356 lines
31 KiB
XML
356 lines
31 KiB
XML
<document ID-ISSN="2305-2562" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683667246730" checkinUser="felipe" docAuthor="., D] D, Qgudnrwr, Csuzdi, Cs. & Blanchart, E." docDate="2011" docId="03DC87D4FF8DFF98FE25FACBFC3A46D6" docLanguage="en" docName="AfrInvertebr.52.2.285-294.pdf" docOrigin="African Invertebrates 52 (2)" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.4:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="4" docTitle="Kynotus proboscideus . & D]D & Qgudnrwr & Csuzdi & Blanchart 2011, sp. n." docType="treatment" docVersion="1" lastPageNumber="293" masterDocId="FFE5FFACFF8BFF91FFD5FF85FF8F434E" masterDocTitle="New and little known giant earthworms from Madagascar (Oligochaeta: Kynotidae)" masterLastPageNumber="294" masterPageNumber="285" pageNumber="290" updateTime="1683667307928" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New and little known giant earthworms from Madagascar (Oligochaeta: Kynotidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>D] D</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Qgudnrwr</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Csuzdi, Cs.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Blanchart, E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2011-12-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>52</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>285</mods:start>
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<mods:end>294</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="ISSN">2305-2562</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03DC87D4FF8DFF98FE25FACBFC3A46D6" httpUri="http://treatment.plazi.org/id/03DC87D4FF8DFF98FE25FACBFC3A46D6" lastPageId="9" lastPageNumber="293" pageId="6" pageNumber="290">
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<subSubSection box="[496,827,1358,1384]" pageId="6" pageNumber="290" type="nomenclature">
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<paragraph blockId="6.[496,827,1358,1384]" box="[496,827,1358,1384]" pageId="6" pageNumber="290">
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<heading box="[496,827,1358,1384]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="290" reason="2">
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<taxonomicName authority=". & D]D & Qgudnrwr & Csuzdi & Blanchart, 2011" authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[496,761,1358,1384]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="290" phylum="Annelida" rank="species" species="proboscideus" status="sp. n.">
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<emphasis bold="true" box="[496,761,1358,1384]" pageId="6" pageNumber="290">Kynotus proboscideus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[767,827,1358,1384]" pageId="6" pageNumber="290" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="9" lastPageNumber="293" pageId="6" pageNumber="290" type="description">
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<paragraph blockId="6.[611,713,1400,1426]" box="[611,713,1400,1426]" pageId="6" pageNumber="290">
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<figureCitation box="[611,713,1400,1426]" captionStart-0="Figs 6, 7" captionStart-1="Figs 8, 9" captionStartId-0="7.[144,183,1501,1522]" captionStartId-1="8.[183,222,1630,1651]" captionTargetBox-0="[152,1098,204,1496]" captionTargetBox-1="[187,1132,442,1612]" captionTargetId-0="figure-59@7.[153,1098,204,968]" captionTargetId-1="figure-110@8.[187,1132,950,1612]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figs 6, 7. K. proboscideus sp. n.: (6) general view (A) and enlarged anterior part of the body (B); (7) prostomium." captionText-1="Figs 8, 9. K. proboscideus sp. n.: (8) spermathecae; (9) penial seta: (A) whole seta, (B) tip of the seta, (C) ornamentation around the tip, (D) ornamentation under the tip." pageId="6" pageNumber="290">Figs 6–9</figureCitation>
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</paragraph>
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<paragraph blockId="6.[181,1141,1443,1502]" pageId="6" pageNumber="290">
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Etymology: From Latin
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<emphasis box="[455,563,1446,1470]" italics="true" pageId="6" pageNumber="290">proboscis</emphasis>
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, referring to the presence of two protrusions on the prostomium.
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</paragraph>
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<paragraph blockId="6.[181,1142,1518,1674]" lastBlockId="7.[144,1103,1584,1674]" lastPageId="7" lastPageNumber="291" pageId="6" pageNumber="290">
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Diagnosis: Length alive
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<emphasis box="[455,481,1521,1545]" italics="true" pageId="6" pageNumber="290">ca</emphasis>
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<quantity box="[487,641,1518,1545]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="6" pageNumber="290" unit="mm" value="550.0" valueMax="600.0" valueMin="500.0">500–600 mm</quantity>
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, diameter
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<quantity box="[756,881,1518,1545]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="6" pageNumber="290" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
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; preserved length
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<quantity metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="6" pageNumber="290" unit="mm" value="300.0" valueMax="320.0" valueMin="280.0">280– 320 mm</quantity>
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diameter
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<quantity box="[397,526,1551,1577]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="6" pageNumber="290" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">18–20 mm</quantity>
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. Colour dark red on dorsum, pale on ventrum. Head with two protrusions. Segments 1–3 simple, 3–13 double ringed. Setae
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<emphasis box="[1024,1052,1585,1609]" italics="true" pageId="6" pageNumber="290">ab</emphasis>
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and
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<emphasis box="[1115,1141,1585,1609]" italics="true" pageId="6" pageNumber="290">cd</emphasis>
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appear from segment 17. Male pore on 16.
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<taxonomicName box="[694,800,1615,1641]" class="Ascidiacea" family="Didemnidae" genus="Clitellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aplousobranchia" pageId="6" pageNumber="293" phylum="Chordata" rank="genus">Clitellum</taxonomicName>
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circular on 22–47, 48. Setae closely paired, both
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<emphasis box="[416,444,1650,1674]" italics="true" pageId="6" pageNumber="290">ab</emphasis>
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and
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<emphasis box="[503,529,1650,1674]" italics="true" pageId="6" pageNumber="290">cd</emphasis>
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lateral. Spermathecal pores lateral, 6–9 on each side in 13/14, 14/15, 15/16. Spermathecae large, irregular sac-shaped, duct short. Genital setal glands in 13, 14, 15. Genital setae lanceolate, length 3.2 mm, diameter 0.06 mm, ornamentation dense serrations.
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</paragraph>
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<caption pageId="7" pageNumber="291" startId="7.[144,183,1501,1522]" targetBox="[152,1098,204,1496]" targetPageId="7" targetType="figure">
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<paragraph blockId="7.[144,1103,1501,1546]" pageId="7" pageNumber="291">
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Figs 6, 7.
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<taxonomicName authority=". & D]D & Qgudnrwr & Csuzdi & Blanchart, 2011" authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[242,391,1503,1522]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="291" phylum="Annelida" rank="species" species="proboscideus" status="sp. n.">
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<emphasis box="[242,391,1503,1522]" italics="true" pageId="7" pageNumber="291">K. proboscideus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[399,453,1501,1522]" pageId="7" pageNumber="291" rank="species">sp. n.</taxonomicNameLabel>
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: (6) general view (A) and enlarged anterior part of the body (B); (7) prostomium.
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</paragraph>
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</caption>
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<paragraph blockId="8.[181,1142,203,398]" box="[181,321,203,229]" pageId="8" pageNumber="292">Description:</paragraph>
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<paragraph blockId="8.[181,1142,203,398]" lastBlockId="9.[144,1104,203,1051]" lastPageId="9" lastPageNumber="293" pageId="8" pageNumber="292">
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<emphasis box="[181,284,246,270]" italics="true" pageId="8" pageNumber="292">
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<typeStatus box="[181,284,246,270]" pageId="8" pageNumber="292">Holotype</typeStatus>
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</emphasis>
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: Contracted length
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<quantity box="[498,590,243,270]" metricMagnitude="-1" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="292" unit="mm" value="320.0">320 mm</quantity>
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, diameter after clitellum
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<quantity box="[862,941,243,270]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="292" unit="mm" value="20.0">20 mm</quantity>
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.
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<emphasis box="[949,1134,246,270]" italics="true" pageId="8" pageNumber="292">Segment number</emphasis>
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: 734 (
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<figureCitation box="[244,317,275,301]" captionStart="Figs 6, 7" captionStartId="7.[144,183,1501,1522]" captionTargetBox="[152,1098,204,1496]" captionTargetId="figure-59@7.[153,1098,204,968]" captionTargetPageId="7" captionText="Figs 6, 7. K. proboscideus sp. n.: (6) general view (A) and enlarged anterior part of the body (B); (7) prostomium." pageId="8" pageNumber="292">Fig. 6</figureCitation>
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).
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<emphasis box="[343,457,278,302]" italics="true" pageId="8" pageNumber="292">
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<typeStatus box="[343,457,278,302]" pageId="8" pageNumber="292">Paratypes</typeStatus>
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</emphasis>
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: Contracted length
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<quantity box="[693,850,275,302]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="8" pageNumber="292" unit="mm" value="300.0" valueMax="320.0" valueMin="280.0">280–320 mm</quantity>
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,
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<quantity box="[867,993,275,302]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.0" metricValueMin="1.3" pageId="8" pageNumber="292" unit="mm" value="16.5" valueMax="20.0" valueMin="13.0">13–20 mm</quantity>
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in diameter.
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<emphasis box="[181,371,310,334]" italics="true" pageId="8" pageNumber="292">Segment number</emphasis>
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: 532–601. The weight of the preserved worms ranges between
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<quantity metricMagnitude="-2" metricUnit="kg" metricValue="5.1" metricValueMax="5.8" metricValueMin="4.4" pageId="8" pageNumber="292" unit="g" value="51.0" valueMax="58.0" valueMin="44.0">44– 58 g</quantity>
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.
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<emphasis box="[250,329,342,366]" italics="true" pageId="8" pageNumber="292">Colour</emphasis>
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: Alive dark red on dorsum becoming pale abruptly ventrally from
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<emphasis box="[1108,1134,342,366]" italics="true" pageId="8" pageNumber="292">cd</emphasis>
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, preserved dark brown on dorsum and lighter on ventrum.
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<emphasis box="[842,903,374,398]" italics="true" pageId="8" pageNumber="292">Head</emphasis>
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: Prolobous with two prominent protrusions (
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<figureCitation box="[415,487,203,229]" captionStart="Figs 6, 7" captionStartId="7.[144,183,1501,1522]" captionTargetBox="[152,1098,204,1496]" captionTargetId="figure-59@7.[153,1098,204,968]" captionTargetPageId="7" captionText="Figs 6, 7. K. proboscideus sp. n.: (6) general view (A) and enlarged anterior part of the body (B); (7) prostomium." pageId="9" pageNumber="293">Fig. 7</figureCitation>
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).
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<emphasis box="[512,618,206,230]" italics="true" pageId="9" pageNumber="293">Segments</emphasis>
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: 1–3 simple, 4–13 clearly double ringed.
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<emphasis box="[144,291,238,262]" italics="true" pageId="9" pageNumber="293">Dorsal pores</emphasis>
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: Lacking.
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<emphasis box="[414,475,238,262]" italics="true" pageId="9" pageNumber="293">Setae</emphasis>
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: Small, both
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<emphasis box="[630,658,238,262]" italics="true" pageId="9" pageNumber="293">ab</emphasis>
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and
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<emphasis box="[715,741,238,262]" italics="true" pageId="9" pageNumber="293">bc</emphasis>
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became observable on segment 17. All setae lateral, setal ratio
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<emphasis box="[489,517,270,294]" italics="true" pageId="9" pageNumber="293">aa</emphasis>
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:
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<emphasis box="[525,553,270,294]" italics="true" pageId="9" pageNumber="293">ab</emphasis>
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:
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<emphasis box="[561,587,270,294]" italics="true" pageId="9" pageNumber="293">bc</emphasis>
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:
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<emphasis box="[595,621,270,294]" italics="true" pageId="9" pageNumber="293">cd</emphasis>
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:
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<emphasis box="[629,657,270,294]" italics="true" pageId="9" pageNumber="293">dd</emphasis>
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= 40:1.25:7.5:1:37.5.
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<emphasis box="[907,1096,270,294]" italics="true" pageId="9" pageNumber="293">Nephridial pores</emphasis>
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: Begin on segment 2, somewhat dorsal to setal line
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<emphasis box="[713,741,303,327]" italics="true" pageId="9" pageNumber="293">ab</emphasis>
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.
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</paragraph>
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<caption pageId="8" pageNumber="292" startId="8.[183,222,1630,1651]" targetBox="[187,1132,442,1612]" targetPageId="8" targetType="figure">
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<paragraph blockId="8.[183,1142,1630,1675]" pageId="8" pageNumber="292">
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Figs 8, 9.
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<taxonomicName authority=". & D]D & Qgudnrwr & Csuzdi & Blanchart, 2011" authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[272,419,1632,1651]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="292" phylum="Annelida" rank="species" species="proboscideus" status="sp. n.">
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<emphasis box="[272,419,1632,1651]" italics="true" pageId="8" pageNumber="292">K. proboscideus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[425,476,1630,1651]" pageId="8" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
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: (8) spermathecae; (9) penial seta: (A) whole seta, (B) tip of the seta, (C) ornamentation around the tip, (D) ornamentation under the tip.
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</paragraph>
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</caption>
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<paragraph blockId="9.[144,1104,203,1051]" pageId="9" pageNumber="293">
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<taxonomicName authority=": Circular" authorityName="Circular" box="[144,355,335,362]" class="Ascidiacea" family="Didemnidae" genus="Clitellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aplousobranchia" pageId="9" pageNumber="293" phylum="Chordata" rank="genus">
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<emphasis box="[144,248,338,362]" italics="true" pageId="9" pageNumber="293">Clitellum</emphasis>
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: Circular
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</taxonomicName>
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on segments 22–47, 48.
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<emphasis box="[638,763,338,362]" italics="true" pageId="9" pageNumber="293">Male pores</emphasis>
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: Ventral on 16.
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<emphasis box="[943,1096,338,362]" italics="true" pageId="9" pageNumber="293">Female pores</emphasis>
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: Not observable.
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<emphasis box="[339,566,370,394]" italics="true" pageId="9" pageNumber="293">Spermathecal pores</emphasis>
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: Lateral, 6–9 on each side in intersegmental furrows 13/14, 14/15, 15/16.
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<emphasis box="[476,691,402,426]" italics="true" pageId="9" pageNumber="293">Genital setal pores</emphasis>
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: Segmental, irregularly occur on lateral sides of segments 13, 14 and 15.
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</paragraph>
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<paragraph blockId="9.[144,1104,203,1051]" pageId="9" pageNumber="293">
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Internal characters:
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<emphasis box="[370,568,469,493]" italics="true" pageId="9" pageNumber="293">Muscular gizzard</emphasis>
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: In 5, well developed, cylindrical.
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<taxonomicName authorityName="Perry" authorityYear="1810" box="[973,1035,469,493]" class="Gastropoda" family="Cymatiidae" genus="Septa" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="293" phylum="Mollusca" rank="genus">
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<emphasis box="[973,1035,469,493]" italics="true" pageId="9" pageNumber="293">Septa</emphasis>
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</taxonomicName>
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: 5/6– 12/13 strongly thickened.
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<emphasis box="[446,660,501,525]" italics="true" pageId="9" pageNumber="293">Calciferous glands</emphasis>
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: Lacking, even lamellar thickening of oesophagus missing.
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<emphasis box="[383,606,534,558]" italics="true" pageId="9" pageNumber="293">Dorsal blood vessel</emphasis>
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: Simple.
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<emphasis box="[714,790,534,558]" italics="true" pageId="9" pageNumber="293">Hearts</emphasis>
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: In 6–11, large moniliform, especially those in 10, 11.
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<emphasis box="[447,514,566,590]" italics="true" pageId="9" pageNumber="293">Testes</emphasis>
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and
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<emphasis box="[571,729,566,590]" italics="true" pageId="9" pageNumber="293">sperm funnels</emphasis>
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: In 10, 11 enclosed in peritoneal sack.
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<emphasis box="[207,390,598,622]" italics="true" pageId="9" pageNumber="293">Seminal vesicles</emphasis>
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: Lacking.
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||
<emphasis box="[509,598,598,622]" italics="true" pageId="9" pageNumber="293">Ovaries</emphasis>
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: In 13.
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<emphasis box="[683,914,598,622]" italics="true" pageId="9" pageNumber="293">Copulatory chamber</emphasis>
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: Large, oval, occupies ventral part of segments 15–20. Each copulatory chamber bears irregular prostate-like gland (pseudoprostate) bulging up to segment 25.
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<emphasis box="[824,983,663,687]" italics="true" pageId="9" pageNumber="293">Spermathecae</emphasis>
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: In 13/14, 14/15, 15/16, 6–9 per side, irregular sac-shaped, their size increases toward ventral pairs (
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<figureCitation box="[215,285,725,751]" captionStart="Figs 8, 9" captionStartId="8.[183,222,1630,1651]" captionTargetBox="[187,1132,442,1612]" captionTargetId="figure-110@8.[187,1132,950,1612]" captionTargetPageId="8" captionText="Figs 8, 9. K. proboscideus sp. n.: (8) spermathecae; (9) penial seta: (A) whole seta, (B) tip of the seta, (C) ornamentation around the tip, (D) ornamentation under the tip." pageId="9" pageNumber="293">Fig. 8</figureCitation>
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).
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<emphasis box="[309,529,727,751]" italics="true" pageId="9" pageNumber="293">Genital setal organ</emphasis>
|
||
: Three pairs in segments 13, 14, 15, consisting of slightly spiral gland similar in shape to pseudoprostates and genital setal sac containing several genital setae.
|
||
<emphasis box="[376,522,792,816]" italics="true" pageId="9" pageNumber="293">Genital setae</emphasis>
|
||
: Lanceolate, tip slightly depressed, 3.2 mm in length, 0.06mm in diameter, with unequal serrated ornamentation. Serrations on dorsal side small and scattered, on ventral side dense and long (
|
||
<figureCitation box="[728,796,854,880]" captionStart="Figs 8, 9" captionStartId="8.[183,222,1630,1651]" captionTargetBox="[187,1132,442,1612]" captionTargetId="figure-110@8.[187,1132,950,1612]" captionTargetPageId="8" captionText="Figs 8, 9. K. proboscideus sp. n.: (8) spermathecae; (9) penial seta: (A) whole seta, (B) tip of the seta, (C) ornamentation around the tip, (D) ornamentation under the tip." pageId="9" pageNumber="293">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="9.[144,1104,203,1051]" pageId="9" pageNumber="293">
|
||
<materialsCitation country="Madagascar" latitude="-17.308971" location="Mangalaza" municipality="Ambatondrazaka district" pageId="9" pageNumber="293" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
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||
<typeStatus box="[144,232,895,916]" pageId="9" pageNumber="293">Holotype</typeStatus>
|
||
:
|
||
<collectingCountry box="[248,413,895,916]" name="Madagascar" pageId="9" pageNumber="293">MADAGASCAR</collectingCountry>
|
||
:
|
||
<collectingMunicipality box="[428,662,895,916]" pageId="9" pageNumber="293">Ambatondrazaka district</collectingMunicipality>
|
||
,
|
||
<collectingRegion box="[679,777,895,916]" country="Madagascar" name="Toamasina" pageId="9" pageNumber="293">Toamasina</collectingRegion>
|
||
town,
|
||
<location LSID="urn:lsid:plazi:treatment:03DC87D4FF8DFF98FE25FACBFC3A46D6:8EAA6019FF82FF98FC8DFCFAFC3440DA" box="[856,955,895,916]" country="Madagascar" latitude="-17.308971" municipality="Ambatondrazaka district" name="Mangalaza" pageId="9" pageNumber="293" stateProvince="Toamasina">Mangalaza</location>
|
||
(
|
||
<geoCoordinate box="[977,1101,895,916]" degrees="17" direction="south" minutes="18" orientation="latitude" pageId="9" pageNumber="293" precision="1" seconds="32.3" value="-17.308971">17°18'32.3"S</geoCoordinate>
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||
:
|
||
<emphasis bold="true" box="[144,817,918,942]" pageId="9" pageNumber="293">48°46‵4Ɵ.6̎(), 8ƟƟ P, l7.LLL.2ƟlƟ, 0. 5D]D¿QGUDNRWR (+1+0 $)/ƽƽll).</emphasis>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="9.[144,1104,203,1051]" pageId="9" pageNumber="293">
|
||
<materialsCitation collectionCode="HNHM, RDPDVLQD, WRZQ" country="Madagascar" county="Same" latitude="-17.44925" location="Ambatondrazaka district" longLatPrecision="1" longitude="48.564526" municipality="Ambatondrazaka district" pageId="9" pageNumber="293" specimenCount="3" stateProvince="Toamasina" typeStatus="paratype">
|
||
<typeStatus box="[144,236,957,978]" pageId="9" pageNumber="293">Paratypes</typeStatus>
|
||
:
|
||
<collectingCounty box="[244,294,957,978]" pageId="9" pageNumber="293">Same</collectingCounty>
|
||
data as holotype (
|
||
<collectionCode box="[458,529,957,978]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="293">HNHM</collectionCode>
|
||
AF/5512 1 ex, ZMUA-027: 2+
|
||
<specimenCount box="[813,860,957,978]" pageId="9" pageNumber="293" type="generic">2 ex.</specimenCount>
|
||
);
|
||
<collectingMunicipality box="[875,1100,957,978]" pageId="9" pageNumber="293">Ambatondrazaka district</collectingMunicipality>
|
||
,
|
||
<emphasis bold="true" box="[144,1104,979,1003]" pageId="9" pageNumber="293">
|
||
7
|
||
<collectionCode box="[152,301,980,1003]" pageId="9" pageNumber="293">RDPDVLQD WRZQ</collectionCode>
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||
, 7VDUDKRQHQDQD (l7°l8‵4Ɵ.l̎6:48°36‵ƽ8.4̎(), 93ƽ P, 2ƽ.LLL.2ƟlƟ, 0. 5D]D¿QGUDNRWR
|
||
</emphasis>
|
||
(ZMUA-029
|
||
<specimenCount box="[269,319,1005,1026]" pageId="9" pageNumber="293" type="generic">1 ex.</specimenCount>
|
||
);
|
||
<location LSID="urn:lsid:plazi:treatment:03DC87D4FF8DFF98FE25FACBFC3A46D6:8EAA6019FF82FF98FE87FC68FDB8474C" box="[338,567,1005,1026]" county="Same" latitude="-17.44925" longLatPrecision="1" longitude="48.564526" municipality="Ambatondrazaka district" name="Ambatondrazaka district" pageId="9" pageNumber="293" stateProvince="Toamasina">Ambatondrazaka district</location>
|
||
,
|
||
<collectingRegion box="[580,678,1005,1026]" country="Madagascar" name="Toamasina" pageId="9" pageNumber="293">Toamasina</collectingRegion>
|
||
town,
|
||
<location LSID="urn:lsid:plazi:treatment:03DC87D4FF8DFF98FE25FACBFC3A46D6:8EAA6019FF82FF98FD3EFC68FCB5474C" box="[747,826,1005,1026]" county="Same" latitude="-17.44925" longLatPrecision="1" longitude="48.564526" municipality="Ambatondrazaka district" name="Vohitsoa" pageId="9" pageNumber="293" stateProvince="Toamasina">Vohitsoa</location>
|
||
(
|
||
<geoCoordinate box="[842,964,1005,1026]" degrees="17" direction="south" minutes="26" orientation="latitude" pageId="9" pageNumber="293" precision="1" seconds="57.3" value="-17.44925">17°26'57.3"S</geoCoordinate>
|
||
:
|
||
<geoCoordinate box="[967,1093,1005,1026]" degrees="48" direction="east" minutes="33" orientation="longitude" pageId="9" pageNumber="293" precision="1" seconds="52.3" value="48.564526">48°33'52.3"E</geoCoordinate>
|
||
),
|
||
<emphasis bold="true" box="[144,672,1027,1051]" pageId="9" pageNumber="293">76ƽ P, 2ƽ.LLL.2ƟlƟ, 0. 5D]D¿QGUDNRWR (=08$-Ɵ28 l H[.)</emphasis>
|
||
</materialsCitation>
|
||
<emphasis bold="true" box="[672,676,1027,1050]" pageId="9" pageNumber="293">.</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="9.[144,1104,1070,1194]" pageId="9" pageNumber="293">
|
||
Habitat: At all places the vegetation is formed by bush composed of invasive species (
|
||
<taxonomicName box="[153,371,1105,1129]" class="Magnoliopsida" family="Phytolaccaceae" genus="Phytolacca" kingdom="Plantae" order="Caryophyllales" pageId="9" pageNumber="293" phylum="Tracheophyta" rank="species" species="acinosa">
|
||
<emphasis box="[153,371,1105,1129]" italics="true" pageId="9" pageNumber="293">Phytolacca acinosa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName box="[384,558,1105,1129]" class="Magnoliopsida" family="Lamiaceae" genus="Salvia" kingdom="Plantae" order="Lamiales" pageId="9" pageNumber="293" phylum="Tracheophyta" rank="species" species="coccinea">
|
||
<emphasis box="[384,558,1105,1129]" italics="true" pageId="9" pageNumber="293">Salvia coccinea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName box="[571,810,1105,1129]" class="Magnoliopsida" family="Asteraceae" genus="Ageratum" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="293" phylum="Tracheophyta" rank="species" species="conyzoides">
|
||
<emphasis box="[571,810,1105,1129]" italics="true" pageId="9" pageNumber="293">Ageratum conyzoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName box="[823,1008,1105,1129]" class="Magnoliopsida" family="Verbenaceae" genus="Lantana" kingdom="Plantae" order="Lamiales" pageId="9" pageNumber="293" phylum="Tracheophyta" rank="species" species="camara">
|
||
<emphasis box="[823,1008,1105,1129]" italics="true" pageId="9" pageNumber="293">Lantana camara</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Ricinus" kingdom="Plantae" order="Malpighiales" pageId="9" pageNumber="293" phylum="Tracheophyta" rank="species" species="communis">
|
||
<emphasis italics="true" pageId="9" pageNumber="293">Ricinus communis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName box="[271,460,1137,1161]" class="Magnoliopsida" family="Malvaceae" genus="Sida" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="293" phylum="Tracheophyta" rank="species" species="rhombifolia">
|
||
<emphasis box="[271,460,1137,1161]" italics="true" pageId="9" pageNumber="293">Sida rhombifolia</emphasis>
|
||
</taxonomicName>
|
||
), with many dead leaves on the soil and some other plant
|
||
<emphasis bold="true" box="[144,527,1165,1194]" pageId="9" pageNumber="293">PDWHULDO WKDW VWDUWV WR EH KXPL¿HG.</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="9.[144,1104,1213,1433]" pageId="9" pageNumber="293">
|
||
Remarks: The new species is similar to
|
||
<taxonomicName authority="Michaelsen, 1892" authorityName="Michaelsen" authorityYear="1892" box="[589,900,1213,1239]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="293" phylum="Annelida" rank="species" species="kelleri">
|
||
<emphasis box="[589,694,1215,1239]" italics="true" pageId="9" pageNumber="293">K. kelleri</emphasis>
|
||
Michaelsen,
|
||
<quantity box="[844,900,1213,1239]" metricMagnitude="1" metricUnit="m" metricValue="4.80568" pageId="9" pageNumber="293" unit="in" value="1892.0">1892</quantity>
|
||
</taxonomicName>
|
||
in its size and colour, but clearly differs from it by the higher number of spermathecae (1–3 per side in
|
||
<taxonomicName authorityName="Michaelsen" authorityYear="1892" box="[174,246,1280,1304]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="293" phylum="Annelida" rank="species" species="kelleri">
|
||
<emphasis box="[174,246,1280,1304]" italics="true" pageId="9" pageNumber="293">kelleri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity box="[261,334,1277,1303]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.286" metricValueMin="1.524" pageId="9" pageNumber="293" unit="in" value="7.5" valueMax="9.0" valueMin="6.0">6–9 in</quantity>
|
||
<taxonomicName authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[344,492,1280,1304]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="293" phylum="Annelida" rank="species" species="proboscideus">
|
||
<emphasis box="[344,492,1280,1304]" italics="true" pageId="9" pageNumber="293">proboscideus</emphasis>
|
||
</taxonomicName>
|
||
) and the presence of two prominent probosces on the prostomium. There is only one other species with similar head structure,
|
||
<taxonomicName authority="Michaelsen, 1897" authorityName="Michaelsen" authorityYear="1897" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="293" phylum="Annelida" rank="species" species="schistocephalus">
|
||
<emphasis italics="true" pageId="9" pageNumber="293">K. schistocephalus</emphasis>
|
||
<bibRefCitation author="MICHAELSEN, W." box="[252,456,1342,1368]" pageId="9" pageNumber="293" pagination="217 - 252" refId="ref4304" refString="MICHAELSEN, W. 1897. Die Terricolen des Madagassischen Inselgebiets. Abhandlungen herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft 21: 217 - 252." type="journal article" year="1897">Michaelsen, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
, but the new species is easy to distinguish because of its longer clitellum (
|
||
<quantity box="[335,437,1374,1400]" metricMagnitude="-1" metricUnit="m" metricValue="7.747" metricValueMax="10.16" metricValueMin="5.334" pageId="9" pageNumber="293" unit="in" value="30.5" valueMax="40.0" valueMin="21.0">21–40 in</quantity>
|
||
<taxonomicName authorityName="Michaelsen" authorityYear="1897" box="[443,620,1376,1400]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="293" phylum="Annelida" rank="species" species="schistocephalus">
|
||
<emphasis box="[443,620,1376,1400]" italics="true" pageId="9" pageNumber="293">schistocephalus</emphasis>
|
||
</taxonomicName>
|
||
, 22–27,
|
||
<quantity box="[718,775,1374,1400]" metricMagnitude="0" metricUnit="m" metricValue="1.2192" pageId="9" pageNumber="293" unit="in" value="48.0">48 in</quantity>
|
||
<taxonomicName authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[781,929,1376,1400]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="293" phylum="Annelida" rank="species" species="proboscideus">
|
||
<emphasis box="[781,929,1376,1400]" italics="true" pageId="9" pageNumber="293">proboscideus</emphasis>
|
||
</taxonomicName>
|
||
) and the higher number of spermathecae (
|
||
<quantity box="[435,506,1406,1432]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="9" pageNumber="293" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">1–3 in</quantity>
|
||
<taxonomicName authorityName="Michaelsen" authorityYear="1897" box="[513,690,1409,1433]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="293" phylum="Annelida" rank="species" species="schistocephalus">
|
||
<emphasis box="[513,690,1409,1433]" italics="true" pageId="9" pageNumber="293">schistocephalus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity box="[704,778,1406,1432]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.286" metricValueMin="1.524" pageId="9" pageNumber="293" unit="in" value="7.5" valueMax="9.0" valueMin="6.0">6–9 in</quantity>
|
||
<taxonomicName authorityName=". & D]D & Qgudnrwr & Csuzdi & Blanchart" authorityYear="2011" box="[785,933,1409,1433]" class="Clitellata" family="Glossoscolecidae" genus="Kynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="293" phylum="Annelida" rank="species" species="proboscideus">
|
||
<emphasis box="[785,933,1409,1433]" italics="true" pageId="9" pageNumber="293">proboscideus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |