315 lines
30 KiB
XML
315 lines
30 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5131771" ID-GBIF-Dataset="09f66541-18e3-4044-b553-7aab04763396" ID-GBIF-Taxon="183733965" ID-ISSN="1982-0224" ID-Zenodo-Dep="5131771" checkinTime="1616760176976" checkinUser="carolina" docAuthor="Menezes, Naércio A. & de Lucena, Carlos Alberto S." docDate="2014" docId="03C4D93834643F62FC2FF89BFC32678D" docLanguage="en" docName="NeotropIchthyol.12.2.193-228.pdf" docOrigin="Neotropical Ichthyology 12 (2)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252014000200193&lng=en&tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Charax tectifer" docType="treatment" docVersion="5" lastPageId="32" lastPageNumber="225" masterDocId="FFFDA140347A3F42FFFBFE69FFCB6623" masterDocTitle="A taxonomic review of the species of Charax Scopoli, 1777 (Teleostei: Characidae: Characinae) with description of a new species from the rio Negro bearing superficial neuromasts on body scales, Amazon basin, Brazil" masterLastPageNumber="228" masterPageNumber="193" pageId="30" pageNumber="223" updateTime="1632022012887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic review of the species of Charax Scopoli, 1777 (Teleostei: Characidae: Characinae) with description of a new species from the rio Negro bearing superficial neuromasts on body scales, Amazon basin, Brazil</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Menezes, Naércio A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>de Lucena, Carlos Alberto S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Neotropical Ichthyology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>12</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>193</mods:start>
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<mods:end>228</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252014000200193&lng=en&tlng=en</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1590/1982-0224-20130175</mods:identifier>
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<mods:identifier type="GBIF-Dataset">09f66541-18e3-4044-b553-7aab04763396</mods:identifier>
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<mods:identifier type="ISSN">1982-0224</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4638995</mods:identifier>
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<mods:identifier type="ZooBank">2CFAB1CE-D283-483B-8C2B-87D95AEADE70</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5131771" ID-GBIF-Taxon="183733965" ID-Zenodo-Dep="5131771" LSID="urn:lsid:plazi:treatment:03C4D93834643F62FC2FF89BFC32678D" httpUri="http://treatment.plazi.org/id/03C4D93834643F62FC2FF89BFC32678D" lastPageId="32" lastPageNumber="225" pageId="30" pageNumber="223">
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<subSubSection box="[980,1296,1778,1802]" pageId="30" pageNumber="223" type="nomenclature">
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<paragraph blockId="30.[980,1296,1778,1834]" box="[980,1296,1778,1802]" pageId="30" pageNumber="223">
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<heading box="[980,1296,1778,1802]" centered="true" fontSize="10" level="2" pageId="30" pageNumber="223" reason="2">
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<taxonomicName authority="(Cope, 1870)" baseAuthorityName="Cope" baseAuthorityYear="1870" box="[980,1296,1778,1802]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="30" pageNumber="223" phylum="Chordata" rank="species" species="tectifer">
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<emphasis bold="true" box="[980,1296,1778,1802]" pageId="30" pageNumber="223">
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<emphasis bold="true" box="[980,1147,1778,1802]" italics="true" pageId="30" pageNumber="223">Charax tectifer</emphasis>
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(Cope, 1870)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[1103,1173,1810,1834]" pageId="30" pageNumber="223" type="description">
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<paragraph blockId="30.[980,1296,1778,1834]" box="[1103,1173,1810,1834]" pageId="30" pageNumber="223">
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<figureCitation box="[1103,1173,1810,1834]" captionStart="Fig" captionStartId="31.[129,167,744,768]" captionTargetBox="[129,1452,176,723]" captionTargetId="figure-619@31.[129,1452,176,723]" captionTargetPageId="31" captionText="Fig. 26. Charax tectifer, MUSM 38670, male, 85 mm SL, Peru, Satipo, río Tambo, Quebrada Pukakunga." figureDoi="http://doi.org/10.5281/zenodo.4639046" httpUri="https://zenodo.org/record/4639046/files/figure.png" pageId="30" pageNumber="223">Fig. 26</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="31" lastPageNumber="224" pageId="30" pageNumber="223" type="reference_group">
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<paragraph blockId="30.[816,1459,1874,1963]" pageId="30" pageNumber="223">
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<taxonomicName authority="Cope, 1870: 565" authorityName="Cope" authorityPageNumber="565" authorityYear="1870" box="[816,1183,1874,1899]" class="Actinopterygii" family="Characidae" genus="Anacyrtus" kingdom="Animalia" order="Characiformes" pageId="30" pageNumber="223" phylum="Chordata" rank="species" species="tectifer">
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<emphasis box="[816,1004,1875,1899]" italics="true" pageId="30" pageNumber="223">Anacyrtus tectifer</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3326898380" box="[1010,1183,1874,1899]" country="Ecuador" location="Ecuador" pageId="30" pageNumber="223" specimenCount="2">Cope, 1870: 565</materialsCitation>
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</taxonomicName>
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[original description,
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<typeStatus box="[1413,1459,1875,1899]" pageId="30" pageNumber="223">type</typeStatus>
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locality:
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<collectingCountry box="[947,1038,1906,1930]" name="Ecuador" pageId="30" pageNumber="223">Ecuador</collectingCountry>
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, Pebas (actually Pebas,
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<collectingCountry box="[1287,1343,1907,1931]" name="Peru" pageId="30" pageNumber="223">Peru</collectingCountry>
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)]. -Böhlke, 1984: 55 (listed in
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<typeStatus box="[1049,1095,1939,1963]" pageId="30" pageNumber="223">type</typeStatus>
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catalog).
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4639046" ID-Zenodo-Dep="4639046" box="[129,1214,744,769]" httpUri="https://zenodo.org/record/4639046/files/figure.png" pageId="31" pageNumber="224" startId="31.[129,167,744,768]" targetBox="[129,1452,176,723]" targetPageId="31">
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<paragraph blockId="31.[129,1214,744,769]" box="[129,1214,744,769]" pageId="31" pageNumber="224">
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<emphasis bold="true" box="[129,210,744,768]" pageId="31" pageNumber="224">Fig. 26.</emphasis>
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<taxonomicName authority=", MUSM" authorityName="MUSM" box="[214,464,745,769]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="tectifer">
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<emphasis box="[214,372,745,769]" italics="true" pageId="31" pageNumber="224">Charax tectifer</emphasis>
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, MUSM
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</taxonomicName>
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38670, male, 85 mm SL, Peru, Satipo, río Tambo, Quebrada Pukakunga.
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</paragraph>
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</caption>
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<paragraph blockId="31.[129,772,816,1481]" pageId="31" pageNumber="224">
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<taxonomicName authority="Gill, 1870: 266" authorityName="Gill" authorityPageNumber="266" authorityYear="1870" box="[129,484,816,841]" class="Actinopterygii" family="Cynodontidae" genus="Hydrolicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="copei">
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<emphasis box="[129,313,817,841]" italics="true" pageId="31" pageNumber="224">Hydrolicus copei</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3326898344" box="[321,484,816,841]" collectorName="Vari & Howe" country="Peru" location="rio Maranon" municipality="rio Napo" pageId="31" pageNumber="224" specimenCount="2">Gill, 1870: 266</materialsCitation>
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</taxonomicName>
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(original description,
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<typeStatus box="[725,771,817,841]" pageId="31" pageNumber="224">type</typeStatus>
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locality:
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<location LSID="urn:lsid:plazi:treatment:03C4D93834643F62FC2FF89BFC32678D:8EB23EF534653F5DFEFEFD38FE40654B" box="[261,395,849,873]" country="Peru" municipality="rio Napo" name="rio Maranon" pageId="31" pageNumber="224">río Marañon</location>
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or
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<collectingMunicipality box="[430,527,849,873]" pageId="31" pageNumber="224">río Napo</collectingMunicipality>
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, Amazon system,
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<collectingCountry box="[722,771,849,872]" name="Peru" pageId="31" pageNumber="224">Peru</collectingCountry>
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or
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<collectingCountry box="[192,282,881,905]" name="Ecuador" pageId="31" pageNumber="224">Ecuador</collectingCountry>
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). -
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<collectorName box="[303,348,881,905]" pageId="31" pageNumber="224">Vari</collectorName>
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&
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<collectorName box="[377,441,881,904]" pageId="31" pageNumber="224">Howe</collectorName>
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, 1991: 26 (listed in
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<typeStatus box="[634,678,881,905]" pageId="31" pageNumber="224">type</typeStatus>
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catalog). -Toledo Piza
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<emphasis box="[303,348,913,937]" italics="true" pageId="31" pageNumber="224">et al</emphasis>
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., 1999: 256 [synonym of
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<taxonomicName authority="(Cope, 1870)" baseAuthorityName="Cope" baseAuthorityYear="1870" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="tectifer">
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<emphasis box="[612,772,913,937]" italics="true" pageId="31" pageNumber="224">Charax tectifer</emphasis>
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(Cope, 1870)
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</taxonomicName>
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].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="224" type="description">
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<paragraph blockId="31.[129,772,816,1481]" pageId="31" pageNumber="224">
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<emphasis box="[129,363,977,1000]" italics="true" pageId="31" pageNumber="224">Anacyrtus sanguineus</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3326898310" collectorName="Lucena & Menezes" country="Peru" county="Fowler" location="Cope" municipality="Bohlke" pageId="31" pageNumber="224" specimenCount="1" typeStatus="lectotype">
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<location LSID="urn:lsid:plazi:treatment:03C4D93834643F62FC2FF89BFC32678D:8EB23EF534653F5DFE8BFDB8FE6065CA" box="[368,427,977,1001]" country="Peru" county="Fowler" municipality="Bohlke" name="Cope" pageId="31" pageNumber="224">Cope</location>
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, 1872: 266 (original description, type locality:
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<collectingCountry box="[303,353,1009,1032]" name="Peru" pageId="31" pageNumber="224">Peru</collectingCountry>
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, ríoAmbyiacu). -
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<collectingMunicipality box="[527,606,1009,1033]" pageId="31" pageNumber="224">Böhlke</collectingMunicipality>
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, 1984: 53 (listed in type catalog). -
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<collectingCounty box="[340,416,1041,1065]" pageId="31" pageNumber="224">Fowler</collectingCounty>
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, 1907: 453 (
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<typeStatus box="[538,634,1041,1065]" pageId="31" pageNumber="224">lectotype</typeStatus>
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designation). -
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<collectorName box="[173,255,1073,1096]" pageId="31" pageNumber="224">Lucena</collectorName>
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&
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<collectorName box="[298,398,1073,1096]" pageId="31" pageNumber="224">Menezes</collectorName>
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, 2003: 202 [synonym of
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<taxonomicName authority="(Cope, 1870)" baseAuthorityName="Cope" baseAuthorityYear="1870" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="tectifer">
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<emphasis italics="true" pageId="31" pageNumber="224">Charax tectifer</emphasis>
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(Cope, 1870)
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</taxonomicName>
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]
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="31.[129,772,816,1481]" pageId="31" pageNumber="224">
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<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1870" box="[129,345,1137,1161]" genus="Moralesia" kingdom="Animalia" pageId="31" pageNumber="224" phylum="Ascomycota" rank="species" species="tectifer">
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<emphasis box="[129,345,1137,1161]" italics="true" pageId="31" pageNumber="224">Moralesia tectifer</emphasis>
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</taxonomicName>
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. -Fowler, 1943: 96 (
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<collectingCountry box="[604,663,1137,1161]" name="Peru" pageId="31" pageNumber="224">Peru</collectingCountry>
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; original description).
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</paragraph>
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<paragraph blockId="31.[129,772,816,1481]" pageId="31" pageNumber="224">
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<taxonomicName box="[129,352,1201,1225]" class="Actinopterygii" family="Characidae" genus="Moralesicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="tectifera">
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<emphasis box="[129,352,1201,1225]" italics="true" pageId="31" pageNumber="224">Moralesicus tectifera</emphasis>
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</taxonomicName>
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. -Fowler, 1958: 9. -Ortega & Vari, 1986: 91 (
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<collectingCountry box="[205,257,1233,1257]" name="Peru" pageId="31" pageNumber="224">Peru</collectingCountry>
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, listed).
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</paragraph>
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<paragraph blockId="31.[129,772,816,1481]" pageId="31" pageNumber="224">
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<taxonomicName authority="tectifer" box="[129,518,1265,1289]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="subSpecies" species="tectifer" subGenus="Moralesia" subSpecies="tectifer">
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<emphasis box="[129,210,1265,1289]" italics="true" pageId="31" pageNumber="224">Charax</emphasis>
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(
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<emphasis box="[227,338,1265,1289]" italics="true" pageId="31" pageNumber="224">Moralesia</emphasis>
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)
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<emphasis box="[356,518,1265,1289]" italics="true" pageId="31" pageNumber="224">tectifer tectifer</emphasis>
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</taxonomicName>
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. -Géry & Vu-Tan-Tuê, 1963: 243 (diagnostic characters in key).
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</paragraph>
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<paragraph blockId="31.[129,772,816,1481]" pageId="31" pageNumber="224">
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<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1870" box="[129,294,1329,1353]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="tectifer">
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<emphasis box="[129,294,1329,1353]" italics="true" pageId="31" pageNumber="224">Charax tectifer</emphasis>
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</taxonomicName>
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. -Lucena, 1987: 60 (diagnosis; description; distribution; geographic variation). -Lucena, 1989: 104 (in key to species; Chang & Ortega, 1995: 5 (
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<collectingCountry box="[617,671,1393,1417]" name="Peru" pageId="31" pageNumber="224">Peru</collectingCountry>
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, listed). - Lucena & Menezes, 2003: 202 (maximum length; distribution).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="224" type="diagnosis">
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<paragraph blockId="31.[129,772,1520,1865]" pageId="31" pageNumber="224">
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<emphasis bold="true" box="[129,246,1520,1544]" pageId="31" pageNumber="224">Diagnosis.</emphasis>
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<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1870" box="[252,416,1521,1545]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="tectifer">
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<emphasis box="[252,416,1521,1545]" italics="true" pageId="31" pageNumber="224">Charax tectifer</emphasis>
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</taxonomicName>
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differs from congeners except
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<taxonomicName authorityName="Menezes & de Lucena" authorityYear="2014" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="delimai">
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<emphasis italics="true" pageId="31" pageNumber="224">C. delimai</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Eigenmann" authorityYear="1922" box="[233,336,1553,1576]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="metae">
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<emphasis box="[233,336,1553,1576]" italics="true" pageId="31" pageNumber="224">C. metae</emphasis>
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</taxonomicName>
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, some specimens of
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<taxonomicName authorityName=", Lucena" authorityYear="1987" box="[589,726,1553,1577]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="gibbosus">
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<emphasis box="[589,726,1553,1577]" italics="true" pageId="31" pageNumber="224">C. gibbosus</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Eigenmann" authorityYear="1912" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="rupununi">
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<emphasis italics="true" pageId="31" pageNumber="224">C. rupununi</emphasis>
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</taxonomicName>
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, and
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<taxonomicName box="[324,447,1585,1609]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="condei">
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<emphasis box="[324,342,1585,1608]" italics="true" pageId="31" pageNumber="224">C</emphasis>
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.
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<emphasis box="[367,447,1585,1609]" italics="true" pageId="31" pageNumber="224">condei</emphasis>
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</taxonomicName>
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in bearing teeth on the ectopterygoid.
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<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1870" box="[299,472,1617,1641]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="tectifer">
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<emphasis box="[299,472,1617,1641]" italics="true" pageId="31" pageNumber="224">Charax tectifer</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName box="[629,739,1617,1641]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="condei">
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<emphasis box="[629,647,1617,1640]" italics="true" pageId="31" pageNumber="224">C</emphasis>
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.
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<emphasis box="[665,739,1617,1641]" italics="true" pageId="31" pageNumber="224">condei</emphasis>
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</taxonomicName>
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in having the lateral line complete (
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<emphasis box="[472,540,1649,1672]" italics="true" pageId="31" pageNumber="224">versus</emphasis>
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incomplete) and from
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<taxonomicName authorityName=", Lucena" authorityYear="1987" box="[129,306,1681,1705]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="gibbosus">
|
|
<emphasis box="[129,306,1681,1705]" italics="true" pageId="31" pageNumber="224">Charax gibbosus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName authorityName="Eigenmann" authorityYear="1912" box="[353,478,1681,1704]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="rupununi">
|
|
<emphasis box="[353,478,1681,1704]" italics="true" pageId="31" pageNumber="224">C. rupununi</emphasis>
|
|
</taxonomicName>
|
|
in having the anal-fin origin along, or slightly posterior to, the vertical through the dorsalfin origin (
|
|
<emphasis box="[239,268,1745,1768]" italics="true" pageId="31" pageNumber="224">vs.</emphasis>
|
|
anterior to the dorsal-fin origin).
|
|
<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1870" box="[612,772,1745,1769]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="tectifer">
|
|
<emphasis box="[612,772,1745,1769]" italics="true" pageId="31" pageNumber="224">Charax tectifer</emphasis>
|
|
</taxonomicName>
|
|
can be distinguished from
|
|
<taxonomicName authorityName="Menezes & de Lucena" authorityYear="2014" box="[424,538,1777,1801]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="delimai">
|
|
<emphasis box="[424,442,1777,1800]" italics="true" pageId="31" pageNumber="224">C</emphasis>
|
|
.
|
|
<emphasis box="[457,538,1777,1801]" italics="true" pageId="31" pageNumber="224">delimai</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName authorityName="Eigenmann" authorityYear="1922" box="[595,693,1777,1800]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="224" phylum="Chordata" rank="species" species="metae">
|
|
<emphasis box="[595,693,1777,1800]" italics="true" pageId="31" pageNumber="224">C. metae</emphasis>
|
|
</taxonomicName>
|
|
by the absence (
|
|
<emphasis box="[228,258,1809,1832]" italics="true" pageId="31" pageNumber="224">vs.</emphasis>
|
|
presence) of the superficial neuromasts on body scales present in those two species (
|
|
<figureCitation box="[516,591,1841,1865]" captionStart="Fig" captionStartId="3.[129,168,636,660]" captionTargetBox="[129,1452,174,615]" captionTargetId="figure-6@3.[129,1452,174,615]" captionTargetPageId="3" captionText="Fig. 1. Superficial neuromasts (black arrows) on lateral body scales of Charax delimai, male (a), MZUSP 81325, paratype, 102 mm SL, Brazil, Amazonas, igarapé Umari, tributary of rio Tiquié in community of São Pedro and female, (b), MZUSP 84988, paratype, 102 mm SL, Brazil, Amazonas, near old community of São Pedro." figureDoi="http://doi.org/10.5281/zenodo.4638997" httpUri="https://zenodo.org/record/4638997/files/figure.png" pageId="31" pageNumber="224">Figs. 1</figureCitation>
|
|
and
|
|
<figureCitation box="[643,659,1841,1865]" captionStart="Fig" captionStartId="3.[129,168,1194,1218]" captionTargetBox="[129,788,734,1181]" captionTargetId="figure-60@3.[129,772,745,1174]" captionText="Fig. 2. Superficial neuromasts (black arrow) on ventral body scales of Charax metae, CAS 69117, 59 mm SL." figureDoi="http://doi.org/10.5281/zenodo.5131735" httpUri="https://zenodo.org/record/5131735/files/figure.png" pageId="31" pageNumber="224">2</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="32" lastPageNumber="225" pageId="31" pageNumber="224" type="description">
|
|
<paragraph blockId="31.[129,771,1904,1961]" pageId="31" pageNumber="224">
|
|
<emphasis bold="true" box="[129,282,1904,1928]" pageId="31" pageNumber="224">Description.</emphasis>
|
|
Morphometrics of examined specimens presented in
|
|
<tableCitation box="[265,360,1936,1961]" captionStart="Table 15" captionStartId="32.[136,199,1287,1311]" captionTargetPageId="32" captionText="Table 15. Morphometrics of Charax tectifer. Specimens are from MZUSP 38709,MUSM38670,37932, 38759." pageId="31" pageNumber="224">Table 15</tableCitation>
|
|
. Body elongate, moderately large (38-
|
|
</paragraph>
|
|
<paragraph blockId="31.[809,1452,817,1961]" pageId="31" pageNumber="224">94 mm SL), compressed and moderately deep. Greatest body depth slightly in advance of dorsal-fin origin. Dorsal profile of head and body convex from tip of snout to anterior region of fontanel, concave from that point to base of supraoccipital spine, convex from that point to dorsal-fin origin, nearly straight along dorsal-fin base and straight to slightly convex from end of dorsal-fin base to caudal peduncle and slightly concave along caudal peduncle. Ventral profile of head and body convex from tip of lower jaw to anal-fin origin, nearly straight along anal-fin base and slightly concave from end of anal-fin base to beginning of procurrent rays. Snout pointed. Lower jaw included in upper jaw when mouth closed. Maxilla extending to vertical through posterior border of orbit.</paragraph>
|
|
<paragraph blockId="31.[809,1452,817,1961]" pageId="31" pageNumber="224">
|
|
Dorsal-fin rays ii,
|
|
<quantity box="[1046,1087,1233,1257]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="31" pageNumber="224" unit="in" value="9.0">9 in</quantity>
|
|
all specimens, posterior most ray unbranched. Adipose fin present. Unbranched anal-fin rays iv or v, usually iv, branched rays 38-43, 40.3. Anterior anal-fin rays not bearing bilateral hooks in examined mature males. Pectoral-fin rays i, 12-16, 13.8. Tips of longest pectoral-fin rays reaching about to vertical through middle of pelvic fin length. Pelvic-fin rays i, 7. No hooks on pelvic-fin rays of all adult males examined. Tips of longest pelvic-fin rays reaching slightly beyond vertical through anal-fin origin. Principal caudal-fin ray count 10/
|
|
<quantity box="[1063,1102,1521,1545]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="31" pageNumber="224" unit="in" value="9.0">9 in</quantity>
|
|
all specimens.
|
|
</paragraph>
|
|
<paragraph blockId="31.[809,1452,817,1961]" pageId="31" pageNumber="224">Lateral line complete; perforated scales 50-56, 53.6. Horizontal scale rows from dorsal-fin origin to lateral line 13- 14, 13.4. Horizontal scale rows from pelvic-fin origin to lateral line 9-10, 9.5. Horizontal scale rows from anal-fin origin to lateral line 11-13, 12.3. Predorsal scales 29-37, 32.8. Scale rows around caudal peduncle 19-21, 19.5. One scale row along analfin base, extending for about two thirds of its length.</paragraph>
|
|
<paragraph blockId="31.[809,1452,817,1961]" lastBlockId="32.[136,779,177,361]" lastPageId="32" lastPageNumber="225" pageId="31" pageNumber="224">Premaxillary with one anterior canine-like tooth followed by set of smaller conical teeth and another canine-like tooth followed by one or two small conical teeth. Total number of premaxillary teeth 11-17, 13.5. Maxillary teeth conical, 34-54, 44.3. Dentary with one canine-like tooth followed by 3-7, 4.5 conical teeth, another canine-like tooth and posterior row of 18-25, 20.7 conical teeth. Left ectopterygoid bone with 3-14, 7.7, right with 2-14 conical teeth. Total number of ectopterygoid teeth 5-25, 15.2. Vertebrae 33 and 34 (2). Gill-rakers on lower limb of first gill-arch 9-10, (9.3). Branchiostegal rays 4; 3 rays originating from anterior ceratohyal and 1 from posterior ceratohyal.</paragraph>
|
|
<paragraph blockId="32.[136,780,401,1225]" pageId="32" pageNumber="225">
|
|
<emphasis bold="true" box="[136,328,401,425]" pageId="32" pageNumber="225">Color in alcohol.</emphasis>
|
|
Body pale to light yellow, slightly darker dorsally than ventrally. Dark chromatophores mostly concentrated on basal portions of scales leaving posterior borders and creating alternating pattern of longitudinal dark and light spots along body. Lateral and ventral portions of body lighter. Dorsal portion of head, snout and tip of lower jaw darker, dark coloration extending over anterior part of maxilla, first and second infraorbitals and between latter bone and anterior border of third infraorbital and defining conspicuous subocular blotch extending ventrally to ventral border of third infraorbital. Remaining portion of third infraorbital, fourth, fifth and sixth infraorbitals, opercle, subopercle and interopercle with scattered dark chromatophores. Irregularly shaped dark humeral blotch, much closer to dorsal-fin origin than to posterior border of opercle; extending two or three scales vertically and horizontally. Dark midlateral stripe extending from humeral blotch to caudal base where it joins oval-shaped dark blotch extending to basal portions of central caudal-fin rays. Vshaped darker lines of chromatophores follow miosepta between musculature. Anal fin with faint basal dark stripe separated by light stripe from wider dark marginal dark stripe. Pelvic and pectoral fins hyaline with very few scattered dark chromatophores. Dorsal and caudal fins hyaline with scattered dark chromatophores mostly located on interradial membranes.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="32" pageNumber="225" type="distribution">
|
|
<paragraph blockId="32.[816,1459,182,271]" pageId="32" pageNumber="225">
|
|
<emphasis bold="true" box="[816,964,182,206]" pageId="32" pageNumber="225">Distribution.</emphasis>
|
|
<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1870" box="[970,1134,183,207]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="32" pageNumber="225" phylum="Chordata" rank="species" species="tectifer">
|
|
<emphasis box="[970,1134,183,207]" italics="true" pageId="32" pageNumber="225">Charax tectifer</emphasis>
|
|
</taxonomicName>
|
|
is known from the
|
|
<collectingRegion box="[1339,1459,183,207]" country="Peru" name="Ucayali" pageId="32" pageNumber="225">río Ucayali</collectingRegion>
|
|
and río Tambo systems,
|
|
<collectingCountry box="[1079,1131,215,238]" name="Peru" pageId="32" pageNumber="225">Peru</collectingCountry>
|
|
, rivers from the río Amazonas drainage in
|
|
<collectingCountry box="[938,1046,247,271]" name="Colombia" pageId="32" pageNumber="225">Colombia</collectingCountry>
|
|
, and río
|
|
<collectingRegion box="[1133,1191,247,270]" country="Ecuador" name="Napo" pageId="32" pageNumber="225">Napo</collectingRegion>
|
|
system,
|
|
<collectingCountry box="[1281,1370,247,271]" name="Ecuador" pageId="32" pageNumber="225">Ecuador</collectingCountry>
|
|
(
|
|
<figureCitation box="[1382,1446,247,271]" captionStart="Fig" captionStartId="5.[809,847,1648,1672]" captionTargetBox="[809,1452,819,1618]" captionTargetId="figure-305@5.[809,1452,815,1618]" captionTargetPageId="5" captionText="Fig.7. Map of SouthAmerica showing the collecting sites of the Charax species, and the type locality of Charax delimai (purple circle). Charax apurensis - yellow square; C. caudimaculatus - white circle; C. condei - white square; C. gibbosus - purple triangle; C. hemigrammus - red star; C. leticiae - black circle;C. macrolepis-black triangle;C.metae-yellow triangle;C.michaeli - yellow circle;C.niger -blue triangle; C.notulatus - blue circle; C. pauciradiatus - red triangle; C. rupununi - red circle; C. stenopterus - green triangle; C. tectifer - green circle. Some symbols may represent more than one locality." figureDoi="http://doi.org/10.5281/zenodo.4639009" httpUri="https://zenodo.org/record/4639009/files/figure.png" pageId="32" pageNumber="225">Fig. 7</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="32" pageNumber="225" type="discussion">
|
|
<paragraph blockId="32.[816,1459,311,431]" pageId="32" pageNumber="225">
|
|
<emphasis bold="true" box="[816,928,311,335]" pageId="32" pageNumber="225">Remarks.</emphasis>
|
|
No significant statistical differences in meristics and morphometrics and no differences in other morphological features were found among the samples from
|
|
<collectingCountry box="[1344,1455,375,399]" name="Colombia" pageId="32" pageNumber="225">Colombia</collectingCountry>
|
|
,
|
|
<collectingCountry box="[816,907,407,431]" name="Ecuador" pageId="32" pageNumber="225">Ecuador</collectingCountry>
|
|
, and
|
|
<collectingCountry box="[960,1013,407,430]" name="Peru" pageId="32" pageNumber="225">Peru</collectingCountry>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |