286 lines
32 KiB
XML
286 lines
32 KiB
XML
<document id="A4DE5CDFD44FAE8FF449476DC9BD2C39" ID-CLB-Dataset="30243" ID-DOI="10.11646/zootaxa.4410.2.7" ID-GBIF-Dataset="3c516d90-f9f4-4003-9329-fd4fe506f0b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1221670" ID-ZooBank="14C3B3C0-2DF9-4893-998F-23D96450C1FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1524196580908" checkinUser="plazi" docAuthor="Ekins, Merrick, Debitus, Cécile, Erpenbeck, Dirk & Hooper, John N. A." docDate="2018" docId="03C4D641FFFB6424A8E1F930010CFC41" docLanguage="en" docName="zootaxa.4410.2.7.pdf" docOrigin="Zootaxa 4410 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Raspailia (Raspaxilla) frondosa Ekins & Debitus & Erpenbeck & Hooper 2018, sp. nov." docType="treatment" docUuid="58E578EB-F6B7-421D-AD4D-922591113883" docUuidSource="ZooBank" docVersion="8" lastPageNumber="383" masterDocId="FFFDAE39FFFA6420A876FFFE007CFFE6" masterDocTitle="A new species of the sponge Raspailia (Raspaxilla) (Porifera: Demospongiae: Axinellida: Raspailiidae) from deep seamounts of the Western Pacific" masterLastPageNumber="386" masterPageNumber="379" pageNumber="380" updateTime="1698538358648" updateUser="ExternalLinkService">
|
||
<mods:mods id="9728490CC3CF149188E1482636A01FEA" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="A2CE0EA3DCC02B965AC6D0F21922D27B">
|
||
<mods:title id="5DE26FBB4CBE89DC281563F617683F5C">A new species of the sponge Raspailia (Raspaxilla) (Porifera: Demospongiae: Axinellida: Raspailiidae) from deep seamounts of the Western Pacific</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="9291656C8BB73495AD8752DF0EC7EAB5" type="personal">
|
||
<mods:role id="396E3264E36F4A22CF4DAF2246533F24">
|
||
<mods:roleTerm id="8A1DD129E26EF9518C78BB1535611046">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="808591E6FD98814793B4CBDCDAB2C6E1">Ekins, Merrick</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="0FFC95EE10E6FA119F66CA72A27DC3C2" type="personal">
|
||
<mods:role id="410BB94644F0F206B677484CC2AB56A4">
|
||
<mods:roleTerm id="A1C064EE1C58806C0E4F6A51C5D33EA4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C3A6B6B8113C4A1D91D6C33FB6E00A29">Debitus, Cécile</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="FDE965D838311910007394A30074D43F" type="personal">
|
||
<mods:role id="4D35CD91D38213FB2F6CD3F99C60CDC6">
|
||
<mods:roleTerm id="F64391989EA757913714BFB5A0CEB8F9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="834D6218DF3EAEEE3CAAC62691F8FE2B">Erpenbeck, Dirk</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="CFB27F79F5E16D4318E8FE93208048C7" type="personal">
|
||
<mods:role id="189EC03E3EBBF6DC52837AF225CDABE0">
|
||
<mods:roleTerm id="747705B38EB25C08C3424716C0FE25EE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A6553E387F76847BC55000477051FB83">Hooper, John N. A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="FDE74E5509E9CE34F5F28E8DE415C907">text</mods:typeOfResource>
|
||
<mods:relatedItem id="D7677BFEA397AF0E71E3AF2B11895BD6" type="host">
|
||
<mods:titleInfo id="5A9641973138DC69EED0AE13BD365B47">
|
||
<mods:title id="4ADAC0B1A57FB8A93CD8E2484DA98A81">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="15F7668E10A54A788A033F83389FF40E">
|
||
<mods:date id="D200B6DE3A6CCDF5390C958DC601A0A8">2018</mods:date>
|
||
<mods:detail id="86719B489D9D044F3BFAE414354F92B2" type="pubDate">
|
||
<mods:number id="EF9F08ACC12CC278AA5680028B43A03B">2018-04-17</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="E6E14543788FE3DEBA15A5946ED96D9C" type="volume">
|
||
<mods:number id="BD199CE989E9B265596678B286133B0A">4410</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="209ADB569622090B670A634EB4016F99" type="issue">
|
||
<mods:number id="9A7B7600E405F2EEEDE637C472FDCCA3">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="4AD4E9EFDCFC4CA3D7F57B8B472EFC7B" unit="page">
|
||
<mods:start id="4E8A73CA9F0DFA1643C6C4ADEEA7BFEF">379</mods:start>
|
||
<mods:end id="248DF2C3051F9754072522C485BB55F7">386</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="7BBF7952EBBE270DF0BA22924F6DD995">journal article</mods:classification>
|
||
<mods:identifier id="0DE2C8D345D625A509931965330A39CB" type="CLB-Dataset">30243</mods:identifier>
|
||
<mods:identifier id="8F3AB76CCA2194106EF90BC649462020" type="DOI">10.11646/zootaxa.4410.2.7</mods:identifier>
|
||
<mods:identifier id="45D949E39F3E01F95C14B14A047F9227" type="GBIF-Dataset">3c516d90-f9f4-4003-9329-fd4fe506f0b8</mods:identifier>
|
||
<mods:identifier id="8EA61D3D71B55AC6A220CD6C98FC5AEC" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="71A4964ECC563D911D4253CD4B3C5157" type="Zenodo-Dep">1221670</mods:identifier>
|
||
<mods:identifier id="6A102F9F60B4F427AB52944233BC60CE" type="ZooBank">14C3B3C0-2DF9-4893-998F-23D96450C1FD</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03C4D641FFFB6424A8E1F930010CFC41" ID-DOI="http://doi.org/10.5281/zenodo.5952262" ID-GBIF-Taxon="143618802" ID-Zenodo-Dep="5952262" LSID="urn:lsid:zoobank.org:act:58E578EB-F6B7-421D-AD4D-922591113883" httpUri="http://treatment.plazi.org/id/03C4D641FFFB6424A8E1F930010CFC41" lastPageId="4" lastPageNumber="383" pageId="1" pageNumber="380">
|
||
<subSubSection id="C37734DCFFFB6421A8E1F9300117F8EC" pageId="1" pageNumber="380" type="nomenclature">
|
||
<paragraph id="8BD26757FFFB6421A8E1F9300200F90E" blockId="1.[151,636,1742,1802]" box="[151,636,1742,1768]" pageId="1" pageNumber="380">
|
||
<heading id="D09AD03BFFFB6421A8E1F9300200F90E" bold="true" box="[151,636,1742,1768]" fontSize="11" level="1" pageId="1" pageNumber="380" reason="1">
|
||
<emphasis id="B919BB45FFFB6421A8E1F9300200F90E" bold="true" box="[151,636,1742,1768]" pageId="1" pageNumber="380">
|
||
<taxonomicName id="4C6D1CD4FFFB6421A8E1F9300264F90E" ID-CoL="4RLMW" authority="Ekins & Debitus & Erpenbeck & Hooper, 2018" authorityName="Ekins & Debitus & Erpenbeck & Hooper" authorityYear="2018" box="[151,536,1742,1768]" class="Demospongiae" family="Raspailiidae" genus="Raspailia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Axinellida" pageId="1" pageNumber="380" phylum="Porifera" rank="species" species="frondosa" status="sp. nov." subGenus="Raspaxilla">
|
||
<emphasis id="B919BB45FFFB6421A8E1F9300264F90E" bold="true" box="[151,536,1742,1768]" italics="true" pageId="1" pageNumber="380">Raspailia (Raspaxilla) frondosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22A063EFFFB6421AA68F9300200F90E" box="[542,636,1742,1768]" pageId="1" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD26757FFFB6421A8E1F90F0117F8EC" blockId="1.[151,636,1742,1802]" box="[151,363,1777,1802]" pageId="1" pageNumber="380">
|
||
(
|
||
<figureCitation id="13567BD2FFFB6421A8E9F90F017FF8EC" box="[159,259,1777,1802]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1506,1528]" captionStartId-1="3.[151,250,1148,1170]" captionStartId-2="3.[151,250,1840,1862]" captionStartId-3="4.[151,250,1732,1754]" captionTargetBox-0="[151,1435,270,1482]" captionTargetBox-1="[152,1435,373,1126]" captionTargetBox-2="[425,1162,1250,1818]" captionTargetBox-3="[306,1281,990,1711]" captionTargetId-0="figure@2.[151,1435,270,1482]" captionTargetId-1="figure@3.[152,1435,373,1126]" captionTargetId-2="figure@3.[425,1162,1250,1818]" captionTargetId-3="figure@4.[306,1281,990,1711]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Fixed specimens of R. (R.) frondosa sp. nov., clockwise from the top left: paratype QM G335144; holotype MNHN-IP-2015-1402 (fragment QM G335124); paratype QM G318701." captionText-1="FIGURE 2. Spicules of the holotype R. (R.) frondosa sp. nov., MNHN-IP-2015-1402 (fragment QM G335124). A subectosomal style, B terminationsof subectosomal style, C–G choanosomal rhabdostyles" captionText-2="FIGURE 3. An undigested cross section of the holotype of R. (R.) frondosa sp. nov., MNHN-IP-2015-1402 (fragment QM G335124), showing the choanosomal rhabdostyles supporting the fragile ectosome and the subectosomal styles penetrating the ectosome." captionText-3="FIGURE 4. Transmitted light micrograph of a crossection of the paratype QM G318701 of R. (R.) frondosa sp. nov., showing the well-differentiated axial and extra axial regions of the skeleton, and the restriction of rhabdostyles and styles to the subectosomal and ectosomal regions." httpUri-0="https://zenodo.org/record/1221672/files/figure.png" httpUri-1="https://zenodo.org/record/1221674/files/figure.png" httpUri-2="https://zenodo.org/record/1221676/files/figure.png" httpUri-3="https://zenodo.org/record/1221678/files/figure.png" pageId="1" pageNumber="380">Figs 1–4</figureCitation>
|
||
,
|
||
<tableCitation id="C6EF52ECFFFB6421A979F90F011CF8EC" box="[271,352,1777,1802]" captionStart="TABLE 1" captionStartId="4.[151,239,260,283]" captionText="TABLE 1. Spicule measurements of the type specimens of Raspailia (Raspaxilla)frondosa sp.nov." pageId="1" pageNumber="380">Table 1</tableCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37734DCFFFB6422A8E1F8C70252FF31" lastPageId="2" lastPageNumber="381" pageId="1" pageNumber="380" type="etymology">
|
||
<paragraph id="8BD26757FFFB6421A8E1F8C70261F825" blockId="1.[151,1437,1849,2024]" pageId="1" pageNumber="380">
|
||
<emphasis id="B919BB45FFFB6421A8E1F8C70162F8B4" bold="true" box="[151,286,1849,1874]" pageId="1" pageNumber="380">Etymology.</emphasis>
|
||
Latin for leafy, reflecting the gross morphology of the sponge resembling the frond of a fern.
|
||
<emphasis id="B919BB45FFFB6421A8B1F8A301C4F890" bold="true" box="[199,440,1885,1910]" pageId="1" pageNumber="380">Material examined.</emphasis>
|
||
<typeStatus id="54D6D9F5FFFB6421A9B0F8A3024FF890" box="[454,563,1885,1910]" pageId="1" pageNumber="380" type="holotype">Holotype</typeStatus>
|
||
: MNHN-IP-2015-1402 (fragment of the
|
||
<typeStatus id="54D6D9F5FFFB6421AC57F8A304FFF890" box="[1057,1155,1885,1910]" pageId="1" pageNumber="380" type="holotype">holotype</typeStatus>
|
||
QM G335124) Station DW3110,
|
||
<collectingCountry id="F37A27C7FFFB6421A966F87B01BFF87B" box="[272,451,1924,1949]" name="New Caledonia" pageId="1" pageNumber="380">New Caledonia</collectingCountry>
|
||
Ridge
|
||
<geoCoordinate id="EE590190FFFB6421AA68F87A02E4F87A" box="[542,664,1920,1949]" direction="south" orientation="latitude" pageId="1" pageNumber="380" precision="9" value="-23.0225">23o1.35’ S</geoCoordinate>
|
||
<geoCoordinate id="EE590190FFFB6421AAD2F87A0340F87A" box="[676,828,1920,1949]" direction="east" orientation="longitude" pageId="1" pageNumber="380" precision="9" value="168.28317">168o16.99’ E</geoCoordinate>
|
||
to
|
||
<geoCoordinate id="EE590190FFFB6421AB1CF87A03A4F87A" box="[874,984,1920,1948]" direction="south" orientation="latitude" pageId="1" pageNumber="380" precision="9" value="-23.0285">23o1.71’S</geoCoordinate>
|
||
<geoCoordinate id="EE590190FFFB6421AB92F87A0415F87B" box="[996,1129,1920,1949]" direction="east" orientation="longitude" pageId="1" pageNumber="380" precision="9" value="168.26433">168o15.86E</geoCoordinate>
|
||
,
|
||
<quantity id="4C95CAB2FFFB6421AC0FF87A057CF87A" box="[1145,1280,1924,1948]" metricMagnitude="2" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="1" pageNumber="380" unit="m" value="290.0" valueMax="310.0" valueMin="270.0">270–310 m</quantity>
|
||
, Coll. Cécile Debitus,
|
||
<date id="FFD34197FFFB6421A88FF8550115F825" box="[249,361,1963,1987]" pageId="1" pageNumber="380" value="2008-10-28">28.x.2008</date>
|
||
(
|
||
<bibRefCitation id="EFFC1AA6FFFB6421A901F855026DF825" box="[375,529,1963,1988]" pageId="1" pageNumber="380" refId="ref3716" refString="Samadi, S. (2008) Terrasse cruise, RV Alis. Available from: https: // doi. org / 10.17600 / 8100100 (accessed 18 September 2017)" type="journal article">Samadi 2008</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD26757FFFB6422A8B1F82E0252FF31" blockId="1.[151,1437,1849,2024]" lastBlockId="2.[151,1436,151,216]" lastPageId="2" lastPageNumber="381" pageId="1" pageNumber="380">
|
||
<materialsCitation id="3B056D0AFFFB6421A8B1F82E0548F801" ID-GBIF-Occurrence="1838187609" box="[199,1332,1998,2024]" collectorName="Station DW" pageId="1" pageNumber="380" specimenCount="1" stateProvince="NORFOLK" typeStatus="paratype">
|
||
<typeStatus id="54D6D9F5FFFB6421A8B1F82E0145F80E" box="[199,313,2000,2024]" pageId="1" pageNumber="380" type="paratype">Paratypes</typeStatus>
|
||
: QM G335144 (same data as holotype). QM G318701
|
||
<collectingRegion id="49A9A9B5FFFB6421ABBEF8310430F801" box="[968,1100,1999,2023]" country="United Kingdom" name="Norfolk" pageId="1" pageNumber="380">NORFOLK</collectingRegion>
|
||
1
|
||
<collectorName id="26980281FFFB6421AC06F8300480F801" box="[1136,1276,1998,2023]" pageId="1" pageNumber="380">Station DW</collectorName>
|
||
1724
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B056D0AFFFB6422AD32F8300252FF31" ID-GBIF-Occurrence="1838187610" collectingDate="2001-06-27" collectorName="Coll. Bertrand Richer & Richer" elevation="246" lastPageId="2" lastPageNumber="381" latitude="-23.284166" location="Coll. Bertrand Richer de Forges" longLatPrecision="12" longitude="168.238" municipality="Dredge" pageId="1" pageNumber="380" specimenCount="1" stateProvince="Norfolk" typeStatus="paratype">
|
||
<collectingRegion id="49A9A9B5FFFB6421AD32F83005E1F801" box="[1348,1437,1998,2023]" country="United Kingdom" name="Norfolk" pageId="1" pageNumber="380">Norfolk</collectingRegion>
|
||
Ridge Seamount Banc.no.1.
|
||
<geoCoordinate id="EE590190FFF86422A99BFF650214FF55" box="[493,616,151,179]" direction="south" orientation="latitude" pageId="2" pageNumber="381" precision="9" value="-23.284166">23o17.05’S</geoCoordinate>
|
||
<geoCoordinate id="EE590190FFF86422AA00FF640378FF55" box="[630,772,151,179]" direction="east" orientation="longitude" pageId="2" pageNumber="381" precision="9" value="168.238">168o14.28’E</geoCoordinate>
|
||
,
|
||
<collectingMunicipality id="6BB6FD2DFFF86422AB61FF640315FF55" box="[791,873,154,179]" pageId="2" pageNumber="381">Dredge</collectingMunicipality>
|
||
<elevation id="00408064FFF86422AB01FF650383FF54" box="[887,1023,155,179]" metricMagnitude="2" metricUnit="m" metricValue="2.455" metricValueMax="2.91" metricValueMin="2.0" pageId="2" pageNumber="381" unit="m" value="245.5" valueMax="291.0" valueMin="200.0">
|
||
<quantity id="4C95CAB2FFF86422AB01FF650383FF54" box="[887,1023,155,179]" metricMagnitude="2" metricUnit="m" metricValue="2.455" metricValueMax="2.91" metricValueMin="2.0" pageId="2" pageNumber="381" unit="m" value="245.5" valueMax="291.0" valueMin="200.0">200–291 m</quantity>
|
||
</elevation>
|
||
,
|
||
<location id="8EB2318CFFF86422AC64FF6405EBFF55" LSID="urn:lsid:plazi:treatment:03C4D641FFFB6424A8E1F930010CFC41:8EB2318CFFF86422AC64FF6405EBFF55" box="[1042,1431,154,179]" latitude="-23.284166" longLatPrecision="12" longitude="168.238" municipality="Dredge" name="Coll. Bertrand Richer de Forges" pageId="2" pageNumber="381" stateProvince="Norfolk">
|
||
<collectorName id="26980281FFF86422AC64FF64056FFF55" box="[1042,1299,154,179]" pageId="2" pageNumber="381">Coll. Bertrand Richer</collectorName>
|
||
de Forges
|
||
</location>
|
||
,
|
||
<collectingDate id="EF97B87FFFF86422A8E1FF400172FF31" box="[151,270,190,215]" pageId="2" pageNumber="381" value="2001-06-27">
|
||
<date id="FFD34197FFF86422A8E1FF400172FF31" box="[151,270,190,215]" pageId="2" pageNumber="381" value="2001-06-27">27.vi.2001</date>
|
||
</collectingDate>
|
||
(
|
||
<collectorName id="26980281FFF86422A96BFF400116FF31" box="[285,362,190,215]" pageId="2" pageNumber="381">Richer</collectorName>
|
||
de
|
||
<bibRefCitation id="EFFC1AA6FFF86422A9E4FF3E025EFF31" box="[402,546,191,216]" pageId="2" pageNumber="381" refId="ref3626" refString="Richer de Forges, B. (2001) Norfolk 1 cruise, RV Alis https: // doi. org / 10.17600 / 1100050" type="journal volume">Forges 2001</bibRefCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1237DFFFF86422A8E1FA1C03F8F9F0" httpUri="https://zenodo.org/record/1221672/files/figure.png" pageId="2" pageNumber="381" startId="2.[151,250,1506,1528]" targetBox="[151,1435,270,1482]" targetPageId="2">
|
||
<paragraph id="8BD26757FFF86422A8E1FA1C03F8F9F0" blockId="2.[151,1436,1506,1558]" pageId="2" pageNumber="381">
|
||
<emphasis id="B919BB45FFF86422A8E1FA1C016AFA11" bold="true" box="[151,278,1506,1528]" pageId="2" pageNumber="381">FIGURE 1.</emphasis>
|
||
Fixed specimens of
|
||
<taxonomicName id="4C6D1CD4FFF86422A981FA1D02DCFA1E" authority="Ekins & Debitus & Erpenbeck & Hooper, 2018" authorityName="Ekins & Debitus & Erpenbeck & Hooper" authorityYear="2018" box="[503,672,1506,1528]" class="Demospongiae" family="Raspailiidae" genus="Raspailia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Axinellida" pageId="2" pageNumber="381" phylum="Porifera" rank="species" species="frondosa" status="sp. nov." subGenus="Raspaxilla">
|
||
<emphasis id="B919BB45FFF86422A981FA1D02DCFA1E" box="[503,672,1506,1528]" italics="true" pageId="2" pageNumber="381">R. (R.) frondosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22A063EFFF86422AADCFA1C0282FA11" box="[682,766,1506,1527]" pageId="2" pageNumber="381" rank="species">
|
||
<emphasis id="B919BB45FFF86422AADCFA1C0282FA11" bold="true" box="[682,766,1506,1527]" pageId="2" pageNumber="381">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, clockwise from the top left: paratype QM G335144; holotype
|
||
<emphasis id="B919BB45FFF86422A8E1F9FE0103F9F3" bold="true" box="[151,383,1536,1557]" pageId="2" pageNumber="381">MNHN-IP-2015-1402</emphasis>
|
||
(fragment QM G335124); paratype QM G318701.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37734DCFFF86424A8B1F9BD010CFC41" lastPageId="4" lastPageNumber="383" pageId="2" pageNumber="381" type="description">
|
||
<paragraph id="8BD26757FFF86422A8B1F9BD02F5F92E" blockId="2.[151,1437,1603,2024]" pageId="2" pageNumber="381">
|
||
<emphasis id="B919BB45FFF86422A8B1F9BD0125F9BA" bold="true" box="[199,345,1603,1628]" pageId="2" pageNumber="381">Description.</emphasis>
|
||
<emphasis id="B919BB45FFF86422A917F9BD0188F9BA" box="[353,500,1603,1628]" italics="true" pageId="2" pageNumber="381">Growth form</emphasis>
|
||
: A stalked foliaceous, flabellate to reniform sponge growing mainly in one plane but showing some undulations. The blades are between 55 and
|
||
<quantity id="4C95CAB2FFF86422ABFBF999039DF999" box="[909,993,1639,1664]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="381" unit="mm" value="65.0">65 mm</quantity>
|
||
high and
|
||
<quantity id="4C95CAB2FFF86422AC29F9990481F999" box="[1119,1277,1639,1664]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="8.5" metricValueMin="5.5" pageId="2" pageNumber="381" unit="mm" value="70.0" valueMax="85.0" valueMin="55.0">55 to 85 mm</quantity>
|
||
wide, with a maximum thickness of
|
||
<quantity id="4C95CAB2FFF86422A9E8F97201A3F945" box="[414,479,1676,1700]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="381" unit="mm" value="3.0">3 mm</quantity>
|
||
at the base sloping to less than
|
||
<quantity id="4C95CAB2FFF86422AB34F97203F8F945" box="[834,900,1676,1700]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="381" unit="mm" value="1.0">1 mm</quantity>
|
||
at the apex. It has a circular stalked base with a diameter of
|
||
<quantity id="4C95CAB2FFF86422A968F94E0111F921" box="[286,365,1712,1736]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="381" unit="mm" value="10.0">10 mm</quantity>
|
||
and small basal holdfast.
|
||
</paragraph>
|
||
<paragraph id="8BD26757FFF86422A8B1F92D03B0F90A" blockId="2.[151,1437,1603,2024]" box="[199,972,1747,1772]" pageId="2" pageNumber="381">
|
||
<emphasis id="B919BB45FFF86422A8B1F92D016AF90A" box="[199,278,1747,1772]" italics="true" pageId="2" pageNumber="381">Colour</emphasis>
|
||
: Pale cream to orange on deck and beige to brown in ethanol.
|
||
</paragraph>
|
||
<paragraph id="8BD26757FFF86422A8B1F9060348F8E9" blockId="2.[151,1437,1603,2024]" box="[199,820,1783,1808]" pageId="2" pageNumber="381">
|
||
<emphasis id="B919BB45FFF86422A8B1F906016BF8E9" box="[199,279,1784,1807]" italics="true" pageId="2" pageNumber="381">Texture</emphasis>
|
||
: Firm and barely compressible, but easy to tear.
|
||
</paragraph>
|
||
<paragraph id="8BD26757FFF86422A8B1F8E20288F89A" blockId="2.[151,1437,1603,2024]" pageId="2" pageNumber="381">
|
||
<emphasis id="B919BB45FFF86422A8B1F8E20160F8D2" box="[199,284,1820,1844]" italics="true" pageId="2" pageNumber="381">Surface</emphasis>
|
||
: Oscules are not visible, however on the surface the absence of parchment-like ectosome over the aquiferous channels resembles leaf veins (this is could also be due to friction or predation). Otherwise the surface is highly hispid and velvety from protruding long styles.
|
||
</paragraph>
|
||
<paragraph id="8BD26757FFF86422A8B1F879052AF825" blockId="2.[151,1437,1603,2024]" pageId="2" pageNumber="381">
|
||
<emphasis id="B919BB45FFF86422A8B1F87901DBF846" box="[199,423,1927,1952]" italics="true" pageId="2" pageNumber="381">Ectosomal skeleton</emphasis>
|
||
: Membranous, parchment-like, without a specialised ‘raspailiid’ skeleton of spiculebouquets, but with a thick palisade of subectosomal styles protruding through the ectosome up to
|
||
<quantity id="4C95CAB2FFF86422ACABF855052EF825" box="[1245,1362,1963,1988]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="2" pageNumber="381" unit="mm" value="1.25" valueMax="2.0" valueMin="0.5">0.5–2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD26757FFF86423A8B1F8310399FEA6" blockId="2.[151,1437,1603,2024]" lastBlockId="3.[151,1437,151,320]" lastPageId="3" lastPageNumber="382" pageId="2" pageNumber="381">
|
||
<emphasis id="B919BB45FFF86422A8B1F83101B3F80E" box="[199,463,1999,2024]" italics="true" pageId="2" pageNumber="381">Choanosomal skeleton</emphasis>
|
||
: With well-differentiated axial and extra-axial skeletons. The axial skeleton is compressed, lacking spicules, but with heavy interlaced spongin fibres forming a wide core up to 50% of the diameter of the skeleton. Long subectosomal styles and shorter choanosomal rhabdostyles have their bases embedded in the axial spongin skeleton, standing perpendicular to the axis forming a radial extra-axial skeleton, with the subectosomal styles protruding through the ectosome and the choanosomal rhabdostyles supporting the parchment-like ectosome on both dorsal and ventral surfaces of the sponge.
|
||
</paragraph>
|
||
<caption id="DF1237DFFFF96423A8E1FB820456FB57" httpUri="https://zenodo.org/record/1221674/files/figure.png" pageId="3" pageNumber="382" startId="3.[151,250,1148,1170]" targetBox="[152,1435,373,1126]" targetPageId="3">
|
||
<paragraph id="8BD26757FFF96423A8E1FB820456FB57" blockId="3.[151,1436,1148,1201]" pageId="3" pageNumber="382">
|
||
<emphasis id="B919BB45FFF96423A8E1FB820166FB77" bold="true" box="[151,282,1148,1170]" pageId="3" pageNumber="382">FIGURE 2.</emphasis>
|
||
Spicules of the holotype
|
||
<taxonomicName id="4C6D1CD4FFF96423AA35FB830288FB74" authority="Ekins & Debitus & Erpenbeck & Hooper, 2018" authorityName="Ekins & Debitus & Erpenbeck & Hooper" authorityYear="2018" box="[579,756,1148,1170]" class="Demospongiae" family="Raspailiidae" genus="Raspailia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Axinellida" pageId="3" pageNumber="382" phylum="Porifera" rank="species" species="frondosa" status="sp. nov." subGenus="Raspaxilla">
|
||
<emphasis id="B919BB45FFF96423AA35FB830288FB74" box="[579,756,1148,1170]" italics="true" pageId="3" pageNumber="382">R. (R.) frondosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22A063EFFF96423AB75FB820326FB77" box="[771,858,1148,1169]" pageId="3" pageNumber="382" rank="species">
|
||
<emphasis id="B919BB45FFF96423AB75FB820326FB77" bold="true" box="[771,858,1148,1169]" pageId="3" pageNumber="382">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B919BB45FFF96423AB06FB820424FB77" bold="true" box="[880,1112,1148,1169]" pageId="3" pageNumber="382">MNHN-IP-2015-1402</emphasis>
|
||
(fragment QM G335124). A subectosomal style, B terminations of subectosomal style, C–G choanosomal rhabdostyles
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1237DFFFF96423A8E1F8CE0082F865" httpUri="https://zenodo.org/record/1221676/files/figure.png" pageId="3" pageNumber="382" startId="3.[151,250,1840,1862]" targetBox="[425,1162,1250,1818]" targetPageId="3">
|
||
<paragraph id="8BD26757FFF96423A8E1F8CE0082F865" blockId="3.[151,1436,1840,1923]" pageId="3" pageNumber="382">
|
||
<emphasis id="B919BB45FFF96423A8E1F8CE0168F8A3" bold="true" box="[151,276,1840,1862]" pageId="3" pageNumber="382">FIGURE 3.</emphasis>
|
||
An undigested cross section of the holotype of
|
||
<taxonomicName id="4C6D1CD4FFF96423AB7EF8CF03D1F8A0" authority="Ekins & Debitus & Erpenbeck & Hooper, 2018" authorityName="Ekins & Debitus & Erpenbeck & Hooper" authorityYear="2018" box="[776,941,1840,1862]" class="Demospongiae" family="Raspailiidae" genus="Raspailia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Axinellida" pageId="3" pageNumber="382" phylum="Porifera" rank="species" species="frondosa" status="sp. nov." subGenus="Raspaxilla">
|
||
<emphasis id="B919BB45FFF96423AB7EF8CF03D1F8A0" box="[776,941,1840,1862]" italics="true" pageId="3" pageNumber="382">R. (R.) frondosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22A063EFFF96423ABC0F8CE0474F8A3" box="[950,1032,1840,1861]" pageId="3" pageNumber="382" rank="species">
|
||
<emphasis id="B919BB45FFF96423ABC0F8CE0474F8A3" bold="true" box="[950,1032,1840,1861]" pageId="3" pageNumber="382">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B919BB45FFF96423AC61F8CE0483F8A3" bold="true" box="[1047,1279,1840,1861]" pageId="3" pageNumber="382">MNHN-IP-2015-1402</emphasis>
|
||
(fragment QM G335124), showing the choanosomal rhabdostyles supporting the fragile ectosome and the subectosomal styles penetrating the ectosome.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD26757FFF96424A8B1F84E0194FF32" blockId="3.[151,1436,1968,2029]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="383" pageId="3" pageNumber="382">
|
||
<emphasis id="B919BB45FFF96423A8B1F84E0128F82F" box="[199,340,1968,1993]" italics="true" pageId="3" pageNumber="382">Megascleres</emphasis>
|
||
(
|
||
<tableCitation id="C6EF52ECFFF96423A910F84E01C2F82F" box="[358,446,1968,1993]" captionStart="TABLE 1" captionStartId="4.[151,239,260,283]" captionText="TABLE 1. Spicule measurements of the type specimens of Raspailia (Raspaxilla)frondosa sp.nov." pageId="3" pageNumber="382">Table 1</tableCitation>
|
||
). There are only two categories of megascleres. Choanosomal rhabdostyles are very smooth, moderately thick, with a smooth non-tylote base, varying from slight to well-developed rhabds (97–469 x 6.2–31.6 µm). Subectosomal styles are rare (<0.2%), smooth, long, thick and either straight or slightly curved (1010–4640 x 10.2–29.1 µm).
|
||
</paragraph>
|
||
<caption id="DF1237DFFFFE6424A8E1FEFA04C6FEFD" box="[151,1210,259,283]" pageId="4" pageNumber="383">
|
||
<paragraph id="8BD26757FFFE6424A8E1FEFA04C6FEFD" blockId="4.[151,1210,259,283]" box="[151,1210,259,283]" pageId="4" pageNumber="383">
|
||
<emphasis id="B919BB45FFFE6424A8E1FEFA0176FEFD" bold="true" box="[151,266,260,283]" pageId="4" pageNumber="383">TABLE 1.</emphasis>
|
||
Spicule measurements of the type specimens of
|
||
<taxonomicName id="4C6D1CD4FFFE6424AB67FEFD041AFEFD" authority="Ekins & Debitus & Erpenbeck & Hooper, 2018" authorityName="Ekins & Debitus & Erpenbeck & Hooper" authorityYear="2018" box="[785,1126,259,283]" class="Demospongiae" family="Raspailiidae" genus="Raspailia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Axinellida" pageId="4" pageNumber="383" phylum="Porifera" rank="species" species="frondosa" status="sp. nov." subGenus="Raspaxilla">
|
||
<emphasis id="B919BB45FFFE6424AB67FEFD041AFEFD" box="[785,1126,259,283]" italics="true" pageId="4" pageNumber="383">Raspailia (Raspaxilla) frondosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22A063EFFFE6424AC1BFEFA04C6FEFD" box="[1133,1210,260,283]" pageId="4" pageNumber="383" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD26757FFFE6424A8E9FEC80535FD66" pageId="4" pageNumber="383">
|
||
<table id="F96D95F7FFFE9BDFA8E9FEC805FEFD58" box="[159,1410,310,702]" gridcols="5" gridrows="4" pageId="4" pageNumber="383">
|
||
<tr id="355D6515FFFE9BDFA8E9FEC805FEFE8D" box="[159,1410,310,363]" gridrow="0" pageId="4" pageNumber="383">
|
||
<th id="768C0C69FFFE9BDFA8E9FEC801FDFE8D" box="[159,385,310,363]" gridcol="0" gridrow="0" pageId="4" pageNumber="383">Specimen</th>
|
||
<th id="768C0C69FFFE9BDFA9E2FEC802EFFE8D" box="[404,659,310,363]" gridcol="1" gridrow="0" pageId="4" pageNumber="383">Choanosomal rhabdostyles length (µm)</th>
|
||
<th id="768C0C69FFFE9BDFAAD1FEC803DCFE8D" box="[679,928,310,363]" gridcol="2" gridrow="0" pageId="4" pageNumber="383">Choanosomal rhabdostyles width (µm)</th>
|
||
<th id="768C0C69FFFE9BDFABC4FEC804FCFE8D" box="[946,1152,310,363]" gridcol="3" gridrow="0" pageId="4" pageNumber="383">Subectosomal styles length (µm)</th>
|
||
<th id="768C0C69FFFE9BDFACC5FEC805FEFE8D" box="[1203,1410,310,363]" gridcol="4" gridrow="0" pageId="4" pageNumber="383">Subectosomal styles width (µm)</th>
|
||
</tr>
|
||
<tr id="355D6515FFFE9BDFA8E9FE8005FEFE16" box="[159,1410,382,496]" gridrow="1" pageId="4" pageNumber="383">
|
||
<th id="768C0C69FFFE9BDFA8E9FE8001FDFE16" box="[159,385,382,496]" gridcol="0" gridrow="1" pageId="4" pageNumber="383">Holotype MNHN-IP-2015-1402 QM G335124 (n=143, 36)</th>
|
||
<td id="768C0C69FFFE9BDFA9E2FE8002EFFE16" box="[404,659,382,496]" gridcol="1" gridrow="1" pageId="4" pageNumber="383">143-366-469</td>
|
||
<td id="768C0C69FFFE9BDFAAD1FE8003DCFE16" box="[679,928,382,496]" gridcol="2" gridrow="1" pageId="4" pageNumber="383">6.2-21.4-31.6</td>
|
||
<td id="768C0C69FFFE9BDFABC4FE8004FCFE16" box="[946,1152,382,496]" gridcol="3" gridrow="1" pageId="4" pageNumber="383">1170-2094-4640</td>
|
||
<td id="768C0C69FFFE9BDFACC5FE8005FEFE16" box="[1203,1410,382,496]" gridcol="4" gridrow="1" pageId="4" pageNumber="383">12-21.7-29.1</td>
|
||
</tr>
|
||
<tr id="355D6515FFFE9BDFA8E9FDFA05FEFDB1" box="[159,1410,516,599]" gridrow="2" pageId="4" pageNumber="383">
|
||
<th id="768C0C69FFFE9BDFA8E9FDFA01FDFDB1" box="[159,385,516,599]" gridcol="0" gridrow="2" pageId="4" pageNumber="383">Paratype QM G335144 (n=97, 31)</th>
|
||
<td id="768C0C69FFFE9BDFA9E2FDFA02EFFDB1" box="[404,659,516,599]" gridcol="1" gridrow="2" pageId="4" pageNumber="383">97-326-422</td>
|
||
<td id="768C0C69FFFE9BDFAAD1FDFA03DCFDB1" box="[679,928,516,599]" gridcol="2" gridrow="2" pageId="4" pageNumber="383">6.8-20.7-28.9</td>
|
||
<td id="768C0C69FFFE9BDFABC4FDFA04FCFDB1" box="[946,1152,516,599]" gridcol="3" gridrow="2" pageId="4" pageNumber="383">1350-2172-2710</td>
|
||
<td id="768C0C69FFFE9BDFACC5FDFA05FEFDB1" box="[1203,1410,516,599]" gridcol="4" gridrow="2" pageId="4" pageNumber="383">10.2-15.9-21.5</td>
|
||
</tr>
|
||
<tr id="355D6515FFFE9BDFA8E9FD9405FEFD58" box="[159,1410,618,702]" gridrow="3" pageId="4" pageNumber="383">
|
||
<th id="768C0C69FFFE9BDFA8E9FD9401FDFD58" box="[159,385,618,702]" gridcol="0" gridrow="3" pageId="4" pageNumber="383">Paratype QM G318701 (n=93, 33)</th>
|
||
<td id="768C0C69FFFE9BDFA9E2FD9402EFFD58" box="[404,659,618,702]" gridcol="1" gridrow="3" pageId="4" pageNumber="383">118-334-444</td>
|
||
<td id="768C0C69FFFE9BDFAAD1FD9403DCFD58" box="[679,928,618,702]" gridcol="2" gridrow="3" pageId="4" pageNumber="383">10.9-19.2-29.2</td>
|
||
<td id="768C0C69FFFE9BDFABC4FD9404FCFD58" box="[946,1152,618,702]" gridcol="3" gridrow="3" pageId="4" pageNumber="383">1010-1749-2350</td>
|
||
<td id="768C0C69FFFE9BDFACC5FD9405FEFD58" box="[1203,1410,618,702]" gridcol="4" gridrow="3" pageId="4" pageNumber="383">10.5-17.9-22.3</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BD26757FFFE6424A8B1FD0001C0FCF1" blockId="4.[151,1437,766,935]" box="[199,444,766,791]" pageId="4" pageNumber="383">
|
||
<emphasis id="B919BB45FFFE6424A8B1FD00012BFCF1" box="[199,343,766,791]" italics="true" pageId="4" pageNumber="383">Microscleres</emphasis>
|
||
: Absent.
|
||
</paragraph>
|
||
<paragraph id="8BD26757FFFE6424A8B1FCDC010CFC41" blockId="4.[151,1437,766,935]" pageId="4" pageNumber="383">
|
||
<emphasis id="B919BB45FFFE6424A8B1FCDC0109FCDD" box="[199,373,802,827]" italics="true" pageId="4" pageNumber="383">Molecular data</emphasis>
|
||
. DNA barcodes for 28S and CO1 could not be retrieved for the
|
||
<typeStatus id="54D6D9F5FFFE6424AC38FCDC04CCFCDD" box="[1102,1200,802,827]" pageId="4" pageNumber="383" type="holotype">holotype</typeStatus>
|
||
or for the
|
||
<typeStatus id="54D6D9F5FFFE6424AD5CFCDD05E5FCDD" box="[1322,1433,803,827]" pageId="4" pageNumber="383" type="paratype">paratypes</typeStatus>
|
||
. All samples underwent two different DNA extractions (see Methods section) and multiple amplification attempts with different deviations from the PCR protocol, but amplification success could not be achieved under the described methods.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |