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<document id="F1DC1520AB5180B55E547FD49B5DD35B" ID-CLB-Dataset="22007" ID-DOI="10.11646/zootaxa.4779.3.1" ID-GBIF-Dataset="712a6136-5d6c-4505-8a8a-3d0556e146c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3835357" ID-ZooBank="0B6F641C-424E-4042-A9B6-A130C58935AB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589959206004" checkinUser="plazi" docAuthor="Ganesh, S. R., Achyuthan, N. S., Chandramouli, S. R. &amp; Vogel, Gernot" docDate="2020" docId="03CB8788EB20FFF8FF047EB3FDE9396C" docLanguage="en" docName="zootaxa.4779.3.1.pdf" docOrigin="Zootaxa 4779 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Boiga beddomei" docType="treatment" docVersion="7" lastPageNumber="310" masterDocId="FFF2FFF0EB27FFF1FF937D3EFFA93A33" masterDocTitle="Taxonomic revision of the Boiga ceylonensis group (Serpentes: Colubridae): reexamination of type specimens, redefinition of nominate taxa and an updated key" masterLastPageNumber="332" masterPageNumber="301" pageNumber="308" updateTime="1732493194171" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D64D3CC1CBD6B0208EC616ADDD592A56">Taxonomic revision of the Boiga ceylonensis group (Serpentes: Colubridae): reexamination of type specimens, redefinition of nominate taxa and an updated key</mods:title>
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<mods:name id="9B28007D9972B67C0BA10D159A22AFFE" type="personal">
<mods:role id="EC854BE75196690B17E1F34EEA6B0B39">
<mods:roleTerm id="FD84F2B32EDA15DB04755AC19A552EC3">Author</mods:roleTerm>
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<mods:namePart id="30620E4B28AEAD71DBC7642E2BDACCBB">Ganesh, S. R.</mods:namePart>
<mods:affiliation id="44B3ED03EFDAB8A04A9985CBA1E4C6F2">Chennai Snake Park, Chennai, Tamil Nadu, India</mods:affiliation>
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<mods:namePart id="D24E7DAE156456582B8D232A26CBAB54">Achyuthan, N. S.</mods:namePart>
<mods:affiliation id="EAAFDE7B5CB97E2C7E21B9AEC0EBBC0A">Centre for Ecological Sciences, Indian Inst. of Science, Bangalore, India</mods:affiliation>
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<mods:namePart id="4676A52CFE6861F39A1DC8D4F7A061D3">Chandramouli, S. R.</mods:namePart>
<mods:affiliation id="1FFAC8028A3A77C75FCF95AC830784D9">Dept. of Ecology &amp; Environmental Sciences, Pondicherry University, India</mods:affiliation>
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<mods:namePart id="9EA7AD6C85DAF01FFC9DD540D25D68A6">Vogel, Gernot</mods:namePart>
<mods:affiliation id="68783E813127C2164E2BD4F9E6CBDD9A">Society for South East Asian Herpetology, Im Sand- 3, Heidelberg, Germany</mods:affiliation>
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<mods:title id="DCB641FF3654B7C85924C327CAD163C6">Zootaxa</mods:title>
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<mods:date id="D1683161C5DD863E74C4DD0BA1F2115C">2020</mods:date>
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<mods:number id="4684B1285EE123B86CB34CBBB1D08392">2020-05-20</mods:number>
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<mods:number id="6793A72F0F4EE688DD0CAEA64B10EB54">4779</mods:number>
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<treatment id="03CB8788EB20FFF8FF047EB3FDE9396C" ID-DOI="http://doi.org/10.5281/zenodo.3851852" ID-GBIF-Taxon="164243322" ID-Zenodo-Dep="3851852" LSID="urn:lsid:plazi:treatment:03CB8788EB20FFF8FF047EB3FDE9396C" httpUri="http://treatment.plazi.org/id/03CB8788EB20FFF8FF047EB3FDE9396C" lastPageId="9" lastPageNumber="310" pageId="7" pageNumber="308" scope_family="Colubridae">
<subSubSection id="C3786515EB20FFF6FF047EB3FE0239F8" pageId="7" pageNumber="308" type="nomenclature">
<paragraph id="8BDD369EEB20FFF6FF047EB3FE503994" blockId="7.[151,505,909,971]" box="[151,505,909,936]" pageId="7" pageNumber="308">
<heading id="D09581F2EB20FFF6FF047EB3FE503994" bold="true" box="[151,505,909,936]" fontSize="11" level="1" pageId="7" pageNumber="308" reason="1">
<taxonomicName id="4C624D1DEB20FFF6FF047EB3FE503994" ID-CoL="5WNGZ" authority="(Wall, 1909)" baseAuthorityName="Wall" baseAuthorityYear="1909" box="[151,505,909,936]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB20FFF6FF047EB3FE503994" bold="true" box="[151,505,909,936]" pageId="7" pageNumber="308">
<emphasis id="B916EA8CEB20FFF6FF047EB3FEF3399B" bold="true" box="[151,346,909,936]" italics="true" pageId="7" pageNumber="308">Boiga beddomei</emphasis>
(
<bibRefCitation id="EFF34B6FEB20FFF6FEFA7EB3FE593994" author="Wall, F." box="[361,496,909,935]" pageId="7" pageNumber="308" pagination="151 - 155" refId="ref14306" refString="Wall, F. (1909) Remarks on some forms of Dipsadomorphus. Records of the Indian Museum, 3, 151 - 155." type="journal article" year="1909">Wall, 1909</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDD369EEB20FFF6FF047E8EFE0239F8" blockId="7.[151,505,909,971]" box="[151,427,944,971]" pageId="7" pageNumber="308">
<heading id="D09581F2EB20FFF6FF047E8EFE0239F8" bold="true" box="[151,427,944,971]" fontSize="11" level="1" pageId="7" pageNumber="308" reason="1">
<emphasis id="B916EA8CEB20FFF6FF047E8EFE0239F8" bold="true" box="[151,427,944,971]" pageId="7" pageNumber="308">
(
<tableCitation id="C6E00325EB20FFF6FF0C7E8EFE8839F8" box="[159,289,944,971]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStartId-0="5.[151,239,478,502]" captionStartId-1="5.[151,239,1290,1314]" captionText-0="TABLE 3. Diagnostic characters of the members of the Boiga ceylonensis group (including B. barnesii; * rarely crown" captionText-1="TABLE 4. Selected morphological characters of the member in the Boiga ceylonensis group (including B. barnesii)" pageId="7" pageNumber="308">Tables 3–4</tableCitation>
;
<figureCitation id="13592A1BEB20FFF6FEBC7E8EFEDA39F8" box="[303,371,944,971]" captionStart="FIGURE 4" captionStartId="8.[151,250,1915,1940]" captionTargetBox="[151,1436,488,1890]" captionTargetId="figure@8.[151,1436,488,1890]" captionTargetPageId="8" captionText="FIGURE 4. Lectotype of Boiga beddomei BMNH 69.8.28.123 from Matheran. (A) entire—dorsal, (B) entire—ventral, (C) ventral pattern close-up, (D) head—ventral, (E) head—lateral, (F) head—dorsal views." figureDoi="http://doi.org/10.5281/zenodo.3835365" httpUri="https://zenodo.org/record/3835365/files/figure.png" pageId="7" pageNumber="308">Fig. 4</figureCitation>
,
<figureCitation id="13592A1BEB20FFF6FE137E8EFE0A39F8" box="[384,419,944,971]" captionStart="FIGURE 9" captionStartId="16.[151,250,1733,1758]" captionTargetBox="[158,1430,181,1707]" captionTargetId="figure@16.[158,1430,181,1707]" captionTargetPageId="16" captionText="FIGURE 9. (A&amp;B) Boiga ceylonensis from Kanneliya &amp; Hunasgiriya, Sri Lanka; (C) Boiga beddomei from Pune, India; (D) Boiga nuchalis from Anaimalai, India; (E) Boiga barnesii from Kandy, Sri Lanka; (F) Boiga andamanensis from Andaman Islands, India." figureDoi="http://doi.org/10.5281/zenodo.3835375" httpUri="https://zenodo.org/record/3835375/files/figure.png" pageId="7" pageNumber="308">9C</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3786515EB20FFF6FF047ECBFD9E3E1F" pageId="7" pageNumber="308" type="reference_group">
<paragraph id="8BDD369EEB20FFF6FF047ECBFD8A3E3E" blockId="7.[151,653,1012,1099]" box="[151,547,1012,1037]" pageId="7" pageNumber="308">
<taxonomicName id="4C624D1DEB20FFF6FF047ECBFD8A3E3E" authority="Wall, 1909" authorityName="Wall" authorityYear="1909" box="[151,547,1012,1037]" class="Squamata" family="Colubridae" genus="Dipsadomorphus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB20FFF6FF047ECBFE073E3E" box="[151,430,1013,1037]" italics="true" pageId="7" pageNumber="308">Dipsadomorphus beddomei</emphasis>
<bibRefCitation id="EFF34B6FEB20FFF6FE277ECAFD8A3E3E" author="Wall, F." box="[436,547,1012,1037]" pageId="7" pageNumber="308" pagination="151 - 155" refId="ref14306" refString="Wall, F. (1909) Remarks on some forms of Dipsadomorphus. Records of the Indian Museum, 3, 151 - 155." type="journal article" year="1909">Wall, 1909</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BDD369EEB20FFF6FF04792AFD9E3E1F" blockId="7.[151,653,1012,1099]" box="[151,567,1043,1068]" pageId="7" pageNumber="308">
<taxonomicName id="4C624D1DEB20FFF6FF04792AFE073E1F" authorityName="Wall" authorityYear="1909" box="[151,430,1044,1068]" class="Squamata" family="Colubridae" genus="Dipsadomorphus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB20FFF6FF04792AFE073E1F" box="[151,430,1044,1068]" italics="true" pageId="7" pageNumber="308">Dipsadomorphus beddomei</emphasis>
</taxonomicName>
—
<bibRefCitation id="EFF34B6FEB20FFF6FE5A792DFD9E3E1F" author="Wall, F." box="[457,567,1043,1068]" pageId="7" pageNumber="308" refId="ref14332" refString="Wall, F. (1919) Notes on a collection of snakes made in the Nilgiri hills and the adjacent Wayanad. Journal of the Bombay" type="book" year="1919">Wall, 1919</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3786515EB20FFF6FF04790DFBBA3C54" pageId="7" pageNumber="308" type="description">
<paragraph id="8BDD369EEB20FFF6FF04790DFD243E79" blockId="7.[151,653,1012,1099]" box="[151,653,1074,1099]" pageId="7" pageNumber="308">
<emphasis id="B916EA8CEB20FFF6FF04790DFEFA3E79" box="[151,339,1074,1098]" italics="true" pageId="7" pageNumber="308">
<taxonomicName id="4C624D1DEB20FFF6FF04790DFF7D3E79" authorityName="Fitzinger" authorityYear="1826" box="[151,212,1075,1098]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="genus">Boiga</taxonomicName>
beddomei—
</emphasis>
<bibRefCitation id="EFF34B6FEB20FFF6FEC0790CFDC93E78" author="Whitaker, R. &amp; Captain, A." box="[339,608,1074,1099]" pageId="7" pageNumber="308" refId="ref14471" refString="Whitaker, R. &amp; Captain, A. (2004) Snakes of India-the field guide. Draco Books, Chennai, 500 pp." type="book" year="2004">Whitaker &amp; Captain, 2004</bibRefCitation>
part
</paragraph>
<paragraph id="8BDD369EEB20FFF6FF047946FB233E84" blockId="7.[151,1437,1144,1999]" pageId="7" pageNumber="308">
<emphasis id="B916EA8CEB20FFF6FF047946FE333EA0" bold="true" box="[151,410,1144,1171]" pageId="7" pageNumber="308">Specimens examined.</emphasis>
<materialsCitation id="3B0A3CC3EB20FFF6FE367947FB233E84" ID-GBIF-Occurrence="2621056324" collectingDate="1848" collectingDateMax="1919" collectingDateMin="1848" collectionCode="BMNH" collectorName="Dr. Leith &amp; Andrew Henderson Leith" country="India" location="Females" pageId="7" pageNumber="308" specimenCode="BMNH 69.8" specimenCount="1" stateProvince="Maharashtra" typeStatus="lectotype">
<location id="8EBD6045EB20FFF6FE367947FDAB3EA0" LSID="urn:lsid:plazi:treatment:03CB8788EB20FFF8FF047EB3FDE9396C:8EBD6045EB20FFF6FE367947FDAB3EA0" box="[421,514,1145,1171]" country="India" name="Females" pageId="7" pageNumber="308" stateProvince="Maharashtra">Females</location>
(n=4).
<typeStatus id="54D9883CEB20FFF6FDCD7947FD673EA0" box="[606,718,1145,1171]" pageId="7" pageNumber="308" type="lectotype">Lectotype</typeStatus>
(designated herein):
<collectingCountry id="F375760EEB20FFF6FC5B7947FBAC3EA0" box="[968,1029,1145,1171]" name="India" pageId="7" pageNumber="308">India</collectingCountry>
.
<collectingRegion id="49A6F87CEB20FFF6FB807947FB0A3EA0" box="[1043,1187,1145,1171]" country="India" name="Maharashtra" pageId="7" pageNumber="308">Maharashtra</collectingRegion>
.
<specimenCode id="DBC49EE5EB20FFF6FB227947FAEA3EA0" box="[1201,1347,1145,1171]" collectionCode="BMNH" pageId="7" pageNumber="308">BMNH 69.8</specimenCode>
.28.123, from “Matheran”; collected by
<collectorName id="26975348EB20FFF6FE6479A3FDF63E84" box="[503,607,1181,1207]" pageId="7" pageNumber="308">Dr. Leith</collectorName>
, probably
<collectorName id="26975348EB20FFF6FD4079A3FC463E84" box="[723,1007,1181,1207]" pageId="7" pageNumber="308">Andrew Henderson Leith</collectorName>
(
<date id="FFDC105EEB20FFF6FC6C79A3FBD73E84" box="[1023,1150,1181,1207]" pageId="7" pageNumber="308" value="1848" valueMax="1919">
<collectingDate id="EF98E9B6EB20FFF6FC6C79A3FBD73E84" box="[1023,1150,1181,1207]" pageId="7" pageNumber="308" value="1848" valueMax="1919">1848-1919</collectingDate>
</date>
).
</materialsCitation>
</paragraph>
<paragraph id="8BDD369EEB20FFF6FF5479FFFBA73EE8" blockId="7.[151,1437,1144,1999]" box="[199,1038,1216,1243]" pageId="7" pageNumber="308">Other specimens: BNHS 1854 and 1854 Matheran; BNHS 1896 Khandala.</paragraph>
<paragraph id="8BDD369EEB20FFF6FF5479DAFEF13F80" blockId="7.[151,1437,1144,1999]" pageId="7" pageNumber="308">
<emphasis id="B916EA8CEB20FFF6FF5479DAFE1B3ECC" bold="true" box="[199,434,1252,1279]" pageId="7" pageNumber="308">Taxonomic History.</emphasis>
<taxonomicName id="4C624D1DEB20FFF6FE2A79D8FDD83ECC" baseAuthorityName="Wall" baseAuthorityYear="1909" box="[441,625,1253,1279]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB20FFF6FE2A79D8FDD83ECC" box="[441,625,1253,1279]" italics="true" pageId="7" pageNumber="308">Boiga beddomei</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFF34B6FEB20FFF6FCD279DBFC6E3ECC" author="Wall, F." box="[833,967,1253,1279]" pageId="7" pageNumber="308" pagination="151 - 155" refId="ref14306" refString="Wall, F. (1909) Remarks on some forms of Dipsadomorphus. Records of the Indian Museum, 3, 151 - 155." type="journal article" year="1909">Wall (1909)</bibRefCitation>
as
<taxonomicName id="4C624D1DEB20FFF6FC7D79DBFA8A3ECC" authorityName="Wall" authorityYear="1909" box="[1006,1315,1253,1279]" class="Squamata" family="Colubridae" genus="Dipsadomorphus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB20FFF6FC7D79DBFA8A3ECC" box="[1006,1315,1253,1279]" italics="true" pageId="7" pageNumber="308">Dipsadomorphus beddomei</emphasis>
</taxonomicName>
during his revision of the
<taxonomicName id="4C624D1DEB20FFF6FED27834FE483F10" baseAuthorityName="Gunther" baseAuthorityYear="1858" box="[321,481,1289,1315]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="ceylonensis">
<emphasis id="B916EA8CEB20FFF6FED27834FE483F10" box="[321,481,1289,1315]" italics="true" pageId="7" pageNumber="308">B. ceylonensis</emphasis>
</taxonomicName>
group. The description was based on a series of specimens from
<collectingCountry id="F375760EEB20FFF6FB2D7837FB513F10" box="[1214,1272,1289,1315]" name="India" pageId="7" pageNumber="308">India</collectingCountry>
and
<collectingCountry id="F375760EEB20FFF6FABD7837FA353F10" box="[1326,1436,1289,1315]" name="Sri Lanka" pageId="7" pageNumber="308">Sri Lanka</collectingCountry>
(
<bibRefCitation id="EFF34B6FEB20FFF6FF0D7813FE093F74" author="Vogel, G. &amp; Ganesh, S. R." box="[158,416,1325,1351]" pageId="7" pageNumber="308" pagination="158 - 168" refId="ref14246" refString="Vogel, G. &amp; Ganesh, S. R. (2013) A new species of cat snake (Reptilia: Serpentes: Colubridae: Boiga) from dry forests of eastern Peninsular India. Zootaxa, 3637 (2), 158 - 168. https: // doi. org / 10.11646 / zootaxa. 3637.2.6" type="journal article" year="2013">Vogel &amp; Ganesh 2013</bibRefCitation>
;
<bibRefCitation id="EFF34B6FEB20FFF6FE3E7813FD263F74" author="Wallach, V. &amp; Williams, K. L. &amp; Boundy, J." box="[429,655,1325,1351]" pageId="7" pageNumber="308" refId="ref14417" refString="Wallach, V., Williams, K. L. &amp; Boundy, J. (2014) Snakes of the World: A Catalogue of Living and Extinct Species. CRC Press, Boca Raton, Florida, 1237 pp. https: // doi. org / 10.1201 / b 16901" type="book" year="2014">
Wallach
<emphasis id="B916EA8CEB20FFF6FD837810FDEA3F74" box="[528,579,1325,1351]" italics="true" pageId="7" pageNumber="308">et al</emphasis>
. 2014
</bibRefCitation>
).
<bibRefCitation id="EFF34B6FEB20FFF6FD317813FCC73F74" author="Annandale, N." box="[674,878,1325,1351]" pageId="7" pageNumber="308" pagination="281 - 282" refId="ref12217" refString="Annandale, N. (1909) Major Wall on some forms of Dipsadomorphus. Records of the Indian Museum, 3 (3), 281 - 282." type="journal article" year="1909">Annandale (1909)</bibRefCitation>
and
<bibRefCitation id="EFF34B6FEB20FFF6FC3B7813FB973F74" author="Smith, M. A." box="[936,1086,1325,1351]" pageId="7" pageNumber="308" refId="ref14068" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London, 583 pp." type="book" year="1943">Smith (1943)</bibRefCitation>
synonymized it with
<taxonomicName id="4C624D1DEB20FFF6FAA47810FF743F58" baseAuthorityName="Gunther" baseAuthorityYear="1858" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="ceylonensis">
<emphasis id="B916EA8CEB20FFF6FAA47810FF743F58" italics="true" pageId="7" pageNumber="308">B. ceylonensis</emphasis>
</taxonomicName>
. However later authors (Taylor, 1950) considered it to be valid.
<bibRefCitation id="EFF34B6FEB20FFF6FC58786FFACE3F58" author="Manamendra-Arachchi, K. &amp; Pethiyagoda, R." box="[971,1383,1361,1387]" pageId="7" pageNumber="308" pagination="123 - 124" refId="ref13515" refString="Manamendra-Arachchi, K. &amp; Pethiyagoda, R. (2007) Boiga rawanawei, a junior synonym of Boiga beddomei. Zeylanica, 7 (1), 123 - 124." type="journal article" year="2007">
Manamendra-Arachchi
<emphasis id="B916EA8CEB20FFF6FB4B786CFAA53F58" box="[1240,1292,1361,1387]" italics="true" pageId="7" pageNumber="308">et al</emphasis>
. (2007)
</bibRefCitation>
was misled that its
<typeStatus id="54D9883CEB20FFF6FEA8784BFEDF3FBC" box="[315,374,1397,1423]" pageId="7" pageNumber="308">types</typeStatus>
were lost and synonymised
<taxonomicName id="4C624D1DEB20FFF6FD267848FCDE3FBC" authorityName="Samarawickrama, Samarawickrama, Wijesena &amp; Orlov" authorityYear="2005" box="[693,887,1398,1423]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="ranawanei">
<emphasis id="B916EA8CEB20FFF6FD267848FCDE3FBC" box="[693,887,1398,1423]" italics="true" pageId="7" pageNumber="308">Boiga ranawanei</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFF34B6FEB20FFF6FC21784BFB5D3FBC" author="Samarawickrama, V. A. M. P. K. &amp; Samarawickrama, V. A. P. &amp; Wijesena, N. M. &amp; Orlov, N. L." box="[946,1268,1396,1423]" pageId="7" pageNumber="308" pagination="213 - 222" refId="ref13922" refString="Samarawickrama, V. A. M. P. K., Samarawickrama, V. A. P., Wijesena, N. M. &amp; Orlov, N. L. (2005) A new species of genus Boiga (Serpentes: Colubridae: Colubrinae) from Sri Lanka. Russian Journal of Herpetology, 12 (3), 213 - 222." type="journal article" year="2005">
Samarawickrama
<emphasis id="B916EA8CEB20FFF6FBE87848FB053FBC" box="[1147,1196,1397,1423]" italics="true" pageId="7" pageNumber="308">et al</emphasis>
. 2005
</bibRefCitation>
) with it (
<bibRefCitation id="EFF34B6FEB20FFF6FACA784BFEE53F80" author="Vogel, G. &amp; Ganesh, S. R." pageId="7" pageNumber="308" pagination="158 - 168" refId="ref14246" refString="Vogel, G. &amp; Ganesh, S. R. (2013) A new species of cat snake (Reptilia: Serpentes: Colubridae: Boiga) from dry forests of eastern Peninsular India. Zootaxa, 3637 (2), 158 - 168. https: // doi. org / 10.11646 / zootaxa. 3637.2.6" type="journal article" year="2013">Vogel &amp; Ganesh 2013</bibRefCitation>
).
</paragraph>
<paragraph id="8BDD369EEB20FFF6FF547883FBBA3C54" blockId="7.[151,1437,1144,1999]" pageId="7" pageNumber="308">
As the type series of
<taxonomicName id="4C624D1DEB20FFF6FE297883FCDD3FE4" authority="Wall, 1909" authorityName="Wall" authorityYear="1909" box="[442,884,1469,1495]" class="Squamata" family="Colubridae" genus="Dipsadomorphus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB20FFF6FE297883FD593FE4" box="[442,752,1469,1495]" italics="true" pageId="7" pageNumber="308">Dipsadomorphus beddomei</emphasis>
<bibRefCitation id="EFF34B6FEB20FFF6FD6B7883FCDD3FE4" author="Wall, F." box="[760,884,1469,1495]" pageId="7" pageNumber="308" pagination="151 - 155" refId="ref14306" refString="Wall, F. (1909) Remarks on some forms of Dipsadomorphus. Records of the Indian Museum, 3, 151 - 155." type="journal article" year="1909">Wall, 1909</bibRefCitation>
</taxonomicName>
is a mixture of several populations (
<bibRefCitation id="EFF34B6FEB20FFF6FA857883FA3D3FE4" author="Wall, F." box="[1302,1428,1469,1495]" pageId="7" pageNumber="308" pagination="151 - 155" refId="ref14306" refString="Wall, F. (1909) Remarks on some forms of Dipsadomorphus. Records of the Indian Museum, 3, 151 - 155." type="journal article" year="1909">Wall, 1909</bibRefCitation>
) and as it is part of a species complex (
<bibRefCitation id="EFF34B6FEB20FFF6FDD278DFFCEF3FC8" author="Vogel, G. &amp; Ganesh, S. R." box="[577,838,1505,1531]" pageId="7" pageNumber="308" pagination="158 - 168" refId="ref14246" refString="Vogel, G. &amp; Ganesh, S. R. (2013) A new species of cat snake (Reptilia: Serpentes: Colubridae: Boiga) from dry forests of eastern Peninsular India. Zootaxa, 3637 (2), 158 - 168. https: // doi. org / 10.11646 / zootaxa. 3637.2.6" type="journal article" year="2013">Vogel &amp; Ganesh, 2013</bibRefCitation>
), it is necessary to designate a
<typeStatus id="54D9883CEB20FFF6FB0C78DFFAAF3FC8" box="[1183,1286,1505,1531]" pageId="7" pageNumber="308" type="lectotype">lectotype</typeStatus>
to clarify the taxonomic status. Thus, as per Art. 74.7 of the Code (ICZN, 1999), we hereby designate an adult specimen BMNH 69.8.28.123 from a precisely named Indian locality “Matheran” as the
<typeStatus id="54D9883CEB20FFF6FC357B17FBA43C70" box="[934,1037,1577,1603]" pageId="7" pageNumber="308" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C624D1DEB20FFF6FBA37B17FACD3C70" authorityName="Wall" authorityYear="1909" box="[1072,1380,1577,1603]" class="Squamata" family="Colubridae" genus="Dipsadomorphus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB20FFF6FBA37B17FACD3C70" box="[1072,1380,1577,1603]" italics="true" pageId="7" pageNumber="308">Dipsadomorphus beddomei</emphasis>
</taxonomicName>
. The chosen
<typeStatus id="54D9883CEB20FFF6FF7F7B73FEFF3C54" box="[236,342,1613,1639]" pageId="7" pageNumber="308" type="lectotype">lectotype</typeStatus>
, is the only specimen of the type series originating from
<collectingCountry id="F375760EEB20FFF6FC417B73FBA63C54" box="[978,1039,1613,1639]" name="India" pageId="7" pageNumber="308">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3786515EB20FFF6FF547B4FFCA33C9C" pageId="7" pageNumber="308" type="etymology">
<paragraph id="8BDD369EEB20FFF6FF547B4FFCA33C9C" blockId="7.[151,1437,1144,1999]" pageId="7" pageNumber="308">
<emphasis id="B916EA8CEB20FFF6FF547B4FFEE73CB8" bold="true" box="[199,334,1649,1675]" pageId="7" pageNumber="308">Etymology.</emphasis>
Patronym named in genitive singular case honouring Col. Richard Henry Beddome (
<date id="FFDC105EEB20FFF6FA9D7B4FFA393CB8" box="[1294,1424,1649,1675]" pageId="7" pageNumber="308" value="1830" valueMax="1911">1830–1911</date>
), one of the most prominent herpetologists of South Asia.
</paragraph>
</subSubSection>
<subSubSection id="C3786515EB20FFF6FF547B86FCAB3DFC" pageId="7" pageNumber="308" type="diagnosis">
<paragraph id="8BDD369EEB20FFF6FF547B86FCAB3DFC" blockId="7.[151,1437,1144,1999]" pageId="7" pageNumber="308">
<emphasis id="B916EA8CEB20FFF6FF547B86FDB43CE0" bold="true" box="[199,541,1720,1747]" pageId="7" pageNumber="308">Diagnosis (redefined herein).</emphasis>
A species of
<taxonomicName id="4C624D1DEB20FFF6FD227B84FD5D3CE0" authorityName="Fitzinger" authorityYear="1826" box="[689,756,1722,1747]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="genus">
<emphasis id="B916EA8CEB20FFF6FD227B84FD5D3CE0" box="[689,756,1722,1747]" italics="true" pageId="7" pageNumber="308">Boiga</emphasis>
</taxonomicName>
inhabiting the Western Ghats, defined by the following combination of characters: 19 midbody scale rows (vs.
<quantity id="4C9A9B7BEB20FFF6FD4B7BE3FC953CC4" box="[728,828,1757,1783]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" metricValueMax="5.842" metricValueMin="5.334" pageId="7" pageNumber="308" unit="in" value="22.0" valueMax="23.0" valueMin="21.0">21–23 in</quantity>
<emphasis id="B916EA8CEB20FFF6FCD77BE0FAB43CC4" box="[836,1309,1757,1783]" italics="true" pageId="7" pageNumber="308">
<taxonomicName id="4C624D1DEB20FFF6FCD77BE0FC6C3CC4" baseAuthorityName="Gunther" baseAuthorityYear="1875" box="[836,965,1757,1783]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="nuchalis">B. nuchalis</taxonomicName>
,
<taxonomicName id="4C624D1DEB20FFF6FC427BE0FBF83CC4" baseAuthorityName="Boulenger" baseAuthorityYear="1894" box="[977,1105,1757,1783]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="dightoni">B. dightoni</taxonomicName>
,
<taxonomicName id="4C624D1DEB20FFF6FBCE7BE0FAB43CC4" baseAuthorityName="Wall" baseAuthorityYear="1909" box="[1117,1309,1757,1783]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="andamanensis">B. andamanensis</taxonomicName>
</emphasis>
); predominantly yellowish-brown dorsum (vs. greenish in
<taxonomicName id="4C624D1DEB20FFF6FD517A3CFCF53D28" authorityName="Vogel &amp; Ganesh" authorityYear="2013" box="[706,860,1793,1819]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="flaviviridis">
<emphasis id="B916EA8CEB20FFF6FD517A3CFCF53D28" box="[706,860,1793,1819]" italics="true" pageId="7" pageNumber="308">B. flaviviridis</emphasis>
</taxonomicName>
; variable in
<taxonomicName id="4C624D1DEB20FFF6FC7D7A3CFB063D28" baseAuthorityName="Wall" baseAuthorityYear="1909" box="[1006,1199,1793,1819]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="andamanensis">
<emphasis id="B916EA8CEB20FFF6FC7D7A3CFB063D28" box="[1006,1199,1793,1819]" italics="true" pageId="7" pageNumber="308">B. andamanensis</emphasis>
</taxonomicName>
); vertebrals strongly enlarged (vs. mildly enlarged in
<taxonomicName id="4C624D1DEB20FFF6FE6D7A18FDD13D0C" baseAuthorityName="Gunther" baseAuthorityYear="1869" box="[510,632,1829,1855]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="barnesii">
<emphasis id="B916EA8CEB20FFF6FE6D7A18FDD13D0C" box="[510,632,1829,1855]" italics="true" pageId="7" pageNumber="308">B. barnesii</emphasis>
</taxonomicName>
); crown unpatterned (vs. with blackish-brown blotches in
<taxonomicName id="4C624D1DEB20FFF6FA927A18FA3C3D0C" box="[1281,1429,1829,1855]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="thackerayi">
<emphasis id="B916EA8CEB20FFF6FA927A18FA3C3D0C" box="[1281,1429,1829,1855]" italics="true" pageId="7" pageNumber="308">B. thackerayi</emphasis>
</taxonomicName>
, brownish blotches in
<taxonomicName id="4C624D1DEB20FFF6FE1B7A74FD813D50" baseAuthorityName="Gunther" baseAuthorityYear="1858" box="[392,552,1865,1891]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="ceylonensis">
<emphasis id="B916EA8CEB20FFF6FE1B7A74FD813D50" box="[392,552,1865,1891]" italics="true" pageId="7" pageNumber="308">B. ceylonensis</emphasis>
</taxonomicName>
); subcaudals&gt; 110 (vs. &lt;
<quantity id="4C9A9B7BEB20FFF6FCC27A77FC3E3D50" box="[849,919,1865,1891]" metricMagnitude="0" metricUnit="m" metricValue="2.7939999999999996" pageId="7" pageNumber="308" unit="in" value="110.0">110 in</quantity>
<emphasis id="B916EA8CEB20FFF6FC0D7A74FB483D50" box="[926,1249,1865,1891]" italics="true" pageId="7" pageNumber="308">
<taxonomicName id="4C624D1DEB20FFF6FC0D7A74FBE83D50" baseAuthorityName="Gunther" baseAuthorityYear="1858" box="[926,1089,1865,1891]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="ceylonensis">B. ceylonensis</taxonomicName>
,
<taxonomicName id="4C624D1DEB20FFF6FBDF7A74FB483D50" box="[1100,1249,1865,1891]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="thackerayi">B. thackerayi</taxonomicName>
</emphasis>
); venter densely speckled with dark brown (vs. uniform in
<taxonomicName id="4C624D1DEB20FFF6FD147A50FC8A3DB4" authorityName="Vogel &amp; Ganesh" authorityYear="2013" box="[647,803,1901,1927]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="flaviviridis">
<emphasis id="B916EA8CEB20FFF6FD147A50FC8A3DB4" box="[647,803,1901,1927]" italics="true" pageId="7" pageNumber="308">B. flaviviridis</emphasis>
</taxonomicName>
); ventrolateral region devoid of large white and black blotches (vs. present in
<taxonomicName id="4C624D1DEB20FFF6FE377AACFD923D98" box="[420,571,1937,1963]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="thackerayi">
<emphasis id="B916EA8CEB20FFF6FE377AACFD923D98" box="[420,571,1937,1963]" italics="true" pageId="7" pageNumber="308">B. thackerayi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C624D1DEB20FFF6FDD97AACFD6F3D98" baseAuthorityName="Gunther" baseAuthorityYear="1869" box="[586,710,1937,1963]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="barnesii">
<emphasis id="B916EA8CEB20FFF6FDD97AACFD6F3D98" box="[586,710,1937,1963]" italics="true" pageId="7" pageNumber="308">B. barnesii</emphasis>
</taxonomicName>
), but has a series of spots on both tips of each ventral scale (vs. venter unpatterned in
<emphasis id="B916EA8CEB20FFF6FE1F7A88FD5B3DFC" box="[396,754,1973,1999]" italics="true" pageId="7" pageNumber="308">
<taxonomicName id="4C624D1DEB20FFF6FE1F7A88FDE73DFC" baseAuthorityName="Wall" baseAuthorityYear="1909" box="[396,590,1973,1999]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="andamanensis">B. andamanensis</taxonomicName>
,
<taxonomicName id="4C624D1DEB20FFF6FDCA7A88FD5B3DFC" authorityName="Vogel &amp; Ganesh" authorityYear="2013" box="[601,754,1973,1999]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="308" phylum="Chordata" rank="species" species="flaviviridis">B. flaviviridis</taxonomicName>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C3786515EB2FFFF8FF547DA6FB163B2C" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309" type="description">
<paragraph id="8BDD369EEB2FFFF9FF547DA6FB043BE1" blockId="8.[151,1437,152,467]" pageId="8" pageNumber="309">
<emphasis id="B916EA8CEB2FFFF9FF547DA6FDB73A80" bold="true" box="[199,542,152,179]" pageId="8" pageNumber="309">
Description of the
<typeStatus id="54D9883CEB2FFFF9FE387DA7FDB13A80" box="[427,536,153,179]" pageId="8" pageNumber="309" type="lectotype">lectotype</typeStatus>
.
</emphasis>
A medium-sized snake reaching
<quantity id="4C9A9B7BEB2FFFF9FC337DA7FC563A80" box="[928,1023,153,179]" metricMagnitude="-1" metricUnit="m" metricValue="8.09" pageId="8" pageNumber="309" unit="mm" value="809.0">809 mm</quantity>
total length, snout-vent length:
<quantity id="4C9A9B7BEB2FFFF9FAE17DA6FF6C3AE4" metricMagnitude="-1" metricUnit="m" metricValue="6.24" pageId="8" pageNumber="309" unit="mm" value="624.0">624 mm</quantity>
, tail length:
<quantity id="4C9A9B7BEB2FFFF9FEC47D82FE113AE4" box="[343,440,188,215]" metricMagnitude="-1" metricUnit="m" metricValue="1.85" pageId="8" pageNumber="309" unit="mm" value="185.0">185 mm</quantity>
, long tail (relative tail length 22.9%); slender habitus, thin neck, wide head; 19–19–15 scale rows; rostral visible from above; preocular 1, subequal to loreal; postoculars 2; loreal 1; supralabials 8/9, with 3
<superScript id="7C179BD6EB2FFFF9FF367C3DFF6B3B2C" attach="none" box="[165,194,259,287]" fontSize="6" pageId="8" pageNumber="309">rd–</superScript>
5
<superScript id="7C179BD6EB2FFFF9FF5C7C3DFF753B22" attach="left" box="[207,220,259,273]" fontSize="6" pageId="8" pageNumber="309">th</superScript>
ones touching eye; infralabials 11/12, with 1–5 touching chin shields; temporals 10/8; preventral 1; ventrals 247, angulate laterally; cloacal 1; subcaudals 117 pairs. Dorsal colour brownish grey, with 109 (81 on body, 28 on tail) blackish crossbars; crossbars covering 2–3 scales in size, extending either sides up to 2–3 scale rows across; interspaces often with sparse dark dots; crown without any markings on top; a thin postocular stripe up to the jaw angle; labials, chin and venter creamy white, finely spotted with darker hue; venter off-white, with a consequent series of brown spots resembling dotted lines, along the ventrolateral edges on both the sides.
</paragraph>
<caption id="DF1D6616EB2FFFF9FF047A45FBA33D8B" ID-DOI="http://doi.org/10.5281/zenodo.3835365" ID-Zenodo-Dep="3835365" httpUri="https://zenodo.org/record/3835365/files/figure.png" pageId="8" pageNumber="309" startId="8.[151,250,1915,1940]" targetBox="[151,1436,488,1890]" targetPageId="8">
<paragraph id="8BDD369EEB2FFFF9FF047A45FBA33D8B" blockId="8.[151,1436,1915,1976]" pageId="8" pageNumber="309">
<emphasis id="B916EA8CEB2FFFF9FF047A45FEBC3DA7" bold="true" box="[151,277,1915,1940]" pageId="8" pageNumber="309">FIGURE 4.</emphasis>
Lectotype of
<taxonomicName id="4C624D1DEB2FFFF9FE397A43FDFB3DA7" baseAuthorityName="Wall" baseAuthorityYear="1909" box="[426,594,1916,1940]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="309" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB2FFFF9FE397A43FDFB3DA7" box="[426,594,1916,1940]" italics="true" pageId="8" pageNumber="309">Boiga beddomei</emphasis>
</taxonomicName>
BMNH 69.8.28.123 from Matheran. (A) entire—dorsal, (B) entire—ventral, (C) ventral pattern close-up, (D) head—ventral, (E) head—lateral, (F) head—dorsal views.
</paragraph>
</caption>
<paragraph id="8BDD369EEB2EFFF8FF547DA6FB163B2C" blockId="9.[151,1437,152,863]" pageId="9" pageNumber="310">
<emphasis id="B916EA8CEB2EFFF8FF547DA6FE963A80" bold="true" box="[199,319,152,179]" pageId="9" pageNumber="310">Variation.</emphasis>
Other specimens agreeing with the
<typeStatus id="54D9883CEB2EFFF8FD447DA7FC973A80" box="[727,830,153,179]" pageId="9" pageNumber="310" type="lectotype">lectotype</typeStatus>
in most respects and showing the following intraspecific variations: snout-vent length
<quantity id="4C9A9B7BEB2EFFF8FD887D82FD1A3AE4" box="[539,691,188,215]" metricMagnitude="-1" metricUnit="m" metricValue="7.89" metricValueMax="8.22" metricValueMin="7.56" pageId="9" pageNumber="310" unit="mm" value="789.0" valueMax="822.0" valueMin="756.0">756–822 mm</quantity>
; tail length 187–219 (incomplete); (18)19–19–15 (14) scale rows; ventrals 238–252; subcaudals 84–87+? (incomplete); temporals
<quantity id="4C9A9B7BEB2EFFF8FCF17DDFFC323AC8" box="[866,923,225,251]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="310" unit="in" value="10.0">10 in</quantity>
all specimens on both sides; supralabials 8–9, rarely with 4
<superScript id="7C179BD6EB2EFFF8FEB57C3AFEEB3B2C" attach="none" box="[294,322,260,287]" fontSize="6" pageId="9" pageNumber="310">th–</superScript>
6
<superScript id="7C179BD6EB2EFFF8FEDC7C3AFEF53B21" attach="left" box="[335,348,260,274]" fontSize="6" pageId="9" pageNumber="310">th</superScript>
touching eye; infralabials 10–11; dorsal bands on body 76–81, on tail 31–33.
</paragraph>
</subSubSection>
<subSubSection id="C3786515EB2EFFF8FF547C16FEEE3BE7" pageId="9" pageNumber="310" type="distribution">
<paragraph id="8BDD369EEB2EFFF8FF547C16FEEE3BE7" blockId="9.[151,1437,152,863]" pageId="9" pageNumber="310">
<emphasis id="B916EA8CEB2EFFF8FF547C16FDE33B70" bold="true" box="[199,586,296,323]" pageId="9" pageNumber="310">Distribution and natural history.</emphasis>
This species occurs only in the hills of northern Western Ghats, north of Goa Gap up to
<collectingCountry id="F375760EEB2EFFF8FE9A7C73FEED3B54" box="[265,324,333,359]" name="India" pageId="9" pageNumber="310">Surat</collectingCountry>
Dangs, where it inhabits deciduous and semi-evergreen forests (Chikane &amp; Bosle 2012;
<bibRefCitation id="EFF34B6FEB2EFFF8FA8B7C73FE873BB8" author="Whitaker, R. &amp; Captain, A." pageId="9" pageNumber="310" refId="ref14471" refString="Whitaker, R. &amp; Captain, A. (2004) Snakes of India-the field guide. Draco Books, Chennai, 500 pp." type="book" year="2004">Whitaker &amp; Captain 2004</bibRefCitation>
part) (Fig. 13). The status of the population from
<collectingCountry id="F375760EEB2EFFF8FCF17C4FFC793BB8" box="[866,976,369,395]" name="Sri Lanka" pageId="9" pageNumber="310">Sri Lanka</collectingCountry>
(see Taylor 1950;
<bibRefCitation id="EFF34B6FEB2EFFF8FB307C4FFA3E3BB8" author="Das, I. &amp; De Silva, A." box="[1187,1431,368,395]" pageId="9" pageNumber="310" refId="ref12663" refString="Das, I. &amp; De Silva, A. (2005) A photographic guide to snakes and other reptiles of Sri Lanka. New Holland Publ., London, 144 pp." type="book" year="2005">Das &amp; De Silva 2005</bibRefCitation>
;
<bibRefCitation id="EFF34B6FEB2EFFF8FF047CABFE7E3B9C" author="Samarawickrama, V. A. M. P. K. &amp; Samarawickrama, V. A. P. &amp; Wijesena, N. M. &amp; Orlov, N. L." box="[151,471,404,431]" pageId="9" pageNumber="310" pagination="213 - 222" refId="ref13922" refString="Samarawickrama, V. A. M. P. K., Samarawickrama, V. A. P., Wijesena, N. M. &amp; Orlov, N. L. (2005) A new species of genus Boiga (Serpentes: Colubridae: Colubrinae) from Sri Lanka. Russian Journal of Herpetology, 12 (3), 213 - 222." type="journal article" year="2005">
Samarawickrama
<emphasis id="B916EA8CEB2EFFF8FECC7CA8FE263B9C" box="[351,399,405,431]" italics="true" pageId="9" pageNumber="310">et al</emphasis>
. 2005
</bibRefCitation>
;
<bibRefCitation id="EFF34B6FEB2EFFF8FE717CABFD0F3B9C" author="Somaweera, R." box="[482,678,405,431]" pageId="9" pageNumber="310" refId="ref14113" refString="Somaweera, R. (2006) The Snakes of Sri Lanka. Wildlife Heritage Trust of Sri Lanka, Colombo, 270 pp. [in Sinhala]" type="book" year="2006">Somaweera 2006</bibRefCitation>
;
<bibRefCitation id="EFF34B6FEB2EFFF8FD237CABFB363B9C" author="Manamendra-Arachchi, K. &amp; Pethiyagoda, R." box="[688,1183,405,431]" pageId="9" pageNumber="310" pagination="123 - 124" refId="ref13515" refString="Manamendra-Arachchi, K. &amp; Pethiyagoda, R. (2007) Boiga rawanawei, a junior synonym of Boiga beddomei. Zeylanica, 7 (1), 123 - 124." type="journal article" year="2007">Manamendra-Arachchi &amp; Pethiyagoda 2007</bibRefCitation>
) warrants further studies (see below).
</paragraph>
</subSubSection>
<subSubSection id="C3786515EB2EFFF8FF547CE2FDE9396C" pageId="9" pageNumber="310" type="discussion">
<paragraph id="8BDD369EEB2EFFF8FF547CE2FDE9396C" blockId="9.[151,1437,152,863]" pageId="9" pageNumber="310">
<emphasis id="B916EA8CEB2EFFF8FF547CE2FEE63BC4" bold="true" box="[199,335,476,503]" pageId="9" pageNumber="310">Comments.</emphasis>
<bibRefCitation id="EFF34B6FEB2EFFF8FEC57CE3FCF13BC4" author="Manamendra-Arachchi, K. &amp; Pethiyagoda, R." box="[342,856,477,503]" pageId="9" pageNumber="310" pagination="123 - 124" refId="ref13515" refString="Manamendra-Arachchi, K. &amp; Pethiyagoda, R. (2007) Boiga rawanawei, a junior synonym of Boiga beddomei. Zeylanica, 7 (1), 123 - 124." type="journal article" year="2007">Manamendra-Arachchi &amp; Pethiyagoda (2007)</bibRefCitation>
considered
<taxonomicName id="4C624D1DEB2EFFF8FC4D7CE0FBDD3BC4" authorityName="Samarawickrama, Samarawickrama, Wijesena &amp; Orlov" authorityYear="2005" box="[990,1140,478,503]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="ranawanei">
<emphasis id="B916EA8CEB2EFFF8FC4D7CE0FBDD3BC4" box="[990,1140,478,503]" italics="true" pageId="9" pageNumber="310">B. ranawanei</emphasis>
</taxonomicName>
to be synonymous with
<taxonomicName id="4C624D1DEB2EFFF8FA177CE0FEAD3828" baseAuthorityName="Wall" baseAuthorityYear="1909" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB2EFFF8FA177CE0FEAD3828" italics="true" pageId="9" pageNumber="310">B. beddomei</emphasis>
</taxonomicName>
as they conceived this species. They created an objective synonymy between these two taxa by designating the
<typeStatus id="54D9883CEB2EFFF8FF787F1BFEE4380C" box="[235,333,549,575]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C624D1DEB2EFFF8FEE17F18FDA1380C" authorityName="Samarawickrama, Samarawickrama, Wijesena &amp; Orlov" authorityYear="2005" box="[370,520,550,575]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="ranawanei">
<emphasis id="B916EA8CEB2EFFF8FEE17F18FDA1380C" box="[370,520,550,575]" italics="true" pageId="9" pageNumber="310">B. ranawanei</emphasis>
</taxonomicName>
as the
<typeStatus id="54D9883CEB2EFFF8FDC57F1BFD06380C" box="[598,687,549,575]" pageId="9" pageNumber="310" type="neotype">neotype</typeStatus>
of
<taxonomicName id="4C624D1DEB2EFFF8FD477F18FCF6380C" baseAuthorityName="Wall" baseAuthorityYear="1909" box="[724,863,549,575]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB2EFFF8FD477F18FCF6380C" box="[724,863,549,575]" italics="true" pageId="9" pageNumber="310">B. beddomei</emphasis>
</taxonomicName>
. As we located without ambiguity the original type series of
<taxonomicName id="4C624D1DEB2EFFF8FF6E7F74FE203850" baseAuthorityName="Wall" baseAuthorityYear="1909" box="[253,393,585,611]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB2EFFF8FF6E7F74FE203850" box="[253,393,585,611]" italics="true" pageId="9" pageNumber="310">B. beddomei</emphasis>
</taxonomicName>
(see also
<bibRefCitation id="EFF34B6FEB2EFFF8FE6A7F77FD5C3850" author="Vogel, G. &amp; Ganesh, S. R." box="[505,757,585,611]" pageId="9" pageNumber="310" pagination="158 - 168" refId="ref14246" refString="Vogel, G. &amp; Ganesh, S. R. (2013) A new species of cat snake (Reptilia: Serpentes: Colubridae: Boiga) from dry forests of eastern Peninsular India. Zootaxa, 3637 (2), 158 - 168. https: // doi. org / 10.11646 / zootaxa. 3637.2.6" type="journal article" year="2013">Vogel &amp; Ganesh 2013</bibRefCitation>
), the
<typeStatus id="54D9883CEB2EFFF8FCA17F77FC223850" box="[818,907,585,611]" pageId="9" pageNumber="310" type="neotype">neotype</typeStatus>
designation done by
<bibRefCitation id="EFF34B6FEB2EFFF8FBE87F77FEDD38B4" author="Manamendra-Arachchi, K. &amp; Pethiyagoda, R." pageId="9" pageNumber="310" pagination="123 - 124" refId="ref13515" refString="Manamendra-Arachchi, K. &amp; Pethiyagoda, R. (2007) Boiga rawanawei, a junior synonym of Boiga beddomei. Zeylanica, 7 (1), 123 - 124." type="journal article" year="2007">Manamendra-Arachchi &amp; Pethiyagoda (2007)</bibRefCitation>
becomes automatically invalid (
<emphasis id="B916EA8CEB2EFFF8FD767F53FCA638B4" box="[741,783,621,647]" italics="true" pageId="9" pageNumber="310">fide</emphasis>
, Art. 75.8 ICZN 1999). The type series of
<taxonomicName id="4C624D1DEB2EFFF8FB667F50FA2B38B4" baseAuthorityName="Wall" baseAuthorityYear="1909" box="[1269,1410,621,647]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB2EFFF8FB667F50FA2B38B4" box="[1269,1410,621,647]" italics="true" pageId="9" pageNumber="310">B. beddomei</emphasis>
</taxonomicName>
is a mixture of two populations, both from
<collectingCountry id="F375760EEB2EFFF8FDF77FAFFD373898" box="[612,670,657,683]" name="India" pageId="9" pageNumber="310">India</collectingCountry>
and
<collectingCountry id="F375760EEB2EFFF8FD457FAFFCE13898" box="[726,840,657,683]" name="Sri Lanka" pageId="9" pageNumber="310">Sri Lanka</collectingCountry>
. We have here designated the only Indian
<typeStatus id="54D9883CEB2EFFF8FAB57FAFFAD43898" box="[1318,1405,657,683]" pageId="9" pageNumber="310" type="syntype">syntype</typeStatus>
as the
<typeStatus id="54D9883CEB2EFFF8FF537F8BFE8E38FC" box="[192,295,693,719]" pageId="9" pageNumber="310" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C624D1DEB2EFFF8FEDF7F88FE7138FC" baseAuthorityName="Wall" baseAuthorityYear="1909" box="[332,472,693,719]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB2EFFF8FEDF7F88FE7138FC" box="[332,472,693,719]" italics="true" pageId="9" pageNumber="310">B. beddomei</emphasis>
</taxonomicName>
. The status of the remaining, Sri Lankan specimens, i.e. the
<typeStatus id="54D9883CEB2EFFF8FBE87F8BFAB438FC" box="[1147,1309,693,719]" pageId="9" pageNumber="310" type="paralectotype">paralectotypes</typeStatus>
of ‘
<taxonomicName id="4C624D1DEB2EFFF8FADF7F88FF7238C0" baseAuthorityName="Wall" baseAuthorityYear="1909" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="beddomei">
<emphasis id="B916EA8CEB2EFFF8FADF7F88FF7238C0" italics="true" pageId="9" pageNumber="310">B. beddomei</emphasis>
</taxonomicName>
’
<taxonomicNameLabel id="A22557F7EB2EFFF8FF797FE7FEF038C0" box="[234,345,729,755]" pageId="9" pageNumber="310" sensu="lato">sensu lato</taxonomicNameLabel>
in BMNH holdings (BMNH 46.12.2
<collectingCountry id="F375760EEB2EFFF8FC937FE7FCFD38C0" box="[768,852,729,755]" name="Sri Lanka" pageId="9" pageNumber="310">Ceylon</collectingCountry>
, BMNH 93.10.6.3_4, BMNH 93.10.6.3 and BMNH 93.10.6.4, all from “Trincomalu” (now Trincomalee or Tirukonamalai, in
<collectingRegion id="49A6F87CEB2EFFF8FC4B7FC3FB333924" box="[984,1178,765,791]" country="Sri Lanka" name="Eastern" pageId="9" pageNumber="310">Eastern Province</collectingRegion>
,
<collectingCountry id="F375760EEB2EFFF8FB347FC3FAB3392B" box="[1191,1306,765,792]" name="Sri Lanka" pageId="9" pageNumber="310">Sri Lanka</collectingCountry>
.) as well as the nominal Sri Lankan species
<taxonomicName id="4C624D1DEB2EFFF8FD997E1CFD1B3908" box="[522,690,802,827]" class="Reptilia" family="Colubridae" genus="Boiga" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="310" phylum="Chordata" rank="species" species="rananwanei">
<emphasis id="B916EA8CEB2EFFF8FD997E1CFD1B3908" box="[522,690,802,827]" italics="true" pageId="9" pageNumber="310">B. rananwanei</emphasis>
</taxonomicName>
await further taxonomic works, pending better resolution of this group and is deferred to future works.
</paragraph>
</subSubSection>
</treatment>
</document>
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