729 lines
81 KiB
XML
729 lines
81 KiB
XML
<document ID-DOI="10.1038/s41598-021-96221-x" ID-GBIF-Dataset="11088aaa-ac64-4b8f-bb4f-5ab4dd5baa6f" ID-PMC="PMC8421421" ID-PubMed="34489502" ID-Zenodo-Dep="5645747" approvalRequired="62" approvalRequired_for_document="1" approvalRequired_for_taxonomicNames="29" approvalRequired_for_treatments="32" checkinTime="1636046587746" checkinUser="valdenar" docAuthor="Mennecart, Bastien, Aiglstorfer, Manuela, Li, Yikun, Li, Chunxiao & Wang, ShiQi" docDate="2021" docId="E97287E44C427A0C1FEAFB6CFADACDC2" docLanguage="en" docName="SciRep.11.17710.pdf" docOrigin="Scientific Reports 1" docTitle="Krabimeryx Metais, Chaimanee, Jaeger, and Ducroq" docType="treatment" docVersion="4" lastPageNumber="4" masterDocId="154BFF9C4C437A0F1E4AFFE2FFB5C812" masterDocTitle="Ruminants reveal Eocene Asiatic palaeobiogeographical provinces as the origin of diachronous mammalian Oligocene dispersals into Europe" masterLastPageNumber="12" masterPageNumber="1" pageNumber="2" updateTime="1668144176883" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Ruminants reveal Eocene Asiatic palaeobiogeographical provinces as the origin of diachronous mammalian Oligocene dispersals into Europe</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mennecart, Bastien</mods:namePart>
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<mods:affiliation>Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria & Naturhistorisches Museum Basel, Augustinergasse 2, 4001 Basel, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="email">mennecartbastien@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Aiglstorfer, Manuela</mods:namePart>
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<mods:affiliation>Naturhistorisches Museum Mainz / Landessammlung Für Naturkunde Rheinland-Pfalz, Reichklarastrasse 10, 55116 Mainz, Germany.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Yikun</mods:namePart>
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<mods:affiliation>Center for Research and Education On Biological Evolution and Environment, Nanjing Univeristy, Nanjing 210023, China.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Chunxiao</mods:namePart>
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<mods:affiliation>Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China & CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wang, ShiQi</mods:namePart>
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<mods:affiliation>Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China & CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China</mods:affiliation>
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<mods:nameIdentifier type="email">wangshiqi@ivpp.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Scientific Reports</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-09-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>12</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1038/s41598-021-96221-x</mods:identifier>
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<mods:identifier type="GBIF-Dataset">11088aaa-ac64-4b8f-bb4f-5ab4dd5baa6f</mods:identifier>
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<mods:identifier type="PMC">PMC8421421</mods:identifier>
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<mods:identifier type="PubMed">34489502</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5645747</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5645755" ID-GBIF-Taxon="189990182" ID-Zenodo-Dep="5645755" LSID="urn:lsid:plazi:treatment:E97287E44C427A0C1FEAFB6CFADACDC2" httpUri="http://treatment.plazi.org/id/E97287E44C427A0C1FEAFB6CFADACDC2" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection box="[416,1059,1163,1188]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph blockId="1.[416,1059,1163,1188]" box="[416,1059,1163,1188]" pageId="1" pageNumber="2">
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Genus
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<taxonomicName authority="Metais, Chaimanee, Jaeger, and Ducroq" authorityName="Metais, Chaimanee, Jaeger, and Ducroq" box="[483,983,1166,1188]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis box="[483,596,1166,1187]" italics="true" pageId="1" pageNumber="2">Krabimeryx</emphasis>
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Métais, Chaimanee, Jaeger, and Ducroq
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</taxonomicName>
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,
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<superScript attach="none" box="[1039,1054,1163,1177]" fontSize="6" pageId="1" pageNumber="2">200117</superScript>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="2" type="etymology">
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<paragraph blockId="1.[416,1478,1224,1274]" pageId="1" pageNumber="2">
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<heading bold="true" box="[416,549,1224,1248]" fontSize="10" level="2" pageId="1" pageNumber="2" reason="0">
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<emphasis bold="true" box="[416,549,1224,1248]" pageId="1" pageNumber="2">Etymology.</emphasis>
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</heading>
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<emphasis box="[575,649,1225,1246]" italics="true" pageId="1" pageNumber="2">Krabi—</emphasis>
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from Krabi Basin, where the fossils were found, and—
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<emphasis box="[1193,1253,1226,1247]" italics="true" pageId="1" pageNumber="2">meryx</emphasis>
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is the Greek word for ruminant.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="2" type="diagnosis">
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<paragraph blockId="1.[416,1478,1306,1572]" pageId="1" pageNumber="2">
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<emphasis bold="true" box="[416,858,1309,1333]" pageId="1" pageNumber="2">Diagnosis [modified after Métais et al.</emphasis>
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<bibRefCitation author="Metais, G. & Chaimanee, Y. & Jaeger, J. - J. & Ducrocq, S." box="[858,876,1306,1321]" journalOrPublisher="Zool. Scr." pageId="1" pageNumber="2" pagination="231 - 248" part="30" refId="ref9541" refString="17. Metais, G., Chaimanee, Y., Jaeger, J. - J. & Ducrocq, S. New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia. Zool. Scr. 30, 231 - 248 (2001)." title="New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia" type="journal article" year="2001">
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<emphasis bold="true" box="[858,876,1306,1321]" pageId="1" pageNumber="2">
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<superScript attach="right" box="[858,876,1306,1321]" fontSize="7" pageId="1" pageNumber="2">17</superScript>
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</emphasis>
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</bibRefCitation>
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<emphasis bold="true" box="[876,892,1309,1333]" pageId="1" pageNumber="2">].</emphasis>
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Small primitive ruminant with lower molars morphologically close to those of
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<taxonomicName box="[631,746,1337,1358]" class="Mammalia" family="Lophiomerycidae" genus="Zhailimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis box="[631,746,1337,1358]" italics="true" pageId="1" pageNumber="2">Zhailimeryx</emphasis>
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</taxonomicName>
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.
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<taxonomicName box="[758,870,1337,1358]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis box="[758,870,1337,1358]" italics="true" pageId="1" pageNumber="2">Krabimeryx</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName box="[998,1113,1337,1358]" class="Mammalia" family="Lophiomerycidae" genus="Zhailimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis box="[998,1113,1337,1358]" italics="true" pageId="1" pageNumber="2">Zhailimeryx</emphasis>
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</taxonomicName>
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in: more laterally compressed lingual cuspids in the lower molars; an entoconid displaced to anterior with respect to the hypoconid; the lack of both a paraconid and a hypoconulid in m1 and m2; a p4 with a mesolingual conid that is located more posterior and less individualized; a p4 without a distinct posterolingual conid.
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<taxonomicName box="[1043,1155,1417,1438]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis box="[1043,1155,1417,1438]" italics="true" pageId="1" pageNumber="2">Krabimeryx</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName box="[1283,1407,1417,1438]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis box="[1283,1407,1417,1438]" italics="true" pageId="1" pageNumber="2">Lophiomeryx</emphasis>
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</taxonomicName>
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by less selenodont labial cuspids in the lower molars, the presence of a developed external postmetacristid, and by a distinct groove on the anterior side of the entoconid, the entoconidian groove.
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<taxonomicName box="[1165,1277,1470,1491]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis box="[1165,1277,1470,1491]" italics="true" pageId="1" pageNumber="2">Krabimeryx</emphasis>
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</taxonomicName>
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can be distinguished from
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<taxonomicName box="[469,577,1497,1518]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis box="[469,577,1497,1518]" italics="true" pageId="1" pageNumber="2">Iberomeryx</emphasis>
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</taxonomicName>
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in having a well-marked entoconidian groove; the lack of a clear external postprotocristid; the third lobe of m3 not forming a complete buckle; and a more transversely compressed hypoconulid in the m3.
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<taxonomicName box="[416,528,1550,1571]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
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<emphasis box="[416,528,1550,1571]" italics="true" pageId="1" pageNumber="2">Krabimeryx</emphasis>
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</taxonomicName>
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possesses a huge notch in lingual view between the entoconid and the third lobe in the m3.
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</paragraph>
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</subSubSection>
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<subSubSection box="[416,1271,1606,1632]" pageId="1" pageNumber="2" type="type_taxon">
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<paragraph blockId="1.[416,1271,1606,1632]" box="[416,1271,1606,1632]" pageId="1" pageNumber="2">
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<heading bold="true" box="[416,568,1608,1632]" fontSize="10" level="2" pageId="1" pageNumber="2" reason="0">
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<emphasis bold="true" box="[416,568,1608,1632]" pageId="1" pageNumber="2">Type species.</emphasis>
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</heading>
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<emphasis box="[594,808,1609,1631]" italics="true" pageId="1" pageNumber="2">
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<taxonomicName box="[594,706,1609,1630]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">Krabimeryx</taxonomicName>
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primitivus
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</emphasis>
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Métais, Chaimanne, Jaeger, and Ducroq,
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<superScript attach="none" box="[1251,1266,1606,1620]" fontSize="6" pageId="1" pageNumber="2">200117</superScript>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="2" type="reference_group">
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<paragraph blockId="1.[416,1079,1665,1691]" box="[416,1079,1665,1691]" pageId="1" pageNumber="2">
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<heading bold="true" box="[416,611,1667,1691]" fontSize="10" level="2" pageId="1" pageNumber="2" reason="0">
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<emphasis bold="true" box="[416,611,1667,1691]" pageId="1" pageNumber="2">Included species.</emphasis>
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</heading>
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<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[637,821,1668,1689]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="gracilis" status="nov. comb.">
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<emphasis box="[637,821,1668,1689]" italics="true" pageId="1" pageNumber="2">Krabimeryx gracilis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[826,931,1668,1690]" pageId="1" pageNumber="2" rank="species">nov. comb.</taxonomicNameLabel>
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(Miao,
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<superScript attach="none" box="[1050,1065,1665,1679]" fontSize="6" pageId="1" pageNumber="2">198220</superScript>
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).
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</paragraph>
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<paragraph blockId="1.[416,1162,1725,1963]" box="[416,861,1725,1750]" pageId="1" pageNumber="2">
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<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[416,601,1728,1749]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="gracilis" status="nov. comb.">
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<emphasis box="[416,601,1728,1749]" italics="true" pageId="1" pageNumber="2">Krabimeryx gracilis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[606,712,1728,1750]" pageId="1" pageNumber="2" rank="species">nov. comb.</taxonomicNameLabel>
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(Miao,
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<superScript attach="right" box="[832,847,1725,1739]" fontSize="6" pageId="1" pageNumber="2">198220</superScript>
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).
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</paragraph>
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</subSubSection>
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<subSubSection box="[448,687,1754,1776]" pageId="1" pageNumber="2" type="description">
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<paragraph blockId="1.[416,1162,1725,1963]" box="[448,687,1754,1776]" pageId="1" pageNumber="2">
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<figureCitation box="[448,544,1754,1776]" captionStart="Figure 1" captionStartId="2.[416,470,1525,1544]" captionTargetBox="[422,1363,140,1491]" captionTargetId="figure-31@2.[416,1368,135,1496]" captionTargetPageId="2" captionText="Figure 1. Dentition of Krabimeryx gracilis nov. comb. (Miao, 1982)20 (A, B, G, H), Chiyoumeryx nov. gen. shinaoensis (Miao, 1982)20 (C, D), Chiyoumeryx nov. gen. flavimperatoris nov. sp. (E) and Iberomeryx miaoi nov. sp. (F–I). Krabimeryx gracilis nov. comb.(Miao, 1982)20: (A) IVPP V 6546-1 (holotype), partial skull with right and lef M1–M3; (B) IVPP V 6546-2 (holotype), right fragmented mandible with m2–m3. Chiyoumeryx nov.gen. shinaoensis (Miao, 1982)20: (C) IVPP V 6531 (holotype), right mandible with p2–m3 and tooth socket of p1; (D) IVPP V 6532 (paratype), right fragmented maxillary with P4-M3. Chiyoumeryx nov. gen. flavimperatoris nov. sp.: (E) IVPP V 6547 (holotype), right mandible with p4–m3; Iberomeryx miaoi nov. sp.: (F) IVPP V 6551 (holotype), lef mandible with m1–m3 (mirrored); (G) lower molar Lophiomerycidae dental nomenclature (based on the m3 of IVPP 6546-2): 1 internal postmetacristid, 2 metaconid, 3 external postmetacristid, 4 internal preentocristid, 5 entoconidian groove, 6 external preentocristid, 7 entoconid, 8 posthypoconulidcristid, 9 hypoconulid, 10 prehypoconuldicristid, 11 posthypocristid, 12 hypoconid, 13 prehypocristid, 14 ectostylid, 15 postprotocristid, 16 protoconid, 17 preprotocristid, 18 anterior cingulid; (H) upper molar Lophiomerycidae dental nomenclature (based on the M2 of IVPP 6546-1): 1 postmetacrista, 2 metacone, 3 premetacrista, 4 mesostyle, 5 postparacrista, 6 paracone, 7 paraconid labial groove, 8 preparacrista, 9 parastyle, 10 preprotocrista, 11 anterolingual cingulum, 12 protocone, 13 postprotocrista, 14 entostyle, 15 additional cone, 16 premetaconulecrista, 17 metaconule, 18 postmetaconulecrista; (I) lower molar Tragulidae dental nomenclature (based on the m2 of IVPP V 6551, reversed): 1 metaconid, 2 external postmetacristid, 3 Dorcatherium fold, 4 internal postmetacristid, 5 preentocristid, 6 entoconid, 7 postentocristid, 8 posterior cingulid, 9 posthypocristid, 10 hypoconid, 11 prehypocristid, 12 ectostylid, 13 external postprotocristid, 14 Tragulus fold, 15 internal postprotocristid, 16 protoconid, 17 preprotocristid, 18 paraconid, 19 preparacristid. (J) phylogenetic position and stratigraphie of the Shinao/ Yangjiachong/Xiaerhete ruminants (topology2). a stem Ruminantia, b Archaeomeryx, c Chiyoumeryx nov.gen. and Krabimeryx gracilis, d crown Ruminantia, e Iberomeryx miaoi nov. sp.; 1 lingual view, 2 occlusal view. Scale bare is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5645749" httpUri="https://zenodo.org/record/5645749/files/figure.png" pageId="1" pageNumber="2">Figure 1A</figureCitation>
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and
|
||
<figureCitation box="[591,684,1754,1776]" captionStart="Figure 1" captionStartId="2.[416,470,1525,1544]" captionTargetBox="[422,1363,140,1491]" captionTargetId="figure-31@2.[416,1368,135,1496]" captionTargetPageId="2" captionText="Figure 1. Dentition of Krabimeryx gracilis nov. comb. (Miao, 1982)20 (A, B, G, H), Chiyoumeryx nov. gen. shinaoensis (Miao, 1982)20 (C, D), Chiyoumeryx nov. gen. flavimperatoris nov. sp. (E) and Iberomeryx miaoi nov. sp. (F–I). Krabimeryx gracilis nov. comb.(Miao, 1982)20: (A) IVPP V 6546-1 (holotype), partial skull with right and lef M1–M3; (B) IVPP V 6546-2 (holotype), right fragmented mandible with m2–m3. Chiyoumeryx nov.gen. shinaoensis (Miao, 1982)20: (C) IVPP V 6531 (holotype), right mandible with p2–m3 and tooth socket of p1; (D) IVPP V 6532 (paratype), right fragmented maxillary with P4-M3. Chiyoumeryx nov. gen. flavimperatoris nov. sp.: (E) IVPP V 6547 (holotype), right mandible with p4–m3; Iberomeryx miaoi nov. sp.: (F) IVPP V 6551 (holotype), lef mandible with m1–m3 (mirrored); (G) lower molar Lophiomerycidae dental nomenclature (based on the m3 of IVPP 6546-2): 1 internal postmetacristid, 2 metaconid, 3 external postmetacristid, 4 internal preentocristid, 5 entoconidian groove, 6 external preentocristid, 7 entoconid, 8 posthypoconulidcristid, 9 hypoconulid, 10 prehypoconuldicristid, 11 posthypocristid, 12 hypoconid, 13 prehypocristid, 14 ectostylid, 15 postprotocristid, 16 protoconid, 17 preprotocristid, 18 anterior cingulid; (H) upper molar Lophiomerycidae dental nomenclature (based on the M2 of IVPP 6546-1): 1 postmetacrista, 2 metacone, 3 premetacrista, 4 mesostyle, 5 postparacrista, 6 paracone, 7 paraconid labial groove, 8 preparacrista, 9 parastyle, 10 preprotocrista, 11 anterolingual cingulum, 12 protocone, 13 postprotocrista, 14 entostyle, 15 additional cone, 16 premetaconulecrista, 17 metaconule, 18 postmetaconulecrista; (I) lower molar Tragulidae dental nomenclature (based on the m2 of IVPP V 6551, reversed): 1 metaconid, 2 external postmetacristid, 3 Dorcatherium fold, 4 internal postmetacristid, 5 preentocristid, 6 entoconid, 7 postentocristid, 8 posterior cingulid, 9 posthypocristid, 10 hypoconid, 11 prehypocristid, 12 ectostylid, 13 external postprotocristid, 14 Tragulus fold, 15 internal postprotocristid, 16 protoconid, 17 preprotocristid, 18 paraconid, 19 preparacristid. (J) phylogenetic position and stratigraphie of the Shinao/ Yangjiachong/Xiaerhete ruminants (topology2). a stem Ruminantia, b Archaeomeryx, c Chiyoumeryx nov.gen. and Krabimeryx gracilis, d crown Ruminantia, e Iberomeryx miaoi nov. sp.; 1 lingual view, 2 occlusal view. Scale bare is 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5645749" httpUri="https://zenodo.org/record/5645749/files/figure.png" pageId="1" pageNumber="2">Figure S1</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="reference_group">
|
||
<paragraph blockId="1.[416,1162,1725,1963]" box="[448,1115,1778,1803]" pageId="1" pageNumber="2">
|
||
<treatmentCitationGroup box="[448,1115,1778,1803]" pageId="1" pageNumber="2">
|
||
*v pars1982
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[567,765,1781,1802]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[567,765,1781,1802]" italics="true" pageId="1" pageNumber="2">Lophiomeryx gracilis</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<treatmentCitation author="Miao" box="[790,1115,1778,1803]" page="532" pageId="1" pageNumber="2" year="1982">
|
||
Miao: 532, Table 3, Figs. 6 and
|
||
<superScript attach="none" box="[1095,1110,1778,1792]" fontSize="6" pageId="1" pageNumber="2">720</superScript>
|
||
.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="1.[416,1162,1725,1963]" box="[448,965,1805,1830]" pageId="1" pageNumber="2">
|
||
<treatmentCitationGroup box="[448,965,1805,1830]" pageId="1" pageNumber="2">
|
||
v non1982
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[554,752,1808,1829]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[554,752,1808,1829]" italics="true" pageId="1" pageNumber="2">Lophiomeryx gracilis</emphasis>
|
||
</taxonomicName>
|
||
?—
|
||
<treatmentCitation author="Miao" box="[786,965,1805,1830]" page="536" pageId="1" pageNumber="2" year="1982">
|
||
Miao: 536, Fig.
|
||
<superScript attach="none" box="[945,960,1805,1819]" fontSize="6" pageId="1" pageNumber="2">820</superScript>
|
||
.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="1.[416,1162,1725,1963]" box="[448,783,1831,1856]" pageId="1" pageNumber="2">
|
||
<treatmentCitationGroup box="[448,783,1831,1856]" pageId="1" pageNumber="2">
|
||
v pars1987
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[557,648,1834,1856]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[557,648,1834,1856]" italics="true" pageId="1" pageNumber="2">L. gracilis</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<treatmentCitation author="Janis" box="[674,783,1831,1856]" page="211" pageId="1" pageNumber="2" year="1987">
|
||
Janis:
|
||
<superScript attach="none" box="[762,777,1831,1845]" fontSize="6" pageId="1" pageNumber="2">21133</superScript>
|
||
.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="1.[416,1162,1725,1963]" box="[448,874,1858,1883]" pageId="1" pageNumber="2">
|
||
<treatmentCitationGroup box="[448,874,1858,1883]" pageId="1" pageNumber="2">
|
||
v pars
|
||
<emphasis box="[505,648,1861,1883]" italics="true" pageId="1" pageNumber="2">
|
||
1997
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[557,648,1861,1883]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
|
||
</emphasis>
|
||
—
|
||
<treatmentCitation box="[672,874,1858,1883]" pageId="1" pageNumber="2">
|
||
Vislobokova: Fig.
|
||
<superScript attach="none" box="[854,869,1858,1872]" fontSize="6" pageId="1" pageNumber="2">321</superScript>
|
||
.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="1.[416,1162,1725,1963]" box="[448,1056,1885,1910]" pageId="1" pageNumber="2">
|
||
<treatmentCitationGroup box="[448,1056,1885,1910]" pageId="1" pageNumber="2">
|
||
v pars
|
||
<emphasis box="[505,648,1888,1909]" italics="true" pageId="1" pageNumber="2">
|
||
2000
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[557,648,1888,1909]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
|
||
</emphasis>
|
||
—
|
||
<treatmentCitation author="Guo, Dawson, and Beard" box="[673,1056,1885,1910]" page="247" pageId="1" pageNumber="2" year="2000">
|
||
Guo, Dawson, and Beard: 247, Table
|
||
<superScript attach="none" box="[1036,1051,1885,1899]" fontSize="6" pageId="1" pageNumber="2">214</superScript>
|
||
.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="1.[416,1162,1725,1963]" box="[448,1162,1911,1936]" pageId="1" pageNumber="2">
|
||
<treatmentCitationGroup box="[448,1162,1911,1936]" pageId="1" pageNumber="2">
|
||
v pars
|
||
<emphasis box="[505,648,1914,1936]" italics="true" pageId="1" pageNumber="2">
|
||
2001
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[557,648,1914,1936]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
|
||
</emphasis>
|
||
—
|
||
<treatmentCitation author="Métais, Chaimanee, Jaeger, and Ducroq" box="[672,1162,1911,1936]" page="239" pageId="1" pageNumber="2" year="2001">
|
||
Métais, Chaimanee, Jaeger, and Ducroq: 239,
|
||
<superScript attach="none" box="[1142,1157,1911,1925]" fontSize="6" pageId="1" pageNumber="2">24117</superScript>
|
||
.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="1.[416,1162,1725,1963]" box="[448,830,1938,1963]" pageId="1" pageNumber="2">
|
||
<treatmentCitationGroup box="[448,830,1938,1963]" pageId="1" pageNumber="2">
|
||
v pars
|
||
<emphasis box="[505,648,1941,1963]" italics="true" pageId="1" pageNumber="2">
|
||
2012
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[557,648,1941,1963]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
|
||
</emphasis>
|
||
—
|
||
<treatmentCitation author="Mennecart" box="[674,830,1938,1963]" page="62" pageId="1" pageNumber="2" year="2012">
|
||
Mennecart:
|
||
<superScript attach="right" box="[810,825,1938,1952]" fontSize="6" pageId="1" pageNumber="2">6234</superScript>
|
||
.
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<subSubSection box="[110,280,2023,2043]" pageId="1" pageNumber="2" type="description">
|
||
<paragraph blockId="1.[110,280,2023,2043]" box="[110,280,2023,2043]" pageId="1" pageNumber="2">
|
||
<heading bold="true" box="[110,280,2023,2043]" fontSize="8" level="4" pageId="1" pageNumber="2" reason="2">
|
||
<emphasis bold="true" box="[110,271,2023,2042]" pageId="1" pageNumber="2">Scientific Reports</emphasis>
|
||
|
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.5645749" ID-Zenodo-Dep="5645749" httpUri="https://zenodo.org/record/5645749/files/figure.png" pageId="2" pageNumber="3" startId="2.[416,470,1525,1544]" targetBox="[422,1363,140,1491]" targetPageId="2">
|
||
<paragraph blockId="2.[416,1475,1524,1977]" pageId="2" pageNumber="3">
|
||
<emphasis bold="true" box="[416,489,1525,1544]" pageId="2" pageNumber="3">Figure 1.</emphasis>
|
||
Dentition of
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[602,753,1525,1544]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="gracilis" status="nov. comb.">
|
||
<emphasis box="[602,753,1525,1544]" italics="true" pageId="2" pageNumber="3">Krabimeryx gracilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[758,845,1525,1545]" pageId="2" pageNumber="3" rank="species">nov. comb.</taxonomicNameLabel>
|
||
(Miao, 1982)
|
||
<bibRefCitation author="Miao, D." box="[949,961,1524,1536]" journalOrPublisher="Acta Palaeontol. Sinica" pageId="2" pageNumber="3" pagination="526 - 536" part="21" refId="ref9719" refString="20. Miao, D. Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province. Acta Palaeontol. Sinica 21, 526 - 536 (1982)." title="Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province" type="journal article" year="1982">
|
||
<superScript attach="left" box="[949,961,1524,1536]" fontSize="5" pageId="2" pageNumber="3">20</superScript>
|
||
</bibRefCitation>
|
||
(
|
||
<emphasis bold="true" box="[973,987,1525,1544]" pageId="2" pageNumber="3">A</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[996,1009,1526,1545]" pageId="2" pageNumber="3">B</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1017,1031,1525,1544]" pageId="2" pageNumber="3">G</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1041,1057,1526,1545]" pageId="2" pageNumber="3">H</emphasis>
|
||
),
|
||
<taxonomicName authority="Mennecart & Aiglstorfer & Li & Li & Wang, 2021" authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" box="[1072,1177,1525,1544]" class="Mammalia" family="Lophiomerycidae" genus="Chiyoumeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="genus" status="nov. gen.">
|
||
<emphasis box="[1072,1177,1525,1544]" italics="true" pageId="2" pageNumber="3">Chiyoumeryx</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1181,1252,1526,1545]" pageId="2" pageNumber="3" rank="genus">nov. gen.</taxonomicNameLabel>
|
||
<emphasis box="[1255,1342,1525,1544]" italics="true" pageId="2" pageNumber="3">shinaoensis</emphasis>
|
||
(Miao, 1982)
|
||
<bibRefCitation author="Miao, D." box="[1449,1461,1524,1536]" journalOrPublisher="Acta Palaeontol. Sinica" pageId="2" pageNumber="3" pagination="526 - 536" part="21" refId="ref9719" refString="20. Miao, D. Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province. Acta Palaeontol. Sinica 21, 526 - 536 (1982)." title="Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province" type="journal article" year="1982">
|
||
<superScript attach="none" box="[1449,1461,1524,1536]" fontSize="5" pageId="2" pageNumber="3">20</superScript>
|
||
</bibRefCitation>
|
||
(
|
||
<emphasis bold="true" box="[423,436,1549,1568]" pageId="2" pageNumber="3">C</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[445,460,1550,1569]" pageId="2" pageNumber="3">D</emphasis>
|
||
),
|
||
<taxonomicName authority="Mennecart & Aiglstorfer & Li & Li & Wang, 2021" authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" box="[476,581,1549,1568]" class="Mammalia" family="Lophiomerycidae" genus="Chiyoumeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="genus" status="nov. gen.">
|
||
<emphasis box="[476,581,1549,1568]" italics="true" pageId="2" pageNumber="3">Chiyoumeryx</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[585,656,1550,1569]" pageId="2" pageNumber="3" rank="genus">nov. gen.</taxonomicNameLabel>
|
||
<emphasis box="[659,776,1549,1568]" italics="true" pageId="2" pageNumber="3">flavimperatoris</emphasis>
|
||
nov. sp. (
|
||
<emphasis bold="true" box="[850,862,1550,1569]" pageId="2" pageNumber="3">E</emphasis>
|
||
) and
|
||
<taxonomicName authority="Mennecart & Aiglstorfer & Li & Li & Wang, 2021" authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" box="[907,1046,1549,1568]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="miaoi" status="nov. sp.">
|
||
<emphasis box="[907,1046,1549,1568]" italics="true" pageId="2" pageNumber="3">Iberomeryx miaoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1050,1110,1550,1569]" pageId="2" pageNumber="3" rank="species">nov. sp.</taxonomicNameLabel>
|
||
(
|
||
<emphasis bold="true" box="[1120,1148,1550,1569]" pageId="2" pageNumber="3">F–I</emphasis>
|
||
).
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[1164,1315,1549,1568]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="gracilis" status="nov. comb.">
|
||
<emphasis box="[1164,1315,1549,1568]" italics="true" pageId="2" pageNumber="3">Krabimeryx gracilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1319,1407,1549,1569]" pageId="2" pageNumber="3" rank="species">nov. comb.</taxonomicNameLabel>
|
||
(Miao, 1982)
|
||
<bibRefCitation author="Miao, D." box="[461,473,1572,1584]" journalOrPublisher="Acta Palaeontol. Sinica" pageId="2" pageNumber="3" pagination="526 - 536" part="21" refId="ref9719" refString="20. Miao, D. Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province. Acta Palaeontol. Sinica 21, 526 - 536 (1982)." title="Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province" type="journal article" year="1982">
|
||
<superScript attach="right" box="[461,473,1572,1584]" fontSize="5" pageId="2" pageNumber="3">20</superScript>
|
||
</bibRefCitation>
|
||
: (
|
||
<emphasis bold="true" box="[489,503,1573,1592]" pageId="2" pageNumber="3">A</emphasis>
|
||
) IVPP V 6546-1 (holotype), partial skull with right and lef M1–M3; (
|
||
<emphasis bold="true" box="[1055,1068,1574,1593]" pageId="2" pageNumber="3">B</emphasis>
|
||
) IVPP V 6546-2 (holotype), right fragmented mandible with m2–m3.
|
||
<taxonomicName authority="Mennecart & Aiglstorfer & Li & Li & Wang, 2021" authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" box="[605,710,1597,1616]" class="Mammalia" family="Lophiomerycidae" genus="Chiyoumeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="genus" status="nov.gen.">
|
||
<emphasis box="[605,710,1597,1616]" italics="true" pageId="2" pageNumber="3">Chiyoumeryx</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[714,786,1598,1617]" pageId="2" pageNumber="3" rank="genus">nov.gen.</taxonomicNameLabel>
|
||
<emphasis box="[789,877,1597,1616]" italics="true" pageId="2" pageNumber="3">shinaoensis</emphasis>
|
||
(Miao, 1982)
|
||
<bibRefCitation author="Miao, D." box="[982,994,1596,1608]" journalOrPublisher="Acta Palaeontol. Sinica" pageId="2" pageNumber="3" pagination="526 - 536" part="21" refId="ref9719" refString="20. Miao, D. Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province. Acta Palaeontol. Sinica 21, 526 - 536 (1982)." title="Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province" type="journal article" year="1982">
|
||
<superScript attach="none" box="[982,994,1596,1608]" fontSize="5" pageId="2" pageNumber="3">20</superScript>
|
||
</bibRefCitation>
|
||
: (
|
||
<emphasis bold="true" box="[1010,1023,1597,1616]" pageId="2" pageNumber="3">C</emphasis>
|
||
) IVPP V 6531 (holotype), right mandible with p2–m3 and tooth socket of p1; (
|
||
<emphasis bold="true" box="[607,622,1622,1641]" pageId="2" pageNumber="3">D</emphasis>
|
||
) IVPP V 6532 (paratype), right fragmented maxillary with P4-M3.
|
||
<taxonomicName authority="Mennecart & Aiglstorfer & Li & Li & Wang, 2021" authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" box="[1155,1260,1621,1640]" class="Mammalia" family="Lophiomerycidae" genus="Chiyoumeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="genus" status="nov. gen.">
|
||
<emphasis box="[1155,1260,1621,1640]" italics="true" pageId="2" pageNumber="3">Chiyoumeryx</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1264,1335,1622,1641]" pageId="2" pageNumber="3" rank="genus">nov. gen.</taxonomicNameLabel>
|
||
<emphasis box="[1338,1455,1621,1640]" italics="true" pageId="2" pageNumber="3">flavimperatoris</emphasis>
|
||
nov. sp.: (
|
||
<emphasis bold="true" box="[490,502,1646,1665]" pageId="2" pageNumber="3">E</emphasis>
|
||
) IVPP V 6547 (holotype), right mandible with p4–m3;
|
||
<taxonomicName authority="Mennecart & Aiglstorfer & Li & Li & Wang, 2021" authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" box="[939,1078,1645,1664]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="miaoi" status="nov. sp.">
|
||
<emphasis box="[939,1078,1645,1664]" italics="true" pageId="2" pageNumber="3">Iberomeryx miaoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1082,1143,1646,1665]" pageId="2" pageNumber="3" rank="species">nov. sp.</taxonomicNameLabel>
|
||
: (
|
||
<emphasis bold="true" box="[1156,1167,1646,1665]" pageId="2" pageNumber="3">F</emphasis>
|
||
) IVPP V 6551 (holotype), lef mandible with m1–m3 (mirrored); (
|
||
<emphasis bold="true" box="[702,716,1669,1688]" pageId="2" pageNumber="3">G</emphasis>
|
||
) lower molar
|
||
<taxonomicName authorityName="Janis" authorityYear="1987" box="[827,966,1669,1688]" class="Mammalia" family="Lophiomerycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="family">Lophiomerycidae</taxonomicName>
|
||
dental nomenclature (based on the m3 of IVPP 6546-2): 1 internal postmetacristid, 2 metaconid, 3 external postmetacristid, 4 internal preentocristid, 5 entoconidian groove, 6 external preentocristid, 7 entoconid, 8 posthypoconulidcristid, 9 hypoconulid, 10 prehypoconuldicristid, 11 posthypocristid, 12 hypoconid, 13 prehypocristid, 14 ectostylid, 15 postprotocristid, 16 protoconid, 17 preprotocristid, 18 anterior cingulid; (
|
||
<emphasis bold="true" box="[1274,1290,1742,1761]" pageId="2" pageNumber="3">H</emphasis>
|
||
) upper molar
|
||
<taxonomicName authorityName="Janis" authorityYear="1987" box="[416,555,1765,1784]" class="Mammalia" family="Lophiomerycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="family">Lophiomerycidae</taxonomicName>
|
||
dental nomenclature (based on the M2 of IVPP 6546-1): 1 postmetacrista, 2 metacone, 3 premetacrista, 4 mesostyle, 5 postparacrista, 6 paracone, 7 paraconid labial groove, 8 preparacrista, 9 parastyle, 10 preprotocrista, 11 anterolingual cingulum, 12 protocone, 13 postprotocrista, 14 entostyle, 15 additional cone, 16 premetaconulecrista, 17 metaconule, 18 postmetaconulecrista; (
|
||
<emphasis bold="true" box="[1440,1447,1814,1833]" pageId="2" pageNumber="3">I</emphasis>
|
||
) lower molar
|
||
<taxonomicName authorityName="Milne-Edwards" authorityYear="1864" box="[516,600,1837,1856]" class="Mammalia" family="Tragulidae" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="family">Tragulidae</taxonomicName>
|
||
dental nomenclature (based on the m2 of IVPP V 6551, reversed): 1 metaconid, 2 external postmetacristid, 3
|
||
<taxonomicName box="[416,525,1861,1880]" class="Mammalia" family="Tragulidae" genus="Dorcatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis box="[416,525,1861,1880]" italics="true" pageId="2" pageNumber="3">Dorcatherium</emphasis>
|
||
</taxonomicName>
|
||
fold, 4 internal postmetacristid, 5 preentocristid, 6 entoconid, 7 postentocristid, 8 posterior cingulid, 9 posthypocristid, 10 hypoconid, 11 prehypocristid, 12 ectostylid, 13 external postprotocristid, 14
|
||
<taxonomicName authorityName="Brisson" authorityYear="1762" box="[1040,1108,1885,1904]" class="Mammalia" family="Tragulidae" genus="Tragulus" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1040,1108,1885,1904]" italics="true" pageId="2" pageNumber="3">Tragulus</emphasis>
|
||
</taxonomicName>
|
||
fold, 15 internal postprotocristid, 16 protoconid, 17 preprotocristid, 18 paraconid, 19 preparacristid. (
|
||
<emphasis bold="true" box="[949,956,1910,1929]" pageId="2" pageNumber="3">J</emphasis>
|
||
) phylogenetic position and stratigraphie of the Shinao/ Yangjiachong/Xiaerhete ruminants (topology
|
||
<bibRefCitation author="Metais, G. & Vislobokova, I." box="[773,779,1932,1944]" editor="Prothero, D. R. & Foss, S. C." journalOrPublisher="Te Johns Hopkins University Press" pageId="2" pageNumber="3" pagination="189 - 212" refId="ref8834" refString="2. Metais, G. & Vislobokova, I. Basal ruminants. In Te Evolution of Artiodactyls (eds Prothero, D. R. & Foss, S. C.) 189 - 212 (Te Johns Hopkins University Press, 2007)." title="Basal ruminants" type="book chapter" volumeTitle="Te Evolution of Artiodactyls" year="2007">
|
||
<superScript attach="left" box="[773,779,1932,1944]" fontSize="5" pageId="2" pageNumber="3">2</superScript>
|
||
</bibRefCitation>
|
||
). a stem
|
||
<taxonomicName authorityName="Scopoli" authorityYear="1777" box="[850,945,1933,1952]" pageId="2" pageNumber="10" rank="subOrder" subOrder="Ruminantia">Ruminantia</taxonomicName>
|
||
, b
|
||
<taxonomicName authorityName="Matthew & Granger" authorityYear="1925" box="[967,1079,1933,1952]" class="Mammalia" family="Hypertragulidae" genus="Archaeomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis box="[967,1079,1933,1952]" italics="true" pageId="2" pageNumber="3">Archaeomeryx</emphasis>
|
||
</taxonomicName>
|
||
, c
|
||
<taxonomicName authority="Mennecart & Aiglstorfer & Li & Li & Wang, 2021" authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" box="[1101,1206,1933,1952]" class="Mammalia" family="Lophiomerycidae" genus="Chiyoumeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="genus" status="nov.gen.">
|
||
<emphasis box="[1101,1206,1933,1952]" italics="true" pageId="2" pageNumber="3">Chiyoumeryx</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1210,1282,1934,1953]" pageId="2" pageNumber="3" rank="genus">nov.gen.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[1319,1469,1933,1952]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[1319,1469,1933,1952]" italics="true" pageId="2" pageNumber="3">Krabimeryx gracilis</emphasis>
|
||
</taxonomicName>
|
||
, d crown
|
||
<taxonomicName authorityName="Scopoli" authorityYear="1777" box="[485,580,1957,1976]" pageId="2" pageNumber="10" rank="subOrder" subOrder="Ruminantia">Ruminantia</taxonomicName>
|
||
, e
|
||
<taxonomicName authority="Mennecart & Aiglstorfer & Li & Li & Wang, 2021" authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" box="[600,739,1957,1976]" class="Mammalia" family="Lophiomerycidae" genus="Iberomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="miaoi" status="nov. sp.">
|
||
<emphasis box="[600,739,1957,1976]" italics="true" pageId="2" pageNumber="3">Iberomeryx miaoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[743,804,1958,1977]" pageId="2" pageNumber="3" rank="species">nov. sp.</taxonomicNameLabel>
|
||
; 1 lingual view, 2 occlusal view. Scale bare is 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="2.[1467,1478,2024,2043]" box="[1467,1478,2024,2043]" pageId="2" pageNumber="3">3</paragraph>
|
||
<paragraph blockId="3.[416,1479,137,240]" pageId="3" pageNumber="4">
|
||
<emphasis bold="true" box="[416,580,137,161]" pageId="3" pageNumber="4">Neodiagnosis.</emphasis>
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[607,791,138,159]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[607,791,138,159]" italics="true" pageId="3" pageNumber="4">Krabimeryx gracilis</emphasis>
|
||
</taxonomicName>
|
||
has an m2 that is wider than the m3; this is the other way round in
|
||
<emphasis italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
. Moreover, the entoconid is less anterior relative to the hypoconid in
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[1173,1266,165,186]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[1173,1266,165,186]" italics="true" pageId="3" pageNumber="4">K. gracilis</emphasis>
|
||
</taxonomicName>
|
||
than it is in
|
||
<emphasis italics="true" pageId="3" pageNumber="4">K primitivus</emphasis>
|
||
. Te ectostylid is large in
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[695,787,191,213]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[695,787,191,213]" italics="true" pageId="3" pageNumber="4">K. gracilis</emphasis>
|
||
</taxonomicName>
|
||
, while it is absent in
|
||
<emphasis box="[987,1110,192,213]" italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
. Te cingulum on the upper molars in
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[416,509,218,240]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[416,509,218,240]" italics="true" pageId="3" pageNumber="4">K. gracilis</emphasis>
|
||
</taxonomicName>
|
||
is more developed than in
|
||
<emphasis box="[768,892,219,240]" italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="4" type="materials_examined">
|
||
<paragraph blockId="3.[416,1478,276,325]" pageId="3" pageNumber="4">
|
||
<emphasis bold="true" box="[416,531,276,300]" pageId="3" pageNumber="4">Holotype.</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3398234309" collectionCode="IVPP" pageId="3" pageNumber="4" specimenCode="IVPP V 6546, IVPP V 6546-1, IVPP V 6546-2" specimenCount="1">
|
||
<specimenCode box="[557,687,277,299]" pageId="3" pageNumber="4">IVPP V 6546</specimenCode>
|
||
, partial skull with right and lef M1–M3 (
|
||
<specimenCode box="[1091,1242,277,299]" pageId="3" pageNumber="4">IVPP V 6546-1</specimenCode>
|
||
) and an associated right fragmented mandible with m2–m3 (
|
||
<specimenCode box="[763,912,303,325]" pageId="3" pageNumber="4">IVPP V 6546-2</specimenCode>
|
||
) found in occlusion with the skull.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="3.[416,1478,361,411]" pageId="3" pageNumber="4">
|
||
<emphasis bold="true" box="[416,646,361,385]" pageId="3" pageNumber="4">Additional material.</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3398234318" box="[673,1139,362,384]" collectionCode="IVPP" pageId="3" pageNumber="4" specimenCode="IVPP V 6549" specimenCount="1">
|
||
<specimenCode box="[673,800,362,384]" pageId="3" pageNumber="4">IVPP V 6549</specimenCode>
|
||
, right m3 on fragmented mandible;
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3398234321" collectionCode="IVPP" pageId="3" pageNumber="4" specimenCode="IVPP V 6550" specimenCount="1">
|
||
<specimenCode box="[1144,1269,362,384]" pageId="3" pageNumber="4">IVPP V 6550</specimenCode>
|
||
lef fragmented mandible with m1–m2;
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="3398234317" box="[603,1193,389,411]" collectionCode="IVPP" pageId="3" pageNumber="4" specimenCode="IVPP V 26638" specimenCount="1">
|
||
<specimenCode box="[603,742,389,411]" pageId="3" pageNumber="4">IVPP V 26638</specimenCode>
|
||
, right m1. Measurements are given in Table S1.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="3.[416,1478,446,496]" pageId="3" pageNumber="4">
|
||
<emphasis bold="true" box="[416,533,446,470]" pageId="3" pageNumber="4">Localities.</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3398234301" country="China" county="Panxian County" location="Shinao Basin" pageId="3" pageNumber="4" specimenCount="1">
|
||
<location LSID="urn:lsid:plazi:treatment:E97287E44C427A0C1FEAFB6CFADACDC2:640460294C407A0C1C7AFE5DFD18C9C7" box="[560,685,447,469]" country="China" county="Panxian County" name="Shinao Basin" pageId="3" pageNumber="4">Shinao Basin</location>
|
||
,
|
||
<collectingCounty box="[693,847,447,469]" pageId="3" pageNumber="4">Panxian County</collectingCounty>
|
||
,
|
||
<locationDeviation box="[854,1074,447,469]" pageId="3" pageNumber="4">Southwestern Guizhou</locationDeviation>
|
||
,
|
||
<collectingCountry box="[1082,1143,447,469]" name="China" pageId="3" pageNumber="4">China</collectingCountry>
|
||
;
|
||
<location LSID="urn:lsid:plazi:treatment:E97287E44C427A0C1FEAFB6CFADACDC2:640460294C407A0C1A34FE5DFA91C9C7" box="[1150,1316,447,469]" country="China" county="Panxian County" name="Xiaerhete locality" pageId="3" pageNumber="4">Xiaerhete locality</location>
|
||
,
|
||
<collectingCounty box="[1324,1474,447,469]" pageId="3" pageNumber="4">Jiminay County</collectingCounty>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:E97287E44C427A0C1FEAFB6CFADACDC2:640460294C407A0C1FEAFE38FDB5C9E2" box="[416,512,474,496]" country="China" county="Panxian County" name="Xingjiang" pageId="3" pageNumber="4">Xingjiang</location>
|
||
, China.
|
||
<geologicalTimeScale box="[590,708,475,496]" pageId="3" pageNumber="4">Late Eocene</geologicalTimeScale>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="4" type="description">
|
||
<paragraph blockId="3.[416,1479,532,1514]" pageId="3" pageNumber="4">
|
||
<emphasis bold="true" box="[416,707,532,556]" pageId="3" pageNumber="4">Taxonomical attribution.</emphasis>
|
||
Te herein described specimens were first attributed to the genus
|
||
<taxonomicName class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="4">Lophiomeryx</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation author="Miao, D." box="[486,501,557,571]" journalOrPublisher="Acta Palaeontol. Sinica" pageId="3" pageNumber="4" pagination="526 - 536" part="21" refId="ref9719" refString="20. Miao, D. Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province. Acta Palaeontol. Sinica 21, 526 - 536 (1982)." title="Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province" type="journal article" year="1982">
|
||
<superScript attach="left" box="[486,501,557,571]" fontSize="6" pageId="3" pageNumber="4">20</superScript>
|
||
</bibRefCitation>
|
||
. However, the thorough reassessment of the specimens now leads to the conclusion that
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis italics="true" pageId="3" pageNumber="4">Lophiomeryx gracilis</emphasis>
|
||
</taxonomicName>
|
||
sensuMiao
|
||
<bibRefCitation author="Miao, D." box="[601,616,583,597]" journalOrPublisher="Acta Palaeontol. Sinica" pageId="3" pageNumber="4" pagination="526 - 536" part="21" refId="ref9719" refString="20. Miao, D. Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province. Acta Palaeontol. Sinica 21, 526 - 536 (1982)." title="Early Tertiary fossil mammals from the Shinao Basin, Panxian County, Guizhou Province" type="journal article" year="1982">
|
||
<superScript attach="left" box="[601,616,583,597]" fontSize="6" pageId="3" pageNumber="4">20</superScript>
|
||
</bibRefCitation>
|
||
contains three different species and genera, but none of them can be assigned to
|
||
<taxonomicName class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="4">Lophiomeryx</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="3.[416,1479,532,1514]" pageId="3" pageNumber="4">
|
||
Based on the presence of a strong lingual cingulum in upper molars and a short anteroposteriorly oriented postprotocrista, as well as the absence of a premetacristid and an anterior fossa widely open in the lower molars, we can conclude that the specimens, IVPP V 6546-1, IVPP V 6546-2, IVPP V 6549, and IVPP V 6550, belong to
|
||
<taxonomicName authorityName="Janis" authorityYear="1987" box="[416,586,719,741]" class="Mammalia" family="Lophiomerycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="family">Lophiomerycidae</taxonomicName>
|
||
or
|
||
<taxonomicName authorityName="Milne-Edwards" authorityYear="1864" box="[618,721,719,741]" class="Mammalia" family="Tragulidae" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="family">Tragulidae</taxonomicName>
|
||
<superScript attach="left" box="[721,755,716,731]" fontSize="6" pageId="3" pageNumber="4">
|
||
<bibRefCitation author="Metais, G. & Welcomme, J. - L. & Ducrocq, S." box="[721,736,716,730]" journalOrPublisher="J. Vertebr. Paleontol." pageId="3" pageNumber="4" pagination="231 - 241" part="29" refId="ref10379" refString="35. Metais, G., Welcomme, J. - L. & Ducrocq, S. New lophiomerycid ruminants from the Oligocene of the Bugti Hills (Balochistan, Pakistan). J. Vertebr. Paleontol. 29, 231 - 241 (2009)." title="New lophiomerycid ruminants from the Oligocene of the Bugti Hills (Balochistan, Pakistan)" type="journal article" year="2009">35</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[740,755,716,730]" journalOrPublisher="Swiss J. Geosci." pageId="3" pageNumber="4" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">36</bibRefCitation>
|
||
</superScript>
|
||
. However, the absence of a large paraconid and the absence of an elongated external postmetacristid distinguish the specimens from primitive
|
||
<taxonomicName authorityName="Milne-Edwards" authorityYear="1864" box="[1059,1162,746,768]" class="Mammalia" family="Tragulidae" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="family">Tragulidae</taxonomicName>
|
||
<superScript attach="left" box="[1162,1196,743,758]" fontSize="6" pageId="3" pageNumber="4">
|
||
<bibRefCitation author="Metais, G. & Chaimanee, Y. & Jaeger, J. - J. & Ducrocq, S." box="[1162,1177,743,757]" journalOrPublisher="Zool. Scr." pageId="3" pageNumber="4" pagination="231 - 248" part="30" refId="ref9541" refString="17. Metais, G., Chaimanee, Y., Jaeger, J. - J. & Ducrocq, S. New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia. Zool. Scr. 30, 231 - 248 (2001)." title="New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia" type="journal article" year="2001">17</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Mennecart, B. & Becker, D. & Berger, J. - P." box="[1181,1196,743,757]" journalOrPublisher="Swiss J. Geosci." pageId="3" pageNumber="4" pagination="S 115 - S 132" part="104" refId="ref10429" refString="36. Mennecart, B., Becker, D. & Berger, J. - P. Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet. Swiss J. Geosci. 104, S 115 - S 132 (2011)." title="Iberomeryx minor (Mammalia, Artiodactyla) from the Early Oligocene of Soulce (Canton Jura, NW Switzerland): Systematics and palaeodiet" type="journal article" year="2011">36</bibRefCitation>
|
||
</superScript>
|
||
. In
|
||
<emphasis box="[1234,1437,746,768]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName box="[1234,1351,746,767]" class="Mammalia" family="Lophiomerycidae" genus="Zhailimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">Zhailimeryx</taxonomicName>
|
||
jingweni
|
||
</emphasis>
|
||
, the cuspids are more slender than in the herein described specimens
|
||
<bibRefCitation author="Guo, J. & Dawson, M. & Beard, K. C." box="[1046,1061,770,784]" journalOrPublisher="J. Mamm. Evol." pageId="3" pageNumber="4" pagination="239 - 258" part="7" refId="ref9394" refString="14. Guo, J., Dawson, M. & Beard, K. C. Zhailimeryx, a new Lophiomerycid Artiodactyl (Mammalia) from the Late Middle Eocene of Central China and early evolution of ruminants. J. Mamm. Evol. 7, 239 - 258 (2000)." title="Zhailimeryx, a new Lophiomerycid Artiodactyl (Mammalia) from the Late Middle Eocene of Central China and early evolution of ruminants" type="journal article" year="2000">
|
||
<superScript attach="left" box="[1046,1061,770,784]" fontSize="6" pageId="3" pageNumber="4">14</superScript>
|
||
</bibRefCitation>
|
||
, a feature the taxon shares with
|
||
<emphasis italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
. In
|
||
<emphasis box="[487,594,800,821]" italics="true" pageId="3" pageNumber="4">Z. jingweni</emphasis>
|
||
, m1 and m2 are of relative similar width
|
||
<bibRefCitation author="Guo, J. & Dawson, M. & Beard, K. C." box="[992,1007,797,811]" journalOrPublisher="J. Mamm. Evol." pageId="3" pageNumber="4" pagination="239 - 258" part="7" refId="ref9394" refString="14. Guo, J., Dawson, M. & Beard, K. C. Zhailimeryx, a new Lophiomerycid Artiodactyl (Mammalia) from the Late Middle Eocene of Central China and early evolution of ruminants. J. Mamm. Evol. 7, 239 - 258 (2000)." title="Zhailimeryx, a new Lophiomerycid Artiodactyl (Mammalia) from the Late Middle Eocene of Central China and early evolution of ruminants" type="journal article" year="2000">
|
||
<superScript attach="left" box="[992,1007,797,811]" fontSize="6" pageId="3" pageNumber="4">14</superScript>
|
||
</bibRefCitation>
|
||
, while in
|
||
<emphasis box="[1104,1230,800,821]" italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
and the herein described specimens from Shinao the m2 is clearly bigger than the m
|
||
<superScript attach="right" box="[998,1013,823,837]" fontSize="6" pageId="3" pageNumber="4">117</superScript>
|
||
. Similarly to
|
||
<emphasis box="[1143,1268,827,848]" italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
, the herein described specimens differ from
|
||
<emphasis box="[635,742,853,874]" italics="true" pageId="3" pageNumber="4">Z. jingweni</emphasis>
|
||
in its lower molar lingual cusps being more laterally compressed, and in an entoconid that is slightly shifed to anterior with respect to the hypoconid, while it is more posterior in
|
||
<emphasis italics="true" pageId="3" pageNumber="4">Z. jingweni</emphasis>
|
||
<superScript attach="left" box="[470,503,903,918]" fontSize="6" pageId="3" pageNumber="4">
|
||
<bibRefCitation author="Guo, J. & Dawson, M. & Beard, K. C." box="[470,485,903,917]" journalOrPublisher="J. Mamm. Evol." pageId="3" pageNumber="4" pagination="239 - 258" part="7" refId="ref9394" refString="14. Guo, J., Dawson, M. & Beard, K. C. Zhailimeryx, a new Lophiomerycid Artiodactyl (Mammalia) from the Late Middle Eocene of Central China and early evolution of ruminants. J. Mamm. Evol. 7, 239 - 258 (2000)." title="Zhailimeryx, a new Lophiomerycid Artiodactyl (Mammalia) from the Late Middle Eocene of Central China and early evolution of ruminants" type="journal article" year="2000">14</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Metais, G. & Chaimanee, Y. & Jaeger, J. - J. & Ducrocq, S." box="[488,503,903,917]" journalOrPublisher="Zool. Scr." pageId="3" pageNumber="4" pagination="231 - 248" part="30" refId="ref9541" refString="17. Metais, G., Chaimanee, Y., Jaeger, J. - J. & Ducrocq, S. New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia. Zool. Scr. 30, 231 - 248 (2001)." title="New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia" type="journal article" year="2001">17</bibRefCitation>
|
||
</superScript>
|
||
. Furthermore,
|
||
<emphasis box="[645,766,907,928]" italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
and the herein described specimens from Shinao both lack the rudimentary paraconid present in
|
||
<emphasis box="[620,726,933,954]" italics="true" pageId="3" pageNumber="4">Z. jingweni</emphasis>
|
||
<superScript attach="left" box="[726,760,930,944]" fontSize="6" pageId="3" pageNumber="4">
|
||
<bibRefCitation author="Guo, J. & Dawson, M. & Beard, K. C." box="[726,741,930,944]" journalOrPublisher="J. Mamm. Evol." pageId="3" pageNumber="4" pagination="239 - 258" part="7" refId="ref9394" refString="14. Guo, J., Dawson, M. & Beard, K. C. Zhailimeryx, a new Lophiomerycid Artiodactyl (Mammalia) from the Late Middle Eocene of Central China and early evolution of ruminants. J. Mamm. Evol. 7, 239 - 258 (2000)." title="Zhailimeryx, a new Lophiomerycid Artiodactyl (Mammalia) from the Late Middle Eocene of Central China and early evolution of ruminants" type="journal article" year="2000">14</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Metais, G. & Chaimanee, Y. & Jaeger, J. - J. & Ducrocq, S." box="[745,760,930,944]" journalOrPublisher="Zool. Scr." pageId="3" pageNumber="4" pagination="231 - 248" part="30" refId="ref9541" refString="17. Metais, G., Chaimanee, Y., Jaeger, J. - J. & Ducrocq, S. New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia. Zool. Scr. 30, 231 - 248 (2001)." title="New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia" type="journal article" year="2001">17</bibRefCitation>
|
||
</superScript>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="3.[416,1479,532,1514]" pageId="3" pageNumber="4">
|
||
Like
|
||
<emphasis box="[493,614,960,981]" italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
, the here-described specimens differ from
|
||
<taxonomicName authority="Mennecart & Aiglstorfer & Li & Li & Wang, 2021" authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" box="[1016,1141,959,980]" class="Mammalia" family="Lophiomerycidae" genus="Chiyoumeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus" status="nov. gen.">
|
||
<emphasis box="[1016,1141,959,980]" italics="true" pageId="3" pageNumber="4">Chiyoumeryx</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1146,1230,960,981]" pageId="3" pageNumber="4" rank="genus">nov. gen.</taxonomicNameLabel>
|
||
(described below) and the
|
||
<taxonomicName box="[416,614,986,1008]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis box="[416,541,986,1007]" italics="true" pageId="3" pageNumber="4">Lophiomeryx</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
<emphasis box="[620,747,986,1008]" italics="true" pageId="3" pageNumber="4">L. mouchelini</emphasis>
|
||
,
|
||
<emphasis box="[759,877,986,1008]" italics="true" pageId="3" pageNumber="4">L. chalaniati</emphasis>
|
||
and
|
||
<emphasis box="[924,1023,987,1008]" italics="true" pageId="3" pageNumber="4">L. angarae</emphasis>
|
||
by having more massive and more bunomorph lowermolars
|
||
<superScript attach="left" box="[542,632,1010,1024]" fontSize="6" pageId="3" pageNumber="4">
|
||
<bibRefCitation author="Mennecart, B. Te" box="[542,557,1010,1024]" journalOrPublisher="PLoS ONE" pageId="3" pageNumber="4" pagination="0116830" part="10" refId="ref9496" refString="16. Mennecart, B. Te European ruminants during the " Microbunodon Event " (MP 28, latest Oligocene): Impact of climate changes and faunal event on the ruminant evolution. PLoS ONE 10, e 0116830 (2015)." title="European ruminants during the " Microbunodon Event " (MP 28, latest Oligocene): Impact of climate changes and faunal event on the ruminant evolution" type="journal article" year="2015">16</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Metais, G. & Chaimanee, Y. & Jaeger, J. - J. & Ducrocq, S." box="[560,575,1010,1024]" journalOrPublisher="Zool. Scr." pageId="3" pageNumber="4" pagination="231 - 248" part="30" refId="ref9541" refString="17. Metais, G., Chaimanee, Y., Jaeger, J. - J. & Ducrocq, S. New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia. Zool. Scr. 30, 231 - 248 (2001)." title="New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia" type="journal article" year="2001">17</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Matthew, W. D. & Granger, W." box="[579,594,1010,1024]" journalOrPublisher="Am. Mus. Novit." pageId="3" pageNumber="4" pagination="1 - 12" part="195" refId="ref9878" refString="24. Matthew, W. D. & Granger, W. New ungulates from the Ardyn Obo Formation of Mongolia. Am. Mus. Novit. 195, 1 - 12 (1925)." title="New ungulates from the Ardyn Obo Formation of Mongolia" type="journal article" year="1925">24</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Mennecart, B. Te" box="[598,613,1010,1024]" journalOrPublisher="GeoFocus" pageId="3" pageNumber="4" pagination="1 - 263" part="32" refId="ref10339" refString="34. Mennecart, B. Te Ruminantia (Mammalia, Cetartiodactyla) from the Oligocene to the Early Miocene of Western Europe: Systematics, palaeoecology and palaeobiogeography. GeoFocus 32, 1 - 263 (2012)." title="Ruminantia (Mammalia, Cetartiodactyla) from the Oligocene to the Early Miocene of Western Europe: Systematics, palaeoecology and palaeobiogeography" type="journal article" year="2012">34</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Brunet, M. & Sudre, J." box="[617,632,1010,1024]" journalOrPublisher="Munch. Geowiss. Abh." pageId="3" pageNumber="4" pagination="225 - 242" part="10" refId="ref10489" refString="37. Brunet, M. & Sudre, J. Evolution et systematique du genre Lophiomeryx POMEL 1853 (Mammalia, Artiodactyla). Munch. Geowiss. Abh. 10, 225 - 242 (1987)." title="Evolution et systematique du genre Lophiomeryx POMEL 1853 (Mammalia, Artiodactyla)" type="journal article" year="1987">37</bibRefCitation>
|
||
</superScript>
|
||
. Furthermore,
|
||
<emphasis box="[780,983,1013,1034]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName box="[780,897,1013,1034]" class="Mammalia" family="Lophiomerycidae" genus="Zhailimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">Zhailimeryx</taxonomicName>
|
||
jingweni
|
||
</emphasis>
|
||
,
|
||
<emphasis box="[994,1120,1013,1034]" italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
, and the herein described specimens differ from
|
||
<taxonomicName box="[525,650,1039,1060]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis box="[525,650,1039,1060]" italics="true" pageId="3" pageNumber="4">Lophiomeryx</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a developed external postmetacristid and by a distinct entoconidian groove on the anterior side of the compressed entoconid
|
||
<superScript attach="left" box="[972,1006,1063,1078]" fontSize="6" pageId="3" pageNumber="4">
|
||
<bibRefCitation author="Guo, J. & Dawson, M. & Beard, K. C." box="[972,987,1063,1077]" journalOrPublisher="J. Mamm. Evol." pageId="3" pageNumber="4" pagination="239 - 258" part="7" refId="ref9394" refString="14. Guo, J., Dawson, M. & Beard, K. C. Zhailimeryx, a new Lophiomerycid Artiodactyl (Mammalia) from the Late Middle Eocene of Central China and early evolution of ruminants. J. Mamm. Evol. 7, 239 - 258 (2000)." title="Zhailimeryx, a new Lophiomerycid Artiodactyl (Mammalia) from the Late Middle Eocene of Central China and early evolution of ruminants" type="journal article" year="2000">14</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Metais, G. & Chaimanee, Y. & Jaeger, J. - J. & Ducrocq, S." box="[991,1006,1063,1077]" journalOrPublisher="Zool. Scr." pageId="3" pageNumber="4" pagination="231 - 248" part="30" refId="ref9541" refString="17. Metais, G., Chaimanee, Y., Jaeger, J. - J. & Ducrocq, S. New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia. Zool. Scr. 30, 231 - 248 (2001)." title="New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia" type="journal article" year="2001">17</bibRefCitation>
|
||
</superScript>
|
||
. In
|
||
<taxonomicName box="[1046,1173,1066,1087]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1046,1173,1066,1087]" italics="true" pageId="3" pageNumber="4">Lophiomeryx</emphasis>
|
||
</taxonomicName>
|
||
, the back fossa of m3 is widely open due to the strong reduction of the posthypoconulidcristid
|
||
<superScript attach="left" box="[1031,1064,1090,1104]" fontSize="6" pageId="3" pageNumber="4">
|
||
<bibRefCitation author="Mennecart, B. Te" box="[1031,1046,1090,1104]" journalOrPublisher="GeoFocus" pageId="3" pageNumber="4" pagination="1 - 263" part="32" refId="ref10339" refString="34. Mennecart, B. Te Ruminantia (Mammalia, Cetartiodactyla) from the Oligocene to the Early Miocene of Western Europe: Systematics, palaeoecology and palaeobiogeography. GeoFocus 32, 1 - 263 (2012)." title="Ruminantia (Mammalia, Cetartiodactyla) from the Oligocene to the Early Miocene of Western Europe: Systematics, palaeoecology and palaeobiogeography" type="journal article" year="2012">34</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Brunet, M. & Sudre, J." box="[1049,1064,1090,1104]" journalOrPublisher="Munch. Geowiss. Abh." pageId="3" pageNumber="4" pagination="225 - 242" part="10" refId="ref10489" refString="37. Brunet, M. & Sudre, J. Evolution et systematique du genre Lophiomeryx POMEL 1853 (Mammalia, Artiodactyla). Munch. Geowiss. Abh. 10, 225 - 242 (1987)." title="Evolution et systematique du genre Lophiomeryx POMEL 1853 (Mammalia, Artiodactyla)" type="journal article" year="1987">37</bibRefCitation>
|
||
</superScript>
|
||
. In contrast to this,
|
||
<emphasis box="[1261,1478,1093,1114]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName box="[1261,1374,1093,1114]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">Krabimeryx</taxonomicName>
|
||
primitivus
|
||
</emphasis>
|
||
possesses a clearly developed posthypoconulidcristid forming a buckle on the m3 back basin
|
||
<bibRefCitation author="Metais, G. & Chaimanee, Y. & Jaeger, J. - J. & Ducrocq, S." box="[1306,1321,1117,1131]" journalOrPublisher="Zool. Scr." pageId="3" pageNumber="4" pagination="231 - 248" part="30" refId="ref9541" refString="17. Metais, G., Chaimanee, Y., Jaeger, J. - J. & Ducrocq, S. New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia. Zool. Scr. 30, 231 - 248 (2001)." title="New remains of primitive ruminants from Tailand: evidence of the early evolution of the Ruminantia in Asia" type="journal article" year="2001">
|
||
<superScript attach="left" box="[1306,1321,1117,1131]" fontSize="6" pageId="3" pageNumber="4">17</superScript>
|
||
</bibRefCitation>
|
||
, similarly to the specimens from Shinao described here.
|
||
</paragraph>
|
||
<paragraph blockId="3.[416,1479,532,1514]" pageId="3" pageNumber="4">
|
||
Summing up, the general morphology of the teeth in the herein described specimens is most similar to the one observed in
|
||
<emphasis box="[573,696,1200,1221]" italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
. Tey both share a similar huge notch in lateral view between the third lobe of m3 and the entoconid and the entoconidian groove, features that clearly distinguishing them both from
|
||
<taxonomicName box="[1354,1477,1226,1247]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1354,1477,1226,1247]" italics="true" pageId="3" pageNumber="4">Lophiomeryx</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[458,575,1253,1274]" class="Mammalia" family="Lophiomerycidae" genus="Zhailimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis box="[458,575,1253,1274]" italics="true" pageId="3" pageNumber="4">Zhailimeryx</emphasis>
|
||
</taxonomicName>
|
||
. Tus, we attribute the specimens IVPP V 6546-1, IVPP V 6546-2, IVPP V 6549, and IVPP V 6550 to the genus
|
||
<taxonomicName box="[590,703,1279,1300]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis box="[590,703,1279,1300]" italics="true" pageId="3" pageNumber="4">Krabimeryx</emphasis>
|
||
</taxonomicName>
|
||
. However, significant differences occur with the type species, ruling out the synonymisation of
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[566,660,1306,1328]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gracilis" status="nov. comb.">
|
||
<emphasis box="[566,660,1306,1328]" italics="true" pageId="3" pageNumber="4">K. gracilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[665,770,1306,1328]" pageId="3" pageNumber="4" rank="species">nov. comb.</taxonomicNameLabel>
|
||
and
|
||
<emphasis box="[816,1032,1306,1328]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName box="[816,929,1306,1327]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">Krabimeryx</taxonomicName>
|
||
primitivus
|
||
</emphasis>
|
||
. While both species are very similar in size,
|
||
<emphasis italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
has an m3 wider than m2, while it is the converse for
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[1031,1125,1333,1354]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gracilis" status="nov. comb.">
|
||
<emphasis box="[1031,1125,1333,1354]" italics="true" pageId="3" pageNumber="4">K. gracilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1130,1235,1333,1355]" pageId="3" pageNumber="4" rank="species">nov. comb.</taxonomicNameLabel>
|
||
Moreover, the entoconid is less shifed to the anterior with respect to the hypoconid in
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[998,1090,1359,1381]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gracilis" status="nov. comb.">
|
||
<emphasis box="[998,1090,1359,1381]" italics="true" pageId="3" pageNumber="4">K. gracilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1095,1199,1359,1381]" pageId="3" pageNumber="4" rank="species">nov. comb.</taxonomicNameLabel>
|
||
than in
|
||
<emphasis box="[1276,1393,1360,1381]" italics="true" pageId="3" pageNumber="4">K primitivus</emphasis>
|
||
. Tere is no ectostylid in
|
||
<emphasis box="[570,694,1387,1408]" italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
, while it is large in
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[878,972,1386,1408]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gracilis" status="nov. comb.">
|
||
<emphasis box="[878,972,1386,1408]" italics="true" pageId="3" pageNumber="4">K. gracilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[977,1084,1386,1408]" pageId="3" pageNumber="4" rank="species">nov. comb.</taxonomicNameLabel>
|
||
, forming a transverse cristid between the protoconid and the hypoconid. Te cingulum on the upper molars is more developed in
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[1272,1367,1413,1434]" class="Mammalia" family="Lophiomerycidae" genus="Lophiomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gracilis" status="nov. comb.">
|
||
<emphasis box="[1272,1367,1413,1434]" italics="true" pageId="3" pageNumber="4">K. gracilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1372,1478,1413,1435]" pageId="3" pageNumber="4" rank="species">nov. comb.</taxonomicNameLabel>
|
||
than in
|
||
<emphasis box="[490,615,1440,1461]" italics="true" pageId="3" pageNumber="4">K. primitivus</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="3.[416,1479,532,1514]" box="[448,1391,1466,1488]" pageId="3" pageNumber="4">
|
||
Due to these differences we decided to create the new combination
|
||
<taxonomicName authorityName="Mennecart & Aiglstorfer & Li & Li & Wang" authorityYear="2021" baseAuthorityName="Mennecart & Aiglstorfer & Li & Li & Wang" baseAuthorityYear="2021" box="[1095,1280,1466,1487]" class="Mammalia" family="Lophiomerycidae" genus="Krabimeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gracilis" status="nov. comb.">
|
||
<emphasis box="[1095,1280,1466,1487]" italics="true" pageId="3" pageNumber="4">Krabimeryx gracilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1285,1391,1466,1488]" pageId="3" pageNumber="4" rank="species">nov. comb.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |