treatments-xml/data/03/B5/87/03B587EFFFA7FF937CD609CF0173F81A.xml

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<mods:title id="519E850D825DDC956FBA9F88F4835C3E">Combretum xylocarpum (Combretaceae), a new liana species from China</mods:title>
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<mods:namePart id="3C6D09771DACAF96BF27DEBD2C0881D8">Liang, Dan</mods:namePart>
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<mods:affiliation id="B10E6B13B723FC5267056E25B7A2B96A">Key Laboratory of Plant Resources Conservation and Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China. &amp; University of Chinese Academy of Sciences, Beijing 100049, China. &amp; Nanning Botanical Garden, Nanning 530021, China. &amp; jordanliangdan @ 163. com; https: // orcid. org / 0000 - 0002 - 9709 - 0510</mods:affiliation>
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<mods:namePart id="6514C09DA7F1DA9DEECE72DEB1357BAB">Yuan, Lang-Xing</mods:namePart>
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<mods:affiliation id="597B488034FBA27F2BC3292F244C355A">Tropical Crops Genetic Resources Institute, Chinese Academy of Tropical Agricultural Sciences, Haikou 571101, China. &amp; yuanlangxing @ 163. com; https: // orcid. org / 0000 - 0003 - 2102 - 9075</mods:affiliation>
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<mods:namePart id="E111C74D48EF2C70B25DD6C5CF05E1F5">Wang, Rui-Jiang</mods:namePart>
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<mods:affiliation id="5F2E036B6CE0087E81A57DF0BC6C4123">Key Laboratory of Plant Resources Conservation and Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China. &amp; University of Chinese Academy of Sciences, Beijing 100049, China. &amp; wangrj @ scbg. ac. cn; https: // orcid. org / 0000 - 0002 - 4985 - 8281</mods:affiliation>
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<paragraph id="8BA336F9FFA7FF907CD609CF03A1FD30" blockId="1.[136,957,668,695]" box="[136,957,668,695]" pageId="1" pageNumber="289">
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<taxonomicName id="4C1C4D7AFFA7FF907CD609CF02A5FD31" ID-CoL="9VHTY" authority="Dan Liang &amp; R.J. Wang" authorityName="Dan Liang &amp; R. J. Wang" authorityYear="2022" box="[136,697,668,695]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="289" phylum="Tracheophyta" rank="species" species="xylocarpum" status="sp. nov.">
<emphasis id="B968EAEBFFA7FF907CD609CF01BDFD31" bold="true" box="[136,417,668,695]" italics="true" pageId="1" pageNumber="289">Combretum xylocarpum</emphasis>
Dan Liang &amp; R.J. Wang
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, 
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<taxonomicNameLabel id="A25B5790FFA7FF907E9D09CE0304FD30" box="[707,792,669,694]" pageId="1" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13272A7CFFA7FF907F7809CF038DFD30" box="[806,913,668,694]" captionStart="FIGURE 1" captionStartId="3.[136,234,1871,1893]" captionTargetBox="[202,1386,200,1847]" captionTargetId="figure-33@3.[183,1403,190,1847]" captionTargetPageId="3" captionText="FIGURE 1. Combretum xylocarpum Dan Liang &amp; R.J. Wang, sp. nov. A: Habit; B: Adaxial (left) and abaxial (right) side of leaf lamina; C: Dome-shaped sessile glands at adaxial side of leaf; D: Orificed pits at abaxial side of leaf; E: Adaxial side of the midrib, showing the pubescence and scales; F: Spike and flower arrangement (by Lang-Xing Yuan); G: Longitudinally dissected flower, showing the stamens, adnation of style, and a ring of hairs at throat; H: Dissected proximal hypanthium, showing the four pendulous ovules; I: Infructescence, with young fruits; J: Transversely dissected young fruit, showing the 4 rounded ridges; K: Mature fruit (leaf) and seeds (right, by Lei Zhao)." figureDoi="http://doi.org/10.5281/zenodo.7095422" httpUri="https://zenodo.org/record/7095422/files/figure.png" pageId="1" pageNumber="289">Figures 1</figureCitation>
, 
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).
</heading>
</paragraph>
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<subSubSection id="C3066572FFA7FF907CD609BB0514FCA4" pageId="1" pageNumber="289" type="materials_examined">
<paragraph id="8BA336F9FFA7FF907CD609BB0514FCA4" blockId="1.[136,1452,744,946]" pageId="1" pageNumber="289">
<emphasis id="B968EAEBFFA7FF907CD609BB00A6FD7B" bold="true" box="[136,186,744,765]" pageId="1" pageNumber="289">Type</emphasis>
:— 
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<collectingCountry id="F30B7669FFA7FF907C8909BB0134FD7B" box="[215,296,744,765]" name="China" pageId="1" pageNumber="289">CHINA</collectingCountry>
. 
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: 
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, 
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, 
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the 
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, roadsides, 
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, 
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, elevation ca. 
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<elevation id="0031D1CAFFA7FF9079DE09BB05B4FD7B" box="[1408,1448,744,765]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="289" unit="m" value="4.0">4 m</elevation>
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, fruiting, 
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<collectingDate id="EFE6E9D1FFA7FF907D4F085F0183FCA7" box="[273,415,780,802]" pageId="1" pageNumber="289" value="2018-08-05">5 August 2018</collectingDate>
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, 
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<collectorName id="26E9532FFFA7FF907DF7085F020DFCA7" box="[425,529,780,801]" pageId="1" pageNumber="289">Dan Liang</collectorName>
&amp; 
<collectorName id="26E9532FFFA7FF907E71085F02F9FCA7" box="[559,741,780,801]" pageId="1" pageNumber="289">Guo-Bin Jiang WP</collectorName>
0569
</emphasis>
(
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!;
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isotype 
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!)
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.
</paragraph>
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<subSubSection id="C3066572FFA7FF907CD6087C0106FC37" pageId="1" pageNumber="289" type="diagnosis">
<paragraph id="8BA336F9FFA7FF907CD6087C0106FC37" blockId="1.[136,1452,744,946]" pageId="1" pageNumber="289">
<emphasis id="B968EAEBFFA7FF907CD6087C00F7FCC3" bold="true" box="[136,235,815,837]" pageId="1" pageNumber="289">Diagnosis</emphasis>
:—This species is similar to 
<taxonomicName id="4C1C4D7AFFA7FF907DA00863028DFCC3" authorityName="Roxburgh" authorityYear="1832" box="[510,657,816,837]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="289" phylum="Tracheophyta" rank="species" species="acuminatum">
<emphasis id="B968EAEBFFA7FF907DA00863028DFCC3" box="[510,657,816,837]" italics="true" pageId="1" pageNumber="289">C. acuminatum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1C4D7AFFA7FF907E9E08630470FCC3" authority="Roxburg ex G. Don (1827: 417)" authorityName="G. Don" authorityPageNumber="417" authorityYear="1827" box="[704,1132,816,838]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="289" phylum="Tracheophyta" rank="species" species="chinense">
<emphasis id="B968EAEBFFA7FF907E9E0863032CFCC3" box="[704,816,816,837]" italics="true" pageId="1" pageNumber="289">C. chinense</emphasis>
Roxburg ex G. 
<bibRefCitation id="EF8D4B08FFA7FF907F9508630470FCC3" author="Don, G." box="[971,1132,816,837]" pageId="1" pageNumber="289" pagination="412 - 441" refId="ref2970" refString="Don, G. (1827) A review of the Genus Combretum. Transactions of the Linnean Society of London 15 (2): 412 - 441. https: // doi. org / 10.1111 / j. 1095 - 8339.1826. tb 00124. x" type="journal article" year="1827">Don (1827: 417)</bibRefCitation>
</taxonomicName>
in woody scandent habit, elliptic leaves, axillary or terminal spikes, and infundibuliform calyx shape, but differs from the former by its fewer secondary veins and flowers with pale whitish petals and from the latter by its longer spikes and woody and small ellipsoidal fruit with 4 rounded ridges (
<tableCitation id="C69E0342FFA7FF907C9A08CF0117FC37" box="[196,267,924,946]" captionStart="TABLE 1" captionStartId="1.[137,234,992,1018]" captionTargetPageId="1" captionText="TABLE 1. Morphological comparison between Combretum xylocarpum and its morphological similar species." httpUri="http://table.plazi.org/id/DF636671FFA7FF907CD708B3057AFC7C" pageId="1" pageNumber="289" tableUuid="DF636671FFA7FF907CD708B3057AFC7C">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<caption id="DF636671FFA7FF907CD708B3057AFC7C" ID-Table-UUID="DF636671FFA7FF907CD708B3057AFC7C" box="[137,1382,992,1019]" httpUri="http://table.plazi.org/id/DF636671FFA7FF907CD708B3057AFC7C" pageId="1" pageNumber="289" startId="1.[137,234,992,1018]" targetBox="[144,1440,1055,1998]" targetIsTable="true" targetPageId="1">
<paragraph id="8BA336F9FFA7FF907CD708B3057AFC7C" blockId="1.[137,1382,992,1019]" box="[137,1382,992,1019]" pageId="1" pageNumber="289">
<emphasis id="B968EAEBFFA7FF907CD708B300E3FC7C" bold="true" box="[137,255,992,1018]" pageId="1" pageNumber="289">TABLE 1</emphasis>
. Morphological comparison between 
<taxonomicName id="4C1C4D7AFFA7FF907EF608B303AAFC7C" authorityName="Dan Liang &amp; R. J. Wang" authorityYear="2022" box="[680,950,992,1018]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="289" phylum="Tracheophyta" rank="species" species="xylocarpum">
<emphasis id="B968EAEBFFA7FF907EF608B303AAFC7C" box="[680,950,992,1018]" italics="true" pageId="1" pageNumber="289">Combretum xylocarpum</emphasis>
</taxonomicName>
and its morphological similar species.
</paragraph>
</caption>
<paragraph id="8BA336F9FFA7FF907CCE0F4C04EBF82C" pageId="1" pageNumber="289">
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<th id="76FD5DC7FFA7006E7CCE0F4C0132FBB3" box="[144,302,1055,1077]" gridcol="0" gridrow="0" pageId="1" pageNumber="289">Characters</th>
<th id="76FD5DC7FFA7006E7D010F4C02BAFBB3" box="[351,678,1055,1077]" gridcol="1" gridrow="0" pageId="1" pageNumber="289">
<taxonomicName id="4C1C4D7AFFA7FF907D010F4C025BFBB2" authorityName="Dan Liang &amp; R. J. Wang" authorityYear="2022" box="[351,583,1055,1076]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="289" phylum="Tracheophyta" rank="species" species="xylocarpum">
<emphasis id="B968EAEBFFA7FF907D010F4C025BFBB2" box="[351,583,1055,1076]" italics="true" pageId="1" pageNumber="289">Combretum xylocarpum</emphasis>
</taxonomicName>
</th>
<th id="76FD5DC7FFA7006E7EE20F4C043BFBB3" box="[700,1063,1055,1077]" gridcol="2" gridrow="0" pageId="1" pageNumber="289">
<taxonomicName id="4C1C4D7AFFA7FF907EE20F4C0353FBB2" authorityName="Roxburgh" authorityYear="1832" box="[700,847,1055,1076]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="289" phylum="Tracheophyta" rank="species" species="acuminatum">
<emphasis id="B968EAEBFFA7FF907EE20F4C0353FBB2" box="[700,847,1055,1076]" italics="true" pageId="1" pageNumber="289">C. acuminatum</emphasis>
</taxonomicName>
</th>
<th id="76FD5DC7FFA7006E78610F4C05BCFBB3" box="[1087,1440,1055,1077]" gridcol="3" gridrow="0" pageId="1" pageNumber="289">
<emphasis id="B968EAEBFFA7FF9078610F4C04B3FBB2" box="[1087,1199,1055,1076]" italics="true" pageId="1" pageNumber="289">C. chinense</emphasis>
</th>
</tr>
<tr id="352C34BBFFA7006E7CCE0F3805BCFB07" box="[144,1440,1131,1153]" gridrow="1" pageId="1" pageNumber="289">
<th id="76FD5DC7FFA7006E7CCE0F380132FB07" box="[144,302,1131,1153]" gridcol="0" gridrow="1" pageId="1" pageNumber="289">Habit</th>
<td id="76FD5DC7FFA7006E7D010F3802BAFB07" box="[351,678,1131,1153]" gridcol="1" gridrow="1" pageId="1" pageNumber="289">Liana woody or scandent shrub</td>
<td id="76FD5DC7FFA7006E7EE20F38043BFB07" box="[700,1063,1131,1153]" gridcol="2" gridrow="1" pageId="1" pageNumber="289">Climbing shrub</td>
<td id="76FD5DC7FFA7006E78610F3805BCFB07" box="[1087,1440,1131,1153]" gridcol="3" gridrow="1" pageId="1" pageNumber="289">Scandent shrub</td>
</tr>
<tr id="352C34BBFFA7006E7CCE0FE605BCFB4D" box="[144,1440,1205,1227]" gridrow="2" pageId="1" pageNumber="289">
<th id="76FD5DC7FFA7006E7CCE0FE60132FB4D" box="[144,302,1205,1227]" gridcol="0" gridrow="2" pageId="1" pageNumber="289">Branchlet</th>
<td id="76FD5DC7FFA7006E7D010FE602BAFB4D" box="[351,678,1205,1227]" gridcol="1" gridrow="2" pageId="1" pageNumber="289">Cylindrical</td>
<td id="76FD5DC7FFA7006E7EE20FE6043BFB4D" box="[700,1063,1205,1227]" gridcol="2" gridrow="2" pageId="1" pageNumber="289">Obtusely quadrangular</td>
<td id="76FD5DC7FFA7006E78610FE605BCFB4D" box="[1087,1440,1205,1227]" gridcol="3" gridrow="2" pageId="1" pageNumber="289">Obtusely quadrangular</td>
</tr>
<tr id="352C34BBFFA7006E7CCE0E5305BCFA07" box="[144,1440,1280,1409]" gridrow="3" pageId="1" pageNumber="289">
<th id="76FD5DC7FFA7006E7CCE0E530132FA07" box="[144,302,1280,1409]" gridcol="0" gridrow="3" pageId="1" pageNumber="289">Leaf shape</th>
<td id="76FD5DC7FFA7006E7D010E5302BAFA07" box="[351,678,1280,1409]" gridcol="1" gridrow="3" pageId="1" pageNumber="289">Oblanceolate to elliptic; apex acuminate, obtuse or slightly emarginate, base subrounded to cuneate</td>
<td id="76FD5DC7FFA7006E7EE20E53043BFA07" box="[700,1063,1280,1409]" gridcol="2" gridrow="3" pageId="1" pageNumber="289">Elliptic to oblong elliptic; apex acuminate, base cuneate, rounded or obtuse</td>
<td id="76FD5DC7FFA7006E78610E5305BCFA07" box="[1087,1440,1280,1409]" gridcol="3" gridrow="3" pageId="1" pageNumber="289">Elliptic; apex rounded or obtuse, base cuneate</td>
</tr>
<tr id="352C34BBFFA7006E7CCE0EE605BCFA4A" box="[144,1440,1461,1484]" gridrow="4" pageId="1" pageNumber="289">
<th id="76FD5DC7FFA7006E7CCE0EE60132FA4A" box="[144,302,1461,1484]" gridcol="0" gridrow="4" pageId="1" pageNumber="289">Secondary veins</th>
<td id="76FD5DC7FFA7006E7D010EE602BAFA4A" box="[351,678,1461,1484]" gridcol="1" gridrow="4" pageId="1" pageNumber="289">4–5(–7) on each side</td>
<td id="76FD5DC7FFA7006E7EE20EE6043BFA4A" box="[700,1063,1461,1484]" gridcol="2" gridrow="4" pageId="1" pageNumber="289">9–10 on each side</td>
<td id="76FD5DC7FFA7006E78610EE605BCFA4A" box="[1087,1440,1461,1484]" gridcol="3" gridrow="4" pageId="1" pageNumber="289">5–7(–9) on each side</td>
</tr>
<tr id="352C34BBFFA7006E7CCE0D5305BCF990" box="[144,1440,1536,1558]" gridrow="5" pageId="1" pageNumber="289">
<th id="76FD5DC7FFA7006E7CCE0D530132F990" box="[144,302,1536,1558]" gridcol="0" gridrow="5" pageId="1" pageNumber="289">Spike</th>
<td id="76FD5DC7FFA7006E7D010D5302BAF990" box="[351,678,1536,1558]" gridcol="1" gridrow="5" pageId="1" pageNumber="289">Terminal or axillary, 6–8 cm long</td>
<td id="76FD5DC7FFA7006E7EE20D53043BF990" box="[700,1063,1536,1558]" gridcol="2" gridrow="5" pageId="1" pageNumber="289">Axillary and terminal, to 10 cm long</td>
<td id="76FD5DC7FFA7006E78610D5305BCF990" box="[1087,1440,1536,1558]" gridcol="3" gridrow="5" pageId="1" pageNumber="289">Axillary, half length of the leaves</td>
</tr>
<tr id="352C34BBFFA7006E7CCE0D1905BCF9E6" box="[144,1440,1610,1632]" gridrow="6" pageId="1" pageNumber="289">
<th id="76FD5DC7FFA7006E7CCE0D190132F9E6" box="[144,302,1610,1632]" gridcol="0" gridrow="6" pageId="1" pageNumber="289">Calyx lobes</th>
<td id="76FD5DC7FFA7006E7D010D1902BAF9E6" box="[351,678,1610,1632]" gridcol="1" gridrow="6" pageId="1" pageNumber="289">4, triangular, ca. 2.2 mm long</td>
<td id="76FD5DC7FFA7006E7EE20D19043BF9E6" box="[700,1063,1610,1632]" gridcol="2" gridrow="6" pageId="1" pageNumber="289">4, ovate to triangular, ca. 1 mm long</td>
<td id="76FD5DC7FFA7006E78610D1905BCF9E6" box="[1087,1440,1610,1632]" gridcol="3" gridrow="6" pageId="1" pageNumber="289">4, triangular, up to 1 mm long</td>
</tr>
<tr id="352C34BBFFA7006E7CCE0DC705BCF948" box="[144,1440,1684,1742]" gridrow="7" pageId="1" pageNumber="289">
<th id="76FD5DC7FFA7006E7CCE0DC70132F948" box="[144,302,1684,1742]" gridcol="0" gridrow="7" pageId="1" pageNumber="289">Flower petals</th>
<td id="76FD5DC7FFA7006E7D010DC702BAF948" box="[351,678,1684,1742]" gridcol="1" gridrow="7" pageId="1" pageNumber="289">4, obovate or spatulate, 3–3.5 mm long, pale white to pale yellow</td>
<td id="76FD5DC7FFA7006E7EE20DC7043BF948" box="[700,1063,1684,1742]" gridcol="2" gridrow="7" pageId="1" pageNumber="289">4, subrounded to narrowly spathulate, 1.5–2 mm long, yellow</td>
<td id="76FD5DC7FFA7006E78610DC705BCF948" box="[1087,1440,1684,1742]" gridcol="3" gridrow="7" pageId="1" pageNumber="289">Obovate, twice longer than calyx lobes, pale white to cream</td>
</tr>
<tr id="352C34BBFFA7006E7CCE0C5105BCF89E" box="[144,1440,1794,1816]" gridrow="8" pageId="1" pageNumber="289">
<th id="76FD5DC7FFA7006E7CCE0C510132F89E" box="[144,302,1794,1816]" gridcol="0" gridrow="8" pageId="1" pageNumber="289">Stamen</th>
<td id="76FD5DC7FFA7006E7D010C5102BAF89E" box="[351,678,1794,1816]" gridcol="1" gridrow="8" pageId="1" pageNumber="289">8, exserted, ca. 9 mm long</td>
<td id="76FD5DC7FFA7006E7EE20C51043BF89E" box="[700,1063,1794,1816]" gridcol="2" gridrow="8" pageId="1" pageNumber="289">8, exserted, 5–7 mm long</td>
<td id="76FD5DC7FFA7006E78610C5105BCF89E" box="[1087,1440,1794,1816]" gridcol="3" gridrow="8" pageId="1" pageNumber="289">8, exserted, a little longer than petals</td>
</tr>
<tr id="352C34BBFFA7006E7CCE0C1F05BCF848" box="[144,1440,1868,1998]" gridrow="9" pageId="1" pageNumber="289">
<th id="76FD5DC7FFA7006E7CCE0C1F0132F848" box="[144,302,1868,1998]" gridcol="0" gridrow="9" pageId="1" pageNumber="289">Fruit</th>
<td id="76FD5DC7FFA7006E7D010C1F02BAF848" box="[351,678,1868,1998]" gridcol="1" gridrow="9" pageId="1" pageNumber="289">ca. 1.5×0.7 cm, ellipsoidal, with 4 longitudinal and rounded ridges</td>
<td id="76FD5DC7FFA7006E7EE20C1F043BF848" box="[700,1063,1868,1998]" gridcol="2" gridrow="9" pageId="1" pageNumber="289">3–6.5×1–2 cm, fusiform, oblongellipsoidal to ellipsoidal, with 4 longitudinal and rounded to sharp ridges</td>
<td id="76FD5DC7FFA7006E78610C1F05BCF848" box="[1087,1440,1868,1998]" gridcol="3" gridrow="9" pageId="1" pageNumber="289">ca. 4×3 cm, subglobose or ellipsoid, with 4 broad wings</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3066572FFA4FF937CD60BCE01C7FC0E" pageId="2" pageNumber="290" type="description">
<paragraph id="8BA336F9FFA4FF937CD60BCE010EFF31" blockId="2.[136,274,157,183]" box="[136,274,157,183]" pageId="2" pageNumber="290">
<heading id="D0EB8195FFA4FF937CD60BCE010EFF31" bold="true" box="[136,274,157,183]" fontSize="11" level="1" pageId="2" pageNumber="290" reason="1">
<emphasis id="B968EAEBFFA4FF937CD60BCE010EFF31" bold="true" box="[136,274,157,183]" pageId="2" pageNumber="290">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA336F9FFA4FF937CD60BB501C7FC0E" blockId="2.[136,1452,230,1948]" pageId="2" pageNumber="290">
Woody liana or large scandent shrub. 
<emphasis id="B968EAEBFFA4FF937E740BB502F5FE86" bold="true" box="[554,745,230,256]" pageId="2" pageNumber="290">Young branches</emphasis>
densely pubescent and lepidote, ferruginous in buds, becoming glabrous with developing. 
<emphasis id="B968EAEBFFA4FF937DEC0A590218FEA2" bold="true" box="[434,516,266,292]" pageId="2" pageNumber="290">Leaves</emphasis>
simple, opposite or more or less alternate, papery to subcoriaceous, oblanceolate to elliptic, (3.5–)5.8–9.6 × (1.9–) 
<quantity id="4CE49B1CFFA4FF937D860A7D0246FECE" box="[472,602,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="4.8" metricValueMin="2.5" pageId="2" pageNumber="290" unit="cm" value="3.65" valueMax="4.8" valueMin="2.5">2.5–4.8 cm</quantity>
, apex acuminate, obtuse or slightly emarginate; base subrounded to cuneate; glabrous and with dome-shaped sessile glands adaxially, glabrous to sparsely lepidote and with orificed pits abaxially; midrib prominent at base and then depressed toward apex, and glabrous or sparsely pubescent adaxially, prominent, glabrous or sparsely pubescent and lepidote abaxially; secondary veins brochidodromous, usually 4–5(–7) on each side; tertiary veins dichotomizing; margin entire. 
<emphasis id="B968EAEBFFA4FF937EE50AEE030BFE51" bold="true" box="[699,791,445,471]" pageId="2" pageNumber="290">Petioles</emphasis>
0.3–1.0 cm long, puberulous and scaly at surface. 
<emphasis id="B968EAEBFFA4FF9379000AEE05B0FE51" bold="true" box="[1374,1452,445,471]" pageId="2" pageNumber="290">Spikes</emphasis>
terminal and axillary, cylindrical, 6.0–8.0 cm long; peduncles up to 3.0 cm long, rusty pubescent and scaly, brownish. 
<emphasis id="B968EAEBFFA4FF937CD6095500FBFDA6" bold="true" box="[136,231,518,544]" pageId="2" pageNumber="290">Flowers</emphasis>
bisexual, numerous, sessile, white or yellowish-white, ca. 1.0 cm long, actinomorphic. 
<emphasis id="B968EAEBFFA4FF93789A09560540FD99" bold="true" box="[1220,1372,517,543]" pageId="2" pageNumber="290">Hypanthium</emphasis>
in two distinct parts, the proximal part tubular in 3.0– 
<quantity id="4CE49B1CFFA4FF937EC9097902F1FDC2" box="[663,749,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="290" unit="mm" value="3.5">3.5 mm</quantity>
long, the distal one infundibuliform in 2.5–3.0 mm long, with a ring of hairs directed upwards at the mouth, slightly pubescent and densely ferruginous scaly abaxially. 
<emphasis id="B968EAEBFFA4FF93793B091E00D9FD0A" bold="true" pageId="2" pageNumber="290">Calyx lobes</emphasis>
4, triangular, ca. 
<quantity id="4CE49B1CFFA4FF937DD6092101C2FD0A" box="[392,478,626,652]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="290" unit="mm" value="2.2">2.2 mm</quantity>
long, sparsely ferruginous scaly abaxially. 
<emphasis id="B968EAEBFFA4FF937F9C09210416FD0A" bold="true" box="[962,1034,626,652]" pageId="2" pageNumber="290">Petals</emphasis>
4, conspicuous, obovate to spatulate, 3.0–3.5 × 
<quantity id="4CE49B1CFFA4FF937CA409C5019EFD36" box="[250,386,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="2" pageNumber="290" unit="mm" value="1.85" valueMax="2.2" valueMin="1.5">1.5–2.2 mm</quantity>
, acute at apex, cuneate at base, pale whitish. 
<emphasis id="B968EAEBFFA4FF937F2409C603C3FD29" bold="true" box="[890,991,661,687]" pageId="2" pageNumber="290">Stamens</emphasis>
8, 2-seriate, alternately antesepalous and alternisepalous, exserted; filaments ca. 8.0 mm long, incurved initially and then exceeding the petals; anthers oblong, ca. 0.8 × 
<quantity id="4CE49B1CFFA4FF937CA9098D014DFD7E" box="[247,337,734,761]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="290" unit="mm" value="0.5">0.5 mm</quantity>
, biloculed, dorsifixed, versatile. 
<emphasis id="B968EAEBFFA4FF937E97098E0318FD71" bold="true" box="[713,772,733,759]" pageId="2" pageNumber="290">Style</emphasis>
simple, filiform, 6.0–8.0 mm long, for ca. 2/3 free and ca. 1/3 attached to the wall of proximal hypanthium, exserted, glabrous. 
<emphasis id="B968EAEBFFA4FF937FC8085203F6FC9D" bold="true" box="[918,1002,769,795]" pageId="2" pageNumber="290">Stigma</emphasis>
punctiform. 
<emphasis id="B968EAEBFFA4FF937820085204D6FC9D" bold="true" box="[1150,1226,769,795]" pageId="2" pageNumber="290">Ovary</emphasis>
inferior, unilocular, with 4 pendulous, anatropous ovules. 
<emphasis id="B968EAEBFFA4FF937E6808760268FCB9" bold="true" box="[566,628,805,831]" pageId="2" pageNumber="290">Fruit</emphasis>
a drupe, shortly stipitate, ellipsoidal, ca. 1.5 × 
<quantity id="4CE49B1CFFA4FF9378D3087504C1FCC6" box="[1165,1245,806,832]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="290" unit="cm" value="0.7">0.7 cm</quantity>
, brownish yellow, muriculate at surface, longitudinally 4 rounded ridges, woody. 
<emphasis id="B968EAEBFFA4FF937F17081A0390FCE5" bold="true" box="[841,908,841,867]" pageId="2" pageNumber="290">Seeds</emphasis>
ca. 6× 
<quantity id="4CE49B1CFFA4FF937F890819042CFCE2" box="[983,1072,842,869]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="290" unit="mm" value="2.5">2.5 mm</quantity>
, ellipsoidal, always single, with 4 longitudinally rounded ridges.
</paragraph>
</subSubSection>
<subSubSection id="C3066572FFA4FF937CE308C1042AFC2A" box="[189,1078,914,940]" pageId="2" pageNumber="290" type="biology_ecology">
<paragraph id="8BA336F9FFA4FF937CE308C1042AFC2A" blockId="2.[136,1452,230,1948]" box="[189,1078,914,940]" pageId="2" pageNumber="290">
<emphasis id="B968EAEBFFA4FF937CE308C10125FC2A" bold="true" box="[189,313,914,940]" pageId="2" pageNumber="290">Phenology</emphasis>
:—Flowering from February to May; Fruiting from June to October.
</paragraph>
</subSubSection>
<subSubSection id="C3066572FFA4FF937CE308E50478FC56" box="[189,1124,950,976]" pageId="2" pageNumber="290" type="etymology">
<paragraph id="8BA336F9FFA4FF937CE308E50478FC56" blockId="2.[136,1452,230,1948]" box="[189,1124,950,976]" pageId="2" pageNumber="290">
<emphasis id="B968EAEBFFA4FF937CE308E50122FC56" bold="true" box="[189,318,950,976]" pageId="2" pageNumber="290">Etymology</emphasis>
:—The species epithet refers to the woody fruit wall of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C3066572FFA4FF937CE3088A027EFBE6" pageId="2" pageNumber="290" type="distribution">
<paragraph id="8BA336F9FFA4FF937CE3088A027EFBE6" blockId="2.[136,1452,230,1948]" pageId="2" pageNumber="290">
<emphasis id="B968EAEBFFA4FF937CE3088A01ECFC75" bold="true" box="[189,496,985,1012]" pageId="2" pageNumber="290">Habitat and Distribution</emphasis>
:— 
<taxonomicName id="4C1C4D7AFFA4FF937E4A0889033BFC72" authorityName="Dan Liang &amp; R. J. Wang" authorityYear="2022" box="[532,807,986,1012]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="species" species="xylocarpum">
<emphasis id="B968EAEBFFA4FF937E4A0889033BFC72" box="[532,807,986,1012]" italics="true" pageId="2" pageNumber="290">Combretum xylocarpum</emphasis>
</taxonomicName>
is currently known from the coastal region of 
<collectingRegion id="49D8F81BFFA4FF937904088900F2FB9E" country="China" name="Hainan" pageId="2" pageNumber="290">Hainan Province</collectingRegion>
, 
<collectingCountry id="F30B7669FFA4FF937CA408AD015DFB9E" box="[250,321,1022,1048]" name="China" pageId="2" pageNumber="290">China</collectingCountry>
. It grows mainly on litoral thickets of roadsides or sometimes close to mangrove margin, at 
<quantity id="4CE49B1CFFA4FF93791608AD05B0FB9E" box="[1352,1452,1022,1048]" metricMagnitude="2" metricUnit="m" metricValue="1.03" metricValueMax="2.02" metricValueMin="0.04" pageId="2" pageNumber="290" unit="m" value="103.0" valueMax="202.0" valueMin="4.0">4–202 m</quantity>
elev., frequently accompanied by species such as 
<taxonomicName id="4C1C4D7AFFA4FF937EED0F710560FBBA" authority="(Linnaeus 1753: 196) Brongniart (1826: 62)" authorityName="Brongniart" authorityPageNumber="62" authorityYear="1826" baseAuthorityName="Linnaeus" baseAuthorityPageNumber="196" baseAuthorityYear="1753" box="[691,1404,1058,1085]" class="Magnoliopsida" family="Rhamnaceae" genus="Colubrina" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="species" species="asiatica">
<emphasis id="B968EAEBFFA4FF937EED0F71039AFBBA" box="[691,902,1058,1084]" italics="true" pageId="2" pageNumber="290">Colubrina asiatica</emphasis>
(
<bibRefCitation id="EF8D4B08FFA4FF937FCB0F71046AFBBA" author="Linnaeus, C." box="[917,1142,1058,1085]" pageId="2" pageNumber="290" refId="ref3600" refString="Linnaeus, C. (1753) Species plantarum. Vol. 1. Imprensis Laurentii Salvii, Holmiae, 560 pp. https: // doi. org / 10.5962 / bhl. title. 669" type="book" year="1753">Linnaeus 1753: 196</bibRefCitation>
) 
<bibRefCitation id="EF8D4B08FFA4FF9378D80F710560FBBA" author="Brongniart, A. T." box="[1158,1404,1058,1084]" pageId="2" pageNumber="290" refId="ref2797" refString="Brongniart, A. T. (1826) Memoire sur la Famille des Rhamnees, ou, Histoire naturelle et medicale des genres qui composent ce groupe de plantes. De l'imprimerie de Didot le Jeune, A Paris, 78 pp. https: // doi. org / 10.5962 / bhl. title. 97298" type="book" year="1826">Brongniart (1826: 62)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C1C4D7AFFA4FF937CD60F150242FBE6" authority="R. Brown (1810: 464)" authorityName="R. Brown" authorityPageNumber="464" authorityYear="1810" box="[136,606,1094,1120]" class="Magnoliopsida" family="Apocynaceae" genus="Secamone" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="species" species="elliptica">
<emphasis id="B968EAEBFFA4FF937CD60F150141FBE6" box="[136,349,1094,1120]" italics="true" pageId="2" pageNumber="290">Secamone elliptica</emphasis>
R. 
<bibRefCitation id="EF8D4B08FFA4FF937DDA0F150242FBE6" author="Brown, R." box="[388,606,1094,1120]" pageId="2" pageNumber="290" refId="ref2855" refString="Brown, R. (1810) Prodromus florae Novae Hollandiae et Insulae Van-Diemen, exhibens characteres plantarum. Typis R. Taylor et socii, Londini, 590 pp. https: // doi. org / 10.5962 / bhl. title. 3678" type="book" year="1810">Brown (1810: 464)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3066572FFA4FF937CE30F3903EFF9B2" pageId="2" pageNumber="290" type="discussion">
<paragraph id="8BA336F9FFA4FF937CE30F390574FADA" blockId="2.[136,1452,230,1948]" pageId="2" pageNumber="290">
<emphasis id="B968EAEBFFA4FF937CE30F39015DFB02" bold="true" box="[189,321,1130,1156]" pageId="2" pageNumber="290">Palynology</emphasis>
:—The pollen grains of 
<taxonomicName id="4C1C4D7AFFA4FF937E0D0F3902EFFB02" box="[595,755,1130,1156]" class="Magnoliopsida" family="Combretaceae" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="family">Combretaceae</taxonomicName>
are quite uniform. The basic characters of the family, as well as in 
<taxonomicName id="4C1C4D7AFFA4FF937C9C0FDD01CCFB2E" authorityName="Dan Liang &amp; R. J. Wang" authorityYear="2022" box="[194,464,1166,1192]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="species" species="xylocarpum">
<emphasis id="B968EAEBFFA4FF937C9C0FDD01CCFB2E" box="[194,464,1166,1192]" italics="true" pageId="2" pageNumber="290">Combretum xylocarpum</emphasis>
</taxonomicName>
, are monads, isopolar, radially symmetrical, and heterocolpate, which possess 
<specimenCount id="9D1AFD70FFA4FF93791E0FDD05B7FB2E" box="[1344,1451,1166,1192]" pageId="2" pageNumber="290" type="generic" typeStatus="types">two types</specimenCount>
of colpi, one with endoaperture and one without, forming a grain with three simple apertures alternating with three composite ones (
<bibRefCitation id="EF8D4B08FFA4FF937D130F8502D2FB76" author="El Ghazali, G. E. B. &amp; Krzywinski, K." box="[333,718,1238,1264]" pageId="2" pageNumber="290" pagination="179 - 186" refId="ref3045" refString="El Ghazali, G. E. B. &amp; Krzywinski, K. (1989) An attempt to clarify the term Heterocolpate. Grana 28 (3): 179 - 186. https: // dx. doi. org / 10.1080 / 00173138909427429" type="journal article" year="1989">El Ghazali and Krzywinski 1989</bibRefCitation>
, 
<bibRefCitation id="EF8D4B08FFA4FF937E830F850485FB77" author="Krachai, P. &amp; Pornpongrungrueng, P." box="[733,1177,1238,1265]" pageId="2" pageNumber="290" pagination="4 - 14" refId="ref3565" refString="Krachai, P. &amp; Pornpongrungrueng, P. (2015) Pollen morphology of Combretaceae from Thailand and its taxonomic significance. Thai Forest Bulletin (Botany) 43: 4 - 14." type="journal article" year="2015">Krachai and Pornpongrungrueng 2015</bibRefCitation>
, 
<bibRefCitation id="EF8D4B08FFA4FF9378F60F850576FB76" author="El Ghazali, G. E. B." box="[1192,1386,1238,1264]" pageId="2" pageNumber="290" pagination="131 - 142" refId="ref3092" refString="El Ghazali, G. E. B. (2016) A study on the pollen morphology of the genus Combretum Loefl. and its taxonomic significance. South Asian Journal Experimental Biology 6: 131 - 142. https: // doi. org / 10.38150 / sajeb. 6 (4). p 131 - 142" type="journal article" year="2016">El Ghazali 2016</bibRefCitation>
). The pollen size of 
<taxonomicName id="4C1C4D7AFFA4FF937D740FA901CCFA92" authorityName="Dan Liang &amp; R. J. Wang" authorityYear="2022" box="[298,464,1274,1300]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="species" species="xylocarpum">
<emphasis id="B968EAEBFFA4FF937D740FA901CCFA92" box="[298,464,1274,1300]" italics="true" pageId="2" pageNumber="290">C. xylocarpum</emphasis>
</taxonomicName>
is 15.85 (13.80–17.70) μm in polar axis (P) and 16.25 (14.57–18.20) μm in equatorial diameters (E), with a P/E value 0.96. According to terminology proposed by 
<bibRefCitation id="EF8D4B08FFA4FF937FB40E4D04AEFABE" author="Punt, W. &amp; Hoen, P. P. &amp; Blackmore, S. &amp; Nilsson, S. &amp; Thomas, A. L." box="[1002,1202,1310,1336]" pageId="2" pageNumber="290" pagination="1 - 81" refId="ref4096" refString="Punt, W., Hoen, P. P., Blackmore, S., Nilsson, S. &amp; Thomas, A. L. (2007) Glossary of pollen and spore terminology. Review of Palaeobotany and Palynology 143: 1 - 81. https: // doi. org / 10.1016 / j. revpalbo. 2006.06.008" type="journal article" year="2007">
Punt 
<emphasis id="B968EAEBFFA4FF93787B0E4C044BFABE" box="[1061,1111,1310,1336]" italics="true" pageId="2" pageNumber="290">et al</emphasis>
. (2007)
</bibRefCitation>
, the pollen is medium in size and oblate spheroidal in shape. The exine ornamentation of the pollen is perforate and psilate (
<figureCitation id="13272A7CFFA4FF9378A60E110547FADA" box="[1272,1371,1346,1372]" captionStart="FIGURE 2" captionStartId="4.[136,234,1361,1383]" captionTargetBox="[193,1399,203,1336]" captionTargetId="figure-19@4.[183,1404,190,1337]" captionTargetPageId="4" captionText="FIGURE 2. Pollen morphology of Combretum xylocarpum under SEM. A: Equatorial view, showing the endoaperture protrusions; B: Polar view, showing the heterocolpate grains; C: Colporate aperture with endoaperture protrusions; D: Perforate and psilate exine. Arrowheads indicate subsidiary colpi." figureDoi="http://doi.org/10.5281/zenodo.7095424" httpUri="https://zenodo.org/record/7095424/files/figure.png" pageId="2" pageNumber="290">Figure 2</figureCitation>
).
</paragraph>
<paragraph id="8BA336F9FFA4FF937CE30E3503EFF9B2" blockId="2.[136,1452,230,1948]" pageId="2" pageNumber="290">
According to the pollen 
<typeStatus id="54A7885BFFA4FF937D910E350216FA06" box="[463,522,1382,1408]" pageId="2" pageNumber="290">types</typeStatus>
identified by 
<bibRefCitation id="EF8D4B08FFA4FF937EF60E350368FA06" author="El Ghazali, G. E. B." box="[680,884,1382,1408]" pageId="2" pageNumber="290" pagination="131 - 142" refId="ref3092" refString="El Ghazali, G. E. B. (2016) A study on the pollen morphology of the genus Combretum Loefl. and its taxonomic significance. South Asian Journal Experimental Biology 6: 131 - 142. https: // doi. org / 10.38150 / sajeb. 6 (4). p 131 - 142" type="journal article" year="2016">El Ghazali (2016)</bibRefCitation>
, the pollen of 
<taxonomicName id="4C1C4D7AFFA4FF9378480E3504A7FA06" authorityName="Dan Liang &amp; R. J. Wang" authorityYear="2022" box="[1046,1211,1382,1408]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="species" species="xylocarpum">
<emphasis id="B968EAEBFFA4FF9378480E3504A7FA06" box="[1046,1211,1382,1408]" italics="true" pageId="2" pageNumber="290">C. xylocarpum</emphasis>
</taxonomicName>
seems to be ascribed to 
<emphasis id="B968EAEBFFA4FF937CFA0ED90140FA22" box="[164,348,1418,1444]" italics="true" pageId="2" pageNumber="290">C. primigenium-</emphasis>
<typeStatus id="54A7885BFFA4FF937D020ED90190FA22" box="[348,396,1418,1444]" pageId="2" pageNumber="290">type</typeStatus>
because of the 3-angled polar view and without vestibula, but its endoaperture protrusions also make it closed to 
<emphasis id="B968EAEBFFA4FF937D080EFD0219FA4E" box="[342,517,1454,1480]" italics="true" pageId="2" pageNumber="290">C. bracteosum-</emphasis>
<typeStatus id="54A7885BFFA4FF937E5B0EFD0225FA4E" box="[517,569,1454,1480]" pageId="2" pageNumber="290">type</typeStatus>
. Whereas the pollen grains of 
<taxonomicName id="4C1C4D7AFFA4FF937FC70EFD0459FA4E" authorityName="Roxburgh" authorityYear="1832" box="[921,1093,1454,1480]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="species" species="acuminatum">
<emphasis id="B968EAEBFFA4FF937FC70EFD0459FA4E" box="[921,1093,1454,1480]" italics="true" pageId="2" pageNumber="290">C. acuminatum</emphasis>
</taxonomicName>
was identified to be 
<taxonomicName id="4C1C4D7AFFA4FF9379650EFD05BEFA4E" box="[1339,1442,1454,1480]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="species" species="laxum">
<emphasis id="B968EAEBFFA4FF9379650EFD05BEFA4E" box="[1339,1442,1454,1480]" italics="true" pageId="2" pageNumber="290">C. laxum</emphasis>
</taxonomicName>
- 
<typeStatus id="54A7885BFFA4FF937CD60E8100A4FA6A" box="[136,184,1490,1516]" pageId="2" pageNumber="290">type</typeStatus>
for their 6-lobed polar view and absence of endoaperture protrusions, and those of 
<taxonomicName id="4C1C4D7AFFA4FF9378D50E81050DFA6A" box="[1163,1297,1490,1516]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="species" species="chinense">
<emphasis id="B968EAEBFFA4FF9378D50E81050DFA6A" box="[1163,1297,1490,1516]" italics="true" pageId="2" pageNumber="290">C. chinense</emphasis>
</taxonomicName>
belong to 
<taxonomicName id="4C1C4D7AFFA4FF9379CC0E810110F996" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="species" species="squamosum">
<emphasis id="B968EAEBFFA4FF9379CC0E810110F996" italics="true" pageId="2" pageNumber="290">C. squamosum</emphasis>
</taxonomicName>
- 
<typeStatus id="54A7885BFFA4FF937D4A0EA5015AF996" box="[276,326,1526,1552]" pageId="2" pageNumber="290">type</typeStatus>
due to their 3-angled polar view and presence of vestibula at apertures. This may indicate the unique and undescribed characters of the new species in both fruits and pollen grains.
</paragraph>
</subSubSection>
<subSubSection id="C3066572FFA4FF937CE30D6D04E3F942" pageId="2" pageNumber="290" type="materials_examined">
<paragraph id="8BA336F9FFA4FF937CE30D6D04E3F942" blockId="2.[136,1452,230,1948]" pageId="2" pageNumber="290">
<emphasis id="B968EAEBFFA4FF937CE30D6D0129F9DE" bold="true" box="[189,309,1598,1624]" pageId="2" pageNumber="290">
<typeStatus id="54A7885BFFA4FF937CE30D6D0129F9DE" box="[189,309,1598,1624]" pageId="2" pageNumber="290" type="paratype">Paratypes</typeStatus>
</emphasis>
:— 
<collectingCountry id="F30B7669FFA4FF937D090D6D01A9F9DE" box="[343,437,1598,1624]" name="China" pageId="2" pageNumber="290">CHINA</collectingCountry>
. 
<collectingRegion id="49D8F81BFFA4FF937D9E0D6D0263F9DE" box="[448,639,1598,1624]" country="China" name="Hainan" pageId="2" pageNumber="290">Hainan province</collectingRegion>
: Wenchang City, Longlou Town, Nearby the Mt. Tongguling, roadsides, 
<geoCoordinate id="EE28503EFFA4FF937CD60D3100F8F9FA" box="[136,228,1634,1660]" degrees="111" direction="east" minutes="0" orientation="longitude" pageId="2" pageNumber="290" precision="925" value="111.0">111°0’E</geoCoordinate>
, 
<geoCoordinate id="EE28503EFFA4FF937CAC0D310149F9FA" box="[242,341,1634,1660]" degrees="19" direction="north" minutes="40" orientation="latitude" pageId="2" pageNumber="290" precision="925" value="19.666666">19°40’N</geoCoordinate>
, elevation ca. 
<quantity id="4CE49B1CFFA4FF937DA90D310239F9FA" box="[503,549,1634,1660]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="290" unit="m" value="4.0">4 m</quantity>
, young fruit, 
<date id="FFA21039FFA4FF937EE30D310355F9FA" box="[701,841,1634,1660]" pageId="2" pageNumber="290" value="2019-06-04">4 June 2019</date>
, 
<emphasis id="B968EAEBFFA4FF937F0A0D30051DF9FA" box="[852,1281,1634,1660]" italics="true" pageId="2" pageNumber="290">Dan Liang &amp; Guo-Bin Jiang WP0930</emphasis>
(
<collectionCode id="ED0DAE3CFFA4FF93794F0D310545F9FA" box="[1297,1369,1634,1660]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15606" lsid="urn:lsid:biocol.org:col:15606" name="South China Botanical Garden" pageId="2" pageNumber="290" type="Herbarium">IBSC</collectionCode>
!); same locality, flowering, 
<date id="FFA21039FFA4FF937D300DD5023AF926" box="[366,550,1670,1696]" pageId="2" pageNumber="290" value="2021-03-16">16 March 2021</date>
, 
<emphasis id="B968EAEBFFA4FF937E690DD403F1F926" box="[567,1005,1670,1696]" italics="true" pageId="2" pageNumber="290">Lang-Xing Yuan WP1723 &amp; WP1724</emphasis>
(
<collectionCode id="ED0DAE3CFFA4FF93785C0DD50456F926" box="[1026,1098,1670,1696]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15606" lsid="urn:lsid:biocol.org:col:15606" name="South China Botanical Garden" pageId="2" pageNumber="290" type="Herbarium">IBSC</collectionCode>
!); Qionghai City, Boao Town, 
<geoCoordinate id="EE28503EFFA4FF937CD60DF900EEF942" box="[136,242,1706,1732]" degrees="111" direction="east" minutes="33" orientation="longitude" pageId="2" pageNumber="290" precision="925" value="111.55">111°33’E</geoCoordinate>
, 
<geoCoordinate id="EE28503EFFA4FF937D5E0DF9017FF942" box="[256,355,1706,1732]" degrees="19" direction="north" minutes="06" orientation="latitude" pageId="2" pageNumber="290" precision="925" value="19.1">19°06’N</geoCoordinate>
, elevation ca. 
<quantity id="4CE49B1CFFA4FF937E5D0DF9022CF942" box="[515,560,1706,1733]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="290" unit="m" value="5.0">5 m</quantity>
, 
<date id="FFA21039FFA4FF937E620DF902C8F942" box="[572,724,1706,1732]" pageId="2" pageNumber="290" value="2020-05-27">27 May 2020</date>
, 
<emphasis id="B968EAEBFFA4FF937E810DF8048BF942" box="[735,1175,1706,1732]" italics="true" pageId="2" pageNumber="290">Rui-Jiang Wang &amp; Dan Liang WP1060</emphasis>
(
<collectionCode id="ED0DAE3CFFA4FF9378F90DF904F3F942" box="[1191,1263,1706,1732]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15606" lsid="urn:lsid:biocol.org:col:15606" name="South China Botanical Garden" pageId="2" pageNumber="290" type="Herbarium">IBSC</collectionCode>
!).
</paragraph>
</subSubSection>
<subSubSection id="C3066572FFA4FF937CE30D9E0173F81A" pageId="2" pageNumber="290" type="conservation">
<paragraph id="8BA336F9FFA4FF937CE30D9E0173F81A" blockId="2.[136,1452,230,1948]" pageId="2" pageNumber="290">
<emphasis id="B968EAEBFFA4FF937CE30D9E01B6F96E" bold="true" box="[189,426,1741,1768]" pageId="2" pageNumber="290">Conservation status</emphasis>
:—Until now, 
<taxonomicName id="4C1C4D7AFFA4FF937E140D9D034BF96E" authorityName="Dan Liang &amp; R. J. Wang" authorityYear="2022" box="[586,855,1742,1768]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="species" species="xylocarpum">
<emphasis id="B968EAEBFFA4FF937E140D9D034BF96E" box="[586,855,1742,1768]" italics="true" pageId="2" pageNumber="290">Combretum xylocarpum</emphasis>
</taxonomicName>
is known from several localities in 
<collectingRegion id="49D8F81BFFA4FF9378BD0D9D052BF96E" box="[1251,1335,1742,1768]" country="China" name="Hainan" pageId="2" pageNumber="290">Hainan</collectingRegion>
, including the two subpopulations found by ourselves and two by others. The population at the 
<typeStatus id="54A7885BFFA4FF93783C0DA1048EF88A" box="[1122,1170,1778,1804]" pageId="2" pageNumber="290">type</typeStatus>
locality only includes 2 mature individuals, but at the other sites include more than 100 mature individuals. In addition, the new species grows at road sides and has no any medicinal and ornamental values. According to the related criteria proposed by IUCN Standards and Petitions Subcommittee (2019), the current conservation status of 
<taxonomicName id="4C1C4D7AFFA4FF93787E0C0D04DAF8FE" authorityName="Dan Liang &amp; R. J. Wang" authorityYear="2022" box="[1056,1222,1886,1912]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="290" phylum="Tracheophyta" rank="species" species="xylocarpum">
<emphasis id="B968EAEBFFA4FF93787E0C0D04DAF8FE" box="[1056,1222,1886,1912]" italics="true" pageId="2" pageNumber="290">C. xylocarpum</emphasis>
</taxonomicName>
can be evaluated as Least Concern (LC).
</paragraph>
</subSubSection>
</treatment>
</document>