232 lines
41 KiB
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232 lines
41 KiB
XML
<document id="26847B591FF83256AC62D89E6FAAEBD3" ID-CLB-Dataset="21928" ID-DOI="10.11646/zootaxa.4779.4.1" ID-GBIF-Dataset="d48aecc5-2a44-491e-802f-4556f052a7a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3839329" ID-ZooBank="861A7289-DD34-4174-B6B2-783FBEC8DF36" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590137887931" checkinUser="plazi" docAuthor="Kluge, Nikita J. & Suttinun, Chanaporn" docDate="2020" docId="03B587A2F203FF91ECABFD1E0FBDF965" docLanguage="en" docName="zootaxa.4779.4.1.pdf" docOrigin="Zootaxa 4779 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Indocloeon Muller-Liebenau 1982, s. str." docType="treatment" docVersion="6" lastPageNumber="455" masterDocId="FF8CFFDAF207FF95EC3CFFC90C6FFFB9" masterDocTitle="Review of the Oriental genus Indocloeon Müller-Liebenau 1982 (Ephemeroptera: Baetidae) with descriptions of two new species" masterLastPageNumber="484" masterPageNumber="451" pageNumber="455" updateTime="1698840818980" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E778EDAF4CD54C48B8B3B09694DC06C3">Review of the Oriental genus Indocloeon Müller-Liebenau 1982 (Ephemeroptera: Baetidae) with descriptions of two new species</mods:title>
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<mods:namePart id="11816A9E04EB1AED1775ECE8DEB2DF31">Kluge, Nikita J.</mods:namePart>
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<mods:affiliation id="83B70D6FA1418108AD761D19DA3E799A">Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia Website: http: // www. insecta. bio. spbu. ru. n. kluge @ spbu. ru; https: // orcid. org / 0000 - 0001 - 9741 - 7790</mods:affiliation>
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<mods:namePart id="17EF5C230E15A428860C59F8B82365D0">Suttinun, Chanaporn</mods:namePart>
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<mods:affiliation id="B729256CEA94C388CC1D6AE65ED03E83">Animal Systematics and Ecology Speciality Research Unit (ASESRU), Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand. chana. suttinun @ gmail. com; https: // orcid. org / 0000 - 0002 - 0380 - 1554</mods:affiliation>
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<mods:title id="E279776B1AEB4A8CC9038D3E3C55A4C6">Zootaxa</mods:title>
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<treatment id="03B587A2F203FF91ECABFD1E0FBDF965" ID-DOI="http://doi.org/10.5281/zenodo.3853344" ID-GBIF-Taxon="164248611" ID-Zenodo-Dep="3853344" LSID="urn:lsid:plazi:treatment:03B587A2F203FF91ECABFD1E0FBDF965" httpUri="http://treatment.plazi.org/id/03B587A2F203FF91ECABFD1E0FBDF965" lastPageNumber="455" pageId="4" pageNumber="455">
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<subSubSection id="C306653FF203FF91ECABFD1E0E72FCAC" pageId="4" pageNumber="455" type="nomenclature">
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<paragraph id="8BA336B4F203FF91ECABFD1E0F62FD4B" blockId="4.[151,781,727,826]" box="[151,781,727,754]" pageId="4" pageNumber="455">
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<heading id="D0EB81D8F203FF91ECABFD1E0F62FD4B" bold="true" box="[151,781,727,754]" fontSize="11" level="1" pageId="4" pageNumber="455" reason="1">
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<emphasis id="B968EAA6F203FF91ECABFD1E0F62FD4B" bold="true" box="[151,781,727,754]" pageId="4" pageNumber="455">
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Subgenus
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<taxonomicName id="4C1C4D37F203FF91ED2BFD1E0ED0FD48" ID-CoL="548R" ID-ENA="2017433" authority="Muller-Liebenau 1982" authorityName="Muller-Liebenau" authorityYear="1982" box="[279,703,727,754]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="B968EAA6F203FF91ED2BFD1E0DCFFD4B" bold="true" box="[279,416,727,754]" italics="true" pageId="4" pageNumber="455">Indocloeon</emphasis>
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<bibRefCitation id="EF8D4B45F203FF91ED9BFD1E0ED0FD48" author="Muller-Liebenau, I." box="[423,703,727,753]" pageId="4" pageNumber="455" pagination="125 - 129" refId="ref14931" refString="Muller-Liebenau, I. (1982) A new genus and species of Baetidae from Sri Lanka (Ceylon): Indocloeon primum gen. n., sp. n. (Insecta, Ephemeroptera). Aquatic Insects, 4 (3), 125 - 129. https: // doi. org / 10.1080 / 01650428209361096" type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25B57DDF203FF91EEF0FD110F62FD4B" box="[716,781,728,754]" pageId="4" pageNumber="455" sensu="stricto">s. str.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA336B4F203FF91ECABFD320E72FCAC" blockId="4.[151,781,727,826]" box="[151,541,763,790]" pageId="4" pageNumber="455">
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Hierarchical name:
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<taxonomicName id="4C1C4D37F203FF91ED48FD320D99FCAC" authorityName="Muller-Liebenau" authorityYear="1982" box="[372,502,763,789]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">Indocloeon</taxonomicName>
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/g2.
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</paragraph>
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<subSubSection id="C306653FF203FF91ECABFCD60D76FC83" box="[151,281,799,826]" pageId="4" pageNumber="455" type="description">
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<paragraph id="8BA336B4F203FF91ECABFCD60D76FC83" blockId="4.[151,781,727,826]" box="[151,281,799,826]" pageId="4" pageNumber="455">
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(
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<figureCitation id="13272A31F203FF91ECA3FCD60D7FFC83" box="[159,272,799,826]" captionStart-0="FIGURES 1–5" captionStart-1="FIGURES 6–9" captionStart-10="FIGURES 53–55" captionStart-2="FIGURES 10–14" captionStart-3="FIGURES 15–24" captionStart-4="FIGURES 25–34" captionStart-5="FIGURES 35–39" captionStart-6="FIGURES 40–41" captionStart-7="FIGURES 42–45" captionStart-8="FIGURES 46–49" captionStart-9="FIGURES 50–52" captionStartId-0="6.[151,264,1859,1884]" captionStartId-1="7.[151,264,1838,1863]" captionStartId-10="16.[151,264,1913,1938]" captionStartId-2="8.[151,264,1264,1289]" captionStartId-3="9.[151,264,1843,1868]" captionStartId-4="10.[151,264,1838,1863]" captionStartId-5="11.[151,264,1880,1905]" captionStartId-6="12.[151,264,1186,1211]" captionStartId-7="13.[151,264,1819,1844]" captionStartId-8="14.[151,264,1772,1797]" captionStartId-9="15.[151,264,1805,1830]" captionTargetBox-0="[151,1436,179,1831]" captionTargetBox-1="[152,1431,185,1802]" captionTargetBox-10="[151,1436,453,1888]" captionTargetBox-2="[151,1436,181,1239]" captionTargetBox-3="[151,1430,181,1817]" captionTargetBox-4="[151,1436,181,1812]" captionTargetBox-5="[151,1436,489,1856]" captionTargetBox-6="[151,1437,181,1161]" captionTargetBox-7="[151,1436,181,1794]" captionTargetBox-8="[151,1431,460,1743]" captionTargetBox-9="[151,1436,345,1780]" captionTargetId-0="figure@6.[151,1436,179,1831]" captionTargetId-1="figure@7.[151,1436,181,1813]" captionTargetId-10="figure@16.[151,1436,453,1888]" captionTargetId-2="figure@8.[151,1436,181,1239]" captionTargetId-3="figure@9.[151,1436,181,1817]" captionTargetId-4="figure@10.[151,1436,181,1812]" captionTargetId-5="figure@11.[151,1436,489,1856]" captionTargetId-6="figure@12.[151,1437,181,1161]" captionTargetId-7="figure@13.[151,1436,181,1794]" captionTargetId-8="figure@14.[151,1436,453,1747]" captionTargetId-9="figure@15.[151,1436,345,1780]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-10="16" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="14" captionTargetPageId-9="15" captionText-0="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." captionText-1="FIGURES 6–9. Larvae of Indocloeon (Indocloeon) secundum sp. n. 6, labrum (arc, submarginal arc of setae; s1, s2, setae between submedian seta and submarginal arc; s-med, submedian seta); 7, labial palp, dorsal view; 8, paraglossa, dorsal view (d-m, dorso-median row of setae); 9, paraglossa and glossa, ventral view (v-m, ventro-median row of setae) (6, 8–9, holotype)." captionText-10="FIGURES 53–55. Egg of Indocloeon (Indocloeon) secundum sp. n. 53, general view; 54–55, enlarged areas marked by frames on Fig. 53." captionText-2="FIGURES 10–14. Larva of Indocloeon (Indocloeon) secundum sp. n. (holotype). 10–11, claw; 12, two outer-apical setae on fore tibia; 13, tibia of fore leg; 14, tibia of middle leg." captionText-3="FIGURES 15–24. Larva of Indocloeon (Indocloeon) primum. 15–20, posterior margins of abdominal terga I, II, IV, VII, IX and X (black triangle shows median line on tergum IX); 21, abdominal tergum VI; 22, abdominal sternum III; 23, cercus; 24, scales on abdominal sternum, enlarged." captionText-4="FIGURES 25–34. Larvae of Indocloeon (Indocloeon) secundum sp. n. 25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype)." captionText-5="FIGURES 35–39. Imagines of Indocloeon (Indocloeon) primum. 35, antennae of female imago; 36, abdominal terga and sterna of male imago (genitals removed); 37, male imago; 38, abdominal terga and sterna of female imago; 39, female imago." captionText-6="FIGURES 40–41. Indocloeon (Indocloeon) secundum sp. n. (holotype). 40, larval paraprocts; 41, male imago." captionText-7="FIGURES 42–45. Indocloeon (Indocloeon) secundum sp. n., female imagines." captionText-8="FIGURES 46–49. Male genitals of Indocloeon s. str. 46, imaginal genitals of Indocloeon (Indocloeon) secundum sp. n. (holotype) (1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 47, their subimaginal exuviae; 48, gonostylus of Indocloeon (Indocloeon) primum; 49, its subimaginal exuviae." captionText-9="FIGURES 50–52. Egg of Indocloeon (Indocloeon) primum. 50, general view; 51–52, enlarged areas marked by frames on Fig. 50." figureDoi-0="http://doi.org/10.5281/zenodo.3839331" figureDoi-1="http://doi.org/10.5281/zenodo.3839333" figureDoi-10="http://doi.org/10.5281/zenodo.3839351" figureDoi-2="http://doi.org/10.5281/zenodo.3839335" figureDoi-3="http://doi.org/10.5281/zenodo.3839337" figureDoi-4="http://doi.org/10.5281/zenodo.3839339" figureDoi-5="http://doi.org/10.5281/zenodo.3839341" figureDoi-6="http://doi.org/10.5281/zenodo.3839343" figureDoi-7="http://doi.org/10.5281/zenodo.3839345" figureDoi-8="http://doi.org/10.5281/zenodo.3839347" figureDoi-9="http://doi.org/10.5281/zenodo.3839349" httpUri-0="https://zenodo.org/record/3839331/files/figure.png" httpUri-1="https://zenodo.org/record/3839333/files/figure.png" httpUri-10="https://zenodo.org/record/3839351/files/figure.png" httpUri-2="https://zenodo.org/record/3839335/files/figure.png" httpUri-3="https://zenodo.org/record/3839337/files/figure.png" httpUri-4="https://zenodo.org/record/3839339/files/figure.png" httpUri-5="https://zenodo.org/record/3839341/files/figure.png" httpUri-6="https://zenodo.org/record/3839343/files/figure.png" httpUri-7="https://zenodo.org/record/3839345/files/figure.png" httpUri-8="https://zenodo.org/record/3839347/files/figure.png" httpUri-9="https://zenodo.org/record/3839349/files/figure.png" pageId="4" pageNumber="455">Figs 1–55</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C306653FF203FF91ECABFCAD0E8AFCC5" box="[151,741,867,892]" pageId="4" pageNumber="455" type="type_taxon">
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<paragraph id="8BA336B4F203FF91ECABFCAD0E8AFCC5" blockId="4.[151,741,867,892]" box="[151,741,867,892]" pageId="4" pageNumber="455">
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<emphasis id="B968EAA6F203FF91ECABFCAD0D49FCC5" bold="true" box="[151,294,867,892]" pageId="4" pageNumber="455">
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<typeStatus id="54A78816F203FF91ECABFCAD0CA3FCC5" box="[151,204,868,892]" pageId="4" pageNumber="455">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C1C4D37F203FF91ED10FCAD0E8EFCC5" ID-CoL="3PN48" authority="Muller-Liebenau 1982" authorityName="Muller-Liebenau" authorityYear="1982" box="[300,737,867,892]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="primum">
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<emphasis id="B968EAA6F203FF91ED10FCAD0D9CFCC2" box="[300,499,868,892]" italics="true" pageId="4" pageNumber="455">Indocloeon primum</emphasis>
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<bibRefCitation id="EF8D4B45F203FF91EDC5FCAA0E8EFCC5" author="Muller-Liebenau, I." box="[505,737,867,892]" pageId="4" pageNumber="455" pagination="125 - 129" refId="ref14931" refString="Muller-Liebenau, I. (1982) A new genus and species of Baetidae from Sri Lanka (Ceylon): Indocloeon primum gen. n., sp. n. (Insecta, Ephemeroptera). Aquatic Insects, 4 (3), 125 - 129. https: // doi. org / 10.1080 / 01650428209361096" type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C306653FF203FF91ECABFC600FBDF965" pageId="4" pageNumber="455" type="diagnosis">
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<paragraph id="8BA336B4F203FF91ECABFC600F4DFA95" blockId="4.[151,1437,937,1757]" pageId="4" pageNumber="455">
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<emphasis id="B968EAA6F203FF91ECABFC600DE2FC7D" bold="true" box="[151,397,937,965]" pageId="4" pageNumber="455">
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Diagnosis.
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<emphasis id="B968EAA6F203FF91ED25FC620D0AFC7C" bold="true" box="[281,357,939,965]" italics="true" pageId="4" pageNumber="455">Larva.</emphasis>
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(1)
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</emphasis>
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Labrum with 2 paired setae between submedian setae and submarginal arc of setae (
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<figureCitation id="13272A31F203FF91E96DFC6309F9FC7D" box="[1361,1430,938,964]" captionStart="FIGURES 6–9" captionStartId="7.[151,264,1838,1863]" captionTargetBox="[152,1431,185,1802]" captionTargetId="figure@7.[151,1436,181,1813]" captionTargetPageId="7" captionText="FIGURES 6–9. Larvae of Indocloeon (Indocloeon) secundum sp. n. 6, labrum (arc, submarginal arc of setae; s1, s2, setae between submedian seta and submarginal arc; s-med, submedian seta); 7, labial palp, dorsal view; 8, paraglossa, dorsal view (d-m, dorso-median row of setae); 9, paraglossa and glossa, ventral view (v-m, ventro-median row of setae) (6, 8–9, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839333" httpUri="https://zenodo.org/record/3839333/files/figure.png" pageId="4" pageNumber="455">Fig. 6</figureCitation>
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;
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<bibRefCitation id="EF8D4B45F203FF91ECABFC070DF8FC51" author="Muller-Liebenau, I." box="[151,407,974,1000]" pageId="4" pageNumber="455" pagination="125 - 129" refId="ref14931" refString="Muller-Liebenau, I. (1982) A new genus and species of Baetidae from Sri Lanka (Ceylon): Indocloeon primum gen. n., sp. n. (Insecta, Ephemeroptera). Aquatic Insects, 4 (3), 125 - 129. https: // doi. org / 10.1080 / 01650428209361096" type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
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: fig. 1a).
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<emphasis id="B968EAA6F203FF91EE3CFC040E4EFC51" bold="true" box="[512,545,973,1000]" pageId="4" pageNumber="455">(2)</emphasis>
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Incisors of left and right mandibles terminated by 3 denticles, 2nd denticle longest (
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<figureCitation id="13272A31F203FF91ECEEFC3B0D59FBB5" box="[210,310,1010,1036]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="4" pageNumber="455">Figs 1–2</figureCitation>
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;
|
||
<bibRefCitation id="EF8D4B45F203FF91ED7EFC3B0E2EFBB5" author="Muller-Liebenau, I." box="[322,577,1010,1036]" pageId="4" pageNumber="455" pagination="125 - 129" refId="ref14931" refString="Muller-Liebenau, I. (1982) A new genus and species of Baetidae from Sri Lanka (Ceylon): Indocloeon primum gen. n., sp. n. (Insecta, Ephemeroptera). Aquatic Insects, 4 (3), 125 - 129. https: // doi. org / 10.1080 / 01650428209361096" type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
|
||
: fig. 1h).
|
||
<emphasis id="B968EAA6F203FF91EE95FC380EA5FBB5" bold="true" box="[681,714,1009,1036]" pageId="4" pageNumber="455">(3)</emphasis>
|
||
Left mandible without setae proximad of prostheca (
|
||
<figureCitation id="13272A31F203FF91E929FC3B0938FBB5" box="[1301,1367,1010,1036]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="4" pageNumber="455">Fig. 1</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF8D4B45F203FF91E958FC3B0D07FB89" author="Muller-Liebenau, I." pageId="4" pageNumber="455" pagination="125 - 129" refId="ref14931" refString="Muller-Liebenau, I. (1982) A new genus and species of Baetidae from Sri Lanka (Ceylon): Indocloeon primum gen. n., sp. n. (Insecta, Ephemeroptera). Aquatic Insects, 4 (3), 125 - 129. https: // doi. org / 10.1080 / 01650428209361096" type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
|
||
: fig. 1h).
|
||
<emphasis id="B968EAA6F203FF91EDEFFBDC0D9BFB89" bold="true" box="[467,500,1045,1072]" pageId="4" pageNumber="455">(4)</emphasis>
|
||
Mola of left mandible with seta(e) on proximal angle (as well as on right mandible) (
|
||
<figureCitation id="13272A31F203FF91EC9CFBF30C8CFBED" box="[160,227,1082,1108]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="4" pageNumber="455">Fig. 1</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF8D4B45F203FF91ECCCFBF30D9FFBED" author="Muller-Liebenau, I." box="[240,496,1082,1108]" pageId="4" pageNumber="455" pagination="125 - 129" refId="ref14931" refString="Muller-Liebenau, I. (1982) A new genus and species of Baetidae from Sri Lanka (Ceylon): Indocloeon primum gen. n., sp. n. (Insecta, Ephemeroptera). Aquatic Insects, 4 (3), 125 - 129. https: // doi. org / 10.1080 / 01650428209361096" type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
|
||
: fig. 1h).
|
||
<emphasis id="B968EAA6F203FF91EE67FBF00E13FBED" bold="true" box="[603,636,1081,1108]" pageId="4" pageNumber="455">(5)</emphasis>
|
||
Maxillary palp consists of 2 subequally long segments and a very short 3rd segment (
|
||
<figureCitation id="13272A31F203FF91ED0FFB970D17FBC1" box="[307,376,1117,1144]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="4" pageNumber="455">Fig. 5</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF8D4B45F203FF91EDBAFB970EE7FBC1" author="Muller-Liebenau, I." box="[390,648,1118,1144]" pageId="4" pageNumber="455" pagination="125 - 129" refId="ref14931" refString="Muller-Liebenau, I. (1982) A new genus and species of Baetidae from Sri Lanka (Ceylon): Indocloeon primum gen. n., sp. n. (Insecta, Ephemeroptera). Aquatic Insects, 4 (3), 125 - 129. https: // doi. org / 10.1080 / 01650428209361096" type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
|
||
: fig. 1e).
|
||
<emphasis id="B968EAA6F203FF91EEC9FB940F79FBC1" bold="true" box="[757,790,1117,1144]" pageId="4" pageNumber="455">(6)</emphasis>
|
||
Paraglossae gradually narrowed from base to apex (
|
||
<figureCitation id="13272A31F203FF91E956FB970CAAFB25" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="4" pageNumber="455">Figs 3–4</figureCitation>
|
||
); dorso-median setal row consists of 2 (occasionally 3) stout setae (
|
||
<figureCitation id="13272A31F203FF91EF81FB4B086DFB25" box="[957,1026,1154,1180]" captionStart="FIGURES 6–9" captionStartId="7.[151,264,1838,1863]" captionTargetBox="[152,1431,185,1802]" captionTargetId="figure@7.[151,1436,181,1813]" captionTargetPageId="7" captionText="FIGURES 6–9. Larvae of Indocloeon (Indocloeon) secundum sp. n. 6, labrum (arc, submarginal arc of setae; s1, s2, setae between submedian seta and submarginal arc; s-med, submedian seta); 7, labial palp, dorsal view; 8, paraglossa, dorsal view (d-m, dorso-median row of setae); 9, paraglossa and glossa, ventral view (v-m, ventro-median row of setae) (6, 8–9, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839333" httpUri="https://zenodo.org/record/3839333/files/figure.png" pageId="4" pageNumber="455">Fig. 8</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF8D4B45F203FF91E833FB4B0960FB25" author="Muller-Liebenau, I." box="[1039,1295,1154,1180]" pageId="4" pageNumber="455" pagination="125 - 129" refId="ref14931" refString="Muller-Liebenau, I. (1982) A new genus and species of Baetidae from Sri Lanka (Ceylon): Indocloeon primum gen. n., sp. n. (Insecta, Ephemeroptera). Aquatic Insects, 4 (3), 125 - 129. https: // doi. org / 10.1080 / 01650428209361096" type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
|
||
: fig. 1b).
|
||
<emphasis id="B968EAA6F203FF91E940FB4809F2FB25" bold="true" box="[1404,1437,1153,1180]" pageId="4" pageNumber="455">(7)</emphasis>
|
||
Labial palp with sharp point on apico-median projection of 2nd segment (
|
||
<figureCitation id="13272A31F203FF91EFE5FB6F0850FB79" box="[985,1087,1190,1216]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="4" pageNumber="455">Figs 3, 4</figureCitation>
|
||
,
|
||
<figureCitation id="13272A31F203FF91E870FB6E0833FB78" box="[1100,1116,1191,1217]" captionStart="FIGURES 6–9" captionStartId="7.[151,264,1838,1863]" captionTargetBox="[152,1431,185,1802]" captionTargetId="figure@7.[151,1436,181,1813]" captionTargetPageId="7" captionText="FIGURES 6–9. Larvae of Indocloeon (Indocloeon) secundum sp. n. 6, labrum (arc, submarginal arc of setae; s1, s2, setae between submedian seta and submarginal arc; s-med, submedian seta); 7, labial palp, dorsal view; 8, paraglossa, dorsal view (d-m, dorso-median row of setae); 9, paraglossa and glossa, ventral view (v-m, ventro-median row of setae) (6, 8–9, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839333" httpUri="https://zenodo.org/record/3839333/files/figure.png" pageId="4" pageNumber="455">7</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF8D4B45F203FF91E856FB6F0905FB79" author="Muller-Liebenau, I." box="[1130,1386,1190,1216]" pageId="4" pageNumber="455" pagination="125 - 129" refId="ref14931" refString="Muller-Liebenau, I. (1982) A new genus and species of Baetidae from Sri Lanka (Ceylon): Indocloeon primum gen. n., sp. n. (Insecta, Ephemeroptera). Aquatic Insects, 4 (3), 125 - 129. https: // doi. org / 10.1080 / 01650428209361096" type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
|
||
: fig. 1b–c); exuviae of this segment often crumpled longitudinally, but with point retaining its shape.
|
||
<emphasis id="B968EAA6F203FF91E883FB00088FFB5D" bold="true" box="[1215,1248,1225,1252]" pageId="4" pageNumber="455">(8)</emphasis>
|
||
Tibiae of middle and hind legs have the same setation as tibia of fore leg: with one stout outer-apical seta (occasionally 2 or more such setae), without other stout setae on outer margin (
|
||
<figureCitation id="13272A31F203FF91EEFEFADB0F7AFA95" box="[706,789,1298,1324]" captionStart="FIGURES 10–14" captionStartId="8.[151,264,1264,1289]" captionTargetBox="[151,1436,181,1239]" captionTargetId="figure@8.[151,1436,181,1239]" captionTargetPageId="8" captionText="FIGURES 10–14. Larva of Indocloeon (Indocloeon) secundum sp. n. (holotype). 10–11, claw; 12, two outer-apical setae on fore tibia; 13, tibia of fore leg; 14, tibia of middle leg." figureDoi="http://doi.org/10.5281/zenodo.3839335" httpUri="https://zenodo.org/record/3839335/files/figure.png" pageId="4" pageNumber="455">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA336B4F203FF91ECFBFAFE0F1EFA05" blockId="4.[151,1437,937,1757]" pageId="4" pageNumber="455">
|
||
<emphasis id="B968EAA6F203FF91ECFBFAFE0D9CFAE9" bold="true" box="[199,499,1333,1361]" pageId="4" pageNumber="455">
|
||
<emphasis id="B968EAA6F203FF91ECFBFAFE0DAEFAE9" bold="true" box="[199,449,1333,1361]" italics="true" pageId="4" pageNumber="455">Imago and subimago</emphasis>
|
||
. (9)
|
||
</emphasis>
|
||
Antenna: scapus without hypodermal pigmented spots; proximal part of pedicellus with hypodermal pigmented spots on lateral and median sides (
|
||
<figureCitation id="13272A31F203FF91EFB5FA930FB2FACD" box="[905,989,1369,1396]" captionStart="FIGURES 35–39" captionStartId="11.[151,264,1880,1905]" captionTargetBox="[151,1436,489,1856]" captionTargetId="figure@11.[151,1436,489,1856]" captionTargetPageId="11" captionText="FIGURES 35–39. Imagines of Indocloeon (Indocloeon) primum. 35, antennae of female imago; 36, abdominal terga and sterna of male imago (genitals removed); 37, male imago; 38, abdominal terga and sterna of female imago; 39, female imago." figureDoi="http://doi.org/10.5281/zenodo.3839341" httpUri="https://zenodo.org/record/3839341/files/figure.png" pageId="4" pageNumber="455">Fig. 35</figureCitation>
|
||
).
|
||
<emphasis id="B968EAA6F203FF91EFCEFA90084EFACD" bold="true" box="[1010,1057,1369,1396]" pageId="4" pageNumber="455">(10)</emphasis>
|
||
First abdominal segment without lateral spots ventrad of metathoracic postnotum (
|
||
<figureCitation id="13272A31F203FF91EEFFFAB70F73FA21" box="[707,796,1406,1432]" captionStart="FIGURES 40–41" captionStartId="12.[151,264,1186,1211]" captionTargetBox="[151,1437,181,1161]" captionTargetId="figure@12.[151,1437,181,1161]" captionTargetPageId="12" captionText="FIGURES 40–41. Indocloeon (Indocloeon) secundum sp. n. (holotype). 40, larval paraprocts; 41, male imago." figureDoi="http://doi.org/10.5281/zenodo.3839343" httpUri="https://zenodo.org/record/3839343/files/figure.png" pageId="4" pageNumber="455">Figs 41</figureCitation>
|
||
,
|
||
<figureCitation id="13272A31F203FF91EF15FAB40F26FA21" box="[809,841,1405,1432]" captionStart="FIGURES 42–45" captionStartId="13.[151,264,1819,1844]" captionTargetBox="[151,1436,181,1794]" captionTargetId="figure@13.[151,1436,181,1794]" captionTargetPageId="13" captionText="FIGURES 42–45. Indocloeon (Indocloeon) secundum sp. n., female imagines." figureDoi="http://doi.org/10.5281/zenodo.3839345" httpUri="https://zenodo.org/record/3839345/files/figure.png" pageId="4" pageNumber="455">45</figureCitation>
|
||
).
|
||
<emphasis id="B968EAA6F203FF91EF62FAB40FE4FA21" bold="true" box="[862,907,1405,1432]" pageId="4" pageNumber="455">(11)</emphasis>
|
||
Each abdominal tergum II–VI with transverse brown or reddish stripe on posterior margin (
|
||
<figureCitation id="13272A31F203FF91EEB2FA6B0F60FA05" box="[654,783,1442,1468]" captionStart="FIGURES 35–39" captionStartId="11.[151,264,1880,1905]" captionTargetBox="[151,1436,489,1856]" captionTargetId="figure@11.[151,1436,489,1856]" captionTargetPageId="11" captionText="FIGURES 35–39. Imagines of Indocloeon (Indocloeon) primum. 35, antennae of female imago; 36, abdominal terga and sterna of male imago (genitals removed); 37, male imago; 38, abdominal terga and sterna of female imago; 39, female imago." figureDoi="http://doi.org/10.5281/zenodo.3839341" httpUri="https://zenodo.org/record/3839341/files/figure.png" pageId="4" pageNumber="455">Figs 36–39</figureCitation>
|
||
,
|
||
<figureCitation id="13272A31F203FF91EF27FA680F0AFA05" box="[795,869,1441,1468]" captionStart-0="FIGURES 40–41" captionStart-1="FIGURES 42–45" captionStartId-0="12.[151,264,1186,1211]" captionStartId-1="13.[151,264,1819,1844]" captionTargetBox-0="[151,1437,181,1161]" captionTargetBox-1="[151,1436,181,1794]" captionTargetId-0="figure@12.[151,1437,181,1161]" captionTargetId-1="figure@13.[151,1436,181,1794]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 40–41. Indocloeon (Indocloeon) secundum sp. n. (holotype). 40, larval paraprocts; 41, male imago." captionText-1="FIGURES 42–45. Indocloeon (Indocloeon) secundum sp. n., female imagines." figureDoi-0="http://doi.org/10.5281/zenodo.3839343" figureDoi-1="http://doi.org/10.5281/zenodo.3839345" httpUri-0="https://zenodo.org/record/3839343/files/figure.png" httpUri-1="https://zenodo.org/record/3839345/files/figure.png" pageId="4" pageNumber="455">41–45</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA336B4F203FF91ECFBFA0C0D57F9BD" blockId="4.[151,1437,937,1757]" pageId="4" pageNumber="455">
|
||
<emphasis id="B968EAA6F203FF91ECFBFA0C0D83FA59" bold="true" box="[199,492,1477,1504]" pageId="4" pageNumber="455">
|
||
<emphasis id="B968EAA6F203FF91ECFBFA0C0DD9FA59" bold="true" box="[199,438,1477,1504]" italics="true" pageId="4" pageNumber="455">Subimaginal cuticle.</emphasis>
|
||
(12)
|
||
</emphasis>
|
||
Postsubalar sclerite with postero-dorsal angle stretched to a long projection (
|
||
<bibRefCitation id="EF8D4B45F203FF91E96AFA0F0CBCF9BD" author="Kluge, N. J." pageId="4" pageNumber="455" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
|
||
: fig. 34).
|
||
</paragraph>
|
||
<paragraph id="8BA336B4F203FF91ECFBF9C408FAF9F5" blockId="4.[151,1437,937,1757]" pageId="4" pageNumber="455">
|
||
<emphasis id="B968EAA6F203FF91ECFBF9C40E7EF991" bold="true" box="[199,529,1549,1576]" pageId="4" pageNumber="455">
|
||
<emphasis id="B968EAA6F203FF91ECFBF9C40DB5F991" bold="true" box="[199,474,1549,1576]" italics="true" pageId="4" pageNumber="455">Genitals of male imago.</emphasis>
|
||
(13)
|
||
</emphasis>
|
||
Styliger with sharply outlined median impression (
|
||
<figureCitation id="13272A31F203FF91E86DF9C708CBF991" box="[1105,1188,1550,1576]" captionStart="FIGURES 46–49" captionStartId="14.[151,264,1772,1797]" captionTargetBox="[151,1431,460,1743]" captionTargetId="figure@14.[151,1436,453,1747]" captionTargetPageId="14" captionText="FIGURES 46–49. Male genitals of Indocloeon s. str. 46, imaginal genitals of Indocloeon (Indocloeon) secundum sp. n. (holotype) (1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 47, their subimaginal exuviae; 48, gonostylus of Indocloeon (Indocloeon) primum; 49, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.3839347" httpUri="https://zenodo.org/record/3839347/files/figure.png" pageId="4" pageNumber="455">Fig. 46</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF8D4B45F203FF91E88CF9C70957F991" author="Kluge, N. J." box="[1200,1336,1550,1576]" pageId="4" pageNumber="455" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
|
||
: fig. 32).
|
||
<emphasis id="B968EAA6F203FF91ECABF9F80CAAF9F5" bold="true" box="[151,197,1585,1612]" pageId="4" pageNumber="455">(14)</emphasis>
|
||
Gonovectes without movably articulated distal portions (
|
||
<figureCitation id="13272A31F203FF91EF76F9FB0FF2F9F5" box="[842,925,1586,1612]" captionStart="FIGURES 46–49" captionStartId="14.[151,264,1772,1797]" captionTargetBox="[151,1431,460,1743]" captionTargetId="figure@14.[151,1436,453,1747]" captionTargetPageId="14" captionText="FIGURES 46–49. Male genitals of Indocloeon s. str. 46, imaginal genitals of Indocloeon (Indocloeon) secundum sp. n. (holotype) (1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 47, their subimaginal exuviae; 48, gonostylus of Indocloeon (Indocloeon) primum; 49, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.3839347" httpUri="https://zenodo.org/record/3839347/files/figure.png" pageId="4" pageNumber="455">Fig. 46</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF8D4B45F203FF91EF95F9FB085FF9F5" author="Kluge, N. J." box="[937,1072,1586,1612]" pageId="4" pageNumber="455" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
|
||
: fig. 32).
|
||
</paragraph>
|
||
<paragraph id="8BA336B4F203FF91ECFBF99E0DE4F92D" blockId="4.[151,1437,937,1757]" pageId="4" pageNumber="455">
|
||
<emphasis id="B968EAA6F203FF91ECFBF99E0D5DF9C9" bold="true" box="[199,306,1621,1649]" pageId="4" pageNumber="455">
|
||
<emphasis id="B968EAA6F203FF91ECFBF99E0C93F9C8" bold="true" box="[199,252,1623,1649]" italics="true" pageId="4" pageNumber="455">Egg.</emphasis>
|
||
(15)
|
||
</emphasis>
|
||
One polar area (about ¼ of egg length) with net-like relief; remainder of surface with sharply outlined cavities (
|
||
<figureCitation id="13272A31F203FF91ECC0F9B30D10F92D" box="[252,383,1657,1684]" captionStart-0="FIGURES 50–52" captionStart-1="FIGURES 53–55" captionStartId-0="15.[151,264,1805,1830]" captionStartId-1="16.[151,264,1913,1938]" captionTargetBox-0="[151,1436,345,1780]" captionTargetBox-1="[151,1436,453,1888]" captionTargetId-0="figure@15.[151,1436,345,1780]" captionTargetId-1="figure@16.[151,1436,453,1888]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 50–52. Egg of Indocloeon (Indocloeon) primum. 50, general view; 51–52, enlarged areas marked by frames on Fig. 50." captionText-1="FIGURES 53–55. Egg of Indocloeon (Indocloeon) secundum sp. n. 53, general view; 54–55, enlarged areas marked by frames on Fig. 53." figureDoi-0="http://doi.org/10.5281/zenodo.3839349" figureDoi-1="http://doi.org/10.5281/zenodo.3839351" httpUri-0="https://zenodo.org/record/3839349/files/figure.png" httpUri-1="https://zenodo.org/record/3839351/files/figure.png" pageId="4" pageNumber="455">Figs 50–55</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA336B4F203FF91ECFBF9540FBDF965" blockId="4.[151,1437,937,1757]" pageId="4" pageNumber="455">
|
||
<emphasis id="B968EAA6F203FF91ECFBF9540E57F901" bold="true" box="[199,568,1693,1720]" pageId="4" pageNumber="455">Composition and distribution.</emphasis>
|
||
Two species:
|
||
<taxonomicName id="4C1C4D37F203FF91EEDEF9570906F901" authority="Muller-Liebenau 1982" authorityName="Muller-Liebenau" authorityYear="1982" box="[738,1385,1694,1721]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="primum" subGenus="Indocloeon">
|
||
<emphasis id="B968EAA6F203FF91EEDEF9570F0FF901" box="[738,864,1694,1720]" italics="true" pageId="4" pageNumber="455">Indocloeon</emphasis>
|
||
(
|
||
<emphasis id="B968EAA6F203FF91EF48F9570F9DF901" box="[884,1010,1694,1720]" italics="true" pageId="4" pageNumber="455">Indocloeon</emphasis>
|
||
)
|
||
<emphasis id="B968EAA6F203FF91E83BF9560831F901" box="[1031,1118,1695,1720]" italics="true" pageId="4" pageNumber="455">primum</emphasis>
|
||
<bibRefCitation id="EF8D4B45F203FF91E855F9570906F901" author="Muller-Liebenau, I." box="[1129,1385,1694,1720]" pageId="4" pageNumber="455" pagination="125 - 129" refId="ref14931" refString="Muller-Liebenau, I. (1982) A new genus and species of Baetidae from Sri Lanka (Ceylon): Indocloeon primum gen. n., sp. n. (Insecta, Ephemeroptera). Aquatic Insects, 4 (3), 125 - 129. https: // doi. org / 10.1080 / 01650428209361096" type="journal article" year="1982">Müller-Liebenau 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D37F203FF91ECABF90B0E4BF965" authority="Kluge & Suttinun, 2020" authorityName="Kluge & Suttinun" authorityYear="2020" box="[151,548,1730,1757]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
|
||
<emphasis id="B968EAA6F203FF91ECABF90B0D7AF965" box="[151,277,1730,1756]" italics="true" pageId="4" pageNumber="455">Indocloeon</emphasis>
|
||
(
|
||
<emphasis id="B968EAA6F203FF91ED19F90B0DCCF965" box="[293,419,1730,1756]" italics="true" pageId="4" pageNumber="455">Indocloeon</emphasis>
|
||
)
|
||
<emphasis id="B968EAA6F203FF91ED88F90B0E4BF965" box="[436,548,1730,1756]" italics="true" pageId="4" pageNumber="455">secundum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B968EAA6F203FF91EE17F90B0E05F965" bold="true" box="[555,618,1730,1756]" pageId="4" pageNumber="455">
|
||
<taxonomicNameLabel id="A25B57DDF203FF91EE17F90B0E05F965" box="[555,618,1730,1756]" pageId="4" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; both species in
|
||
<collectingCountry id="F30B7624F203FF91EF1AF90B0FFBF965" box="[806,916,1730,1756]" name="Sri Lanka" pageId="4" pageNumber="455">Sri Lanka</collectingCountry>
|
||
only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |