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<mods:title id="0D4222764E7638B535B7675B8EECF5B3">New insect fossils discovered from the Lower Jurassic Sangonghe Formation at the Turpan Basin, Xinjiang, NW China</mods:title>
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<treatment id="03CBDC45FFF67B0933E7FBEFFBF8DE81" ID-DOI="http://doi.org/10.5281/zenodo.6532869" ID-GBIF-Taxon="195277764" ID-Zenodo-Dep="6532869" LSID="urn:lsid:plazi:treatment:03CBDC45FFF67B0933E7FBEFFBF8DE81" httpUri="http://treatment.plazi.org/id/03CBDC45FFF67B0933E7FBEFFBF8DE81" lastPageNumber="186" pageId="3" pageNumber="186">
<subSubSection id="C3783ED8FFF67B0933E7FBEFFEBAD83E" pageId="3" pageNumber="186" type="nomenclature">
<paragraph id="8BDD6D53FFF67B0933E7FBEFFD9BD81A" blockId="3.[136,631,1089,1115]" box="[136,631,1089,1115]" pageId="3" pageNumber="186">
<heading id="D095DA3FFFF67B0933E7FBEFFD9BD81A" bold="true" box="[136,631,1089,1115]" fontSize="10" level="2" pageId="3" pageNumber="186" reason="3">
<emphasis id="B916B141FFF67B0933E7FBEFFD9BD81A" bold="true" box="[136,631,1089,1115]" pageId="3" pageNumber="186">
Superorder
<taxonomicName id="4C6216D0FFF67B093277FBEFFD9BD81A" authority="Lameere, 1900" authorityName="Lameere" authorityYear="1900" box="[280,631,1089,1115]" kingdom="Animalia" pageId="3" pageNumber="186" rank="superOrder" superOrder="Odonatoptera">Odonatoptera Lameere, 1900</taxonomicName>
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<paragraph id="8BDD6D53FFF67B0933E7FBCBFEBAD83E" blockId="3.[136,342,1125,1151]" box="[136,342,1125,1151]" pageId="3" pageNumber="186">
<heading id="D095DA3FFFF67B0933E7FBCBFEBAD83E" box="[136,342,1125,1151]" fontSize="10" level="2" pageId="3" pageNumber="186" reason="2">
<emphasis id="B916B141FFF67B0933E7FBCBFEBAD83E" bold="true" box="[136,342,1125,1151]" pageId="3" pageNumber="186">Family uncertain</emphasis>
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<subSubSection id="C3783ED8FFF67B0933E7FB24FF1CD8E5" box="[136,240,1162,1188]" pageId="3" pageNumber="186" type="description">
<paragraph id="8BDD6D53FFF67B0933E7FB24FF1CD8E5" blockId="3.[136,240,1162,1188]" box="[136,240,1162,1188]" pageId="3" pageNumber="186">
(
<figureCitation id="135971D6FFF67B0933FFFB24FF04D8E5" box="[144,232,1162,1188]" captionStart="FIGURE 3" captionStartId="3.[136,235,1968,1992]" captionTargetBox="[151,1436,1369,1944]" captionTargetId="figure-398@3.[151,1436,1369,1944]" captionTargetPageId="3" captionText="FIGURE 3. Insect wing fragments from the Sangonghe Formation at Daheyan locality. A, Wing fragment of Odonatoptera, possibly a Protozygoptera of Permagrionidae type, NIGP180148. B, Probably a fragment of auchenorrhynchan hind wing, NIGP180149. C and D, A cockroach forewing without claval part, NIGP180150a, b. Scale = 5 mm in A, 2 mm in BD." figureDoi="http://doi.org/10.5281/zenodo.6530371" httpUri="https://zenodo.org/record/6530371/files/figure.png" pageId="3" pageNumber="186">Fig. 3A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3783ED8FFF67B0933E7FB7DFF24D950" pageId="3" pageNumber="186" type="materials_examined">
<paragraph id="8BDD6D53FFF67B0933E7FB7DFF24D950" blockId="3.[136,777,1235,1334]" pageId="3" pageNumber="186">
<emphasis id="B916B141FFF67B0933E7FB7DFF1BD8AC" bold="true" box="[136,247,1235,1261]" pageId="3" pageNumber="186">Material.</emphasis>
<collectionCode id="ED73F596FFF67B09326FFB7DFEA8D8AC" box="[256,324,1235,1261]" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="3" pageNumber="186">NIGP</collectionCode>
180148, fragment of the costal area of a wing.
</paragraph>
</subSubSection>
<subSubSection id="C3783ED8FFF67B0933D2FAB2FBF8DE81" pageId="3" pageNumber="186" type="discussion">
<paragraph id="8BDD6D53FFF67B0933D2FAB2FBF8DE81" blockId="3.[136,777,1235,1334]" lastBlockId="3.[810,1452,158,704]" pageId="3" pageNumber="186">
<emphasis id="B916B141FFF67B0933D2FAB2FEDDD977" bold="true" box="[189,305,1308,1334]" pageId="3" pageNumber="186">Remarks.</emphasis>
The absence of a kink in ScP and of a distinct nodal furrow strongly suggest that this wing fragment does not belong to Panodonata (
<emphasis id="B916B141FFF67B093660FF6AFAA1DC9C" box="[1295,1357,196,221]" italics="true" pageId="3" pageNumber="186">sensu</emphasis>
<bibRefCitation id="EFF310A2FFF67B093638FF6DFC8ADD42" author="Bechly, G." pageId="3" pageNumber="186" refId="ref6393" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der * Odonata. Petalura, Boblingen, Special Volume 2, 402 pp. (Revised edition of the 1995 publication, with an English appendix including a new phylogenetic system of fossil and Recent Odonata)." type="book" year="1996">Bechly, 1996</bibRefCitation>
). Its nodus fits better with that of a Protozygoptera of the superfamily
<taxonomicName id="4C6216D0FFF67B093093FEA0FAB3DD69" authority="Tillyard, 1928" authorityName="Tillyard" authorityYear="1928" box="[1020,1375,270,296]" pageId="3" pageNumber="186" rank="superFamily" superFamily="Permagrionoidea">Permagrionoidea Tillyard, 1928</taxonomicName>
. These also have aligned nodal crossing and subnodus, numerous postnodal and postsubnodal crossveins, and the base of RP2 distad subnodus, as in this fossil (
<bibRefCitation id="EFF310A2FFF67B093789FED3FA4CDDD6" author="Nel, A. &amp; Prokop, J. &amp; Nel, P. &amp; Grandcolas, P. &amp; Huang, D. Y. &amp; Roques, P. &amp; Guilbert, E. &amp; Dostal, O. &amp; Szwedo, J." box="[1254,1440,381,408]" pageId="3" pageNumber="186" pagination="480 - 506" refId="ref8627" refString="Nel, A., Prokop, J., Nel, P., Grandcolas, P., Huang, D. Y., Roques, P., Guilbert, E., Dostal, O. &amp; Szwedo, J. (2012) Traits and evolution of wing venation pattern in paraneopteran insects. Journal of Morphology, 273, 480 - 506. https: // doi. org / 10.1002 / jmor. 11036" type="journal article" year="2012">
Nel
<emphasis id="B916B141FFF67B093676FED0FAA0DDD6" box="[1305,1356,381,407]" italics="true" pageId="3" pageNumber="186">et al</emphasis>
., 2012
</bibRefCitation>
). But the
<taxonomicName id="4C6216D0FFF67B0930EBFE0CFBAFDDFD" authorityName="Tillyard" authorityYear="1928" box="[900,1091,418,444]" pageId="3" pageNumber="186" rank="superFamily" superFamily="Permagrionoidea">Permagrionoidea</taxonomicName>
are strictly Permian to date, the presence of a representative of this group in the early Jurassic would be quite surprising. This wing could also correspond to a case of reversal in the nodal structures, somewhat similar to the situation in the Mesozoic anisopteran
<taxonomicName id="4C6216D0FFF67B0930D9FDF2FBBADE37" baseAuthorityName="Handlirsch" baseAuthorityYear="1906" box="[950,1110,604,630]" class="Insecta" family="Aeschnidiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="186" phylum="Arthropoda" rank="family">Aeschnidiidae</taxonomicName>
or the Cenozoic zygopteran
<taxonomicName id="4C6216D0FFF67B093045FD2FFC5FDEDA" authorityName="Handlirsch" authorityYear="1907" box="[810,947,641,667]" class="Insecta" family="Sieblosiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="186" phylum="Arthropoda" rank="family">Sieblosiidae</taxonomicName>
that have ScP crossing through the nodus (
<bibRefCitation id="EFF310A2FFF67B09305EFD08FBE4DE81" author="Fleck, G. &amp; Nel, A." box="[817,1032,678,704]" pageId="3" pageNumber="186" pagination="1 - 180" refId="ref7032" refString="Fleck, G. &amp; Nel, A. (2003) Revision of the Mesozoic family Aeschnidiidae (Odonata: Anisoptera). Zoologica, 153, 1 - 180." type="journal article" year="2003">Fleck &amp; Nel, 2003</bibRefCitation>
).
</paragraph>
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</treatment>
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