146 lines
17 KiB
XML
146 lines
17 KiB
XML
<document id="2E609C567739FADE9CD89B66DA4946BB" ID-DOI="10.11646/zootaxa.5382.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10279878" ID-ZooBank="8D24E252-9AAD-49C0-9614-82DC61D49E7E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701937850805" checkinUser="plazi" docAuthor="Heiss, Ernst" docDate="2023" docId="03C4DE0AFFDE8626FF1259D7FAFD4569" docLanguage="en" docName="zootaxa.5382.1.6.pdf" docOrigin="Zootaxa 5382 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5382.1.6/52422" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calisiomorpha herczeki Heiss 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="41" masterDocId="FFFDA672FFDF8624FF85582FFFC74474" masterDocTitle="Calisiomorpha herczeki sp. nov. in Cretaceous Burmese amber (Hemiptera: Heteroptera: Aradidae)" masterLastPageNumber="42" masterPageNumber="39" pageNumber="40" updateTime="1701938211500" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="E816DFBEA6EEF6056B0E491FD0525276">
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<mods:title id="50E50C4C87756025000D3CBAA67BCFD7">Calisiomorpha herczeki sp. nov. in Cretaceous Burmese amber (Hemiptera: Heteroptera: Aradidae)</mods:title>
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<mods:name id="BC847EE2AC2906D4CE25B3627A738C80" type="personal">
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<mods:namePart id="6878823FBA57DB7A6699DF505DFA9018">Heiss, Ernst</mods:namePart>
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<mods:title id="74EFF3B602FED1492ECB6B8EE94BD779">Zootaxa</mods:title>
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<mods:part id="D81719280E4D5393F65F6C946D06EC90">
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<mods:date id="263D7612A3F1AC4894FBA911AD91402F">2023</mods:date>
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<mods:number id="942FDC75ACDCC449875C0C46DCB901F5">2023-12-07</mods:number>
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<mods:number id="BA6D0D0554B757DFBEEE345333B16C66">5382</mods:number>
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<mods:start id="22630E2CCF34043FCC54F0EDC373D1EA">39</mods:start>
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<mods:url id="FF2CDD4C8FE08199ECDB66BBD320F266">https://www.mapress.com/zt/article/download/zootaxa.5382.1.6/52422</mods:url>
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<mods:classification id="541406C47730E29718660B9AF2B05B33">journal article</mods:classification>
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<mods:identifier id="ECD8327EFB1A8268BA0AFAA889A8283F" type="DOI">10.11646/zootaxa.5382.1.6</mods:identifier>
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<mods:identifier id="1F2E4EE2E5A5C3E34F6586E98FFEECFC" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03C4DE0AFFDE8626FF1259D7FAFD4569" ID-DOI="http://doi.org/10.5281/zenodo.10279883" ID-Zenodo-Dep="10279883" LSID="urn:lsid:plazi:treatment:03C4DE0AFFDE8626FF1259D7FAFD4569" httpUri="http://treatment.plazi.org/id/03C4DE0AFFDE8626FF1259D7FAFD4569" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40">
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<subSubSection id="C3773C97FFDE8625FF1259D7FDDB4667" box="[151,540,504,531]" pageId="1" pageNumber="40" type="nomenclature">
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<paragraph id="8BD26F1CFFDE8625FF1259D7FDDB4667" blockId="1.[151,540,504,567]" box="[151,540,504,531]" pageId="1" pageNumber="40">
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<heading id="D09AD870FFDE8625FF1259D7FDDB4667" bold="true" box="[151,540,504,531]" fontSize="11" level="1" pageId="1" pageNumber="40" reason="1">
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<emphasis id="B919B30EFFDE8625FF1259D7FDDB4667" bold="true" box="[151,540,504,531]" pageId="1" pageNumber="40">
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<taxonomicName id="4C6D149FFFDE8625FF1259D7FE704667" authority="Heiss, 2023" authorityName="Heiss" authorityYear="2023" box="[151,439,504,531]" class="Insecta" family="Aradidae" genus="Calisiomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="herczeki" status="sp. nov.">
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<emphasis id="B919B30EFFDE8625FF1259D7FE704667" bold="true" box="[151,439,504,531]" italics="true" pageId="1" pageNumber="40">Calisiomorpha herczeki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22A0E75FFDE8625FE3B59D6FDDB4667" box="[446,540,505,531]" pageId="1" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3773C97FFDE8625FF125A33FECC4643" box="[151,267,540,567]" pageId="1" pageNumber="40" type="description">
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<paragraph id="8BD26F1CFFDE8625FF125A33FECC4643" blockId="1.[151,540,504,567]" box="[151,267,540,567]" pageId="1" pageNumber="40">
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(
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<figureCitation id="13567399FFDE8625FF1A5A33FEC44643" box="[159,259,540,567]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1894,1918]" captionTargetBox="[151,1436,309,1870]" captionTargetId="figure-93@2.[151,1436,309,1870]" captionTargetPageId="2" captionText="FIGURES 1–4. Calisiomorpha herczeki sp. nov. 1. Dorsal view; 2. Head and thorax, dorsal view; 3. Abdomen, dorsal view; 4. Abdomen ventral view. Abbreviations: bif = bifurcate carina; car = lateral carina; cly = egg-shaped clypeus; exo = exocorium; par = paranotum; pyg = pygophore?; sc = separated sclerite; sty = stylate eye; TVIII = tergum VIII; vltg VII = ventral laterotergite VII. Scale bars = 1mm Fig. 1; 0.5mm Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.10279880" httpUri="https://zenodo.org/record/10279880/files/figure.png" pageId="1" pageNumber="40">Figs 1–4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3773C97FFDE8625FF125A4AFE4646B2" pageId="1" pageNumber="40" type="materials_examined">
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<paragraph id="8BD26F1CFFDE8625FF125A4AFE4646B2" blockId="1.[151,1437,612,2042]" pageId="1" pageNumber="40">
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<emphasis id="B919B30EFFDE8625FF125A4AFECC460B" bold="true" box="[151,267,613,639]" pageId="1" pageNumber="40">
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<typeStatus id="54D6D1BEFFDE8625FF125A4AFEC1460B" box="[151,262,613,639]" pageId="1" pageNumber="40" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male specimen in an oval piece of dark honey-coloured Burmese amber (16 x
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<quantity id="4C95C2F9FFDE8625FBF65A4BFB02460B" box="[1139,1221,612,639]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="40" unit="mm" value="16.0">16 mm</quantity>
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), legs and antennae nearly complete, ventral side with head and prosternum obscured by impurities. It is designated as
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<typeStatus id="54D6D1BEFFDE8625FA8C5AA7FAAC46D6" box="[1289,1387,648,674]" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
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and labelled accordingly.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3773C97FFDE8625FF425AFFFCB8470E" pageId="1" pageNumber="40" type="diagnosis">
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<paragraph id="8BD26F1CFFDE8625FF425AFFFCB8470E" blockId="1.[151,1437,612,2042]" pageId="1" pageNumber="40">
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<emphasis id="B919B30EFFDE8625FF425AFFFE85469E" bold="true" box="[199,322,720,746]" pageId="1" pageNumber="40">Diagnosis.</emphasis>
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The new species is the second one tentatively assigned to the genus
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<taxonomicName id="4C6D149FFFDE8625FBE55AFFFACF469E" authorityName="Heiss" authorityYear="2016" box="[1120,1288,720,746]" class="Insecta" family="Aradidae" genus="Calisiomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus">
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<emphasis id="B919B30EFFDE8625FBE55AFFFACF469E" box="[1120,1288,720,746]" italics="true" pageId="1" pageNumber="40">Calisiomorpha</emphasis>
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</taxonomicName>
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. It shares its basic characters but differs from
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<taxonomicName id="4C6D149FFFDE8625FD8D5ADAFD46477A" authorityName="Heiss" authorityYear="2016" box="[520,641,757,782]" class="Insecta" family="Aradidae" genus="Calisiomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="yuripopovi">
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<emphasis id="B919B30EFFDE8625FD8D5ADAFD46477A" box="[520,641,757,782]" italics="true" pageId="1" pageNumber="40">yuripopovi</emphasis>
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</taxonomicName>
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by larger size (5.1 vs
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<quantity id="4C95C2F9FFDE8625FCFC5ADAFC14477B" box="[889,979,757,783]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="40" unit="mm" value="3.8">3.8 mm</quantity>
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), head longer than wide (shorter); longer antennae, ratio length of antennae / width of head 1.08 (0.93); pronotum more stout 2.42 times as wide as long (more transverse 2.94 times), longer scutellum 1.33 times as long as width (shorter 1.23 times); lateral connexival expansions shorter and more triangular (longer, conical tubercles).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3773C97FFDE8626FF425BABFBA944A1" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40" type="description">
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<paragraph id="8BD26F1CFFDE8625FF425BABFA8B4002" blockId="1.[151,1437,612,2042]" pageId="1" pageNumber="40">
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<emphasis id="B919B30EFFDE8625FF425BABFE9E47EA" bold="true" box="[199,345,900,926]" pageId="1" pageNumber="40">Description.</emphasis>
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<emphasis id="B919B30EFFDE8625FEE75BABFE5847EA" box="[354,415,900,926]" italics="true" pageId="1" pageNumber="40">Head</emphasis>
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: Slightly longer than wide across eyes (1.1 / 0.95); clypeus egg-shaped enlarged apically reaching apex of antennal segment II, surface granulate and medially depressed; antenniferous lobes long, slightly twisted, apex rounded; antennae 1.08 times as long as width of head, segment I cylindrical, II longer and thicker, tapering towards base, III shortest, IV fusiform and longest, apex pilose; length of antennal segments I / II / III / IV = 0.25 / 0.30 / 0.15 / 0.325; eyes globular, lightly stylate and laterally produced; postocular lobes straightly converging posteriorly to constricted collar; vertex with longitudinal rows of tubercles, depressed between and laterally, carinate along the eyes; rostrum arising from an open atrium, as long as head, rostral groove wide.
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</paragraph>
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<paragraph id="8BD26F1CFFDE8625FF425CAEFE18415F" blockId="1.[151,1437,612,2042]" pageId="1" pageNumber="40">
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<emphasis id="B919B30EFFDE8625FF425CAEFEF140EE" box="[199,310,1153,1178]" italics="true" pageId="1" pageNumber="40">Pronotum</emphasis>
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: 2.42 times as wide as long (0.65 / 1.57), ring-like collar carinate and raised; expanded paranota wide, lateral margins slightly sinuate and bent upward, converging anteriorly, its border crenulate; disk with strongly elevated granulate lateral carinae delimiting the paranota followed on inner side by a less prominent sublateral carina; disk anterior to sublateral carinae with ovate callosities, these separated medially by a deep sulcus; posterior margin carinate and concave.
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</paragraph>
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<paragraph id="8BD26F1CFFDE8625FF425D1BFD3C4106" blockId="1.[151,1437,612,2042]" pageId="1" pageNumber="40">
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<emphasis id="B919B30EFFDE8625FF425D1BFEF2413A" box="[199,309,1332,1358]" italics="true" pageId="1" pageNumber="40">Scutellum</emphasis>
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: Longer than wide (1.0 / 0.75), basal and lateral margins carinate, apex rounded; disk with a median carina, which is bifurcate anteriorly, depressed laterad.
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</paragraph>
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<paragraph id="8BD26F1CFFDE8625FF425D53FD864276" blockId="1.[151,1437,612,2042]" pageId="1" pageNumber="40">
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<emphasis id="B919B30EFFDE8625FF425D53FEFA41E2" box="[199,317,1404,1430]" italics="true" pageId="1" pageNumber="40">Hemelytra</emphasis>
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: Triangular clavus between scutellum and corium distinct, the latter shorter than scutellum; exocorium delimited by a raised carinate vein, then laterally expanded and bent upward; only a small vestige of a transparent membrane is preserved, which might indicate a subbrachypterous status but may also be a remnant of broken membranes of a macropterous morph.
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</paragraph>
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<paragraph id="8BD26F1CFFDE8625FF425E23FDBF4357" blockId="1.[151,1437,612,2042]" pageId="1" pageNumber="40">
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<emphasis id="B919B30EFFDE8625FF425E23FEF64252" box="[199,305,1548,1574]" italics="true" pageId="1" pageNumber="40">Abdomen</emphasis>
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: Of oval outline, connexival plates subrectangular, plain and separated by sutures; tergal plate comprising mediotergites III–VI each with (1+1) oval depressions, surface with flat granulation; anterior margin of deltgs V–VII with an additional small triangular sclerite separated by sutures; lateral margin of connexival edge of deltg III and deltg VII with a dorsal and vltg III and vltg VII with a ventral row of three, deltgs IV, V, VI and vltgs IV, V, VI with four tooth–like flat lateral expansions, the dorsal ones of smaller size and bent upward, der ventral ones are larger and flat; tergite VIII dorsally strongly transverse with four conical posteriorly directed tubercles, ventral side cup-shaped for the reception of the pygophore, which is concealed by tergite VII but seems accessable through a transverse caudal opening of tergite VIII.
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</paragraph>
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<paragraph id="8BD26F1CFFDE8625FF425F02FB0843C6" blockId="1.[151,1437,612,2042]" pageId="1" pageNumber="40">
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<emphasis id="B919B30EFFDE8625FF425F02FEC84332" box="[199,271,1837,1862]" italics="true" pageId="1" pageNumber="40">Venter</emphasis>
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: Pro-, meso- and metasternum flat at middle, separated by transverse sutures; fused sternites II+III anterolaterally constricted and roundly expanded anteriorly between metacoxae; lamellate exocorium is visible and exposed laterad of meso- and metasternum; subrectangular outline of vltgs III–VII and sutures separating them as of dorsal side; no metathoracic scent gland structures and no spiracles discernible with certainty.
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</paragraph>
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<paragraph id="8BD26F1CFFDE8625FF425F92FD0E438E" blockId="1.[151,1437,612,2042]" pageId="1" pageNumber="40">
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<emphasis id="B919B30EFFDE8625FF425F92FF3B43A2" box="[199,252,1981,2006]" italics="true" pageId="1" pageNumber="40">Legs</emphasis>
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: Femora fused to trochanters, moderately incrassate, surface granulate; tibiae cylindrical shorter than femora, tarsi two-segmented with distinct pulvilli.
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</paragraph>
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<paragraph id="8BD26F1CFFDD8626FF4258B8FBA944A1" blockId="2.[151,1436,150,285]" pageId="2" pageNumber="41">
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<emphasis id="B919B30EFFDD8626FF4258B8FEB144C5" bold="true" box="[199,374,151,177]" pageId="2" pageNumber="41">Measurements</emphasis>
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. Length
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<quantity id="4C95C2F9FFDD8626FE5E58B9FDF344C5" box="[475,564,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="41" unit="mm" value="5.1">5.1 mm</quantity>
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; length of antennae
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<quantity id="4C95C2F9FFDD8626FC9058B9FC4D44C5" box="[789,906,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.025" pageId="2" pageNumber="41" unit="mm" value="1.025">1.025 mm</quantity>
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; width of corium at posterior angles of exocorium
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<quantity id="4C95C2F9FFDD8626FD8C5895FDB744A1" box="[521,624,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="41" unit="mm" value="1.25">1.25 mm</quantity>
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; width of abdomen across tergite V
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||
<quantity id="4C95C2F9FFDD8626FB865895FBAD44A1" box="[1027,1130,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="2" pageNumber="41" unit="mm" value="2.25">2.25 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3773C97FFDD8626FF4258F0FAFD4569" pageId="2" pageNumber="41" type="etymology">
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<paragraph id="8BD26F1CFFDD8626FF4258F0FAFD4569" blockId="2.[151,1436,150,285]" pageId="2" pageNumber="41">
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<emphasis id="B919B30EFFDD8626FF4258F0FE89448D" bold="true" box="[199,334,223,249]" pageId="2" pageNumber="41">Etymology.</emphasis>
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It is a great pleasure to dedicate this unique amber taxon to my friend and eminent palaeontologist Aleksander Herczek (University of
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<collectingRegion id="49A9A1FEFFDD8626FDA2592CFDB54569" box="[551,626,259,285]" country="Poland" name="Slaskie" pageId="2" pageNumber="41">Silesia</collectingRegion>
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in Katowice,
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<collectingCountry id="F37A2F8CFFDD8626FC8A592CFCA64569" box="[783,865,259,285]" name="Poland" pageId="2" pageNumber="41">Poland</collectingCountry>
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), honouring and sharing his 70th birthday.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |