829]" pageId="6" pageNumber="7">
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<emphasis bold="true" box="[416,536,1673,1697]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
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<taxonomicName box="[562,670,1674,1695]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
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<emphasis box="[562,670,1674,1695]" italics="true" pageId="6" pageNumber="7">Iberomeryx</emphasis>
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</taxonomicName>
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with a very large paraconid, which is smaller in
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<emphasis box="[1148,1320,1674,1696]" italics="true" pageId="6" pageNumber="7">
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<taxonomicName box="[1148,1256,1674,1695]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">Iberomeryx</taxonomicName>
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minor
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</emphasis>
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and
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<emphasis italics="true" pageId="6" pageNumber="7">
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<taxonomicName box="[1370,1478,1674,1695]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">Iberomeryx</taxonomicName>
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parvus
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</emphasis>
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. Te metastylid is not strong but is more developed than in the other species. Te ectostylid is big on m1, smaller on m2 and absent on m3, while
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<emphasis box="[807,884,1728,1749]" italics="true" pageId="6" pageNumber="7">I. minor</emphasis>
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displays an ectostylid on all molars and
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<emphasis box="[1279,1364,1728,1749]" italics="true" pageId="6" pageNumber="7">I. parvus</emphasis>
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none at all.
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<taxonomicName authority="Mennecart & Aiglstorfer & Li & Li & Wang, 2021" authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" box="[416,584,1754,1776]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="miaoi" status="nov. sp.">
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<emphasis box="[416,584,1754,1776]" italics="true" pageId="6" pageNumber="7">Iberomeryx miaoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[590,663,1755,1776]" pageId="6" pageNumber="7" rank="species">nov. sp.</taxonomicNameLabel>
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is of similar size to
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<emphasis box="[857,934,1755,1776]" italics="true" pageId="6" pageNumber="7">I. minor</emphasis>
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and its m2 is smaller than the one of
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<emphasis box="[1301,1385,1755,1776]" italics="true" pageId="6" pageNumber="7">I. parvus</emphasis>
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. It differs from
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<emphasis box="[468,544,1781,1802]" italics="true" pageId="6" pageNumber="7">I. minor</emphasis>
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by a thin anterior cingulid. Moreover, its protoconid is positioned slightly more anterior than in
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<emphasis italics="true" pageId="6" pageNumber="7">I. parvus</emphasis>
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. Te molars appear to be more massive and bulkier in this species than in
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<emphasis box="[1191,1267,1808,1829]" italics="true" pageId="6" pageNumber="7">I. minor</emphasis>
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and
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<emphasis box="[1314,1404,1808,1829]" italics="true" pageId="6" pageNumber="7">I. parvus.</emphasis>
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</paragraph>
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<paragraph blockId="6.[416,1478,1865,1914]" pageId="6" pageNumber="7">
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<heading bold="true" box="[416,531,1865,1889]" fontSize="10" level="2" pageId="6" pageNumber="7" reason="0">
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<emphasis bold="true" box="[416,531,1865,1889]" pageId="6" pageNumber="7">Holotype.</emphasis>
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</heading>
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<materialsCitation ID-GBIF-Occurrence="3398234324" collectionCode="IVPP" pageId="6" pageNumber="7" specimenCode="IVPP V 6551" specimenCount="1">
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<specimenCode box="[557,686,1866,1888]" pageId="6" pageNumber="7">IVPP V 6551</specimenCode>
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, lef mandible with m1–m3 (only specimen known). m1 5.1 × 3.5, m2 5.2 × 4.1, m3 8.0× 4.0.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection box="[1467,1478,2024,2043]" pageId="6" pageNumber="7" type="description">
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<paragraph blockId="6.[1467,1478,2024,2043]" box="[1467,1478,2024,2043]" pageId="6" pageNumber="7">7</paragraph>
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</subSubSection>
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<subSubSection box="[416,1450,137,161]" pageId="7" pageNumber="8" type="etymology">
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<paragraph blockId="7.[416,1450,137,161]" box="[416,1450,137,161]" pageId="7" pageNumber="8">
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<heading bold="true" box="[416,549,137,161]" fontSize="10" level="2" pageId="7" pageNumber="8" reason="0">
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<emphasis bold="true" box="[416,549,137,161]" pageId="7" pageNumber="8">Etymology.</emphasis>
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</heading>
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We dedicate this species to Prof. Miao Desui who was the first to describe the Shinao fauna.
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</paragraph>
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</subSubSection>
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<subSubSection box="[416,1407,196,220]" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph blockId="7.[416,1407,196,220]" box="[416,1407,196,220]" pageId="7" pageNumber="8">
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<heading bold="true" box="[416,658,196,220]" fontSize="10" level="2" pageId="7" pageNumber="8" reason="0">
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<emphasis bold="true" box="[416,658,196,220]" pageId="7" pageNumber="8">Locality and horizon.</emphasis>
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</heading>
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<materialsCitation ID-GBIF-Occurrence="3398234311" box="[684,1403,197,219]" country="China" county="Panxian County" location="Shinao Basin" pageId="7" pageNumber="8" specimenCount="1">
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<location LSID="urn:lsid:plazi:treatment:E97287E44C457A081F8AFA3DFA72CBF7:640460294C447A081CE6FF27FC9EC8C9" box="[684,811,197,219]" country="China" county="Panxian County" name="Shinao Basin" pageId="7" pageNumber="8">Shinao Basin</location>
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,
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<collectingCounty box="[820,975,197,219]" pageId="7" pageNumber="8">Panxian County</collectingCounty>
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,
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<locationDeviation box="[984,1205,197,219]" pageId="7" pageNumber="8">Southwestern Guizhou</locationDeviation>
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,
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<collectingCountry box="[1215,1276,197,219]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
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.
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<geologicalTimeScale box="[1284,1403,197,218]" pageId="7" pageNumber="8">Late Eocene</geologicalTimeScale>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="description">
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<paragraph blockId="7.[416,1479,253,997]" pageId="7" pageNumber="8">
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<emphasis bold="true" box="[416,702,254,278]" pageId="7" pageNumber="8">Taxonomical attribution.</emphasis>
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Tis minute ruminant was referred to
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<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[1103,1300,255,276]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="gracilis">
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<emphasis box="[1103,1300,255,276]" italics="true" pageId="7" pageNumber="8">Lophiomeryx gracilis</emphasis>
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</taxonomicName>
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? by Miao
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<bibRefCitation author="Miao, D." box="[1396,1411,253,267]" journalOrPublisher="Acta Palaeontol. Sinica" pageId="7" pageNumber="8" pagination="526 - 536" part="21" refId="ref9719" refString="20. Miao, D. Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province. Acta Palaeontol. Sinica 21, 526 - 536 (1982)." title="Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province" type="journal article" year="1982">
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<superScript attach="right" box="[1396,1411,253,267]" fontSize="6" pageId="7" pageNumber="8">20</superScript>
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</bibRefCitation>
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. However, he already noticed that the size of this individual was smaller than in the other specimens attributed to
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<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[416,612,309,330]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="gracilis">
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||
<emphasis box="[416,612,309,330]" italics="true" pageId="7" pageNumber="8">Lophiomeryx gracilis</emphasis>
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||
</taxonomicName>
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||
.Miao
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||
<bibRefCitation author="Miao, D." box="[674,689,306,320]" journalOrPublisher="Acta Palaeontol. Sinica" pageId="7" pageNumber="8" pagination="526 - 536" part="21" refId="ref9719" refString="20. Miao, D. Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province. Acta Palaeontol. Sinica 21, 526 - 536 (1982)." title="Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province" type="journal article" year="1982">
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<superScript attach="left" box="[674,689,306,320]" fontSize="6" pageId="7" pageNumber="8">20</superScript>
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||
</bibRefCitation>
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||
excluded an attribution of IVPP V 6551 to "
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<emphasis box="[1117,1328,309,330]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName box="[1117,1242,309,330]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Lophiomeryx</taxonomicName>
|
||
" gaudryi
|
||
</emphasis>
|
||
due to a closed posterior section of the posterior fossa on the m3. However, in both teeth, the posterior fossa is still open by the reduction of the postentocristid.
|
||
</paragraph>
|
||
<paragraph blockId="7.[416,1479,253,997]" pageId="7" pageNumber="8">
|
||
Te here-described specimen clearly differs from
|
||
<taxonomicName box="[915,1039,389,410]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[915,1039,389,410]" italics="true" pageId="7" pageNumber="8">Lophiomeryx</emphasis>
|
||
</taxonomicName>
|
||
by the presence of an external postmetacristid forming a slight
|
||
<taxonomicName box="[574,707,415,436]" class="Mammalia" family="Tragulidae" genus="Dorcatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[574,707,415,436]" italics="true" pageId="7" pageNumber="8">Dorcatherium</emphasis>
|
||
</taxonomicName>
|
||
fold, a developed external postprotocristid (clearly visible at least on m2), and a largeparaconid
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[567,582,439,453]" journalOrPublisher="Swiss J. Geosci." pageId="7" pageNumber="8" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">
|
||
<superScript attach="left" box="[567,582,439,453]" fontSize="6" pageId="7" pageNumber="8">36</superScript>
|
||
</bibRefCitation>
|
||
. Furthermore the external postprotocristid and prehypocristid are connected on their distal ends and the third basin of m3 forms a well-formed buckle, unlike the condition in
|
||
<taxonomicName authorityName="Janis" authorityYear="1987" box="[1212,1382,469,491]" class="Mammalia" family="Lophiomerycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Lophiomerycidae</taxonomicName>
|
||
<superScript attach="left" box="[1382,1472,466,480]" fontSize="6" pageId="7" pageNumber="8">
|
||
<bibRefCitation author="Guo, J. & Dawson, M. & Beard, K. C." box="[1382,1397,466,480]" journalOrPublisher="J. Mamm. Evol." pageId="7" pageNumber="8" pagination="239 - 258" part="7" refId="ref9394" refString="14. Guo, J., Dawson, M. & Beard, K. C. Zhailimeryx, a new Lophiomerycid Artiodactyl (Mammalia) from the Late Middle Eocene of Central China and early evolution of ruminants. J. Mamm. Evol. 7, 239 - 258 (2000)." title="Zhailimeryx, a new Lophiomerycid Artiodactyl (Mammalia) from the Late Middle Eocene of Central China and early evolution of ruminants" type="journal article" year="2000">14</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Mennecart, B. Te" box="[1401,1416,466,480]" journalOrPublisher="PLoS ONE" pageId="7" pageNumber="8" pagination="0116830" part="10" refId="ref9496" refString="16. Mennecart, B. Te European ruminants during the " Microbunodon Event " (MP 28, latest Oligocene): Impact of climate changes and faunal event on the ruminant evolution. PLoS ONE 10, e 0116830 (2015)." title="European ruminants during the " Microbunodon Event " (MP 28, latest Oligocene): Impact of climate changes and faunal event on the ruminant evolution" type="journal article" year="2015">16</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Janis, C. M." box="[1420,1435,466,480]" journalOrPublisher="J. Vertebr. Paleontol." pageId="7" pageNumber="8" pagination="200 - 216" part="7" refId="ref10294" refString="33. Janis, C. M. Grades and clades in hornless ruminants evolution: Te reality of the Gelocidae and the systematic position of Lophiomeryx and Bachitherium. J. Vertebr. Paleontol. 7, 200 - 216 (1987)." title="Grades and clades in hornless ruminants evolution: Te reality of the Gelocidae and the systematic position of Lophiomeryx and Bachitherium" type="journal article" year="1987">33</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[1439,1454,466,480]" journalOrPublisher="Swiss J. Geosci." pageId="7" pageNumber="8" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">36</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Brunet, M. & Sudre, J." box="[1457,1472,466,480]" journalOrPublisher="Munch. Geowiss. Abh." pageId="7" pageNumber="8" pagination="225 - 242" part="10" refId="ref10489" refString="37. Brunet, M. & Sudre, J. Evolution et systematique du genre Lophiomeryx POMEL 1853 (Mammalia, Artiodactyla). Munch. Geowiss. Abh. 10, 225 - 242 (1987)." title="Evolution et systematique du genre Lophiomeryx POMEL 1853 (Mammalia, Artiodactyla)" type="journal article" year="1987">37</bibRefCitation>
|
||
</superScript>
|
||
. Te combination of these characters is typical for
|
||
<taxonomicName authorityName="Milne-Edwards" authorityYear="1864" box="[894,997,495,517]" class="Mammalia" family="Tragulidae" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Tragulidae</taxonomicName>
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[997,1012,492,506]" journalOrPublisher="Swiss J. Geosci." pageId="7" pageNumber="8" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">
|
||
<superScript attach="left" box="[997,1012,492,506]" fontSize="6" pageId="7" pageNumber="8">36</superScript>
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="7.[416,1479,253,997]" pageId="7" pageNumber="8">
|
||
Very few taxa are so far known in the early evolution of the
|
||
<taxonomicName authorityName="Milne-Edwards" authorityYear="1864" box="[1001,1104,522,544]" class="Mammalia" family="Tragulidae" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Tragulidae</taxonomicName>
|
||
. Only
|
||
<taxonomicName box="[1165,1316,522,543]" class="Mammalia" family="Tragulidae" genus="Archaeotragulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1165,1316,522,543]" italics="true" pageId="7" pageNumber="8">Archaeotragulus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName box="[1325,1431,522,543]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1325,1431,522,543]" italics="true" pageId="7" pageNumber="8">Iberomeryx</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName box="[416,520,549,570]" class="Mammalia" family="Lophiomerycidae" genus="Nalameryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[416,520,549,570]" italics="true" pageId="7" pageNumber="8">Nalameryx</emphasis>
|
||
</taxonomicName>
|
||
are recognized as potential Paleogene
|
||
<taxonomicName authorityName="Milne-Edwards" authorityYear="1864" box="[883,985,549,571]" class="Mammalia" family="Tragulidae" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Tragulidae</taxonomicName>
|
||
<superScript attach="left" box="[985,1037,546,560]" fontSize="6" pageId="7" pageNumber="8">
|
||
<bibRefCitation author="Metais, G. & Chaimanee, Y. & Jaeger, J. - J. & Ducrocq, S." box="[985,1000,546,560]" journalOrPublisher="Zool. Scr." pageId="7" pageNumber="8" pagination="231 - 248" part="30" refId="ref9541" refString="17. Metais, G., Chaimanee, Y., Jaeger, J. - J. & Ducrocq, S. New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia. Zool. Scr. 30, 231 - 248 (2001)." title="New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia" type="journal article" year="2001">17</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[1003,1018,546,560]" journalOrPublisher="Swiss J. Geosci." pageId="7" pageNumber="8" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">36</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Sudre, J." box="[1022,1037,546,560]" journalOrPublisher="Palaeovertebrata" pageId="7" pageNumber="8" pagination="1 - 31" part="14" refId="ref10833" refString="46. Sudre, J. Cryptomeryx Schlosser, 1886, tragulide de l'Oligocene d'Europe relations du genre et considerations sur l'origine de ruminants. Palaeovertebrata 14, 1 - 31 (1984)." title="Cryptomeryx Schlosser, 1886, tragulide de l'Oligocene d'Europe relations du genre et considerations sur l'origine de ruminants" type="journal article" year="1984">46</bibRefCitation>
|
||
</superScript>
|
||
, of which
|
||
<taxonomicName box="[1134,1286,549,570]" class="Mammalia" family="Tragulidae" genus="Archaeotragulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1134,1286,549,570]" italics="true" pageId="7" pageNumber="8">Archaeotragulus</emphasis>
|
||
</taxonomicName>
|
||
is currently the oldest representative described
|
||
<superScript attach="right" box="[677,710,573,587]" fontSize="6" pageId="7" pageNumber="8">
|
||
<bibRefCitation author="Metais, G. & Chaimanee, Y. & Jaeger, J. - J. & Ducrocq, S." box="[677,692,573,587]" journalOrPublisher="Zool. Scr." pageId="7" pageNumber="8" pagination="231 - 248" part="30" refId="ref9541" refString="17. Metais, G., Chaimanee, Y., Jaeger, J. - J. & Ducrocq, S. New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia. Zool. Scr. 30, 231 - 248 (2001)." title="New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia" type="journal article" year="2001">17</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Rossner, G. E." box="[695,710,573,587]" editor="Prothero, D. R. & Foss, S. C." journalOrPublisher="Te Johns Hopkins University Press" pageId="7" pageNumber="8" pagination="213 - 220" refId="ref10868" refString="47. Rossner, G. E. Family Tragulidae. In Te Evolution of Artiodactyls (eds Prothero, D. R. & Foss, S. C.) 213 - 220 (Te Johns Hopkins University Press, 2007)." title="Family Tragulidae" type="book chapter" volumeTitle="Te Evolution of Artiodactyls" year="2007">47</bibRefCitation>
|
||
</superScript>
|
||
.
|
||
<taxonomicName box="[720,872,575,596]" class="Mammalia" family="Tragulidae" genus="Archaeotragulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[720,872,575,596]" italics="true" pageId="7" pageNumber="8">Archaeotragulus</emphasis>
|
||
</taxonomicName>
|
||
possesses lower molars with a broadened talonid in comparison to the trigonid and displays an entoconidian groove
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[914,929,599,613]" journalOrPublisher="Swiss J. Geosci." pageId="7" pageNumber="8" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">
|
||
<superScript attach="left" box="[914,929,599,613]" fontSize="6" pageId="7" pageNumber="8">36</superScript>
|
||
</bibRefCitation>
|
||
. In the case of IVPP V 6551, the trigonid and talonid are of similar size and no specific entoconidian groove can be observed. Mennecart et al.
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[1240,1255,626,640]" journalOrPublisher="Swiss J. Geosci." pageId="7" pageNumber="8" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">
|
||
<superScript attach="left" box="[1240,1255,626,640]" fontSize="6" pageId="7" pageNumber="8">36</superScript>
|
||
</bibRefCitation>
|
||
considered
|
||
<taxonomicName box="[1372,1477,629,650]" class="Mammalia" family="Lophiomerycidae" genus="Nalameryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1372,1477,629,650]" italics="true" pageId="7" pageNumber="8">Nalameryx</emphasis>
|
||
</taxonomicName>
|
||
a
|
||
<taxonomicName authorityName="Milne-Edwards" authorityYear="1864" box="[432,535,655,677]" class="Mammalia" family="Tragulidae" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="family">Tragulidae</taxonomicName>
|
||
notably based on the presence of the M structure (the external postmetacristid, the internal postmetacristid, the internal postprotocristid, and the external postprotocristid are interconnected forming a M in occlusal view), including the
|
||
<taxonomicName authorityName="Brisson" authorityYear="1762" box="[700,783,709,730]" class="Mammalia" family="Tragulidae" genus="Tragulus" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[700,783,709,730]" italics="true" pageId="7" pageNumber="8">Tragulus</emphasis>
|
||
</taxonomicName>
|
||
fold and
|
||
<taxonomicName box="[877,1011,709,730]" class="Mammalia" family="Tragulidae" genus="Dorcatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[877,1011,709,730]" italics="true" pageId="7" pageNumber="8">Dorcatherium</emphasis>
|
||
</taxonomicName>
|
||
fold, and the absence of a rounded mesolingual conid in the p
|
||
<superScript attach="none" box="[561,576,732,746]" fontSize="6" pageId="7" pageNumber="8">435</superScript>
|
||
. IVPP V 6551 differs from
|
||
<taxonomicName box="[837,942,735,756]" class="Mammalia" family="Lophiomerycidae" genus="Nalameryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[837,942,735,756]" italics="true" pageId="7" pageNumber="8">Nalameryx</emphasis>
|
||
</taxonomicName>
|
||
in having an m3 wider than m1 and similar m1 and m2 widths
|
||
<bibRefCitation author="Metais, G. & Chaimanee, Y. & Jaeger, J. - J. & Ducrocq, S." box="[481,496,759,773]" journalOrPublisher="Zool. Scr." pageId="7" pageNumber="8" pagination="231 - 248" part="30" refId="ref9541" refString="17. Metais, G., Chaimanee, Y., Jaeger, J. - J. & Ducrocq, S. New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia. Zool. Scr. 30, 231 - 248 (2001)." title="New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia" type="journal article" year="2001">
|
||
<superScript attach="left" box="[481,496,759,773]" fontSize="6" pageId="7" pageNumber="8">17</superScript>
|
||
</bibRefCitation>
|
||
. In size proportions and molar morphology, IVPP V 6551 resembles the genus
|
||
<taxonomicName box="[1257,1366,762,783]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1257,1366,762,783]" italics="true" pageId="7" pageNumber="8">Iberomeryx</emphasis>
|
||
</taxonomicName>
|
||
. In IVPP V 6551, the relative size of the m2 is more similar to
|
||
<emphasis box="[893,968,789,810]" italics="true" pageId="7" pageNumber="8">I. minor</emphasis>
|
||
. In
|
||
<emphasis box="[1005,1176,789,810]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName box="[1005,1113,789,810]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Iberomeryx</taxonomicName>
|
||
minor
|
||
</emphasis>
|
||
, the anterior cingulid is big
|
||
<superScript attach="left" box="[1439,1472,786,800]" fontSize="6" pageId="7" pageNumber="8">
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[1439,1454,786,800]" journalOrPublisher="Swiss J. Geosci." pageId="7" pageNumber="8" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">36</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Sudre, J." box="[1457,1472,786,800]" journalOrPublisher="Palaeovertebrata" pageId="7" pageNumber="8" pagination="1 - 31" part="14" refId="ref10833" refString="46. Sudre, J. Cryptomeryx Schlosser, 1886, tragulide de l'Oligocene d'Europe relations du genre et considerations sur l'origine de ruminants. Palaeovertebrata 14, 1 - 31 (1984)." title="Cryptomeryx Schlosser, 1886, tragulide de l'Oligocene d'Europe relations du genre et considerations sur l'origine de ruminants" type="journal article" year="1984">46</bibRefCitation>
|
||
</superScript>
|
||
, while in
|
||
<emphasis box="[501,681,815,837]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName box="[501,610,815,836]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Iberomeryx</taxonomicName>
|
||
parvus
|
||
</emphasis>
|
||
the cingulid is thin
|
||
<bibRefCitation author="Metais, G." box="[872,887,813,827]" journalOrPublisher="Palaeontol. Electron." pageId="7" pageNumber="8" pagination="1 - 23" part="19" refId="ref10913" refString="48. Metais, G. et al. Oligocene ruminants from the Kizilirmak Formation, Cankiri-Corum Basin, Central Anatolia, Turkey. Palaeontol. Electron. 19, 1 - 23 (2016)." title="Oligocene ruminants from the Kizilirmak Formation, Cankiri-Corum Basin, Central Anatolia, Turkey" type="journal article" year="2016">
|
||
<superScript attach="left" box="[872,887,813,827]" fontSize="6" pageId="7" pageNumber="8">48</superScript>
|
||
</bibRefCitation>
|
||
like in IVPP V 6551. Te teeth of IVPP V 6551 appear to be more massive and bulkier than in
|
||
<emphasis box="[736,812,843,864]" italics="true" pageId="7" pageNumber="8">I. minor</emphasis>
|
||
and
|
||
<emphasis box="[857,940,843,864]" italics="true" pageId="7" pageNumber="8">I. parvus</emphasis>
|
||
<superScript attach="left" box="[940,973,839,854]" fontSize="6" pageId="7" pageNumber="8">
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[940,955,839,853]" journalOrPublisher="Swiss J. Geosci." pageId="7" pageNumber="8" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">36</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Metais, G." box="[958,973,839,853]" journalOrPublisher="Palaeontol. Electron." pageId="7" pageNumber="8" pagination="1 - 23" part="19" refId="ref10913" refString="48. Metais, G. et al. Oligocene ruminants from the Kizilirmak Formation, Cankiri-Corum Basin, Central Anatolia, Turkey. Palaeontol. Electron. 19, 1 - 23 (2016)." title="Oligocene ruminants from the Kizilirmak Formation, Cankiri-Corum Basin, Central Anatolia, Turkey" type="journal article" year="2016">48</bibRefCitation>
|
||
</superScript>
|
||
. Similarly to
|
||
<emphasis box="[1098,1173,843,864]" italics="true" pageId="7" pageNumber="8">I. minor</emphasis>
|
||
, the protoconid of IVPP V 6551 is a little more anterior than in
|
||
<emphasis box="[711,795,869,890]" italics="true" pageId="7" pageNumber="8">I. parvus</emphasis>
|
||
<superScript attach="left" box="[794,828,866,880]" fontSize="6" pageId="7" pageNumber="8">
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[794,809,866,880]" journalOrPublisher="Swiss J. Geosci." pageId="7" pageNumber="8" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">36</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Metais, G." box="[813,828,866,880]" journalOrPublisher="Palaeontol. Electron." pageId="7" pageNumber="8" pagination="1 - 23" part="19" refId="ref10913" refString="48. Metais, G. et al. Oligocene ruminants from the Kizilirmak Formation, Cankiri-Corum Basin, Central Anatolia, Turkey. Palaeontol. Electron. 19, 1 - 23 (2016)." title="Oligocene ruminants from the Kizilirmak Formation, Cankiri-Corum Basin, Central Anatolia, Turkey" type="journal article" year="2016">48</bibRefCitation>
|
||
</superScript>
|
||
. IVPP V 6551 clearly differs from
|
||
<emphasis box="[1155,1239,869,890]" italics="true" pageId="7" pageNumber="8">I. parvus</emphasis>
|
||
and
|
||
<emphasis box="[1284,1360,869,890]" italics="true" pageId="7" pageNumber="8">I. minor</emphasis>
|
||
by the presence of a very large paraconid, which is smaller in the two other species
|
||
<superScript attach="left" box="[1108,1141,892,907]" fontSize="6" pageId="7" pageNumber="8">
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[1108,1123,892,906]" journalOrPublisher="Swiss J. Geosci." pageId="7" pageNumber="8" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">36</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Metais, G." box="[1126,1141,893,907]" journalOrPublisher="Palaeontol. Electron." pageId="7" pageNumber="8" pagination="1 - 23" part="19" refId="ref10913" refString="48. Metais, G. et al. Oligocene ruminants from the Kizilirmak Formation, Cankiri-Corum Basin, Central Anatolia, Turkey. Palaeontol. Electron. 19, 1 - 23 (2016)." title="Oligocene ruminants from the Kizilirmak Formation, Cankiri-Corum Basin, Central Anatolia, Turkey" type="journal article" year="2016">48</bibRefCitation>
|
||
</superScript>
|
||
. Moreover, the metastylid in IVPP V 6551 is slightly more developed than in
|
||
<emphasis box="[814,889,923,944]" italics="true" pageId="7" pageNumber="8">I. minor</emphasis>
|
||
and not present in
|
||
<emphasis box="[1072,1155,923,944]" italics="true" pageId="7" pageNumber="8">I. parvus</emphasis>
|
||
<superScript attach="left" box="[1155,1188,919,934]" fontSize="6" pageId="7" pageNumber="8">
|
||
<bibRefCitation author="Gabunia, L. Te" box="[1155,1170,919,933]" journalOrPublisher="Academy of Sciences of URSS" pageId="7" pageNumber="8" refId="ref10717" refString="43. Gabunia, L. Te Oligocene Mammalian Fauna of Benara 268 (Academy of Sciences of URSS, 1964)." title="Oligocene Mammalian Fauna of Benara 268" type="book" year="1964">43</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Metais, G." box="[1173,1188,919,933]" journalOrPublisher="Palaeontol. Electron." pageId="7" pageNumber="8" pagination="1 - 23" part="19" refId="ref10913" refString="48. Metais, G. et al. Oligocene ruminants from the Kizilirmak Formation, Cankiri-Corum Basin, Central Anatolia, Turkey. Palaeontol. Electron. 19, 1 - 23 (2016)." title="Oligocene ruminants from the Kizilirmak Formation, Cankiri-Corum Basin, Central Anatolia, Turkey" type="journal article" year="2016">48</bibRefCitation>
|
||
</superScript>
|
||
.
|
||
<emphasis box="[1199,1368,922,944]" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName box="[1199,1307,922,943]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">Iberomeryx</taxonomicName>
|
||
minor
|
||
</emphasis>
|
||
displays an ectostylid on all molars
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[641,656,946,960]" journalOrPublisher="Swiss J. Geosci." pageId="7" pageNumber="8" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">
|
||
<superScript attach="right" box="[641,656,946,960]" fontSize="6" pageId="7" pageNumber="8">36</superScript>
|
||
</bibRefCitation>
|
||
, while this structure is absent from
|
||
<emphasis box="[1002,1086,949,970]" italics="true" pageId="7" pageNumber="8">I. parvus</emphasis>
|
||
<bibRefCitation author="Metais, G." box="[1086,1101,946,960]" journalOrPublisher="Palaeontol. Electron." pageId="7" pageNumber="8" pagination="1 - 23" part="19" refId="ref10913" refString="48. Metais, G. et al. Oligocene ruminants from the Kizilirmak Formation, Cankiri-Corum Basin, Central Anatolia, Turkey. Palaeontol. Electron. 19, 1 - 23 (2016)." title="Oligocene ruminants from the Kizilirmak Formation, Cankiri-Corum Basin, Central Anatolia, Turkey" type="journal article" year="2016">
|
||
<superScript attach="left" box="[1086,1101,946,960]" fontSize="6" pageId="7" pageNumber="8">48</superScript>
|
||
</bibRefCitation>
|
||
. Te ectostylid in IVPP V 6551 is large on m1 to absent on m3. Based on these differences we decided to erect the new species
|
||
<taxonomicName authority="Mennecart & Aiglstorfer & Li & Li & Wang, 2021" authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" box="[1236,1403,975,997]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="miaoi" status="nov.sp.">
|
||
<emphasis box="[1236,1403,975,997]" italics="true" pageId="7" pageNumber="8">Iberomeryx miaoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1408,1479,976,997]" pageId="7" pageNumber="8" rank="species">nov.sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |