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<document id="A3FCE9B25A526B2E8D77D09D7F90E176" ID-DOI="10.11646/zootaxa.5497.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="13619154" ID-ZooBank="5F28AF84-B715-4F31-82EB-E88DB75786FC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725027189025" checkinUser="plazi" docAuthor="Prado, André Wanderley Do, Pantoja, Paulo &amp; Baptista, Renner Luiz Cerqueira" docDate="2024" docId="03DC87CBFA77996DFF5F041BFBD773F0" docLanguage="en" docName="zootaxa.5497.4.6.pdf" docOrigin="Zootaxa 5497 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5497.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tibelloides Mello-Leitao 1939" docType="treatment" docVersion="2" lastPageNumber="561" masterDocId="FFE5FFB3FA75996FFFC8053CFF9E7729" masterDocTitle="Two new and unusual species of Tibelloides Mello-Leitão, 1939 (Araneae: Philodromidae)" masterLastPageNumber="576" masterPageNumber="559" pageNumber="561" updateTime="1725029231814" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D58F928DB7D70660534E2D806E9DAB4D">Two new and unusual species of Tibelloides Mello-Leitão, 1939 (Araneae: Philodromidae)</mods:title>
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<mods:namePart id="A26CC27A9BC93836B79FFF8335C5DB42">Prado, André Wanderley Do</mods:namePart>
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<mods:affiliation id="7A2693D34CD21D652F41C80CC5BC1A7C">Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Department of Terrestrial Zoology, Western Australian Museum, Locked Bag 49, Welshpool DC, Western Australia</mods:affiliation>
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<mods:namePart id="61DA1F455583AF39EC04F2A3F4561457">Pantoja, Paulo</mods:namePart>
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<mods:affiliation id="C6397CB93D248688A260D9DCF41DDBBC">Programa de Pós-graduação em Zoologia, Universidade Federal do Pará / Museu Paraense Emílio Goeldi</mods:affiliation>
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<mods:namePart id="5912F1F06A646F4D244E400115439837">Baptista, Renner Luiz Cerqueira</mods:namePart>
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<mods:title id="1E04C04E10B112CC09D8C6B1D2C87E6C">Zootaxa</mods:title>
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<mods:date id="2532E0926FE1B6BC0AB1413799F7C567">2024</mods:date>
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<mods:number id="61D094A357618FD3F133D7A9F2D830A4">5497</mods:number>
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<treatment id="03DC87CBFA77996DFF5F041BFBD773F0" ID-DOI="http://doi.org/10.5281/zenodo.13619194" ID-Zenodo-Dep="13619194" LSID="urn:lsid:plazi:treatment:03DC87CBFA77996DFF5F041BFBD773F0" httpUri="http://treatment.plazi.org/id/03DC87CBFA77996DFF5F041BFBD773F0" lastPageNumber="561" pageId="2" pageNumber="561">
<subSubSection id="C36F6556FA77996DFF5F041BFD927668" box="[151,524,295,322]" pageId="2" pageNumber="561" type="nomenclature">
<paragraph id="8BCA36DDFA77996DFF5F041BFD927668" blockId="2.[151,524,295,322]" box="[151,524,295,322]" pageId="2" pageNumber="561">
<heading id="D08281B1FA77996DFF5F041BFD927668" bold="true" box="[151,524,295,322]" fontSize="11" level="1" pageId="2" pageNumber="561" reason="1">
<taxonomicName id="4C754D5EFA77996DFF5F041BFD927668" ID-CoL="C8KBT" authority="Mello-Leitao, 1939" authorityName="Mello-Leitao" authorityYear="1939" box="[151,524,295,322]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B901EACFFA77996DFF5F041BFD927668" bold="true" box="[151,524,295,322]" pageId="2" pageNumber="561">
<emphasis id="B901EACFFA77996DFF5F041BFE84766B" bold="true" box="[151,282,295,322]" italics="true" pageId="2" pageNumber="561">Tibelloides</emphasis>
<bibRefCitation id="EFE44B2CFA77996DFEE9041BFD927668" author="Mello-Leitao, C. F. de" box="[289,524,295,321]" pageId="2" pageNumber="561" pagination="43 - 93" refId="ref10660" refString="Mello-Leitao, C. F. de (1939) Araignees americaines du Musee d'histoire naturelle de Bale. Revue Suisse de Zoologie, 46, 43 - 93. https: // doi. org / 10.5962 / bhl. part. 117928" type="journal article" year="1939">Mello-Leitão, 1939</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F6556FA77996DFF5F0450FD5176E8" pageId="2" pageNumber="561" type="reference_group">
<paragraph id="8BCA36DDFA77996DFF5F0450FDE8768B" blockId="2.[151,1437,363,449]" pageId="2" pageNumber="561">
<treatmentCitationGroup id="AB6511F3FA77996DFF5F0450FDE8768B" pageId="2" pageNumber="561">
<treatmentCitation id="0AD410CCFA77996DFF5F0450FD8076AA" author="Mello-Leitao, C. F. de" box="[151,542,363,388]" page="77" pageId="2" pageNumber="561" year="1939">
<taxonomicName id="4C754D5EFA77996DFF5F0450FD8076AA" ID-CoL="C8KBT" authority="Mello-Leitao, 1939: 76 - 77" authorityName="Mello-Leitao" authorityPageNumber="76 - 77" authorityYear="1939" box="[151,542,363,388]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B901EACFFA77996DFF5F0450FE9876AD" box="[151,262,364,388]" italics="true" pageId="2" pageNumber="561">Tibelloides</emphasis>
<bibRefCitation id="EFE44B2CFA77996DFEC30450FD8076AA" author="Mello-Leitao, C. F. de" box="[267,542,363,388]" pageId="2" pageNumber="561" pagination="43 - 93" refId="ref10660" refString="Mello-Leitao, C. F. de (1939) Araignees americaines du Musee d'histoire naturelle de Bale. Revue Suisse de Zoologie, 46, 43 - 93. https: // doi. org / 10.5962 / bhl. part. 117928" type="journal article" year="1939">Mello-Leitão, 1939: 76–77</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<treatmentCitation id="0AD410CCFA77996DFDEE0450FCBA76AA" author="Prado, A. W. do &amp; Baptista, R. L. C. &amp; Schinelli, H. B. P. &amp; Takiya, D. M." box="[550,804,363,388]" page="15" pageId="2" pageNumber="561" year="2022">
<bibRefCitation id="EFE44B2CFA77996DFDEE0450FCBA76AA" author="Prado, A. W. do &amp; Baptista, R. L. C. &amp; Schinelli, H. B. P. &amp; Takiya, D. M." box="[550,804,363,388]" pageId="2" pageNumber="561" pagination="1 - 35" refId="ref10957" refString="Prado, A. W. do, Baptista, R. L. C., Schinelli, H. B. P. &amp; Takiya, D. M. (2022) Solving a running crab spider puzzle: delimiting Cleocnemis Simon, 1886 with implications on the phylogeny and terminology of genital structures of Philodromidae. BMC Zoology, 7 (51), 1 - 35. https: // doi. org / 10.1186 / s 40850 - 022 - 00136 - 7" type="journal article" year="2022">
Prado
<emphasis id="B901EACFFA77996DFDAE0450FD0C76AD" box="[614,658,364,388]" italics="true" pageId="2" pageNumber="561">et al</emphasis>
., 2022: 15–25
</bibRefCitation>
</treatmentCitation>
, figs. 5a–c, 6a–f, 7a–g, 8a–g, 9a–g (reasoning for revalidation);
<treatmentCitation id="0AD410CCFA77996DFF0F04B6FE0B768B" author="Prado, A. W. do &amp; Baptista, R. L. C. &amp; Schinelli, H. B. P. &amp; Takiya, D. M." box="[199,405,394,418]" page="2" pageId="2" pageNumber="561" year="2023">
<bibRefCitation id="EFE44B2CFA77996DFF0F04B6FE0B768B" author="Prado, A. W. do &amp; Baptista, R. L. C. &amp; Schinelli, H. B. P. &amp; Takiya, D. M." box="[199,405,394,418]" pageId="2" pageNumber="561" pagination="1 - 7" refId="ref11046" refString="Prado, A. W. do, Baptista, R. L. C., Schinelli, H. B. P. &amp; Takiya, D. M. (2023) Correction: Solving a running crab spider puzzle: delimiting Cleocnemis Simon, 1886 with implications on the phylogeny and terminology of genital structures of Philodromidae. BMC Zoology, 8 (2), 1 - 7. https: // doi. org / 10.1186 / s 40850 - 022 - 00136 - 7" type="journal article" year="2023">
Prado
<emphasis id="B901EACFFA77996DFEC104B7FEAB768B" box="[265,309,394,418]" italics="true" pageId="2" pageNumber="561">et al</emphasis>
., 2023: 2
</bibRefCitation>
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(formal revalidation).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA36DDFA77996DFF5F0495FD5176E8" blockId="2.[151,1437,363,449]" box="[151,719,425,449]" pageId="2" pageNumber="561">
<treatmentCitationGroup id="AB6511F3FA77996DFF5F0495FD5176E8" box="[151,719,425,449]" pageId="2" pageNumber="561">
<taxonomicName id="4C754D5EFA77996DFF5F0495FEDA76E8" ID-CoL="7WRZ" authority="(Simon)" baseAuthorityName="Simon" box="[151,324,425,449]" class="Arachnida" family="Philodromidae" genus="Tibellus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B901EACFFA77996DFF5F0495FF7976E8" box="[151,231,425,449]" italics="true" pageId="2" pageNumber="561">Tibellus</emphasis>
(Simon)
</taxonomicName>
:
<treatmentCitation id="0AD410CCFA77996DFE860495FDD976E8" author="Mello-Leitao, C. F. de" box="[334,583,425,449]" page="224" pageId="2" pageNumber="561" year="1945">
<bibRefCitation id="EFE44B2CFA77996DFE860495FDD976E8" author="Mello-Leitao, C. F. de" box="[334,583,425,449]" pageId="2" pageNumber="561" pagination="213 - 302" refId="ref10747" refString="Mello-Leitao, C. F. de (1945) Aranas de Misiones, Corrientes y Entre Rios. Revista del Museo de La Plata, New Series, Zoology, 4, 213 - 302." type="journal article" year="1945">Mello-Leitão, 1945: 224</bibRefCitation>
</treatmentCitation>
(synonymy).
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</paragraph>
</subSubSection>
<subSubSection id="C36F6556FA77996DFF5F04D3FBD773F0" pageId="2" pageNumber="561" type="type_taxon">
<paragraph id="8BCA36DDFA77996DFF5F04D3FC577504" blockId="2.[151,1437,495,1242]" pageId="2" pageNumber="561">
<emphasis id="B901EACFFA77996DFF5F04D3FEAB7520" bold="true" box="[151,309,495,521]" pageId="2" pageNumber="561">
<typeStatus id="54CE887FFA77996DFF5F04D3FF4C7520" box="[151,210,495,521]" pageId="2" pageNumber="561">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C754D5EFA77996DFEF704D3FC897520" ID-CoL="C8Q6L" authority="Mello-Leitao, 1939" authorityName="Mello-Leitao" authorityYear="1939" box="[319,791,495,521]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="spatulifer">
<emphasis id="B901EACFFA77996DFEF704D3FDB17520" box="[319,559,495,521]" italics="true" pageId="2" pageNumber="561">Tibelloides spatulifer</emphasis>
<bibRefCitation id="EFE44B2CFA77996DFDF204D3FC897520" author="Mello-Leitao, C. F. de" box="[570,791,495,521]" pageId="2" pageNumber="561" pagination="43 - 93" refId="ref10660" refString="Mello-Leitao, C. F. de (1939) Araignees americaines du Musee d'histoire naturelle de Bale. Revue Suisse de Zoologie, 46, 43 - 93. https: // doi. org / 10.5962 / bhl. part. 117928" type="journal article" year="1939">Mello-Leitão, 1939</bibRefCitation>
</taxonomicName>
[original spelling
<taxonomicName id="4C754D5EFA77996DFC2604D3FF517504" authority="Mello-Leitao, 1939" authorityName="Mello-Leitao" authorityYear="1939" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="spatuliferus">
<emphasis id="B901EACFFA77996DFC2604D3FB697520" box="[1006,1271,495,521]" italics="true" pageId="2" pageNumber="561">Tibelloides spatuliferus</emphasis>
<bibRefCitation id="EFE44B2CFA77996DFAC904D3FF517504" author="Mello-Leitao, C. F. de" pageId="2" pageNumber="561" pagination="43 - 93" refId="ref10660" refString="Mello-Leitao, C. F. de (1939) Araignees americaines du Musee d'histoire naturelle de Bale. Revue Suisse de Zoologie, 46, 43 - 93. https: // doi. org / 10.5962 / bhl. part. 117928" type="journal article" year="1939">Mello-Leitão, 1939</bibRefCitation>
</taxonomicName>
= junior synonym of
<taxonomicName id="4C754D5EFA77996DFE0B072FFC217504" ID-CoL="C8KBY" authority="(Taczanowski, 1872)" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[451,959,531,557]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B901EACFFA77996DFE0B072FFD567504" box="[451,712,531,557]" italics="true" pageId="2" pageNumber="561">Tibelloides punctulatus</emphasis>
(
<bibRefCitation id="EFE44B2CFA77996DFD1E072FFC287504" author="Taczanowski, L." box="[726,950,531,557]" pageId="2" pageNumber="561" pagination="64 - 112" refId="ref11366" refString="Taczanowski, L. (1872) Les araneides de la Guyane francaise. Horae Societatis Entomologicae Rossicae, 9, 64 - 112." type="journal article" year="1872">Taczanowski, 1872</bibRefCitation>
)
</taxonomicName>
].
</paragraph>
<paragraph id="8BCA36DDFA77996DFF0F070BFE4875B3" blockId="2.[151,1437,495,1242]" pageId="2" pageNumber="561">
<emphasis id="B901EACFFA77996DFF0F070BFEF97578" bold="true" box="[199,359,567,593]" pageId="2" pageNumber="561">Composition.</emphasis>
<taxonomicName id="4C754D5EFA77996DFEBC070BFCB87578" ID-CoL="C8KBV" authority="(Gertsch, 1933)" baseAuthorityName="Gertsch" baseAuthorityYear="1933" box="[372,806,567,593]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="bryantae">
<emphasis id="B901EACFFA77996DFEBC070BFDFE7578" box="[372,608,567,593]" italics="true" pageId="2" pageNumber="561">Tibelloides bryantae</emphasis>
(
<bibRefCitation id="EFE44B2CFA77996DFDBD070BFC837578" author="Gertsch, W. J." box="[629,797,567,593]" pageId="2" pageNumber="561" pagination="1 - 22" refId="ref10355" refString="Gertsch, W. J. (1933) Notes on American spiders of the family Thomisidae. American Museum Novitates, 593, 1 - 22." type="journal article" year="1933">Gertsch, 1933</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C754D5EFA77996DFCF0070BFA917578" ID-CoL="C8KBW" authority="(Gertsch, 1933)" baseAuthorityName="Gertsch" baseAuthorityYear="1933" box="[824,1295,567,593]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="paraguensis">
<emphasis id="B901EACFFA77996DFCF0070BFBD77578" box="[824,1097,567,593]" italics="true" pageId="2" pageNumber="561">Tibelloides paraguensis</emphasis>
(
<bibRefCitation id="EFE44B2CFA77996DFB96070BFA997578" author="Gertsch, W. J." box="[1118,1287,567,593]" pageId="2" pageNumber="561" pagination="1 - 22" refId="ref10355" refString="Gertsch, W. J. (1933) Notes on American spiders of the family Thomisidae. American Museum Novitates, 593, 1 - 22." type="journal article" year="1933">Gertsch, 1933</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C754D5EFA77996DFAE9070BFD83755C" ID-CoL="C8KBY" authority="(Taczanowski, 1872)" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B901EACFFA77996DFAE9070BFE84755C" italics="true" pageId="2" pageNumber="561">Tibelloides punctulatus</emphasis>
(
<bibRefCitation id="EFE44B2CFA77996DFEF80767FD8B755C" author="Taczanowski, L." box="[304,533,603,629]" pageId="2" pageNumber="561" pagination="64 - 112" refId="ref11366" refString="Taczanowski, L. (1872) Les araneides de la Guyane francaise. Horae Societatis Entomologicae Rossicae, 9, 64 - 112." type="journal article" year="1872">Taczanowski, 1872</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C754D5EFA77996DFDF80767FB94755C" ID-CoL="C8KC2" authority="(Keyserling, 1891)" baseAuthorityName="Keyserling" baseAuthorityYear="1891" box="[560,1034,603,629]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="taquarae">
<emphasis id="B901EACFFA77996DFDF80767FC80755C" box="[560,798,603,629]" italics="true" pageId="2" pageNumber="561">Tibelloides taquarae</emphasis>
(
<bibRefCitation id="EFE44B2CFA77996DFCFC0767FB9F755C" author="Keyserling, E." box="[820,1025,603,629]" pageId="2" pageNumber="561" refId="ref10537" refString="Keyserling, E. (1891) Die Spinnen Amerikas. Brasilianische Spinnen. Vol. 3. Bauer &amp; Raspe, Nurnberg, 278 pp. https: // doi. org / 10.5962 / bhl. title. 64832" type="book" year="1891">Keyserling, 1891</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C754D5EFA77996DFBD50767FB68755C" authorityName="Prado &amp; Pantoja &amp; Baptista" authorityYear="2024" box="[1053,1270,603,629]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="castelo" status="sp. nov.">
<emphasis id="B901EACFFA77996DFBD50767FB68755C" box="[1053,1270,603,629]" italics="true" pageId="2" pageNumber="561">Tibelloides castelo</emphasis>
</taxonomicName>
<emphasis id="B901EACFFA77996DFACC0767FAFB755C" bold="true" box="[1284,1381,603,629]" pageId="2" pageNumber="561">
<taxonomicNameLabel id="A23257B4FA77996DFACC0767FAFB755C" box="[1284,1381,603,629]" pageId="2" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C754D5EFA77996DFF5F0743FEF175B0" authorityName="Prado &amp; Pantoja &amp; Baptista" authorityYear="2024" box="[151,367,639,665]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="juatuba" status="sp. nov.">
<emphasis id="B901EACFFA77996DFF5F0743FEF175B0" box="[151,367,639,665]" italics="true" pageId="2" pageNumber="561">Tibelloides juatuba</emphasis>
</taxonomicName>
<emphasis id="B901EACFFA77996DFEBE0743FE5175B0" bold="true" box="[374,463,639,665]" pageId="2" pageNumber="561">
<taxonomicNameLabel id="A23257B4FA77996DFEBE0743FE5175B0" box="[374,463,639,665]" pageId="2" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
<paragraph id="8BCA36DDFA77996DFF0F079FFBD773F0" blockId="2.[151,1437,495,1242]" pageId="2" pageNumber="561">
<emphasis id="B901EACFFA77996DFF0F079FFE097594" bold="true" box="[199,407,675,701]" pageId="2" pageNumber="561">Taxonomic notes</emphasis>
. Unlike other
<taxonomicName id="4C754D5EFA77996DFD86079FFD577594" ID-CoL="C8KBT" authorityName="Mello-Leitao" authorityYear="1939" box="[590,713,675,701]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B901EACFFA77996DFD86079FFD577594" box="[590,713,675,701]" italics="true" pageId="2" pageNumber="561">Tibelloides</emphasis>
</taxonomicName>
, the two new species described herein share many unusual characteristics that set them apart from typical species of the genus.
<taxonomicName id="4C754D5EFA77996DFC7A07FBFB1875C8" authorityName="Prado &amp; Pantoja &amp; Baptista" authorityYear="2024" box="[946,1158,711,737]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="castelo" status="sp. nov.">
<emphasis id="B901EACFFA77996DFC7A07FBFB1875C8" box="[946,1158,711,737]" italics="true" pageId="2" pageNumber="561">Tibelloides castelo</emphasis>
</taxonomicName>
<emphasis id="B901EACFFA77996DFB5807FBFB7275C8" bold="true" box="[1168,1260,711,737]" pageId="2" pageNumber="561">
<taxonomicNameLabel id="A23257B4FA77996DFB5807FBFB7275C8" box="[1168,1260,711,737]" pageId="2" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C754D5EFA77996DFAE007F4FA0375C8" authorityName="Prado &amp; Pantoja &amp; Baptista" authorityYear="2024" box="[1320,1437,711,737]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="juatuba" status="sp. nov.">
<emphasis id="B901EACFFA77996DFAE007F4FA0375C8" box="[1320,1437,711,737]" italics="true" pageId="2" pageNumber="561">T. juatuba</emphasis>
</taxonomicName>
<emphasis id="B901EACFFA77996DFF5F07D7FF6F742C" bold="true" box="[151,241,747,773]" pageId="2" pageNumber="561">
<taxonomicNameLabel id="A23257B4FA77996DFF5F07D7FF6F742C" box="[151,241,747,773]" pageId="2" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
share body and legs very thin and elongated, with abdomen at least four times longer than wide, carapace strongly narrowed at eye region, abdomen dorsally with one longitudinal median dark band, from anterior to posterior margins (
<figureCitation id="134E2A58FA77996DFEA4060FFE727464" box="[364,492,819,845]" captionStart="FIGURES 1" captionStartId="3.[151,264,1925,1949]" captionTargetBox="[159,1428,201,1901]" captionTargetId="figure-19@3.[159,1428,201,1901]" captionTargetPageId="3" captionText="FIGURES 1A–F. Tibelloides castelo sp. nov., male (MPEG.ARA 35202). A dorsal habitus; B ventral habitus; C cephalothorax in dorsal view; D cephalothorax in frontal view; E, F left pedipalp (E ventral, F retrolateral).Abbreviations: C, conductor; RTA, retrolateral tibial apophysis. Scale bars: A, B 1.0 mm; C, D 0.5 mm; E, F 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13619158" httpUri="https://zenodo.org/record/13619158/files/figure.png" pageId="2" pageNumber="561">Figs 1A–C</figureCitation>
;
<figureCitation id="134E2A58FA77996DFE31060FFDA17464" box="[505,575,819,845]" captionStart="FIGURES 2" captionStartId="4.[151,264,1583,1607]" captionTargetBox="[159,1428,181,1559]" captionTargetId="figure-19@4.[159,1428,181,1559]" captionTargetPageId="4" captionText="FIGURES 2A–G. Tibelloides castelo sp. nov., female (A–E MPEG.ARA 35206, F,G UFRJ 235). A dorsal habitus; B ventral habitus; C cephalothorax in dorsal view; D cephalothorax in frontal view; E epigyne; F cleared epigyne in ventral view; G vulva in dorsal view. Abbreviations: CD, copulatory duct; CG, copulatory guide; GH, glandular head of spermatheca; PR, posterior rim. Scale bars: A, B 2.0 mm; C, D 0.5 mm; E–G 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13619160" httpUri="https://zenodo.org/record/13619160/files/figure.png" pageId="2" pageNumber="561">2A–C</figureCitation>
;
<figureCitation id="134E2A58FA77996DFD84060FFD0C7464" box="[588,658,819,845]" captionStart="FIGURES 4" captionStartId="9.[151,264,1905,1929]" captionTargetBox="[159,1428,181,1881]" captionTargetId="figure-19@9.[159,1428,181,1881]" captionTargetPageId="9" captionText="FIGURES 4A–F. Tibelloides juatuba sp. nov., male (IBSP 138220). A dorsal habitus; B ventral habitus; C cephalothorax in dorsal view; D cephalothorax in frontal view; E, F left pedipalp (E ventral, F retrolateral). Abbreviations: C, conductor; RTA, retrolateral tibial apophysis. Scale bars: A, B 2.0 mm; C, D 0.5 mm; E, F 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13619168" httpUri="https://zenodo.org/record/13619168/files/figure.png" pageId="2" pageNumber="561">4A–C</figureCitation>
;
<figureCitation id="134E2A58FA77996DFD56060FFD7A7467" box="[670,740,819,846]" captionStart="FIGURES 5" captionStartId="11.[151,264,1674,1698]" captionTargetBox="[159,1428,273,1651]" captionTargetId="figure-59@11.[159,1428,273,1651]" captionTargetPageId="11" captionText="FIGURES 5A–F. Tibelloides juatuba sp. nov., female (A–E UFMG 12612, F IBSP 81042).A dorsal habitus; B ventral habitus; C cephalothorax in dorsal view; D cephalothorax in frontal view; E epigyne; F vulva in dorsal view.Abbreviations: CD, copulatory duct; CG, copulatory guide; GH, glandular head of spermatheca. Scale bars: A, B 2.0 mm; C, D 0.5 mm; E, F 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13619170" httpUri="https://zenodo.org/record/13619170/files/figure.png" pageId="2" pageNumber="561">5A–C</figureCitation>
), male pedipalp with tenent setae not covering only the apex, but extending through almost all the prolateral face of cymbium, without retrolateral marginal conductor (RMC), and with embolus curved around the inflated rounded conductor and pointing prolaterally (
<figureCitation id="134E2A58FA77996DFB5E0647FA9674BC" box="[1174,1288,891,917]" captionStart="FIGURES 1" captionStartId="3.[151,264,1925,1949]" captionTargetBox="[159,1428,201,1901]" captionTargetId="figure-19@3.[159,1428,201,1901]" captionTargetPageId="3" captionText="FIGURES 1A–F. Tibelloides castelo sp. nov., male (MPEG.ARA 35202). A dorsal habitus; B ventral habitus; C cephalothorax in dorsal view; D cephalothorax in frontal view; E, F left pedipalp (E ventral, F retrolateral).Abbreviations: C, conductor; RTA, retrolateral tibial apophysis. Scale bars: A, B 1.0 mm; C, D 0.5 mm; E, F 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13619158" httpUri="https://zenodo.org/record/13619158/files/figure.png" pageId="2" pageNumber="561">Figs 1E,F</figureCitation>
;
<figureCitation id="134E2A58FA77996DFADC0647FACC74BC" box="[1300,1362,891,917]" captionStart="FIGURES 3" captionStartId="6.[151,257,1911,1935]" captionTargetBox="[210,1377,151,1889]" captionTargetId="figure-19@6.[210,1377,151,1889]" captionTargetPageId="6" captionText="FIGURES 3A–G. Tibelloides castelo sp. nov., male (A–D UFMG 620) and female (E IBSP 345186, F–G MPEG.ARA 35200). A–D left pedipalp (A ventral view, arrows indicate tenent setae, B tegulum apex, C tibial apophyses in retrolateral view, D tibial apophyses in ventral view); E epigyne in ventral view; F, G vulva in dorsal view.Abbreviations: C, conductor; CG, copulatory guide; E, embolus; MF, membranous field; RTA, retrolateral tibial apophysis; VTA, ventral tibial apophysis. Scale bars: A, C, E, F 0.1 mm; B, D, G 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13619164" httpUri="https://zenodo.org/record/13619164/files/figure.png" pageId="2" pageNumber="561">3A,B</figureCitation>
;
<figureCitation id="134E2A58FA77996DFA970647FA0674BC" box="[1375,1432,891,917]" captionStart="FIGURES 4" captionStartId="9.[151,264,1905,1929]" captionTargetBox="[159,1428,181,1881]" captionTargetId="figure-19@9.[159,1428,181,1881]" captionTargetPageId="9" captionText="FIGURES 4A–F. Tibelloides juatuba sp. nov., male (IBSP 138220). A dorsal habitus; B ventral habitus; C cephalothorax in dorsal view; D cephalothorax in frontal view; E, F left pedipalp (E ventral, F retrolateral). Abbreviations: C, conductor; RTA, retrolateral tibial apophysis. Scale bars: A, B 2.0 mm; C, D 0.5 mm; E, F 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13619168" httpUri="https://zenodo.org/record/13619168/files/figure.png" pageId="2" pageNumber="561">4E,F</figureCitation>
;
<figureCitation id="134E2A58FA77996DFF5F06A3FF487490" box="[151,214,927,953]" captionStart="FIGURES 6" captionStartId="12.[151,264,1871,1895]" captionTargetBox="[159,1429,345,1848]" captionTargetId="figure-107@12.[159,1429,345,1848]" captionTargetPageId="12" captionText="FIGURES 6A–E. Tibelloides juatuba sp. nov., male (A–D IBSP 138219, E MPEG.ARA 35230) and female (F MPEG.ARA 35230). A–D left pedipalp (A ventral view, arrows indicate tenent setae, B tegulum apex, C tibial apophyses in ventral view, D tibial apophyses in retrolateral view); E leg II, prolateral claw (mesal); F epigyne. Abbreviations: C, conductor; CG, copulatory guide; E, embolus; MF, membranous field; MS, median septum; RTA, retrolateral tibial apophysis; VTA, ventral tibial apophysis. Scale bars: A, F 0.1 mm; B–E 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13619174" httpUri="https://zenodo.org/record/13619174/files/figure.png" pageId="2" pageNumber="561">6A,B</figureCitation>
), and female genitalia with elongated and tubular compound copulatory ducts (
<figureCitation id="134E2A58FA77996DFBA606A3FB707490" box="[1134,1262,927,953]" captionStart="FIGURES 2" captionStartId="4.[151,264,1583,1607]" captionTargetBox="[159,1428,181,1559]" captionTargetId="figure-19@4.[159,1428,181,1559]" captionTargetPageId="4" captionText="FIGURES 2A–G. Tibelloides castelo sp. nov., female (A–E MPEG.ARA 35206, F,G UFRJ 235). A dorsal habitus; B ventral habitus; C cephalothorax in dorsal view; D cephalothorax in frontal view; E epigyne; F cleared epigyne in ventral view; G vulva in dorsal view. Abbreviations: CD, copulatory duct; CG, copulatory guide; GH, glandular head of spermatheca; PR, posterior rim. Scale bars: A, B 2.0 mm; C, D 0.5 mm; E–G 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13619160" httpUri="https://zenodo.org/record/13619160/files/figure.png" pageId="2" pageNumber="561">Figs 2E–G</figureCitation>
,
<figureCitation id="134E2A58FA77996DFB3406A3FADF7490" box="[1276,1345,927,953]" captionStart="FIGURES 3" captionStartId="6.[151,257,1911,1935]" captionTargetBox="[210,1377,151,1889]" captionTargetId="figure-19@6.[210,1377,151,1889]" captionTargetPageId="6" captionText="FIGURES 3A–G. Tibelloides castelo sp. nov., male (A–D UFMG 620) and female (E IBSP 345186, F–G MPEG.ARA 35200). A–D left pedipalp (A ventral view, arrows indicate tenent setae, B tegulum apex, C tibial apophyses in retrolateral view, D tibial apophyses in ventral view); E epigyne in ventral view; F, G vulva in dorsal view.Abbreviations: C, conductor; CG, copulatory guide; E, embolus; MF, membranous field; RTA, retrolateral tibial apophysis; VTA, ventral tibial apophysis. Scale bars: A, C, E, F 0.1 mm; B, D, G 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.13619164" httpUri="https://zenodo.org/record/13619164/files/figure.png" pageId="2" pageNumber="561">3E–G</figureCitation>
;
<figureCitation id="134E2A58FA77996DFA8706A3FA0E7493" box="[1359,1424,927,954]" captionStart="FIGURES 5" captionStartId="11.[151,264,1674,1698]" captionTargetBox="[159,1428,273,1651]" captionTargetId="figure-59@11.[159,1428,273,1651]" captionTargetPageId="11" captionText="FIGURES 5A–F. Tibelloides juatuba sp. nov., female (A–E UFMG 12612, F IBSP 81042).A dorsal habitus; B ventral habitus; C cephalothorax in dorsal view; D cephalothorax in frontal view; E epigyne; F vulva in dorsal view.Abbreviations: CD, copulatory duct; CG, copulatory guide; GH, glandular head of spermatheca. Scale bars: A, B 2.0 mm; C, D 0.5 mm; E, F 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13619170" httpUri="https://zenodo.org/record/13619170/files/figure.png" pageId="2" pageNumber="561">5E–F</figureCitation>
). Also, the CG is placed at the level of epigyne surface, while most of MS is placed deep inside the median depression of the epigyne, differing from most
<taxonomicName id="4C754D5EFA77996DFDEA06DBFD657328" box="[546,763,999,1025]" class="Arachnida" family="Philodromidae" genus="Tibelloides" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B901EACFFA77996DFDEA06DBFD037328" box="[546,669,999,1025]" italics="true" pageId="2" pageNumber="561">Tibelloides</emphasis>
species.
</taxonomicName>
Despite these distinct features, we recognize that
<taxonomicName id="4C754D5EFA77996DFAE906DBFF78730C" authorityName="Prado &amp; Pantoja &amp; Baptista" authorityYear="2024" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="castelo" status="sp. nov.">
<emphasis id="B901EACFFA77996DFAE906DBFF78730C" italics="true" pageId="2" pageNumber="561">Tibelloides castelo</emphasis>
</taxonomicName>
<emphasis id="B901EACFFA77996DFF270137FED4730C" bold="true" box="[239,330,1035,1061]" pageId="2" pageNumber="561">
<taxonomicNameLabel id="A23257B4FA77996DFF270137FED4730C" box="[239,330,1035,1061]" pageId="2" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C754D5EFA77996DFE4C0137FDC0730C" authorityName="Prado &amp; Pantoja &amp; Baptista" authorityYear="2024" box="[388,606,1035,1061]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="juatuba" status="sp. nov.">
<emphasis id="B901EACFFA77996DFE4C0137FDC0730C" box="[388,606,1035,1061]" italics="true" pageId="2" pageNumber="561">Tibelloides juatuba</emphasis>
</taxonomicName>
<emphasis id="B901EACFFA77996DFDAE0137FD5F730C" bold="true" box="[614,705,1035,1061]" pageId="2" pageNumber="561">
<taxonomicNameLabel id="A23257B4FA77996DFDAE0137FD5F730C" box="[614,705,1035,1061]" pageId="2" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
bear important diagnostic characters of the genus, such as male pedipalp with conspicuous RTA, consisting of an elongated sclerotized rod adjoined with an acute or rounded expanded lobe (VTA), CG placed over the inner margin of the anterior portion of the spermathecae and curved around the CA, compound copulatory ducts (CD) robust and with texture and sclerotization similar to the spermathecae itself, and with stalk of GH originating near the anterior border of spermathecae. These features contrast with those from
<taxonomicName id="4C754D5EFA77996DFF1D0183FEB073F0" ID-CoL="7WRZ" box="[213,302,1215,1241]" class="Arachnida" family="Philodromidae" genus="Tibellus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B901EACFFA77996DFF1D0183FEB073F0" box="[213,302,1215,1241]" italics="true" pageId="2" pageNumber="561">Tibellus</emphasis>
</taxonomicName>
, the genus most similar to
<taxonomicName id="4C754D5EFA77996DFD940183FD4973F0" ID-CoL="C8KBT" authorityName="Mello-Leitao" authorityYear="1939" box="[604,727,1215,1241]" class="Arachnida" family="Philodromidae" genus="Tibelloides" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B901EACFFA77996DFD940183FD4973F0" box="[604,727,1215,1241]" italics="true" pageId="2" pageNumber="561">Tibelloides</emphasis>
</taxonomicName>
according to
<bibRefCitation id="EFE44B2CFA77996DFCB80183FBDB73F0" author="Prado, A. W. do &amp; Baptista, R. L. C. &amp; Schinelli, H. B. P. &amp; Takiya, D. M." box="[880,1093,1215,1241]" pageId="2" pageNumber="561" pagination="1 - 35" refId="ref10957" refString="Prado, A. W. do, Baptista, R. L. C., Schinelli, H. B. P. &amp; Takiya, D. M. (2022) Solving a running crab spider puzzle: delimiting Cleocnemis Simon, 1886 with implications on the phylogeny and terminology of genital structures of Philodromidae. BMC Zoology, 7 (51), 1 - 35. https: // doi. org / 10.1186 / s 40850 - 022 - 00136 - 7" type="journal article" year="2022">
Prado
<emphasis id="B901EACFFA77996DFC7001FCFC6F73F0" box="[952,1009,1215,1241]" italics="true" pageId="2" pageNumber="561">et al.</emphasis>
(2022)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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