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<mods:title id="3D17C92152975E49989C9B2B65D97E88">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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<mods:namePart id="EEFABD02513E688B8D7AD268A053FAB2">Ketwetsuriya, Chatchalerm</mods:namePart>
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<treatment id="03B587AB4F25156DFF517E46FC40FE72" ID-DOI="http://doi.org/10.5281/zenodo.3803873" ID-GBIF-Taxon="163737211" ID-Zenodo-Dep="3803873" LSID="urn:lsid:plazi:treatment:03B587AB4F25156DFF517E46FC40FE72" httpUri="http://treatment.plazi.org/id/03B587AB4F25156DFF517E46FC40FE72" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="C30665364F25156CFF517E46FD84F9D7" box="[151,591,1592,1619]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BA336BD4F25156CFF517E46FD84F9D7" blockId="18.[151,591,1592,1655]" box="[151,591,1592,1619]" pageId="18" pageNumber="19">
<heading id="D0EB81D14F25156CFF517E46FD84F9D7" bold="true" box="[151,591,1592,1619]" fontSize="10" level="2" pageId="18" pageNumber="19" reason="2">
<taxonomicName id="4C1C4D3E4F25156CFF517E46FD84F9D7" authority="Mansuy, 1912" authorityName="Mansuy" authorityYear="1912" box="[151,591,1592,1619]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="pagoda">
<emphasis id="B968EAAF4F25156CFF517E46FD84F9D7" bold="true" box="[151,591,1592,1619]" pageId="18" pageNumber="19">
<emphasis id="B968EAAF4F25156CFF517E46FEDDF9D6" bold="true" box="[151,278,1592,1619]" italics="true" pageId="18" pageNumber="19">Worthenia</emphasis>
cf.
<emphasis id="B968EAAF4F25156CFE857E46FE50F9D6" bold="true" box="[323,411,1592,1619]" italics="true" pageId="18" pageNumber="19">pagoda</emphasis>
<bibRefCitation id="EF8D4B4C4F25156CFE647E47FD84F9D7" author="Mansuy" box="[418,591,1592,1619]" firstAuthor="Mansuy" pageId="18" pageNumber="19" pagination="1 - 146" refId="ref25972" refString="Mansuy, H. (1912) Paleontologie. In: Mansuy, H. &amp; Deprat, J. (Eds.), Etude geologique du Yun-Nan oriental. Actes de la Societe Geologique de l'Indo-Chine, 1 (2), pp. 1 - 146." type="journal article" year="1912">Mansuy, 1912</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30665364F25156CFF517E22FEEDF9F2" box="[151,294,1628,1655]" pageId="18" pageNumber="19" type="description">
<paragraph id="8BA336BD4F25156CFF517E22FEEDF9F2" blockId="18.[151,591,1592,1655]" box="[151,294,1628,1655]" pageId="18" pageNumber="19">
(
<figureCitation id="13272A384F25156CFF667E22FF25F9F2" box="[160,238,1628,1655]" captionStart="FIGURE 14" captionStartId="17.[151,250,1945,1970]" captionTargetBox="[155,1432,181,1920]" captionTargetId="figure@17.[155,1432,181,1920]" captionTargetPageId="17" captionText="FIGURE 14. AI, Worthenia humiligrada sp. nov. AD, ESKU-19-LP 22. EF, ESKU-19-LP 33. GH, ESKU-19-LP 23. I, ESKU-19-LP 34. JN, Worthenia cf. pagoda Mansuy, 1912, J, ESKU-19-LP 17. KN, ESKU-19-LP 66. OS, Worthenia sp., ESKU-19-LP 88. Scale bars represent: 2 mm (A, B, G, H, I, K, L, O); 1 mm (C, D, J, M, N, PS); 5 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.3763935" httpUri="https://zenodo.org/record/3763935/files/figure.png" pageId="18" pageNumber="19">Fig. 14</figureCitation>
JN)
</paragraph>
</subSubSection>
<subSubSection id="C30665364F25156CFF517EDEFC03F93C" box="[151,968,1696,1721]" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="8BA336BD4F25156CFF517EDEFC03F93C" blockId="18.[151,968,1696,1721]" box="[151,968,1696,1721]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="AB0C11934F25156CFF517EDEFC03F93C" box="[151,968,1696,1721]" pageId="18" pageNumber="19">
cf.
<treatmentCitation id="0ABD10AC4F25156CFF7E7EDFFDFEF93C" box="[184,565,1697,1721]" pageId="18" pageNumber="19">
<taxonomicName id="4C1C4D3E4F25156CFF7E7EDFFDFAF93C" authority="Mansuy, 1912: 39" authorityName="Mansuy" authorityPageNumber="39" authorityYear="1912" box="[184,561,1697,1721]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="pagoda">
<emphasis id="B968EAAF4F25156CFF7E7EDFFEB9F93C" box="[184,370,1697,1721]" italics="true" pageId="18" pageNumber="19">Worthenia pagoda</emphasis>
<bibRefCitation id="EF8D4B4C4F25156CFEBF7EDFFDFAF93C" author="Mansuy" box="[377,561,1697,1721]" firstAuthor="Mansuy" pageId="18" pageNumber="19" pagination="1 - 146" refId="ref25972" refString="Mansuy, H. (1912) Paleontologie. In: Mansuy, H. &amp; Deprat, J. (Eds.), Etude geologique du Yun-Nan oriental. Actes de la Societe Geologique de l'Indo-Chine, 1 (2), pp. 1 - 146." type="journal article" year="1912">Mansuy, 1912: 39</bibRefCitation>
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,
</treatmentCitation>
pl. 7, fig. 6;
<treatmentCitation id="0ABD10AC4F25156CFD7E7EDEFCBFF93C" box="[696,884,1696,1721]" pageId="18" pageNumber="19">
<bibRefCitation id="EF8D4B4C4F25156CFD7E7EDEFCBFF93C" author="Delpey" box="[696,884,1696,1721]" firstAuthor="Delpey" pageId="18" pageNumber="19" pagination="345 - 369" refId="ref23861" refString="Delpey, G. (1941) Les Gasteropodes permiens du Cambodge. Journal de Conchyologie, 84, 255 - 278, 345 - 369." type="journal article" year="1941">Delpey, 1941: 353</bibRefCitation>
</treatmentCitation>
, fig. 26.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30665364F25156CFF517E98FEA3F8CC" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BA336BD4F25156CFF517E98FD7FF884" blockId="18.[151,1436,1766,2045]" box="[151,692,1766,1794]" pageId="18" pageNumber="19">
<emphasis id="B968EAAF4F25156CFF517E98FECDF884" bold="true" box="[151,262,1766,1793]" pageId="18" pageNumber="19">Material.</emphasis>
Two specimens: ESKU-19-
<collectionCode id="ED0DAE784F25156CFDF87E99FDABF884" box="[574,608,1767,1793]" country="Russia" name="Laboratory of Palaeontology" pageId="18" pageNumber="19">LP</collectionCode>
17, 66.
</paragraph>
<paragraph id="8BA336BD4F25156CFF017F75FEA3F8CC" blockId="18.[151,1436,1766,2045]" pageId="18" pageNumber="19">
Dimensions (mm): ESKU-19-
<collectionCode id="ED0DAE784F25156CFDD37F75FDFCF8A0" box="[533,567,1803,1829]" country="Russia" name="Laboratory of Palaeontology" pageId="18" pageNumber="19">LP</collectionCode>
17: height =
<emphasis id="B968EAAF4F25156CFD0F7F72FD1EF8A0" box="[713,725,1804,1829]" italics="true" pageId="18" pageNumber="19">c</emphasis>
. 6.6; width = 4.9. ESKU-19-
<collectionCode id="ED0DAE784F25156CFBD07F75FBF3F8A0" box="[1046,1080,1803,1829]" country="Russia" name="Laboratory of Palaeontology" pageId="18" pageNumber="19">LP</collectionCode>
66: height = 5.5; width =
<emphasis id="B968EAAF4F25156CFA9E7F72FAAFF8A0" box="[1368,1380,1804,1829]" italics="true" pageId="18" pageNumber="19">c</emphasis>
. 5.0; apical angle = 75º.
</paragraph>
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<subSubSection id="C30665364F25156DFF017F2CFD67FEC6" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="8BA336BD4F25156DFF017F2CFD67FEC6" blockId="18.[151,1436,1766,2045]" lastBlockId="19.[151,1437,153,503]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B968EAAF4F25156CFF017F2CFE92F8E8" bold="true" box="[199,345,1874,1901]" pageId="18" pageNumber="19">Description.</emphasis>
Shell moderately high-spired, trochiform, consisting of 6 whorls; first three teleoconch whorls smooth, convex, rounded; mid angulation develops starting from the 4th whorl onward; the last quarter of the last whorl slightly deflected; suture slightly impressed; subsutural ramp with faint subsutural convexity, slightly concave on remaining part; ramp inclining at an angle of about 4560°; subsutural ramp ornamented with three spiral ribs starting from 4th whorl and almost straight prosocline growth lines; spiral ribs ornamented with equally spaced nodes; mid-angulation forms periphery where selenizone situated; selenizone prominent, wide, comprising about 1/7 of the whole whorl face of 5th whorl, ornamented with equally spaced strong nodes, bordered by fine spiral ribs; lower whorl face facing abapically, concave, ornamented with two nodular spiral ribs; lowest spiral rib stronger, situated at angulation, forming basal edge; base convex, ornamented by several evenly spaced nodular spiral ribs, narrowly phaneromphalous; aperture unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F24156DFF017933FC40FE72" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8BA336BD4F24156DFF017933FC40FE72" blockId="19.[151,1437,153,503]" pageId="19" pageNumber="20">
<emphasis id="B968EAAF4F24156DFF017933FEF0FEE2" bold="true" box="[199,315,333,359]" pageId="19" pageNumber="20">Remarks.</emphasis>
We tentatively assign the studied specimens to
<taxonomicName id="4C1C4D3E4F24156DFC987933FBE5FEE2" authorityName="Mansuy" authorityYear="1912" box="[862,1070,333,359]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="pagoda">
<emphasis id="B968EAAF4F24156DFC987933FBE5FEE2" box="[862,1070,333,359]" italics="true" pageId="19" pageNumber="20">Worthenia pagoda</emphasis>
</taxonomicName>
, based on the whorl profile, the presence of three nodular spiral ribs on the upper whorl face and the nodular selenizone. Species with a similar whorl profile belonging to
<taxonomicName id="4C1C4D3E4F24156DFE0C79EBFDF5FE2A" authorityName="de Koninck" authorityYear="1883" box="[458,574,405,431]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B968EAAF4F24156DFE0C79EBFDF5FE2A" box="[458,574,405,431]" italics="true" pageId="19" pageNumber="20">Worthenia</emphasis>
</taxonomicName>
are more common in the Triassic e.g.,
<taxonomicName id="4C1C4D3E4F24156DFC3B79E8FA96FE2A" authority="Mansuy, 1913 b" authorityName="Mansuy" authorityYear="1913" box="[1021,1373,405,431]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="annamensis">
<emphasis id="B968EAAF4F24156DFC3B79E8FBDFFE2A" box="[1021,1044,406,431]" italics="true" pageId="19" pageNumber="20">W</emphasis>
.
<emphasis id="B968EAAF4F24156DFBE579E8FB6CFE2A" box="[1059,1191,406,431]" italics="true" pageId="19" pageNumber="20">annamensis</emphasis>
<bibRefCitation id="EF8D4B4C4F24156DFB7679EBFA96FE2A" author="Mansuy" box="[1200,1373,405,431]" firstAuthor="Mansuy" pageId="19" pageNumber="20" pagination="1 - 48" refId="ref26054" refString="Mansuy, H. (1913 b) Paleontologie de L'Annam et du Tonkin. Memoires du Service Geologique de L`Indochine, 2 (3), 1 - 48." type="journal article" year="1913" yearSuffix="b">
Mansuy, 1913
<emphasis id="B968EAAF4F24156DFA8979EBFA96FE2A" box="[1359,1373,405,431]" italics="true" pageId="19" pageNumber="20">b</emphasis>
</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F30B762D4F24156DFF5179C7FF22FE56" box="[151,233,441,467]" name="Vietnam" pageId="19" pageNumber="20">Tonkin</collectingCountry>
,
<collectingCountry id="F30B762D4F24156DFF3279C7FE98FE56" box="[244,339,441,467]" name="Vietnam" pageId="19" pageNumber="20">Vietnam</collectingCountry>
but are very rare in the Permian.
<emphasis id="B968EAAF4F24156DFD0979C4FD2DFE56" box="[719,742,442,467]" italics="true" pageId="19" pageNumber="20">W</emphasis>
.?
<emphasis id="B968EAAF4F24156DFCC779C7FC82FE56" box="[769,841,441,467]" italics="true" pageId="19" pageNumber="20">psiche</emphasis>
(
<bibRefCitation id="EF8D4B4C4F24156DFC9E79C7FBE6FE56" author="Gemmellaro" box="[856,1069,441,467]" firstAuthor="Gemmellaro" pageId="19" pageNumber="20" pagination="97 - 182" refId="ref24122" refString="Gemmellaro, G. G. (1889) La Fauna dei Calcari Con Fusulina della Valle del Fiume Sosio nella Provincia di Palermo, Fasc 2. Nautiloidea e Gastropoda. Giornale di Scienze Naturali e Economiche, 20, 97 - 182. https: // doi. org / 10.5962 / bhl. title. 10774" type="journal article" year="1889">Gemmellaro, 1889</bibRefCitation>
) from the Permian of
<collectingCountry id="F30B762D4F24156DFAEE79C7FA90FE56" box="[1320,1371,441,467]" name="Italy" pageId="19" pageNumber="20">Italy</collectingCountry>
has a similar shape but have only two spiral ribs on the upper whorl face.
</paragraph>
</subSubSection>
</treatment>
</document>