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<document id="FAEAB321F6953A74B7136FD8EB2530D0" ID-CLB-Dataset="22833" ID-DOI="10.11646/zootaxa.4766.1.1" ID-GBIF-Dataset="ed4d1bf6-284a-4898-952a-58943d5a3026" ID-ISSN="1175-5334" ID-Zenodo-Dep="3763907" ID-ZooBank="urn:lsid:zoobank.org:pub:B1B5DA41-5035-4783-8D47-28857B6305AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1587670592452" checkinUser="carolina" docAuthor="Ketwetsuriya, Chatchalerm, Karapunar, Baran, Charoentitirat, Thasinee &amp; Nützel, Al- Exander" docDate="2020" docId="03B587AB4F3A1570FF517D1FFAF9FE2A" docLanguage="en" docName="zootaxa.4766.1.1.pdf.imf" docOrigin="Zootaxa 4766 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Apachella thailandensis Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="15" masterDocId="FF8CFFD34F37157EFFC6787EFFCBFF85" masterDocTitle="Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane" masterLastPageNumber="47" masterPageNumber="1" pageNumber="14" updateTime="1698836971451" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5258C86CC5321BF3FDC07AAC15441397">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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<mods:namePart id="D00824E03C432946454F9F1F71791705">Ketwetsuriya, Chatchalerm</mods:namePart>
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<mods:namePart id="6251A83CF463FF9A1501C6ED183F13F7">Karapunar, Baran</mods:namePart>
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<mods:namePart id="265014A1D2FE0EE8E1C199FA70B511EC">Charoentitirat, Thasinee</mods:namePart>
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<mods:namePart id="2699B3DA4A1E9FAB85CF188B3CDA9553">Nützel, Al- Exander</mods:namePart>
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<mods:number id="8D4569C1FC2DFF44B8CF0F56254314EB">2020-04-17</mods:number>
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<treatment id="03B587AB4F3A1570FF517D1FFAF9FE2A" ID-DOI="http://doi.org/10.5281/zenodo.3803935" ID-GBIF-Taxon="163737207" ID-Zenodo-Dep="3803935" LSID="urn:lsid:plazi:treatment:03B587AB4F3A1570FF517D1FFAF9FE2A" httpUri="http://treatment.plazi.org/id/03B587AB4F3A1570FF517D1FFAF9FE2A" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="C30665364F3A1573FF517D1FFDD4FAFE" box="[151,543,1377,1404]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BA336BD4F3A1573FF517D1FFDD4FAFE" blockId="13.[151,543,1377,1439]" box="[151,543,1377,1404]" pageId="13" pageNumber="14">
<heading id="D0EB81D14F3A1573FF517D1FFDD4FAFE" bold="true" box="[151,543,1377,1404]" fontSize="10" level="2" pageId="13" pageNumber="14" reason="2">
<emphasis id="B968EAAF4F3A1573FF517D1FFDD4FAFE" bold="true" box="[151,543,1377,1404]" pageId="13" pageNumber="14">
<taxonomicName id="4C1C4D3E4F3A1573FF517D1FFE71FAF9" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[151,442,1377,1404]" class="Gastropoda" family="Gosseletinidae" genus="Apachella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="thailandensis" status="sp. nov.">
<emphasis id="B968EAAF4F3A1573FF517D1FFE71FAF9" bold="true" box="[151,442,1377,1404]" italics="true" pageId="13" pageNumber="14">Apachella thailandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F3A1573FE077D1FFDD4FAFE" box="[449,543,1377,1403]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30665364F3A1573FF517DFBFF33FA1A" box="[151,248,1413,1439]" pageId="13" pageNumber="14" type="description">
<paragraph id="8BA336BD4F3A1573FF517DFBFF33FA1A" blockId="13.[151,543,1377,1439]" box="[151,248,1413,1439]" pageId="13" pageNumber="14">
(
<figureCitation id="13272A384F3A1573FF667DFBFF24FA1A" box="[160,239,1413,1439]" captionStart="FIGURE 11" captionStartId="14.[151,250,1467,1492]" captionTargetBox="[151,1436,1036,1442]" captionTargetId="figure@14.[151,1436,1036,1442]" captionTargetPageId="14" captionText="FIGURE 11. Apachella thailandensis sp. nov.AD, ESKU-19-LP 62, holotype. E, ESKU-19-LP 25, paratype. FG, ESKU-19- LP 89, paratype. All scale bars represent 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3763929" httpUri="https://zenodo.org/record/3763929/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30665364F3A1573FF517DB3FDBFFA62" box="[151,628,1485,1511]" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BA336BD4F3A1573FF517DB3FDBFFA62" blockId="13.[151,1437,1485,2015]" box="[151,628,1485,1511]" pageId="13" pageNumber="14">
<emphasis id="B968EAAF4F3A1573FF517DB3FED5FA62" bold="true" box="[151,286,1485,1511]" pageId="13" pageNumber="14">Etymology.</emphasis>
After the country of
<collectingCountry id="F30B762D4F3A1573FDCF7DB3FDBBFA62" box="[521,624,1485,1511]" name="Thailand" pageId="13" pageNumber="14">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F3A1573FF017D8FFBB8F91E" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BA336BD4F3A1573FF017D8FFDCFF98E" blockId="13.[151,1437,1485,2015]" box="[199,516,1521,1547]" pageId="13" pageNumber="14">
<materialsCitation id="3B743CE04F3A1573FF017D8FFDCFF98E" ID-GBIF-Occurrence="2608527138" box="[199,516,1521,1547]" collectionCode="LP" pageId="13" pageNumber="14" specimenCode="ESKU-19-LP 62" specimenCount="1" typeStatus="holotype">
<emphasis id="B968EAAF4F3A1573FF017D8FFEF0F98E" bold="true" box="[199,315,1521,1547]" pageId="13" pageNumber="14">
<typeStatus id="54A7881F4F3A1573FF017D8FFEFDF98E" box="[199,310,1521,1547]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBBA9EC64F3A1573FE847D8FFE34F98E" box="[322,511,1521,1547]" pageId="13" pageNumber="14">ESKU-19-LP 62</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA336BD4F3A1573FF017E6BFC91F9AA" blockId="13.[151,1437,1485,2015]" box="[199,858,1556,1583]" pageId="13" pageNumber="14">
<emphasis id="B968EAAF4F3A1573FF017E6BFE8DF9AA" bold="true" box="[199,326,1557,1583]" pageId="13" pageNumber="14">
<typeStatus id="54A7881F4F3A1573FF017E6BFE8AF9AA" box="[199,321,1557,1583]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Four specimens: ESKU-19-
<collectionCode id="ED0DAE784F3A1573FD447E6BFD6FF9AA" box="[642,676,1557,1583]" country="Russia" name="Laboratory of Palaeontology" pageId="13" pageNumber="14">LP</collectionCode>
25, 60, 89, 100.
</paragraph>
<paragraph id="8BA336BD4F3A1573FF017E47FBB8F91E" blockId="13.[151,1437,1485,2015]" pageId="13" pageNumber="14">
Dimensions (mm): ESKU-19-
<collectionCode id="ED0DAE784F3A1573FDD37E47FDFCF9D6" box="[533,567,1593,1619]" country="Russia" name="Laboratory of Palaeontology" pageId="13" pageNumber="14">LP</collectionCode>
25: height =
<emphasis id="B968EAAF4F3A1573FD0D7E44FD1CF9D6" box="[715,727,1594,1619]" italics="true" pageId="13" pageNumber="14">c</emphasis>
. 9.4; width = 5.5; apical angle = 75º. ESKU-19-
<collectionCode id="ED0DAE784F3A1573FB2B7E47FAC4F9D6" box="[1261,1295,1593,1619]" country="Russia" name="Laboratory of Palaeontology" pageId="13" pageNumber="14">LP</collectionCode>
62: height = 8.5; width = 5.2; apical angle = 60º. ESKU-19-
<collectionCode id="ED0DAE784F3A1573FD6F7E23FD00F9F2" box="[681,715,1629,1655]" country="Russia" name="Laboratory of Palaeontology" pageId="13" pageNumber="14">LP</collectionCode>
60: height =
<emphasis id="B968EAAF4F3A1573FCA47E20FCA5F9F2" box="[866,878,1630,1655]" italics="true" pageId="13" pageNumber="14">c</emphasis>
. 4.6; width =
<emphasis id="B968EAAF4F3A1573FBC87E20FBD1F9F2" box="[1038,1050,1630,1655]" italics="true" pageId="13" pageNumber="14">c</emphasis>
. 4.6. ESKU-19-
<collectionCode id="ED0DAE784F3A1573FB097E23FB3AF9F2" box="[1231,1265,1629,1655]" country="Russia" name="Laboratory of Palaeontology" pageId="13" pageNumber="14">LP</collectionCode>
89: height =
<emphasis id="B968EAAF4F3A1573FA4F7E20FA5EF9F2" box="[1417,1429,1630,1655]" italics="true" pageId="13" pageNumber="14">c</emphasis>
. 7.1; width =
<emphasis id="B968EAAF4F3A1573FEE17EFCFEF8F91E" box="[295,307,1666,1691]" italics="true" pageId="13" pageNumber="14">c</emphasis>
. 4.1; apical angle = 66º. ESKU-19-
<collectionCode id="ED0DAE784F3A1573FD797EFFFD2AF91E" box="[703,737,1665,1691]" country="Russia" name="Laboratory of Palaeontology" pageId="13" pageNumber="14">LP</collectionCode>
100: height =
<emphasis id="B968EAAF4F3A1573FC437EFCFC5AF91E" box="[901,913,1666,1691]" italics="true" pageId="13" pageNumber="14">c</emphasis>
. 6.5; width =
<emphasis id="B968EAAF4F3A1573FBE87EFCFBF1F91E" box="[1070,1082,1666,1691]" italics="true" pageId="13" pageNumber="14">c</emphasis>
. 4.8.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F3A1573FF017EDBFE74F882" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BA336BD4F3A1573FF017EDBFE74F882" blockId="13.[151,1437,1485,2015]" pageId="13" pageNumber="14">
<emphasis id="B968EAAF4F3A1573FF017EDBFD6AF93A" bold="true" box="[199,673,1700,1727]" pageId="13" pageNumber="14">Type locality and stratigraphical range.</emphasis>
Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about
<quantity id="4CE49B584F3A1573FEF17EB7FEB4F966" box="[311,383,1737,1763]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="14" unit="km" value="13.0">13 km</quantity>
east of
<collectingRegion id="49D8F85F4F3A1573FE1C7EB7FD69F966" box="[474,674,1737,1763]" country="Thailand" name="Lop Buri" pageId="13" pageNumber="14">Lopburi Province</collectingRegion>
, Central
<collectingCountry id="F30B762D4F3A1573FCCF7EB7FCA6F966" box="[777,877,1737,1763]" name="Thailand" pageId="13" pageNumber="14">Thailand</collectingCountry>
(
<figureCitation id="13272A384F3A1573FCBB7EB7FC0BF966" box="[893,960,1737,1763]" captionStart="FIGURE" captionStartId="1.[151,168,1975,2001]" captionTargetBox="[243,1345,1164,1941]" captionTargetId="figure@1.[236,1350,1160,1948]" captionTargetPageId="1" captionText="FIGURE 1. Geological map of the study location at Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about 13 km east of Lopburi Province, Central Thailand." figureDoi="http://doi.org/10.5281/zenodo.3763909" httpUri="https://zenodo.org/record/3763909/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
), Khao Khad Formation,
<collectingRegion id="49D8F85F4F3A1573FB217EB7FA8DF966" box="[1255,1350,1737,1763]" country="Thailand" name="Sara Buri" pageId="13" pageNumber="14">Saraburi</collectingRegion>
Group, Middle Permian, Roadian.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F3A1570FF017F6EFAA5FF52" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="8BA336BD4F3A1570FF017F6EFAA5FF52" blockId="13.[151,1437,1485,2015]" lastBlockId="14.[151,1437,153,431]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B968EAAF4F3A1573FF017F6EFE92F8AE" bold="true" box="[199,345,1808,1835]" pageId="13" pageNumber="14">Description.</emphasis>
Shell high-spired, consisting of 8 whorls, first whorl planispiral; first two teleoconch whorls smooth, rounded, third whorl ornamented with spiral threads; selenizone starts within the 4th whorl; suture slightly impressed; subsutural ramp slightly convex, ornamented with 34 spiral ribs that are ornamented with nodes; selenizone concave, ornamented with very faint lunulae, bordered by two straight spiral ribs; lower rib forms periphery; lower whorl face twice as wide as the upper whorl face; lower whorl face slightly convex, ornamented with one spiral rib just above suture, equal in strength to spiral rib bordering the lower edge of selenizone; lower whorl face further ornamented with two spiral threads between two spiral ribs; base convex, ornamented with several evenly spaced spiral ribs of same strength as ribs on lower whorl face; base convex, anomphalous; aperture unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F391570FF01789FFAF9FE2A" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BA336BD4F391570FF01789FFAF9FE2A" blockId="14.[151,1437,153,431]" pageId="14" pageNumber="15">
<emphasis id="B968EAAF4F391570FF01789FFEF0FF7E" bold="true" box="[199,315,225,251]" pageId="14" pageNumber="15">Remarks.</emphasis>
<taxonomicName id="4C1C4D3E4F391570FE87789FFD85FF7E" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[321,590,225,251]" class="Gastropoda" family="Gosseletinidae" genus="Apachella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="thailandensis" status="sp. nov.">
<emphasis id="B968EAAF4F391570FE87789FFD85FF7E" box="[321,590,225,251]" italics="true" pageId="14" pageNumber="15">Apachella thailandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F391570FD93789FFD67FF7E" box="[597,684,225,251]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
has a wider upper whorl face (above selenizone) than other species of
<taxonomicName id="4C1C4D3E4F391570FF73797BFEECFE9A" authorityName="Winters" authorityYear="1956" box="[181,295,261,287]" class="Gastropoda" family="Gosseletinidae" genus="Apachella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="B968EAAF4F391570FF73797BFEECFE9A" box="[181,295,261,287]" italics="true" pageId="14" pageNumber="15">Apachella</emphasis>
</taxonomicName>
. The ratio of the height of the upper and the lower whorl face of
<taxonomicName id="4C1C4D3E4F391570FBC6797BFB7FFE9A" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[1024,1204,261,287]" class="Gastropoda" family="Gosseletinidae" genus="Apachella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="thailandensis" status="sp. nov.">
<emphasis id="B968EAAF4F391570FBC6797BFBDAFE9A" box="[1024,1041,261,287]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.
<emphasis id="B968EAAF4F391570FBD9797BFB7FFE9A" box="[1055,1204,261,287]" italics="true" pageId="14" pageNumber="15">thailandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F391570FB7D797BFAD9FE9A" box="[1211,1298,261,287]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
is similar to that of
<taxonomicName id="4C1C4D3E4F391570FF227957FDC5FEC6" authority="(Girty, 1909)" baseAuthorityName="Girty" baseAuthorityYear="1909" box="[228,526,297,323]" class="Gastropoda" family="Gosseletinidae" genus="Apachella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="alimillana">
<emphasis id="B968EAAF4F391570FF227957FF3EFEC6" box="[228,245,297,323]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.
<emphasis id="B968EAAF4F391570FEC47957FEBEFEC6" box="[258,373,297,323]" italics="true" pageId="14" pageNumber="15">alimillana</emphasis>
(
<bibRefCitation id="EF8D4B4C4F391570FE447957FDCDFEC6" author="Girty" box="[386,518,297,323]" firstAuthor="Girty" pageId="14" pageNumber="15" pagination="40 - 141" refId="ref24181" refString="Girty, G. H. (1909) Paleontology of the Manzano group. In: Lee, W. T. &amp; Girty, G. H. (Eds.), The Manzano group of the Rio Grande Valley, New Mexico. Bulletin of the U. S. Geological Survey. Government Printing Office, Washington, D. C., pp. 40 - 141." type="book chapter" year="1909">Girty, 1909</bibRefCitation>
)
</taxonomicName>
. However, the latter lacks spiral ribs on upper whorl face.
<taxonomicName id="4C1C4D3E4F391570FB487957FF04FEE2" authority="Batten, 1995" authorityName="Batten" authorityYear="1995" class="Gastropoda" family="Gosseletinidae" genus="Apachella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="powwowensis">
<emphasis id="B968EAAF4F391570FB487957FB54FEC6" box="[1166,1183,297,323]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.
<emphasis id="B968EAAF4F391570FB6A7954FA8DFEC6" box="[1196,1350,298,323]" italics="true" pageId="14" pageNumber="15">powwowensis</emphasis>
<bibRefCitation id="EF8D4B4C4F391570FA8A7957FF04FEE2" author="Batten" firstAuthor="Batten" pageId="14" pageNumber="15" pagination="1 - 46" refId="ref23199" refString="Batten, R. L. (1995) Pennsylvanina (Morrowan) gastropods from the Magdalena Formation of the Hueco Mountains, Texas. American Museum Novitates, 3122, 1 - 46." type="journal article" year="1995">Batten, 1995</bibRefCitation>
</taxonomicName>
has a wider selenizone, a median spiral lira on its selenizone and only one spiral rib on its upper whorl face.
<taxonomicName id="4C1C4D3E4F391570FF51790FFE39FE0E" authority="(Chronic, 1952)" baseAuthorityName="Chronic" baseAuthorityYear="1952" box="[151,498,368,395]" class="Gastropoda" family="Gosseletinidae" genus="Apachella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="franciscana">
<emphasis id="B968EAAF4F391570FF51790FFF63FE0E" box="[151,168,369,395]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.
<emphasis id="B968EAAF4F391570FF70790FFEF1FE0E" box="[182,314,369,395]" italics="true" pageId="14" pageNumber="15">franciscana</emphasis>
(
<bibRefCitation id="EF8D4B4C4F391570FE8E790FFE21FE0E" author="Chronic" box="[328,490,368,395]" firstAuthor="Chronic" pageId="14" pageNumber="15" pagination="95 - 166" refId="ref23437" refString="Chronic, H. (1952) Molluscan fauna from the Permian Kaibab Formation, Walnut Canyon, Arizona. Geological Society America Bulletin, 63 (2), 95 - 166. https: // doi. org / 10.1130 / 0016 - 7606 (1952) 63 [95: MFFTPK] 2.0. CO; 2" type="journal article" year="1952">Chronic, 1952</bibRefCitation>
)
</taxonomicName>
differs by having two faint spiral ribs on the upper whorl face adjacent to the suture, more prominent ribs bordering the selenizone, a stepped whorl profile and a higher whorl expansion rate.
</paragraph>
</subSubSection>
</treatment>
</document>