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<document id="45EB4E042913F5F1E3DE5ECEFEE76BA3" ID-DOI="10.11646/phytotaxa.312.2.2" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725576863161" checkinUser="felipe" docAuthor="Domínguez, Yoannis, Valdés, Cristina Mercedes Panfet &amp; Miranda, Vitor Fernandes Oliveira" docDate="2017" docId="03CB87EAFF957C1DFF5050E7FDF3FEE6" docLanguage="en" docName="phytotaxa.312.2.2.pdf" docOrigin="Phytotaxa 312 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.312.2.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Pinguicula calyptrata Kunth" docType="treatment" docVersion="1" lastPageNumber="183" masterDocId="FFF2FF92FF917C18FFD85431FFB2FFE7" masterDocTitle="Typification of names in the genus Pinguicula L. (Lentibulariaceae)" masterLastPageNumber="198" masterPageNumber="179" pageNumber="183" updateTime="1725586476569" updateUser="GgImagineBatch" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:namePart id="499A0A8A12BB8F6B19116A571D88C753">Domínguez, Yoannis</mods:namePart>
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<mods:date id="3B6B983A6AACE5FE2F6750F6668A7382">2017</mods:date>
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<taxonomicName id="4C624D7FFF957C1CFF5050E7FE7DFB16" ID-CoL="4HYGC" authority="Kunth" authorityName="Kunth" box="[136,463,1238,1265]" class="Magnoliopsida" family="Lentibulariaceae" genus="Pinguicula" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="183" phylum="Tracheophyta" rank="species" species="calyptrata">
<emphasis id="B916EAEEFF957C1CFF5050E7FE30FB16" bold="true" box="[136,386,1238,1265]" italics="true" pageId="4" pageNumber="183">Pinguicula calyptrata</emphasis>
Kunth
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in
<treatmentCitation id="0AC310EDFF957C1CFE2B50E9FCCDFB16" author="Bonpland, A. J. A. &amp; Humboldt, F. W. H. A." box="[499,895,1240,1265]" page="226" pageId="4" pageNumber="183" year="1817">
<bibRefCitation id="EFF34B0DFF957C1CFE2B50E9FCCDFB16" author="Bonpland, A. J. A. &amp; Humboldt, F. W. H. A." box="[499,895,1240,1265]" pageId="4" pageNumber="183" refId="ref13099" refString="Bonpland, A. J. A. &amp; Humboldt, F. W. H. A. (1817) Nova genera et species plantarum 2. Sumtibus Librariae Graeco-Latino-Germanicae, Paris, 405 pp. https: // doi. org / 10.5962 / bhl. title. 640" type="book" year="1817">Bonpland &amp; Humboldt (1817: 226)</bibRefCitation>
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<emphasis id="B916EAEEFF957C1CFF5050CFFEFBFAF4" bold="true" box="[136,329,1278,1299]" pageId="4" pageNumber="183">Protologue locality</emphasis>
:—Crescit locis temperatis et frigidis juxta Loxam et in Paramo Saraguru, alt. 6001400 hex, (Regno Novo-Granat.).
</paragraph>
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<paragraph id="8BDD36FCFF957C1CFF505113FE3BFA98" blockId="4.[136,1452,1278,1659]" pageId="4" pageNumber="183">
<emphasis id="B916EAEEFF957C1CFF505113FF08FAD0" bold="true" box="[136,186,1314,1335]" pageId="4" pageNumber="183">Type</emphasis>
(
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, designated by
<bibRefCitation id="EFF34B0DFF957C1CFE195113FDDCFAD0" author="Casper, S. J." box="[449,622,1314,1335]" pageId="4" pageNumber="183" pagination="1 - 209" refId="ref13425" refString="Casper, S. J. (1966) Monographie der Gattung Pinguicula L. Bibliotheca Botanica 127 / 128: 1 - 209." type="journal article" year="1966">Casper 1966: 115</bibRefCitation>
):—
<materialsCitation id="3B0A3CA1FF957C1CFD4A5113FE36FA98" collectionCode="A, J, F, W, H, P" collectorName="von Humboldt" country="Ecuador" location="Loxa" pageId="4" pageNumber="183" specimenCode="P00670050, P00603424" specimenCount="2" typeStatus="isolectotype">
[
<collectingCountry id="F375766CFF957C1CFD405113FCA5FAD0" box="[664,791,1314,1335]" name="Ecuador" pageId="4" pageNumber="183">ECUADOR</collectingCountry>
].
<location id="8EBD6027FF957C1CFCFF5113FCE9FAD0" LSID="urn:lsid:plazi:treatment:03CB87EAFF957C1DFF5050E7FDF3FEE6:8EBD6027FF957C1CFCFF5113FCE9FAD0" box="[807,859,1314,1335]" country="Ecuador" name="Loxa" pageId="4" pageNumber="183">Loxa</location>
, s.d.,
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<collectionCode id="ED73AE39FF957C1CFC4F5113FC1AFAD0" box="[919,936,1314,1335]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="183" type="Herbarium">A</collectionCode>
.
<collectionCode id="ED73AE39FF957C1CFC735113FC0BFAD0" box="[939,953,1314,1335]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="4" pageNumber="183" type="Herbarium">J</collectionCode>
.
<collectionCode id="ED73AE39FF957C1CFC645113FC7FFAD0" box="[956,973,1314,1335]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="183" type="Herbarium">A</collectionCode>
. [Bonpland &amp;
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.
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.
<collectionCode id="ED73AE39FF957C1CFB575113FB11FAD0" box="[1167,1187,1314,1335]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="4" pageNumber="183" type="Herbarium">H</collectionCode>
.
<collectionCode id="ED73AE39FF957C1CFB7E5113FB04FAD0" box="[1190,1206,1314,1335]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="183" type="Herbarium">A</collectionCode>
.
<collectorName id="2697532AFF957C1CFB195113FAFFFAD0" box="[1217,1357,1314,1335]" pageId="4" pageNumber="183">von Humboldt</collectorName>
] 3319
</emphasis>
(
<collectionCode id="ED73AE39FF957C1CFA465113FA1EFAD0" box="[1438,1452,1314,1335]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="183" type="Herbarium">P</collectionCode>
barcode
<specimenCode id="DBC49E87FF957C1CFED75177FE31FABC" box="[271,387,1350,1371]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="183" type="Herbarium">P00670050</specimenCode>
!,
<typeStatus id="54D9885EFF957C1CFE575177FDB1FABC" box="[399,515,1350,1371]" pageId="4" pageNumber="183" type="isolectotype">isolectotype</typeStatus>
<collectionCode id="ED73AE39FF957C1CFDD15177FDA7FABC" box="[521,533,1350,1371]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="183" type="Herbarium">P</collectionCode>
barcode
<specimenCode id="DBC49E87FF957C1CFDB45177FD50FABC" box="[620,738,1350,1371]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="183" type="Herbarium">P00603424</specimenCode>
!; image of the
<typeStatus id="54D9885EFF957C1CFCB15177FC73FABC" box="[873,961,1350,1371]" pageId="4" pageNumber="183" type="lectotype">lectotype</typeStatus>
is available at http://coldb.mnhn.fr/catalognumber/ mnhn/p/p00670050)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3786577FF957C1CFF5051BFFC79F99C" pageId="4" pageNumber="183" type="reference_group">
<paragraph id="8BDD36FCFF957C1CFF5051BFFEECF9B0" blockId="4.[136,1452,1278,1659]" pageId="4" pageNumber="183">
<treatmentCitationGroup id="AB7211D2FF957C1CFF5051BFFEECF9B0" pageId="4" pageNumber="183">
=
<treatmentCitation id="0AC310EDFF957C1CFF4451BFFDE6FA44" author="Cuatrecasas, J." box="[156,596,1422,1443]" page=")" pageId="4" pageNumber="183" year="1945">
<taxonomicName id="4C624D7FFF957C1CFF4451BFFDE6FA44" ID-CoL="6VLLP" authority="Cuatrecasas (1945: 432)" authorityName="Cuatrecasas" authorityPageNumber="432" authorityYear="1945" box="[156,596,1422,1443]" class="Magnoliopsida" family="Lentibulariaceae" genus="Pinguicula" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="183" phylum="Tracheophyta" rank="species" species="huilensis">
<emphasis id="B916EAEEFF957C1CFF4451BFFED3FA44" box="[156,353,1422,1443]" italics="true" pageId="4" pageNumber="183">Pinguicula huilensis</emphasis>
<bibRefCitation id="EFF34B0DFF957C1CFEBF51BFFDE6FA44" author="Cuatrecasas, J." box="[359,596,1422,1443]" pageId="4" pageNumber="183" pagination="421 - 437" refId="ref13657" refString="Cuatrecasas, J. (1945) Estudios sobre plantas andinas, VI. Caldasia 3: 421 - 437." type="journal article" year="1945">Cuatrecasas (1945: 432)</bibRefCitation>
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. Type (
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):—
<collectingCountry id="F375766CFF957C1CFCCC51BFFC28FA44" box="[788,922,1422,1443]" name="Colombia" pageId="4" pageNumber="183">COLOMBIA</collectingCountry>
. Departamento del
<collectingRegion id="49A6F81EFF957C1CFB8C51BFFB20FA44" box="[1108,1170,1422,1443]" country="Colombia" name="Cauca" pageId="4" pageNumber="183">Cauca</collectingRegion>
, Cordillera Central, vertiente occidental del macizo del
<collectingRegion id="49A6F81EFF957C1CFE655183FE47FA20" box="[445,501,1458,1479]" country="Colombia" name="Huila" pageId="4" pageNumber="183">Huila</collectingRegion>
, cabeceras del río Palo en la quebrada del río López, Quebrada del Duende,
<quantity id="4C9A9B19FF957C1CFB395183FAD9FA20" box="[1249,1387,1458,1479]" metricMagnitude="3" metricUnit="m" metricValue="3.425" metricValueMax="3.45" metricValueMin="3.4" pageId="4" pageNumber="183" unit="m" value="3425.0" valueMax="3450.0" valueMin="3400.0">34003450 m</quantity>
. alt.,
<date id="FFDC103CFF957C1CFA475183FEE9FA0C" pageId="4" pageNumber="183" value="1944-12-03">3 December 1944</date>
,
<emphasis id="B916EAEEFF957C1CFEB051E7FD8FFA0C" box="[360,573,1494,1515]" italics="true" pageId="4" pageNumber="183">J. Cuatrecasas 18948</emphasis>
(COL No. 76119 barcode COL000004406 [digital photo!],
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DUKE No. 310256 barcode 10000719 [digital photo!], F No. 1366340 barcode V0061967F [digital photo!], GH barcode 00093360 [digital photo!], MO No. 3521103 barcode MO-716276 [digital photo!], NY barcode 00312393 [digital photo!], US No. 2817244 barcode 00136347 [digital photo!]).
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</paragraph>
<paragraph id="8BDD36FCFF957C1CFF505257FC79F99C" blockId="4.[136,1452,1278,1659]" box="[136,971,1638,1659]" pageId="4" pageNumber="183">
=
<taxonomicName id="4C624D7FFF957C1CFF445257FE09F99C" ID-CoL="4HYKH" authority="Bonpland" authorityName="Bonpland" box="[156,443,1638,1659]" class="Magnoliopsida" family="Lentibulariaceae" genus="Pinguicula" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="183" phylum="Tracheophyta" rank="species" species="loxensis">
<emphasis id="B916EAEEFF957C1CFF445257FEEAF99C" box="[156,344,1638,1659]" italics="true" pageId="4" pageNumber="183">Pinguicula loxensis</emphasis>
Bonpland
</taxonomicName>
in
<bibRefCitation id="EFF34B0DFF957C1CFE025257FD44F99C" author="Humboldt, F. W. H. &amp; Bonpland, A. J. A." box="[474,758,1638,1659]" pageId="4" pageNumber="183" refId="ref14564" refString="Humboldt, F. W. H. A &amp; Bonpland, A. J. A. (1806 - 1823) Voyage aux Regions equinoxiales du Nouveau Continent fait en 1799 - 1804. Chez F. Schoell, Paris, 643 pp." type="book" year="1806">Humboldt &amp; Bonpland (1806</bibRefCitation>
1823: 22),
<emphasis id="B916EAEEFF957C1CFCB25257FC79F99C" box="[874,971,1638,1659]" italics="true" pageId="4" pageNumber="183">nom. nud.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3786577FF957C1DFF50529DFDF3FEE6" lastPageId="5" lastPageNumber="184" pageId="4" pageNumber="183" type="description">
<paragraph id="8BDD36FCFF957C1DFF50529DFDF3FEE6" blockId="4.[136,1452,1708,2093]" lastBlockId="5.[136,1452,160,257]" lastPageId="5" lastPageNumber="184" pageId="4" pageNumber="183">
<emphasis id="B916EAEEFF957C1CFF50529DFF72F922" bold="true" box="[136,192,1708,1733]" pageId="4" pageNumber="183">Note</emphasis>
:—
<taxonomicName id="4C624D7FFF957C1CFF3C529DFE6BF922" box="[228,473,1708,1733]" class="Magnoliopsida" family="Lentibulariaceae" genus="Pinguicula" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="183" phylum="Tracheophyta" rank="species" species="calyptrata">
<emphasis id="B916EAEEFF957C1CFF3C529DFE6BF922" box="[228,473,1708,1733]" italics="true" pageId="4" pageNumber="183">Pinguicula calyptrata</emphasis>
</taxonomicName>
was firstly referred as
<taxonomicName id="4C624D7FFF957C1CFD04529DFBB9F922" authority="Bonpl." authorityName="Bonpl." box="[732,1035,1708,1733]" class="Magnoliopsida" family="Lentibulariaceae" genus="Pinguicula" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="183" phylum="Tracheophyta" rank="species" species="loxensis">
<emphasis id="B916EAEEFF957C1CFD04529DFC0AF922" box="[732,952,1708,1733]" italics="true" pageId="4" pageNumber="183">Pinguicula loxensis</emphasis>
Bonpl.
</taxonomicName>
(
<bibRefCitation id="EFF34B0DFF957C1CFBC2529DFAD2F922" author="Humboldt, F. W. H. &amp; Bonpland, A. J. A." box="[1050,1376,1708,1733]" pageId="4" pageNumber="183" refId="ref14564" refString="Humboldt, F. W. H. A &amp; Bonpland, A. J. A. (1806 - 1823) Voyage aux Regions equinoxiales du Nouveau Continent fait en 1799 - 1804. Chez F. Schoell, Paris, 643 pp." type="book" year="1806">Humboldt &amp; Bonpland 1806</bibRefCitation>
1823: 22) and this name also appears in
<bibRefCitation id="EFF34B0DFF957C1CFDDB52E1FCEBF90E" author="Humboldt, F. W. H. A. &amp; Bonpland, A. J. A." box="[515,857,1744,1769]" pageId="4" pageNumber="183" refId="ref14470" refString="Humboldt, F. W. H. A. &amp; Bonpland, A. J. A. (1805) Voyage de M. M. Alexandre de Humboldt et Aime Bonpland. Essai sur la Geographie des Plantes; acompagne d'un tableau physique des regions equinoxiales, fonde sur des mesures executees, depuis le dixieme degre de latitude boreale jusqu'au dixieme de latitude australe, pendant les annees 1799, 1800, 1801, 1802 et 1803. Chez Levrault, Schoell et Compagnie, Paris, 155 pp." type="book" year="1805">Humboldt &amp; Bonpland (1805)</bibRefCitation>
, but it was never validly published or used otherwise (see
<bibRefCitation id="EFF34B0DFF957C1CFF6452C5FEFDF8EA" author="Casper, S. J." box="[188,335,1780,1805]" pageId="4" pageNumber="183" pagination="14 - 19" refId="ref13450" refString="Casper, S. J. (2003) Pinguicula loxensis (Lentibulariaceae) - A lost name by Humboldt and Bonpland? Carnivorous Plants Newsletter 32: 14 - 19." type="journal article" year="2003">Casper, 2003</bibRefCitation>
for further discussion on this topic). Humboldt and Bonpland seem to have been the first collectors who consistently numbered their plant material in the field (
<bibRefCitation id="EFF34B0DFF957C1CFCE55329FC0EF8D6" author="Lack, H. W." box="[829,956,1816,1841]" pageId="4" pageNumber="183" pagination="218 - 229" refId="ref14805" refString="Lack, H. W. (2001) The plant self impressions prepared by Humboldt and Bonpland in Tropical America. Curtis's Botanical Magazine 18: 218 - 229. https: // doi. org / 10.1111 / 1467 - 8748.00319" type="journal article" year="2001">Lack 2001</bibRefCitation>
); nevertheless, in the work
<emphasis id="B916EAEEFF957C1CFB235329FEE1F8B2" italics="true" pageId="4" pageNumber="183">Nova genera et species plantarum</emphasis>
(
<bibRefCitation id="EFF34B0DFF957C1CFEB8530DFD11F8B2" author="Bonpland, A. J. A. &amp; Humboldt, F. W. H. A." box="[352,675,1852,1877]" pageId="4" pageNumber="183" refId="ref13099" refString="Bonpland, A. J. A. &amp; Humboldt, F. W. H. A. (1817) Nova genera et species plantarum 2. Sumtibus Librariae Graeco-Latino-Germanicae, Paris, 405 pp. https: // doi. org / 10.5962 / bhl. title. 640" type="book" year="1817">Bonpland &amp; Humboldt 1817</bibRefCitation>
), the authors did not link any specimen to the new names published for
<taxonomicName id="4C624D7FFF957C1CFF505351FEB1F89E" authorityName="Linnaeus" authorityYear="1753" box="[136,259,1888,1913]" class="Magnoliopsida" family="Lentibulariaceae" genus="Pinguicula" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="183" phylum="Tracheophyta" rank="genus">
<emphasis id="B916EAEEFF957C1CFF505351FEB1F89E" box="[136,259,1888,1913]" italics="true" pageId="4" pageNumber="183">Pinguicula</emphasis>
</taxonomicName>
. The herbarium of the National Museum of Natural History in Paris (MNHN-P), which keeps the Bonplands private herbarium of the collections made by Humboldt and Bonpland in America (P-Bonpl), holds
<specimenCount id="9D64FD75FF957C1CFB2F53B5FA1EF87A" box="[1271,1452,1924,1949]" count="3" pageId="4" pageNumber="183" type="generic">three specimens</specimenCount>
of
<taxonomicName id="4C624D7FFF957C1CFF7E5399FE86F826" box="[166,308,1960,1985]" class="Magnoliopsida" family="Lentibulariaceae" genus="Pinguicula" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="183" phylum="Tracheophyta" rank="species" species="calyptrata">
<emphasis id="B916EAEEFF957C1CFF7E5399FE86F826" box="[166,308,1960,1985]" italics="true" pageId="4" pageNumber="183">P. calyptrata</emphasis>
</taxonomicName>
from this expedition. Two of these specimens were cited by
<bibRefCitation id="EFF34B0DFF957C1CFC025399FBC9F826" author="Casper, S. J." box="[986,1147,1960,1985]" pageId="4" pageNumber="183" pagination="1 - 209" refId="ref13425" refString="Casper, S. J. (1966) Monographie der Gattung Pinguicula L. Bibliotheca Botanica 127 / 128: 1 - 209." type="journal article" year="1966">Casper (1966)</bibRefCitation>
, the first one as “
<typeStatus id="54D9885EFF957C1CFAE35399FA10F826" box="[1339,1442,1960,1985]" pageId="4" pageNumber="183" type="holotype">holotype</typeStatus>
” (P00670050) kept in the general herbarium P, and the second as “Dupl. in herbaria Bonpland-P” (P00603424). Those specimens certainly resulted from a single gathering according to the locality (Loxa) and collector number (3319); on the other hand, the third specimen (P00603425), in spite of bearing a handwritten label by Bonpland, lacks collector number, and the information on locality is different (
<collectingCountry id="F375766CFF947C1DFD3B5491FC9AFF5E" box="[739,808,160,185]" name="Peru" pageId="5" pageNumber="184">Pérou</collectingCountry>
) which makes it difficult its recognition as
<typeStatus id="54D9885EFF947C1DFAC55491FA1AFF5E" box="[1309,1448,160,185]" pageId="5" pageNumber="184" type="isolectotype">isolectotype</typeStatus>
. Because of this, we agree with Caspers selection and the specimens P00670050 and P00603424 are here corrected to
<typeStatus id="54D9885EFF947C1DFF5054D9FF5DFEE6" box="[136,239,232,257]" pageId="5" pageNumber="184" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54D9885EFF947C1DFEFD54D9FE1EFEE6" box="[293,428,232,257]" pageId="5" pageNumber="184" type="isolectotype">isolectotype</typeStatus>
respectively.
</paragraph>
</subSubSection>
</treatment>
</document>