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<mods:title id="E4F0034EBF3A2717520492A0C04D25D8">Review of the Oriental genus Indocloeon Müller-Liebenau 1982 (Ephemeroptera: Baetidae) with descriptions of two new species</mods:title>
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<mods:namePart id="EF6896CE66631683BEFB533EDD8D989B">Kluge, Nikita J.</mods:namePart>
<mods:affiliation id="9F6BB987F3BA0F5025BB8831F3ACD595">Department of Entomology, Saint Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia Website: http: // www. insecta. bio. spbu. ru. n. kluge @ spbu. ru; https: // orcid. org / 0000 - 0001 - 9741 - 7790</mods:affiliation>
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<mods:namePart id="84AEAA78BC954308615BF0992DA8E041">Suttinun, Chanaporn</mods:namePart>
<mods:affiliation id="2D81D494C988D70FBD68AD8929697F0D">Animal Systematics and Ecology Speciality Research Unit (ASESRU), Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand. chana. suttinun @ gmail. com; https: // orcid. org / 0000 - 0002 - 0380 - 1554</mods:affiliation>
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<taxonomicName id="4C1C4D37F202FF90ECABFAA80E28FAC5" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[151,583,1377,1404]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F202FF90ECABFAA80D4FFAC5" bold="true" box="[151,288,1377,1404]" italics="true" pageId="5" pageNumber="456">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F202FF90ED0DFAA80DD5FAC5" bold="true" box="[305,442,1377,1404]" italics="true" pageId="5" pageNumber="456">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F202FF90EDF7FAA80E28FAC5" bold="true" box="[459,583,1377,1404]" italics="true" pageId="5" pageNumber="456">secundum</emphasis>
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(
<figureCitation id="13272A31F202FF90ECA3FA4C0D6BFA26" box="[159,260,1413,1439]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="5" pageNumber="456">Figs 1–3</figureCitation>
, 
<figureCitation id="13272A31F202FF90ED2CFA4C0D25FA26" box="[272,330,1413,1439]" captionStart-0="FIGURES 6–9" captionStart-1="FIGURES 10–14" captionStartId-0="7.[151,264,1838,1863]" captionStartId-1="8.[151,264,1264,1289]" captionTargetBox-0="[152,1431,185,1802]" captionTargetBox-1="[151,1436,181,1239]" captionTargetId-0="figure@7.[151,1436,181,1813]" captionTargetId-1="figure@8.[151,1436,181,1239]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 6–9. Larvae of Indocloeon (Indocloeon) secundum sp. n. 6, labrum (arc, submarginal arc of setae; s1, s2, setae between submedian seta and submarginal arc; s-med, submedian seta); 7, labial palp, dorsal view; 8, paraglossa, dorsal view (d-m, dorso-median row of setae); 9, paraglossa and glossa, ventral view (v-m, ventro-median row of setae) (6, 8–9, holotype)." captionText-1="FIGURES 10–14. Larva of Indocloeon (Indocloeon) secundum sp. n. (holotype). 10–11, claw; 12, two outer-apical setae on fore tibia; 13, tibia of fore leg; 14, tibia of middle leg." figureDoi-0="http://doi.org/10.5281/zenodo.3839333" figureDoi-1="http://doi.org/10.5281/zenodo.3839335" httpUri-0="https://zenodo.org/record/3839333/files/figure.png" httpUri-1="https://zenodo.org/record/3839335/files/figure.png" pageId="5" pageNumber="456">6–14</figureCitation>
, 
<figureCitation id="13272A31F202FF90ED6AFA4D0DF0FA26" box="[342,415,1412,1439]" captionStart="FIGURES 25–34" captionStartId="10.[151,264,1838,1863]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure@10.[151,1436,181,1812]" captionTargetPageId="10" captionText="FIGURES 25–34. Larvae of Indocloeon (Indocloeon) secundum sp. n. 25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839339" httpUri="https://zenodo.org/record/3839339/files/figure.png" pageId="5" pageNumber="456">25–34</figureCitation>
, 
<figureCitation id="13272A31F202FF90ED96FA4C0D9DFA26" box="[426,498,1413,1439]" captionStart-0="FIGURES 40–41" captionStart-1="FIGURES 42–45" captionStart-2="FIGURES 46–49" captionStartId-0="12.[151,264,1186,1211]" captionStartId-1="13.[151,264,1819,1844]" captionStartId-2="14.[151,264,1772,1797]" captionTargetBox-0="[151,1437,181,1161]" captionTargetBox-1="[151,1436,181,1794]" captionTargetBox-2="[151,1431,460,1743]" captionTargetId-0="figure@12.[151,1437,181,1161]" captionTargetId-1="figure@13.[151,1436,181,1794]" captionTargetId-2="figure@14.[151,1436,453,1747]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURES 40–41. Indocloeon (Indocloeon) secundum sp. n. (holotype). 40, larval paraprocts; 41, male imago." captionText-1="FIGURES 42–45. Indocloeon (Indocloeon) secundum sp. n., female imagines." captionText-2="FIGURES 46–49. Male genitals of Indocloeon s. str. 46, imaginal genitals of Indocloeon (Indocloeon) secundum sp. n. (holotype) (1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 47, their subimaginal exuviae; 48, gonostylus of Indocloeon (Indocloeon) primum; 49, its subimaginal exuviae." figureDoi-0="http://doi.org/10.5281/zenodo.3839343" figureDoi-1="http://doi.org/10.5281/zenodo.3839345" figureDoi-2="http://doi.org/10.5281/zenodo.3839347" httpUri-0="https://zenodo.org/record/3839343/files/figure.png" httpUri-1="https://zenodo.org/record/3839345/files/figure.png" httpUri-2="https://zenodo.org/record/3839347/files/figure.png" pageId="5" pageNumber="456">40–47</figureCitation>
, 
<figureCitation id="13272A31F202FF90EDC2FA4D0E2AFA26" box="[510,581,1412,1439]" captionStart="FIGURES 53–55" captionStartId="16.[151,264,1913,1938]" captionTargetBox="[151,1436,453,1888]" captionTargetId="figure@16.[151,1436,453,1888]" captionTargetPageId="16" captionText="FIGURES 53–55. Egg of Indocloeon (Indocloeon) secundum sp. n. 53, general view; 54–55, enlarged areas marked by frames on Fig. 53." figureDoi="http://doi.org/10.5281/zenodo.3839351" httpUri="https://zenodo.org/record/3839351/files/figure.png" pageId="5" pageNumber="456">53–55</figureCitation>
)
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<treatmentCitation id="0ABD10A5F202FF90ECABFA000E79FA58" author="Kluge, N. J." box="[151,534,1480,1505]" page="367" pageId="5" pageNumber="456" year="2012">
<taxonomicName id="4C1C4D37F202FF90ECABFA000E79FA58" ID-CoL="3PN48" authority=": Kluge 2012: 367" authorityName=": Kluge" authorityPageNumber="367" authorityYear="2012" box="[151,534,1480,1505]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="primum">
<emphasis id="B968EAA6F202FF90ECABFA000D31FA58" box="[151,350,1481,1505]" italics="true" pageId="5" pageNumber="456">Indocloeon primum</emphasis>
: 
<bibRefCitation id="EF8D4B45F202FF90ED50FA010E79FA58" author="Kluge, N. J." box="[364,534,1480,1505]" pageId="5" pageNumber="456" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012: 367</bibRefCitation>
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(partim).
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</paragraph>
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<subSubSection id="C306653FF202FF90ECABF9D90CBDF9F7" pageId="5" pageNumber="456" type="etymology">
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<emphasis id="B968EAA6F202FF90ECABF9D90D71F993" bold="true" box="[151,286,1552,1578]" pageId="5" pageNumber="456">Etymology.</emphasis>
This is the second species of 
<taxonomicName id="4C1C4D37F202FF90EE54F9D90E89F993" authorityName="Muller-Liebenau" authorityYear="1982" box="[616,742,1552,1578]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B968EAA6F202FF90EE54F9D90E89F993" box="[616,742,1552,1578]" italics="true" pageId="5" pageNumber="456">Indocloeon</emphasis>
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in 
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and the second known species of 
<taxonomicName id="4C1C4D37F202FF90E922F9D909F3F993" authorityName="Muller-Liebenau" authorityYear="1982" box="[1310,1436,1552,1578]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F202FF90E922F9D909F3F993" box="[1310,1436,1552,1578]" italics="true" pageId="5" pageNumber="456">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF202FF90ECABF9FD0CBDF9F7" box="[151,210,1588,1614]" pageId="5" pageNumber="456" sensu="stricto">s. str.</taxonomicNameLabel>
</paragraph>
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<emphasis id="B968EAA6F202FF90ECFBF99E0DC4F9CB" bold="true" box="[199,427,1623,1650]" pageId="5" pageNumber="456">Material examined</emphasis>
(
<collectionCode id="ED0DAE71F202FF90ED87F9910D80F9CB" box="[443,495,1624,1650]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="456" type="Museum">ZIN</collectionCode>
). 
<materialsCitation id="3B743CE9F202FF90EE3FF9910F8CF900" ID-GBIF-Occurrence="2622593316" collectingDate="2020-02-04" collectingDateMax="2020-02-05" collectingDateMin="2020-02-04" collectorName="N. Kluge &amp; L. Sheyko" country="Sri Lanka" county="river Seethagangula" latitude="6.826111" location="Central province" longLatPrecision="21" longitude="80.52473" municipality="Delhausie" pageId="5" pageNumber="456" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B968EAA6F202FF90EE3FF9910E15F9CB" bold="true" box="[515,634,1624,1650]" pageId="5" pageNumber="456">
<typeStatus id="54A78816F202FF90EE3FF9910E1BF9CB" box="[515,628,1624,1650]" pageId="5" pageNumber="456" type="holotype">Holotype</typeStatus>
:
</emphasis>
L-S-I 
<specimenCount id="9D1AFD3DF202FF90EEFDF9910EBBF9CB" box="[705,724,1624,1650]" pageId="5" pageNumber="456" type="male">♂</specimenCount>
{specimen [XVI] (16) 2020}, 
<collectingCountry id="F30B7624F202FF90E808F99108A3F9CB" box="[1076,1228,1624,1650]" name="Sri Lanka" pageId="5" pageNumber="456">SRI LANKA</collectingCountry>
, 
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, 
<location id="8EC3606FF202FF90ECABF9B50DE9F92C" LSID="urn:lsid:plazi:treatment:03B587A2F202FF85ECABFAA80DB0FE16:8EC3606FF202FF90ECABF9B50DE9F92C" box="[151,390,1659,1686]" country="Sri Lanka" county="river Seethagangula" latitude="6.826111" longLatPrecision="21" longitude="80.52473" municipality="Delhausie" name="Nuwara Elya District" pageId="5" pageNumber="456">Nuwara Elya District</location>
, foot of the 
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, 
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, 
<collectingCounty id="62C24E38F202FF90EF62F9B20853F92C" box="[862,1084,1659,1685]" pageId="5" pageNumber="456">river Seethagangula</collectingCounty>
close to guesthouse «Vegetable Garden», 
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, 
<date id="FFA21074F202FF90EE27F9560ECFF903" box="[539,672,1695,1722]" pageId="5" pageNumber="456" value="2020-02-04" valueMax="2020-02-05" valueMin="2020-02-04">
<collectingDate id="EFE6E99CF202FF90EE27F9560ECFF903" box="[539,672,1695,1722]" pageId="5" pageNumber="456" value="2020-02-04" valueMax="2020-02-05" valueMin="2020-02-04">4–5.II.2020</collectingDate>
</date>
, coll. 
<collectorName id="26E95362F202FF90EEDFF9690F26F900" box="[739,841,1695,1722]" pageId="5" pageNumber="456">N. Kluge</collectorName>
&amp; 
<collectorName id="26E95362F202FF90EF50F9690F8CF900" box="[876,995,1695,1722]" pageId="5" pageNumber="456">L. Sheyko</collectorName>
</materialsCitation>
: 
<materialsCitation id="3B743CE9F202FF90EFD2F9690C8EF8B8" ID-GBIF-Occurrence="2622593318" collectingDate="2011-01-24" collectingDateMax="2011-01-26" collectingDateMin="2011-01-24" collectorName="N. Kluge &amp; L. Sheyko" country="Sri Lanka" county="river Seethagangula" latitude="6.826111" location="Central province" longLatPrecision="21" longitude="80.52473" municipality="Delhausie" pageId="5" pageNumber="456" specimenCount="10" specimenCount-female="2" specimenCount-larva="5" specimenCount-male="3" typeStatus="paratype">
<emphasis id="B968EAA6F202FF90EFD2F9690800F903" bold="true" box="[1006,1135,1696,1722]" pageId="5" pageNumber="456">
<typeStatus id="54A78816F202FF90EFD2F9690805F903" box="[1006,1130,1696,1722]" pageId="5" pageNumber="456" type="paratype">Paratypes</typeStatus>
:
</emphasis>
the same locality, date and collectors, 2 L-S-I 
<specimenCount id="9D1AFD3DF202FF90ED51F90D0DEFF967" box="[365,384,1732,1758]" pageId="5" pageNumber="456" type="male">♂</specimenCount>
, 1 L-S/I 
<specimenCount id="9D1AFD3DF202FF90EDD8F90D0D97F967" box="[484,504,1732,1758]" pageId="5" pageNumber="456" type="male">♂</specimenCount>
, 2 L-S 
<specimenCount id="9D1AFD3DF202FF90EE75F90D0E32F967" box="[585,605,1732,1758]" pageId="5" pageNumber="456" type="male">♂</specimenCount>
, 8 L-S-I 
<specimenCount id="9D1AFD3DF202FF90EEF4F90D0EB9F967" box="[712,726,1732,1758]" pageId="5" pageNumber="456" type="female">♀</specimenCount>
, 
<specimenCount id="9D1AFD3DF202FF90EED8F90A0F2CF964" box="[740,835,1731,1758]" pageId="5" pageNumber="456" type="larva">5 larvae</specimenCount>
; the same locality and collectors, 
<date id="FFA21074F202FF90E8F7F90A090BF964" box="[1227,1380,1731,1757]" pageId="5" pageNumber="456" value="2011-01-24" valueMax="2011-01-26" valueMin="2011-01-24">
<collectingDate id="EFE6E99CF202FF90E8F7F90A090BF964" box="[1227,1380,1731,1757]" pageId="5" pageNumber="456" value="2011-01-24" valueMax="2011-01-26" valueMin="2011-01-24">24–26.I.2011</collectingDate>
</date>
: 
<specimenCount id="9D1AFD3DF202FF90E94CF90D09F3F967" box="[1392,1436,1732,1758]" pageId="5" pageNumber="456" type="female">1 ♀</specimenCount>
imago
</materialsCitation>
.
</paragraph>
<paragraph id="8BA336B4F202FF90ECFBF8C20818F80C" blockId="5.[151,1436,1551,1974]" pageId="5" pageNumber="456">
<emphasis id="B968EAA6F202FF90ECFBF8C20DD8F89F" bold="true" box="[199,439,1803,1830]" pageId="5" pageNumber="456">
Descriptions. 
<emphasis id="B968EAA6F202FF90ED50F8C50DDFF89F" bold="true" box="[364,432,1804,1830]" italics="true" pageId="5" pageNumber="456">Larva</emphasis>
.
</emphasis>
CUTICULAR COLOURATION. Cuticle of head mostly light brownish. Pronotum and mesonotum light brownish, with diffusive lighter and darker markings; mesonotum with diffusive darker wide longitudinal median stripe; protoptera with darker and lighter stripes along some longitudinal veins. Thoracic pleura from brown to nearly colourless, sterna lighter. Legs lighter or darker brownish. All abdominal terga I–X nearly uniformly brown. Abdominal sterna lighter than terga. Caudalii nearly uniformly brown.
</paragraph>
<caption id="DF63663CF201FF93ECABF88A0EC8F81D" ID-DOI="http://doi.org/10.5281/zenodo.3839331" ID-Zenodo-Dep="3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="6" pageNumber="457" startId="6.[151,264,1859,1884]" targetBox="[151,1436,179,1831]" targetPageId="6">
<paragraph id="8BA336B4F201FF93ECABF88A0EC8F81D" blockId="6.[151,1436,1859,1956]" pageId="6" pageNumber="457">
<emphasis id="B968EAA6F201FF93ECABF88A0D53F8E5" bold="true" box="[151,316,1859,1884]" pageId="6" pageNumber="457">FIGURES 1–5.</emphasis>
Larvae of 
<taxonomicName id="4C1C4D37F201FF93ED93F88A0E4EF8E2" authorityName="Muller-Liebenau" authorityYear="1982" box="[431,545,1859,1883]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F201FF93ED93F88A0E4EF8E2" box="[431,545,1859,1883]" italics="true" pageId="6" pageNumber="457">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF201FF93EE14F88D0E0FF8E2" box="[552,608,1860,1883]" pageId="6" pageNumber="457" sensu="stricto">s. str.</taxonomicNameLabel>
1–3, 
<taxonomicName id="4C1C4D37F201FF93EEA7F88A086BF8E2" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[667,1028,1859,1883]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F201FF93EEA7F88A0F62F8E2" box="[667,781,1859,1883]" italics="true" pageId="6" pageNumber="457">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F201FF93EF21F88A0FE0F8E2" box="[797,911,1859,1883]" italics="true" pageId="6" pageNumber="457">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F201FF93EFA3F88A086BF8E2" box="[927,1028,1859,1883]" italics="true" pageId="6" pageNumber="457">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F201FF93E830F88A0828F8E2" bold="true" box="[1036,1095,1859,1883]" pageId="6" pageNumber="457">
<taxonomicNameLabel id="A25B57DDF201FF93E830F88A0828F8E2" box="[1036,1095,1859,1883]" pageId="6" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, 
<taxonomicName id="4C1C4D37F201FF93E89CF8AE0C89F81A" authorityName=": Kluge" authorityYear="2012" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="primum" subGenus="Indocloeon">
<emphasis id="B968EAA6F201FF93E89CF8AE097DF8C6" box="[1184,1298,1895,1919]" italics="true" pageId="6" pageNumber="457">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F201FF93E91EF8AE09FBF8C6" box="[1314,1428,1895,1919]" italics="true" pageId="6" pageNumber="457">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F201FF93ECABF8450C89F81A" box="[151,230,1932,1955]" italics="true" pageId="6" pageNumber="457">primum</emphasis>
</taxonomicName>
, labium and maxilla (focus on ventral side).
</paragraph>
</caption>
<caption id="DF63663CF200FF92ECABF8E709F2F836" ID-DOI="http://doi.org/10.5281/zenodo.3839333" ID-Zenodo-Dep="3839333" httpUri="https://zenodo.org/record/3839333/files/figure.png" pageId="7" pageNumber="458" startId="7.[151,264,1838,1863]" targetBox="[152,1431,185,1802]" targetPageId="7">
<paragraph id="8BA336B4F200FF92ECABF8E709F2F836" blockId="7.[151,1437,1838,1935]" pageId="7" pageNumber="458">
<emphasis id="B968EAA6F200FF92ECABF8E70D52F8FE" bold="true" box="[151,317,1838,1863]" pageId="7" pageNumber="458">FIGURES 6–9.</emphasis>
Larvae of 
<taxonomicName id="4C1C4D37F200FF92ED8FF8E60F70F8FE" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[435,799,1839,1863]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F200FF92ED8FF8E60E4AF8FE" box="[435,549,1839,1863]" italics="true" pageId="7" pageNumber="458">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F200FF92EE0BF8E60EC6F8FE" box="[567,681,1839,1863]" italics="true" pageId="7" pageNumber="458">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F200FF92EE86F8E60F70F8FE" box="[698,799,1839,1863]" italics="true" pageId="7" pageNumber="458">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F200FF92EF14F8E60F0BF8FE" bold="true" box="[808,868,1839,1863]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="A25B57DDF200FF92EF14F8E60F0BF8FE" box="[808,868,1839,1863]" pageId="7" pageNumber="458" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
6, labrum (arc, submarginal arc of setae; s1, s2, setae between submedian seta and submarginal arc; s-med, submedian seta); 7, labial palp, dorsal view; 8, paraglossa, dorsal view (d-m, dorso-median row of setae); 9, paraglossa and glossa, ventral view (v-m, ventro-median row of setae) (6, 8–9, holotype).
</paragraph>
</caption>
<caption id="DF63663CF20FFF9DECABFB390ED6FA94" ID-DOI="http://doi.org/10.5281/zenodo.3839335" ID-Zenodo-Dep="3839335" httpUri="https://zenodo.org/record/3839335/files/figure.png" pageId="8" pageNumber="459" startId="8.[151,264,1264,1289]" targetBox="[151,1436,181,1239]" targetPageId="8">
<paragraph id="8BA336B4F20FFF9DECABFB390ED6FA94" blockId="8.[151,1436,1264,1325]" pageId="8" pageNumber="459">
<emphasis id="B968EAA6F20FFF9DECABFB390D39FAB0" bold="true" box="[151,342,1264,1289]" pageId="8" pageNumber="459">FIGURES 10–14.</emphasis>
Larva of 
<taxonomicName id="4C1C4D37F20FFF9DED81FB380F49FAB0" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[445,806,1265,1289]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F20FFF9DED81FB380E40FAB0" box="[445,559,1265,1289]" italics="true" pageId="8" pageNumber="459">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F20FFF9DEE03FB380EDEFAB0" box="[575,689,1265,1289]" italics="true" pageId="8" pageNumber="459">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F20FFF9DEEFDFB380F49FAB0" box="[705,806,1265,1289]" italics="true" pageId="8" pageNumber="459">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F20FFF9DEF12FB380F06FAB0" bold="true" box="[814,873,1265,1289]" pageId="8" pageNumber="459">
<taxonomicNameLabel id="A25B57DDF20FFF9DEF12FB380F06FAB0" box="[814,873,1265,1289]" pageId="8" pageNumber="459" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 10–11, claw; 12, two outer-apical setae on fore tibia; 13, tibia of fore leg; 14, tibia of middle leg.
</paragraph>
</caption>
<paragraph id="8BA336B4F20FFF9DECFBFA920D14FA20" blockId="8.[151,1437,1370,2046]" pageId="8" pageNumber="459">
<emphasis id="B968EAA6F20FFF9DECFBFA920D98FACC" bold="true" box="[199,503,1370,1397]" pageId="8" pageNumber="459">Hypodermal colouration.</emphasis>
Each abdominal tergum can have more or less expressed reddish transverse band on posterior margin.
</paragraph>
<paragraph id="8BA336B4F20FFF9EECFBFA6B080FFE6D" blockId="8.[151,1437,1370,2046]" lastBlockId="11.[151,1437,152,468]" lastPageId="11" lastPageNumber="462" pageId="8" pageNumber="459">
<emphasis id="B968EAA6F20FFF9DECFBFA6B0DDFFA04" bold="true" box="[199,432,1442,1469]" pageId="8" pageNumber="459">Shape and setation.</emphasis>
Antenna: anterior side of flagellum with pointed denticles at apex of each segment, without enlarged spines (as in 
<figureCitation id="13272A31F20FFF9DED82FA0E0E7DFA5B" box="[446,530,1479,1506]" captionStart="FIGURES 68–70" captionStartId="21.[151,264,1414,1439]" captionTargetBox="[151,1436,181,1388]" captionTargetId="figure@21.[151,1436,181,1388]" captionTargetPageId="21" captionText="FIGURES 68–70. Larvae of Indocloeon (Hindocloeon) continentale sp. n. 68, antenna; 69–70, maxilla, focus on ventral side (arrows show two apical setae under crown)." figureDoi="http://doi.org/10.5281/zenodo.3839359" httpUri="https://zenodo.org/record/3839359/files/figure.png" pageId="8" pageNumber="459">Fig. 68</figureCitation>
). Dorsal side of labrum with paired row of setae stretching from submedian seta to anterolateral margin and including two setae between submedian seta and submarginal setal arc (
<figureCitation id="13272A31F20FFF9DE92BFA220932F9BF" box="[1303,1373,1515,1542]" captionStart="FIGURES 6–9" captionStartId="7.[151,264,1838,1863]" captionTargetBox="[152,1431,185,1802]" captionTargetId="figure@7.[151,1436,181,1813]" captionTargetPageId="7" captionText="FIGURES 6–9. Larvae of Indocloeon (Indocloeon) secundum sp. n. 6, labrum (arc, submarginal arc of setae; s1, s2, setae between submedian seta and submarginal arc; s-med, submedian seta); 7, labial palp, dorsal view; 8, paraglossa, dorsal view (d-m, dorso-median row of setae); 9, paraglossa and glossa, ventral view (v-m, ventro-median row of setae) (6, 8–9, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839333" httpUri="https://zenodo.org/record/3839333/files/figure.png" pageId="8" pageNumber="459">Fig. 6</figureCitation>
) [see 
<taxonomicName id="4C1C4D37F20FFF9DECABF9C60D7AF990" authorityName="Muller-Liebenau" authorityYear="1982" box="[151,277,1551,1577]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F20FFF9DECABF9C60D7AF990" box="[151,277,1551,1577]" italics="true" pageId="8" pageNumber="459">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF20FFF9DED22F9D90D35F993" box="[286,346,1552,1578]" pageId="8" pageNumber="459" sensu="stricto">s. str.</taxonomicNameLabel>
(1)]. Left mandible: incisor terminated by 3 denticles with 2nd denticle longest; kinetodontium terminated by 3 denticles with 1st denticle longest; setae between prostheca and mola absent; mola with seta(e) on proximal angle (
<figureCitation id="13272A31F20FFF9DED6CF99E0DFEF9C8" box="[336,401,1623,1649]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="8" pageNumber="459">Fig. 1</figureCitation>
) [see 
<taxonomicName id="4C1C4D37F20FFF9DEDE4F99E0E39F9C8" authorityName="Muller-Liebenau" authorityYear="1982" box="[472,598,1623,1649]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F20FFF9DEDE4F99E0E39F9C8" box="[472,598,1623,1649]" italics="true" pageId="8" pageNumber="459">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF20FFF9DEE62F9910EF6F9CB" box="[606,665,1624,1650]" pageId="8" pageNumber="459" sensu="stricto">s. str.</taxonomicNameLabel>
(2), (3), (4)]. Right mandible: incisor terminated by 3 denticles with 2nd denticle longest [see 
<taxonomicName id="4C1C4D37F20FFF9DED89F9B20E5CF92C" authorityName="Muller-Liebenau" authorityYear="1982" box="[437,563,1659,1685]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F20FFF9DED89F9B20E5CF92C" box="[437,563,1659,1685]" italics="true" pageId="8" pageNumber="459">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF20FFF9DEE06F9B50E1AF92F" box="[570,629,1660,1686]" pageId="8" pageNumber="459" sensu="stricto">s. str.</taxonomicNameLabel>
(2)]; kinetodontium terminated by 3 denticles with 1st denticle longest; margin between prostheca and mola with tuft of setae; mola with seta(e) on proximal angle (
<figureCitation id="13272A31F20FFF9DE8A2F956088CF900" box="[1182,1251,1695,1721]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="8" pageNumber="459">Fig. 2</figureCitation>
). Maxillary palp long and slender, indistinctly 3-segmented, with remnant of 3rd segment very short and bent inward (as in 
<figureCitation id="13272A31F20FFF9DE96DF90A09FCF964" box="[1361,1427,1730,1757]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="8" pageNumber="459">Fig. 5</figureCitation>
) [see 
<taxonomicName id="4C1C4D37F20FFF9DECF2F92E0D23F8B8" authorityName="Muller-Liebenau" authorityYear="1982" box="[206,332,1767,1793]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F20FFF9DECF2F92E0D23F8B8" box="[206,332,1767,1793]" italics="true" pageId="8" pageNumber="459">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF20FFF9DED69F9210DFDF8BB" box="[341,402,1768,1794]" pageId="8" pageNumber="459" sensu="stricto">s. str.</taxonomicNameLabel>
(5)]. Labial palp with 2nd segment moderately long, 3rd segment short; projected medioapical angle of 2nd segment sharply pointed (
<figureCitation id="13272A31F20FFF9DEEAAF8C20EB1F89C" box="[662,734,1803,1829]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="8" pageNumber="459">Figs 3</figureCitation>
, 
<figureCitation id="13272A31F20FFF9DEED7F8C50E95F89F" box="[747,762,1804,1830]" captionStart="FIGURES 6–9" captionStartId="7.[151,264,1838,1863]" captionTargetBox="[152,1431,185,1802]" captionTargetId="figure@7.[151,1436,181,1813]" captionTargetPageId="7" captionText="FIGURES 6–9. Larvae of Indocloeon (Indocloeon) secundum sp. n. 6, labrum (arc, submarginal arc of setae; s1, s2, setae between submedian seta and submarginal arc; s-med, submedian seta); 7, labial palp, dorsal view; 8, paraglossa, dorsal view (d-m, dorso-median row of setae); 9, paraglossa and glossa, ventral view (v-m, ventro-median row of setae) (6, 8–9, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839333" httpUri="https://zenodo.org/record/3839333/files/figure.png" pageId="8" pageNumber="459">7</figureCitation>
) [see 
<taxonomicName id="4C1C4D37F20FFF9DEF02F8C20FD3F89C" authorityName="Muller-Liebenau" authorityYear="1982" box="[830,956,1803,1829]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F20FFF9DEF02F8C20FD3F89C" box="[830,956,1803,1829]" italics="true" pageId="8" pageNumber="459">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF20FFF9DEFFFF8C50F91F89F" box="[963,1022,1804,1830]" pageId="8" pageNumber="459" sensu="stricto">s. str.</taxonomicNameLabel>
(7)]. Legs slender, with femur parallel-sided; in 
<typeStatus id="54A78816F20FFF9DED1BF8E60DE3F8F0" box="[295,396,1839,1865]" pageId="8" pageNumber="459" type="holotype">holotype</typeStatus>
, length of femur: tibia: tarsus: claw (mm) of fore leg 0.83: 0.55: 0.57: 0.15, of middle legs 0.92: 0.67: 0.47: 0.15, of hind leg 0.94: 0.67: 0.47: 0.15. Stout setae on outer side of femur non-pectinate, apically blunt (as in 
<figureCitation id="13272A31F20FFF9DED41F8BE0DBFF828" box="[381,464,1911,1937]" captionStart="FIGURES 76–81" captionStartId="23.[151,264,1752,1777]" captionTargetBox="[151,1436,181,1727]" captionTargetId="figure@23.[151,1436,181,1727]" captionTargetPageId="23" captionText="FIGURES 76–81. Larvae of Indocloeon (Hindocloeon) continentale sp. n. 76–77, tibiae of fore and hind legs; 78–79, fore and hind legs; 80, claw; 81, outer margin of femur (76–79, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839363" httpUri="https://zenodo.org/record/3839363/files/figure.png" pageId="8" pageNumber="459">Fig. 81</figureCitation>
). Apex of femur normally with 2 (occasionally up to 4) stout blunt setae (
<figureCitation id="13272A31F20FFF9DE931F8BE09FFF828" box="[1293,1424,1911,1937]" captionStart="FIGURES 10–14" captionStartId="8.[151,264,1264,1289]" captionTargetBox="[151,1436,181,1239]" captionTargetId="figure@8.[151,1436,181,1239]" captionTargetPageId="8" captionText="FIGURES 10–14. Larva of Indocloeon (Indocloeon) secundum sp. n. (holotype). 10–11, claw; 12, two outer-apical setae on fore tibia; 13, tibia of fore leg; 14, tibia of middle leg." figureDoi="http://doi.org/10.5281/zenodo.3839335" httpUri="https://zenodo.org/record/3839335/files/figure.png" pageId="8" pageNumber="459">Figs 13–14</figureCitation>
). Tibiae of fore, middle and hind legs normally with one stout outer-apical seta (occasionally 2 such setae), without other stout setae on outer margin (
<figureCitation id="13272A31F20FFF9DEE20F8760EF1F860" box="[540,670,1983,2009]" captionStart="FIGURES 10–14" captionStartId="8.[151,264,1264,1289]" captionTargetBox="[151,1436,181,1239]" captionTargetId="figure@8.[151,1436,181,1239]" captionTargetPageId="8" captionText="FIGURES 10–14. Larva of Indocloeon (Indocloeon) secundum sp. n. (holotype). 10–11, claw; 12, two outer-apical setae on fore tibia; 13, tibia of fore leg; 14, tibia of middle leg." figureDoi="http://doi.org/10.5281/zenodo.3839335" httpUri="https://zenodo.org/record/3839335/files/figure.png" pageId="8" pageNumber="459">Figs 12–14</figureCitation>
) [see 
<taxonomicName id="4C1C4D37F20FFF9DEEDFF8760F0EF860" authorityName="Muller-Liebenau" authorityYear="1982" box="[739,865,1983,2009]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F20FFF9DEEDFF8760F0EF860" box="[739,865,1983,2009]" italics="true" pageId="8" pageNumber="459">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF20FFF9DEF55F8090FCAF863" box="[873,933,1984,2010]" pageId="8" pageNumber="459" sensu="stricto">s. str.</taxonomicNameLabel>
(8)]. Claw with 2 rows of denticles; in each row 3–4 most distal denticles larger than others (
<figureCitation id="13272A31F20FFF9DEE8FF82A0F5AF844" box="[691,821,2019,2045]" captionStart="FIGURES 10–14" captionStartId="8.[151,264,1264,1289]" captionTargetBox="[151,1436,181,1239]" captionTargetId="figure@8.[151,1436,181,1239]" captionTargetPageId="8" captionText="FIGURES 10–14. Larva of Indocloeon (Indocloeon) secundum sp. n. (holotype). 10–11, claw; 12, two outer-apical setae on fore tibia; 13, tibia of fore leg; 14, tibia of middle leg." figureDoi="http://doi.org/10.5281/zenodo.3839335" httpUri="https://zenodo.org/record/3839335/files/figure.png" pageId="8" pageNumber="459">Figs 10–11</figureCitation>
). Hind margin of tergum I either lacks denticles or with few occasional small denticles (
<figureCitation id="13272A31F20CFF9EEDC0FF500E20FF0A" box="[508,591,152,179]" captionStart="FIGURES 25–34" captionStartId="10.[151,264,1838,1863]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure@10.[151,1436,181,1812]" captionTargetPageId="10" captionText="FIGURES 25–34. Larvae of Indocloeon (Indocloeon) secundum sp. n. 25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839339" httpUri="https://zenodo.org/record/3839339/files/figure.png" pageId="11" pageNumber="462">Fig. 25</figureCitation>
); hind margin of each tergum II–VII bears regular row of denticles fused at their proximal halves (
<figureCitation id="13272A31F20CFF9EEDAFFF740E78FF61" box="[403,535,189,216]" captionStart="FIGURES 25–34" captionStartId="10.[151,264,1838,1863]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure@10.[151,1436,181,1812]" captionTargetPageId="10" captionText="FIGURES 25–34. Larvae of Indocloeon (Indocloeon) secundum sp. n. 25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839339" httpUri="https://zenodo.org/record/3839339/files/figure.png" pageId="11" pageNumber="462">Figs 26–28</figureCitation>
); denticles on terga 
<collectionCode id="ED0DAE71F20CFF9EEEC9FF740F3EFF6E" box="[757,849,189,215]" pageId="11" pageNumber="462">VIII–IX</collectionCode>
either non-fused, or partly fused at bases; on middle of posterior margin of tergum IX (behind submedian pair of setae) denticles shorter (
<figureCitation id="13272A31F20CFF9EE869FF2808C5FF42" box="[1109,1194,225,251]" captionStart="FIGURES 25–34" captionStartId="10.[151,264,1838,1863]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure@10.[151,1436,181,1812]" captionTargetPageId="10" captionText="FIGURES 25–34. Larvae of Indocloeon (Indocloeon) secundum sp. n. 25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839339" httpUri="https://zenodo.org/record/3839339/files/figure.png" pageId="11" pageNumber="462">Fig. 29</figureCitation>
); denticles on tergum X non-fused (
<figureCitation id="13272A31F20CFF9EED0FFECC0DE7FEA6" box="[307,392,261,287]" captionStart="FIGURES 25–34" captionStartId="10.[151,264,1838,1863]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure@10.[151,1436,181,1812]" captionTargetPageId="10" captionText="FIGURES 25–34. Larvae of Indocloeon (Indocloeon) secundum sp. n. 25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839339" httpUri="https://zenodo.org/record/3839339/files/figure.png" pageId="11" pageNumber="462">Fig. 30</figureCitation>
). Hind margins of sterna I–II lack denticles; each sternum III–VIII with fused denticles (
<figureCitation id="13272A31F20CFF9EE953FECC0CD8FEFA" captionStart="FIGURES 25–34" captionStartId="10.[151,264,1838,1863]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure@10.[151,1436,181,1812]" captionTargetPageId="10" captionText="FIGURES 25–34. Larvae of Indocloeon (Indocloeon) secundum sp. n. 25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839339" httpUri="https://zenodo.org/record/3839339/files/figure.png" pageId="11" pageNumber="462">Fig. 32</figureCitation>
); sternum IX of female bears non-fused or partly fused denticles; sternum IX of male bears non-fused or partly fused denticles at sides of protogonostyli and between protogonostyli. Scales on abdominal terga and sterna pointed, forming irregular transverse rows (
<figureCitation id="13272A31F20CFF9EEE1AFEB80E12FE32" box="[550,637,369,395]" captionStart="FIGURES 25–34" captionStartId="10.[151,264,1838,1863]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure@10.[151,1436,181,1812]" captionTargetPageId="10" captionText="FIGURES 25–34. Larvae of Indocloeon (Indocloeon) secundum sp. n. 25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839339" httpUri="https://zenodo.org/record/3839339/files/figure.png" pageId="11" pageNumber="462">Fig. 34</figureCitation>
). Each segment of cercus with regular row of short denticles and few elongated spine-like denticles, especially expressed on each 2nd or each 4th segment (
<figureCitation id="13272A31F20CFF9EE86BFE5C08C6FE16" box="[1111,1193,405,431]" captionStart="FIGURES 25–34" captionStartId="10.[151,264,1838,1863]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure@10.[151,1436,181,1812]" captionTargetPageId="10" captionText="FIGURES 25–34. Larvae of Indocloeon (Indocloeon) secundum sp. n. 25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839339" httpUri="https://zenodo.org/record/3839339/files/figure.png" pageId="11" pageNumber="462">Fig. 33</figureCitation>
). For other characters see diagnoses of 
<taxonomicName id="4C1C4D37F20CFF9EED6AFE700DBBFE6A" authorityName="Muller-Liebenau" authorityYear="1982" box="[342,468,441,467]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F20CFF9EED6AFE700DBBFE6A" box="[342,468,441,467]" italics="true" pageId="11" pageNumber="462">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF20CFF9EEDE7FE700E79FE6A" box="[475,534,441,467]" pageId="11" pageNumber="462" sensu="stricto">s. str.</taxonomicNameLabel>
above and 
<taxonomicName id="4C1C4D37F20CFF9EEEA9FE700F7CFE6A" authorityName="Muller-Liebenau" authorityYear="1982" box="[661,787,441,467]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B968EAA6F20CFF9EEEA9FE700F7CFE6A" box="[661,787,441,467]" italics="true" pageId="11" pageNumber="462">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF20CFF9EEF26FE700F2DFE6A" box="[794,834,441,467]" pageId="11" pageNumber="462" sensu="lato">s. l.</taxonomicNameLabel>
(
<bibRefCitation id="EF8D4B45F20CFF9EEF6DFE700FB9FE6A" author="Kluge, N. J." box="[849,982,441,467]" pageId="11" pageNumber="462" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
and above).
</paragraph>
<caption id="DF63663CF20EFF9CECABF8FA0D8BF82D" ID-DOI="http://doi.org/10.5281/zenodo.3839337" ID-Zenodo-Dep="3839337" httpUri="https://zenodo.org/record/3839337/files/figure.png" pageId="9" pageNumber="460" startId="9.[151,264,1843,1868]" targetBox="[151,1430,181,1817]" targetPageId="9">
<paragraph id="8BA336B4F20EFF9CECABF8FA0D8BF82D" blockId="9.[151,1437,1843,1940]" pageId="9" pageNumber="460">
<emphasis id="B968EAA6F20EFF9CECABF8FA0D3CF8F5" bold="true" box="[151,339,1843,1868]" pageId="9" pageNumber="460">FIGURES 15–24.</emphasis>
Larva of 
<taxonomicName id="4C1C4D37F20EFF9CED88F8FD0F6CF8F5" authorityName=": Kluge" authorityYear="2012" box="[436,771,1844,1868]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="primum" subGenus="Indocloeon">
<emphasis id="B968EAA6F20EFF9CED88F8FD0E49F8F5" box="[436,550,1844,1868]" italics="true" pageId="9" pageNumber="460">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F20EFF9CEE08F8FD0EC9F8F5" box="[564,678,1844,1868]" italics="true" pageId="9" pageNumber="460">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F20EFF9CEE88F8FC0F6CF8F5" box="[692,771,1845,1868]" italics="true" pageId="9" pageNumber="460">primum</emphasis>
</taxonomicName>
. 15–20, posterior margins of abdominal terga I, II, IV, VII, IX and X (black triangle shows median line on tergum IX); 21, abdominal tergum VI; 22, abdominal sternum III; 23, cercus; 24, scales on abdominal sternum, enlarged.
</paragraph>
</caption>
<caption id="DF63663CF20DFF9FECABF8E70F40F836" ID-DOI="http://doi.org/10.5281/zenodo.3839339" ID-Zenodo-Dep="3839339" httpUri="https://zenodo.org/record/3839339/files/figure.png" pageId="10" pageNumber="461" startId="10.[151,264,1838,1863]" targetBox="[151,1436,181,1812]" targetPageId="10">
<paragraph id="8BA336B4F20DFF9FECABF8E70F40F836" blockId="10.[151,1436,1838,1935]" pageId="10" pageNumber="461">
<emphasis id="B968EAA6F20DFF9FECABF8E70D3AF8FE" bold="true" box="[151,341,1838,1863]" pageId="10" pageNumber="461">FIGURES 25–34.</emphasis>
Larvae of 
<taxonomicName id="4C1C4D37F20DFF9FEDFAF8E70F40F8FF" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[454,815,1838,1862]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F20DFF9FEDFAF8E70E57F8FF" box="[454,568,1838,1862]" italics="true" pageId="10" pageNumber="461">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F20DFF9FEE74F8E70ED5F8FF" box="[584,698,1838,1862]" italics="true" pageId="10" pageNumber="461">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F20DFF9FEEF6F8E70F40F8FF" box="[714,815,1838,1862]" italics="true" pageId="10" pageNumber="461">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F20DFF9FEF0BF8E70F1EF8FF" bold="true" box="[823,881,1838,1862]" pageId="10" pageNumber="461">
<taxonomicNameLabel id="A25B57DDF20DFF9FEF0BF8E70F1EF8FF" box="[823,881,1838,1862]" pageId="10" pageNumber="461" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype).
</paragraph>
</caption>
<caption id="DF63663CF20CFF9EECABF8910920F82C" ID-DOI="http://doi.org/10.5281/zenodo.3839341" ID-Zenodo-Dep="3839341" httpUri="https://zenodo.org/record/3839341/files/figure.png" pageId="11" pageNumber="462" startId="11.[151,264,1880,1905]" targetBox="[151,1436,489,1856]" targetPageId="11">
<paragraph id="8BA336B4F20CFF9EECABF8910920F82C" blockId="11.[151,1436,1880,1942]" pageId="11" pageNumber="462">
<emphasis id="B968EAA6F20CFF9EECABF8910D3BF8C8" bold="true" box="[151,340,1880,1905]" pageId="11" pageNumber="462">FIGURES 35–39.</emphasis>
Imagines of 
<taxonomicName id="4C1C4D37F20CFF9EEDEAF8900F4AF8C8" authorityName=": Kluge" authorityYear="2012" box="[470,805,1881,1905]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="primum" subGenus="Indocloeon">
<emphasis id="B968EAA6F20CFF9EEDEAF8900E27F8C8" box="[470,584,1881,1905]" italics="true" pageId="11" pageNumber="462">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F20CFF9EEE6AF8900EA7F8C8" box="[598,712,1881,1905]" italics="true" pageId="11" pageNumber="462">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F20CFF9EEEEAF8930F4AF8C8" box="[726,805,1882,1905]" italics="true" pageId="11" pageNumber="462">primum</emphasis>
</taxonomicName>
. 35, antennae of female imago; 36, abdominal terga and sterna of male imago (genitals removed); 37, male imago; 38, abdominal terga and sterna of female imago; 39, female imago.
</paragraph>
</caption>
<caption id="DF63663CF20BFF99ECABFB6B0891FB02" ID-DOI="http://doi.org/10.5281/zenodo.3839343" ID-Zenodo-Dep="3839343" box="[151,1278,1186,1211]" httpUri="https://zenodo.org/record/3839343/files/figure.png" pageId="12" pageNumber="463" startId="12.[151,264,1186,1211]" targetBox="[151,1437,181,1161]" targetPageId="12">
<paragraph id="8BA336B4F20BFF99ECABFB6B0891FB02" blockId="12.[151,1278,1186,1211]" box="[151,1278,1186,1211]" pageId="12" pageNumber="463">
<emphasis id="B968EAA6F20BFF99ECABFB6B0D3BFB02" bold="true" box="[151,340,1186,1211]" pageId="12" pageNumber="463">FIGURES 40–41.</emphasis>
<taxonomicName id="4C1C4D37F20BFF99ED66FB6A0EAEFB02" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[346,705,1187,1211]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F20BFF99ED66FB6A0DA3FB02" box="[346,460,1187,1211]" italics="true" pageId="12" pageNumber="463">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F20BFF99EDE7FB6A0E22FB02" box="[475,589,1187,1211]" italics="true" pageId="12" pageNumber="463">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F20BFF99EE60FB6A0EAEFB02" box="[604,705,1187,1211]" italics="true" pageId="12" pageNumber="463">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F20BFF99EEFBFB6A0F6FFB02" bold="true" box="[711,768,1187,1211]" pageId="12" pageNumber="463">
<taxonomicNameLabel id="A25B57DDF20BFF99EEFBFB6A0F6FFB02" box="[711,768,1187,1211]" pageId="12" pageNumber="463" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 40, larval paraprocts; 41, male imago.
</paragraph>
</caption>
<paragraph id="8BA336B4F20BFF99ECFBFB200E42FAD6" blockId="12.[151,1437,1257,2004]" pageId="12" pageNumber="463">
<emphasis id="B968EAA6F20BFF99ECFBFB200D2CFABD" bold="true" box="[199,323,1257,1284]" pageId="12" pageNumber="463">
<emphasis id="B968EAA6F20BFF99ECFBFB200D53FABD" bold="true" box="[199,316,1257,1284]" italics="true" pageId="12" pageNumber="463">Subimago</emphasis>
.
</emphasis>
CUTICULAR COLOURATION. Cuticle nearly completely colourless, only margin of scutellum and femur-tiba articulations tinged with brownish (
<figureCitation id="13272A31F20BFF99EF3BFAC40F34FA91" box="[775,859,1293,1320]" captionStart="FIGURES 42–45" captionStartId="13.[151,264,1819,1844]" captionTargetBox="[151,1436,181,1794]" captionTargetId="figure@13.[151,1436,181,1794]" captionTargetPageId="13" captionText="FIGURES 42–45. Indocloeon (Indocloeon) secundum sp. n., female imagines." figureDoi="http://doi.org/10.5281/zenodo.3839345" httpUri="https://zenodo.org/record/3839345/files/figure.png" pageId="12" pageNumber="463">Fig. 42</figureCitation>
). Postsubalar sclerite colourless, with antero-dorsal angle stretched and curved and postero-dorsal angle stretched into long projection (as in 
<bibRefCitation id="EF8D4B45F20BFF99E8BDFAF80966FAF5" author="Kluge, N. J." box="[1153,1289,1329,1356]" pageId="12" pageNumber="463" pagination="361 - 376" refId="ref14683" refString="Kluge, N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012</bibRefCitation>
: fig. 34) [see 
<taxonomicName id="4C1C4D37F20BFF99ECABFA9C0D7AFAD6" authorityName="Muller-Liebenau" authorityYear="1982" box="[151,277,1365,1391]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F20BFF99ECABFA9C0D7AFAD6" box="[151,277,1365,1391]" italics="true" pageId="12" pageNumber="463">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF20BFF99ED20FA9F0D38FAC9" box="[284,343,1366,1392]" pageId="12" pageNumber="463" sensu="stricto">s. str.</taxonomicNameLabel>
(12)]. Wings light.
</paragraph>
<paragraph id="8BA336B4F20BFF99ECFBFAB30F7BFA2D" blockId="12.[151,1437,1257,2004]" box="[199,788,1401,1428]" pageId="12" pageNumber="463">
<emphasis id="B968EAA6F20BFF99ECFBFAB30D9AFA2D" bold="true" box="[199,501,1401,1428]" pageId="12" pageNumber="463">Hypodermal colouration.</emphasis>
As in imago (see below).
</paragraph>
<paragraph id="8BA336B4F20BFF99ECFBFA540DFAF8DE" blockId="12.[151,1437,1257,2004]" pageId="12" pageNumber="463">
<emphasis id="B968EAA6F20BFF99ECFBFA540D3FFA0E" bold="true" box="[199,336,1437,1464]" italics="true" pageId="12" pageNumber="463">Male imago</emphasis>
(
<figureCitation id="13272A31F20BFF99ED62FA540DDEFA01" box="[350,433,1437,1464]" captionStart="FIGURES 40–41" captionStartId="12.[151,264,1186,1211]" captionTargetBox="[151,1437,181,1161]" captionTargetId="figure@12.[151,1437,181,1161]" captionTargetPageId="12" captionText="FIGURES 40–41. Indocloeon (Indocloeon) secundum sp. n. (holotype). 40, larval paraprocts; 41, male imago." figureDoi="http://doi.org/10.5281/zenodo.3839343" httpUri="https://zenodo.org/record/3839343/files/figure.png" pageId="12" pageNumber="463">Fig. 41</figureCitation>
). Head pale (ocher or white). Antenna: scapus without hypodermal pigmented spots; proximal part of pedicellus with hypodermal pigmented spots on lateral and median sides (as in 
<figureCitation id="13272A31F20BFF99E8B1FA0808B3FA65" box="[1165,1244,1473,1500]" captionStart="FIGURES 35–39" captionStartId="11.[151,264,1880,1905]" captionTargetBox="[151,1436,489,1856]" captionTargetId="figure@11.[151,1436,489,1856]" captionTargetPageId="11" captionText="FIGURES 35–39. Imagines of Indocloeon (Indocloeon) primum. 35, antennae of female imago; 36, abdominal terga and sterna of male imago (genitals removed); 37, male imago; 38, abdominal terga and sterna of female imago; 39, female imago." figureDoi="http://doi.org/10.5281/zenodo.3839341" httpUri="https://zenodo.org/record/3839341/files/figure.png" pageId="12" pageNumber="463">Fig. 35</figureCitation>
) [see 
<taxonomicName id="4C1C4D37F20BFF99E922FA0809F3FA62" authorityName="Muller-Liebenau" authorityYear="1982" box="[1310,1436,1473,1499]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F20BFF99E922FA0809F3FA62" box="[1310,1436,1473,1499]" italics="true" pageId="12" pageNumber="463">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF20BFF99ECABFA2F0CBDF9B9" box="[151,210,1510,1536]" pageId="12" pageNumber="463" sensu="stricto">s. str.</taxonomicNameLabel>
(9)]. Turbinate eyes with facetted surface and distal part of stem pale yellowish; middle part of stem grayish, proximal part black. Thorax entirely pale (ocher or white), with few small brown markings. Legs pale ocher. In 
<typeStatus id="54A78816F20BFF99E94BF9C00C8FF9FE" pageId="12" pageNumber="463" type="holotype">holotype</typeStatus>
, length of femur: tibia: tarsomeres (mm) of fore leg 1.00: 1.25: 0.05: 0.52: 0.37: 0,20: 0.14 (in subimago 0.83: 0.80: 0.05: 0.30: 0.22: 12: 14), of middle leg 0.92: 1.00: 0.21: 0.08: 0.04: 0.14 (same in subimago), of hind leg 0.94: 0.98: 0.21: 0.08: 0.04: 0.14 (same in subimago). Wings with longitudinal veins light brownish, only junction of Sc, RA and costal brace tinged with brown, cross veins contrastingly coloured with dark brown; pterostigma with 3–6 oblique cross veins; marginal intercalaries long. Abdominal segments I–VII translucent and whitish, with dark lateral tracheal trunks; each tergum II–VII with contrasting red or brown hypodermal narrow transverse stripe on posterior margin, laterally these stripes pointed and not reaching tracheal trunks; tergum VIII reddish medially, with white lateral parts; segments IX and X whitish [see 
<taxonomicName id="4C1C4D37F20BFF99EFD3F8E00802F8FA" authorityName="Muller-Liebenau" authorityYear="1982" box="[1007,1133,1833,1859]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F20BFF99EFD3F8E00802F8FA" box="[1007,1133,1833,1859]" italics="true" pageId="12" pageNumber="463">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF20BFF99E849F8E308DDF8FD" box="[1141,1202,1834,1860]" pageId="12" pageNumber="463" sensu="stricto">s. str.</taxonomicNameLabel>
(10), (11)]. Genitals whitish. Cerci whitish.
</paragraph>
<paragraph id="8BA336B4F20BFF99ECFBF8B80F96F86D" blockId="12.[151,1437,1257,2004]" pageId="12" pageNumber="463">
Genitals (
<figureCitation id="13272A31F20BFF99ED0BF8B80DE0F835" box="[311,399,1905,1932]" captionStart="FIGURES 46–49" captionStartId="14.[151,264,1772,1797]" captionTargetBox="[151,1431,460,1743]" captionTargetId="figure@14.[151,1436,453,1747]" captionTargetPageId="14" captionText="FIGURES 46–49. Male genitals of Indocloeon s. str. 46, imaginal genitals of Indocloeon (Indocloeon) secundum sp. n. (holotype) (1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 47, their subimaginal exuviae; 48, gonostylus of Indocloeon (Indocloeon) primum; 49, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.3839347" httpUri="https://zenodo.org/record/3839347/files/figure.png" pageId="12" pageNumber="463">Fig. 46</figureCitation>
): Styliger medially with sharply outlined concavity [see 
<taxonomicName id="4C1C4D37F20BFF99E822F8B808F3F832" authorityName="Muller-Liebenau" authorityYear="1982" box="[1054,1180,1905,1931]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F20BFF99E822F8B808F3F832" box="[1054,1180,1905,1931]" italics="true" pageId="12" pageNumber="463">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF20BFF99E89AF8BB088CF835" box="[1190,1251,1906,1932]" pageId="12" pageNumber="463" sensu="stricto">s. str.</taxonomicNameLabel>
(13)]. Proximal (1st+2nd) segment of gonostylus greatly widened at base, with inner margin evenly convex. Distal (3rd) segment of gonostylus with outer side convex and inner side straight or slightly concave.
</paragraph>
<caption id="DF63663CF20AFF98ECABF8D20FD4F88D" ID-DOI="http://doi.org/10.5281/zenodo.3839345" ID-Zenodo-Dep="3839345" box="[151,955,1819,1844]" httpUri="https://zenodo.org/record/3839345/files/figure.png" pageId="13" pageNumber="464" startId="13.[151,264,1819,1844]" targetBox="[151,1436,181,1794]" targetPageId="13">
<paragraph id="8BA336B4F20AFF98ECABF8D20FD4F88D" blockId="13.[151,955,1819,1844]" box="[151,955,1819,1844]" pageId="13" pageNumber="464">
<emphasis id="B968EAA6F20AFF98ECABF8D20D3BF88D" bold="true" box="[151,340,1819,1844]" pageId="13" pageNumber="464">FIGURES 42–45.</emphasis>
<taxonomicName id="4C1C4D37F20AFF98ED66F8D50EAEF88D" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[346,705,1820,1844]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F20AFF98ED66F8D50DA3F88D" box="[346,460,1820,1844]" italics="true" pageId="13" pageNumber="464">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F20AFF98EDE7F8D50E22F88D" box="[475,589,1820,1844]" italics="true" pageId="13" pageNumber="464">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F20AFF98EE60F8D50EAEF88D" box="[604,705,1820,1844]" italics="true" pageId="13" pageNumber="464">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F20AFF98EEFBF8D50F6FF88D" bold="true" box="[711,768,1820,1844]" pageId="13" pageNumber="464">
<taxonomicNameLabel id="A25B57DDF20AFF98EEFBF8D50F6FF88D" box="[711,768,1820,1844]" pageId="13" pageNumber="464" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, female imagines.
</paragraph>
</caption>
<paragraph id="8BA336B4F20AFF9BECFBF8A808C7FEA6" blockId="13.[151,1436,1889,2025]" lastBlockId="14.[151,1437,152,432]" lastPageId="14" lastPageNumber="465" pageId="13" pageNumber="464">
<emphasis id="B968EAA6F20AFF98ECFBF8A80D01F8C5" bold="true" box="[199,366,1889,1916]" italics="true" pageId="13" pageNumber="464">Female imago</emphasis>
(
<figureCitation id="13272A31F20AFF98ED42F8AB0E6DF8C5" box="[382,514,1889,1916]" captionStart="FIGURES 42–45" captionStartId="13.[151,264,1819,1844]" captionTargetBox="[151,1436,181,1794]" captionTargetId="figure@13.[151,1436,181,1794]" captionTargetPageId="13" captionText="FIGURES 42–45. Indocloeon (Indocloeon) secundum sp. n., female imagines." figureDoi="http://doi.org/10.5281/zenodo.3839345" httpUri="https://zenodo.org/record/3839345/files/figure.png" pageId="13" pageNumber="464">Figs 42–45</figureCitation>
). Head and thorax pale (ocher or white); antennae as in male; pronotum with one pair of contrasting brown or reddish maculae. Legs pale ocher or diffusively tinged with reddish or yellowish. Wings as in male. Abdominal segments I–VII pale ocher, with dark lateral tracheal trunks and following hypodermal colouration: each tergum II–VI with contrasting red or brown narrow transverse stripe on posterior margin, laterally these stripes are pointed and do not reach tracheal trunks [see 
<taxonomicName id="4C1C4D37F209FF9BEF68FF500FBDFF0A" authorityName="Muller-Liebenau" authorityYear="1982" box="[852,978,153,179]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F209FF9BEF68FF500FBDFF0A" box="[852,978,153,179]" italics="true" pageId="14" pageNumber="465">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF209FF9BEFE6FF50087AFF0A" box="[986,1045,153,179]" pageId="14" pageNumber="465" sensu="stricto">s. str.</taxonomicNameLabel>
(11)]; each tergum III–VI, besides transverse stripe, with median longitudinal red or brown stripe, either parallel-sided (
<figureCitation id="13272A31F209FF9BE87EFF7408FBFF6E" box="[1090,1172,189,215]" captionStart="FIGURES 42–45" captionStartId="13.[151,264,1819,1844]" captionTargetBox="[151,1436,181,1794]" captionTargetId="figure@13.[151,1436,181,1794]" captionTargetPageId="13" captionText="FIGURES 42–45. Indocloeon (Indocloeon) secundum sp. n., female imagines." figureDoi="http://doi.org/10.5281/zenodo.3839345" httpUri="https://zenodo.org/record/3839345/files/figure.png" pageId="14" pageNumber="465">Fig. 42</figureCitation>
), or triangular, widened posteriorly (
<figureCitation id="13272A31F209FF9BED22FF280DCFFF42" box="[286,416,225,251]" captionStart="FIGURES 42–45" captionStartId="13.[151,264,1819,1844]" captionTargetBox="[151,1436,181,1794]" captionTargetId="figure@13.[151,1436,181,1794]" captionTargetPageId="13" captionText="FIGURES 42–45. Indocloeon (Indocloeon) secundum sp. n., female imagines." figureDoi="http://doi.org/10.5281/zenodo.3839345" httpUri="https://zenodo.org/record/3839345/files/figure.png" pageId="14" pageNumber="465">Figs 43–44</figureCitation>
); tergum VII whitish, with posterior margin narrowly bordered by red or brown; tergum VIII reddish medially, with whitish lateral parts; tergum IX whitish; tergum X ocher with reddish.
</paragraph>
<paragraph id="8BA336B4F209FF9BECFBFEE00DC0FE32" blockId="14.[151,1437,152,432]" pageId="14" pageNumber="465">
<emphasis id="B968EAA6F209FF9BECFBFEE00C92FEFA" bold="true" box="[199,253,297,323]" pageId="14" pageNumber="465">
<emphasis id="B968EAA6F209FF9BECFBFEE00C99FEFA" bold="true" box="[199,246,297,323]" italics="true" pageId="14" pageNumber="465">Egg</emphasis>
.
</emphasis>
Oval, about 
<quantity id="4CE49B51F209FF9BEDB1FEE10D9EFEFA" box="[397,497,296,323]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="465" unit="mm" value="0.15">0.15 mm</quantity>
length. Polar area bearing net-like relief [see 
<taxonomicName id="4C1C4D37F209FF9BEFCCFEE00801FEFA" authorityName="Muller-Liebenau" authorityYear="1982" box="[1008,1134,297,323]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F209FF9BEFCCFEE00801FEFA" box="[1008,1134,297,323]" italics="true" pageId="14" pageNumber="465">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF209FF9BE849FEE008DFFEFA" box="[1141,1200,297,323]" pageId="14" pageNumber="465" sensu="stricto">s. str.</taxonomicNameLabel>
15)] with cells wide, each with irregular convexities on bottom (
<figureCitation id="13272A31F209FF9BEE40FE840EA0FEDE" box="[636,719,332,359]" captionStart="FIGURES 53–55" captionStartId="16.[151,264,1913,1938]" captionTargetBox="[151,1436,453,1888]" captionTargetId="figure@16.[151,1436,453,1888]" captionTargetPageId="16" captionText="FIGURES 53–55. Egg of Indocloeon (Indocloeon) secundum sp. n. 53, general view; 54–55, enlarged areas marked by frames on Fig. 53." figureDoi="http://doi.org/10.5281/zenodo.3839351" httpUri="https://zenodo.org/record/3839351/files/figure.png" pageId="14" pageNumber="465">Fig. 55</figureCitation>
); sharply outlined cavities (see ibid.) located all over remaining part of egg (
<figureCitation id="13272A31F209FF9BED23FEB80DCCFE32" box="[287,419,368,395]" captionStart="FIGURES 53–55" captionStartId="16.[151,264,1913,1938]" captionTargetBox="[151,1436,453,1888]" captionTargetId="figure@16.[151,1436,453,1888]" captionTargetPageId="16" captionText="FIGURES 53–55. Egg of Indocloeon (Indocloeon) secundum sp. n. 53, general view; 54–55, enlarged areas marked by frames on Fig. 53." figureDoi="http://doi.org/10.5281/zenodo.3839351" httpUri="https://zenodo.org/record/3839351/files/figure.png" pageId="14" pageNumber="465">Figs 53–54</figureCitation>
).
</paragraph>
<paragraph id="8BA336B4F209FF9BECFBFE5D0F8FFE16" blockId="14.[151,1437,152,432]" box="[199,992,404,432]" pageId="14" pageNumber="465">
<emphasis id="B968EAA6F209FF9BECFBFE5D0D21FE16" bold="true" box="[199,334,404,431]" pageId="14" pageNumber="465">Dimension.</emphasis>
Fore wing length (and approximate body length) 
<quantity id="4CE49B51F209FF9BEF40FE5D0FB3FE16" box="[892,988,404,431]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="14" pageNumber="465" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
.
</paragraph>
<caption id="DF63663CF209FF9BECABF9250F7EF8C8" ID-DOI="http://doi.org/10.5281/zenodo.3839347" ID-Zenodo-Dep="3839347" httpUri="https://zenodo.org/record/3839347/files/figure.png" pageId="14" pageNumber="465" startId="14.[151,264,1772,1797]" targetBox="[151,1431,460,1743]" targetPageId="14">
<paragraph id="8BA336B4F209FF9BECABF9250F7EF8C8" blockId="14.[151,1436,1772,1905]" pageId="14" pageNumber="465">
<emphasis id="B968EAA6F209FF9BECABF9250D39F8BC" bold="true" box="[151,342,1772,1797]" pageId="14" pageNumber="465">FIGURES 46–49.</emphasis>
Male genitals of 
<taxonomicName id="4C1C4D37F209FF9BEE30F9240E11F8BC" authorityName="Muller-Liebenau" authorityYear="1982" box="[524,638,1773,1797]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B968EAA6F209FF9BEE30F9240E11F8BC" box="[524,638,1773,1797]" italics="true" pageId="14" pageNumber="465">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF209FF9BEEBAF9240ED2F8BD" box="[646,701,1773,1796]" pageId="14" pageNumber="465" sensu="stricto">s. str.</taxonomicNameLabel>
46, imaginal genitals of 
<taxonomicName id="4C1C4D37F209FF9BEF83F9240947F8BC" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[959,1320,1773,1797]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F209FF9BEF83F924085EF8BC" box="[959,1073,1773,1797]" italics="true" pageId="14" pageNumber="465">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F209FF9BE87DF92408DCF8BC" box="[1089,1203,1773,1797]" italics="true" pageId="14" pageNumber="465">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F209FF9BE8FFF9240947F8BC" box="[1219,1320,1773,1797]" italics="true" pageId="14" pageNumber="465">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F209FF9BE90CF9240905F8BC" bold="true" box="[1328,1386,1773,1797]" pageId="14" pageNumber="465">
<taxonomicNameLabel id="A25B57DDF209FF9BE90CF9240905F8BC" box="[1328,1386,1773,1797]" pageId="14" pageNumber="465" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype) (1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 47, their subimaginal exuviae; 48, gonostylus of 
<taxonomicName id="4C1C4D37F209FF9BECABF8900D87F8C8" authorityName=": Kluge" authorityYear="2012" box="[151,488,1881,1905]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="primum" subGenus="Indocloeon">
<emphasis id="B968EAA6F209FF9BECABF8900D66F8C8" box="[151,265,1881,1905]" italics="true" pageId="14" pageNumber="465">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F209FF9BED24F8900DE5F8C8" box="[280,394,1881,1905]" italics="true" pageId="14" pageNumber="465">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F209FF9BEDA5F8930D87F8C8" box="[409,488,1882,1905]" italics="true" pageId="14" pageNumber="465">primum</emphasis>
</taxonomicName>
; 49, its subimaginal exuviae.
</paragraph>
</caption>
<paragraph id="8BA336B4F208FF9AECFBFF510E6EFEFA" blockId="15.[151,1436,152,323]" pageId="15" pageNumber="466">
<emphasis id="B968EAA6F208FF9AECFBFF510D45FF0A" bold="true" box="[199,298,152,179]" pageId="15" pageNumber="466">Habitat.</emphasis>
Larvae of 
<taxonomicName id="4C1C4D37F208FF9AED9AFF530E37FF0A" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[422,600,153,179]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F208FF9AED9AFF530DD9FF0A" box="[422,438,154,179]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
(
<emphasis id="B968EAA6F208FF9AEDFBFF530DB7FF0A" box="[455,472,154,179]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
) 
<emphasis id="B968EAA6F208FF9AEDD4FF500E37FF0A" box="[488,600,153,179]" italics="true" pageId="15" pageNumber="466">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F208FF9AEE63FF500EF0FF0A" bold="true" box="[607,671,153,179]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="A25B57DDF208FF9AEE63FF500EF0FF0A" box="[607,671,153,179]" pageId="15" pageNumber="466" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
were collected in the same area as 
<taxonomicName id="4C1C4D37F208FF9AE812FF5308A7FF0A" authorityName=": Kluge" authorityYear="2012" box="[1070,1224,153,179]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="primum" subGenus="Indocloeon">
<emphasis id="B968EAA6F208FF9AE812FF530850FF0A" box="[1070,1087,154,179]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
(
<emphasis id="B968EAA6F208FF9AE873FF53080FFF0A" box="[1103,1120,154,179]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
) 
<emphasis id="B968EAA6F208FF9AE84DFF5308A7FF0A" box="[1137,1224,154,179]" italics="true" pageId="15" pageNumber="466">primum</emphasis>
</taxonomicName>
(village Delhausie near Sri Pada), but mainly in another biotope: all larvae of 
<taxonomicName id="4C1C4D37F208FF9AEF18FF770FBBFF6E" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[804,980,189,215]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F208FF9AEF18FF770F5AFF6E" box="[804,821,190,215]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
(
<emphasis id="B968EAA6F208FF9AEF78FF770F3AFF6E" box="[836,853,190,215]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
) 
<emphasis id="B968EAA6F208FF9AEF58FF740FBBFF6E" box="[868,980,189,215]" italics="true" pageId="15" pageNumber="466">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F208FF9AEFE7FF740875FF6E" bold="true" box="[987,1050,189,215]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="A25B57DDF208FF9AEFE7FF740875FF6E" box="[987,1050,189,215]" pageId="15" pageNumber="466" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
were collected in the river Seethagangula, while most larvae of 
<taxonomicName id="4C1C4D37F208FF9AEDD0FF2B0EE9FF42" authorityName=": Kluge" authorityYear="2012" box="[492,646,225,251]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="primum" subGenus="Indocloeon">
<emphasis id="B968EAA6F208FF9AEDD0FF2B0D92FF42" box="[492,509,226,251]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
(
<emphasis id="B968EAA6F208FF9AEE31FF2B0E71FF42" box="[525,542,226,251]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
) 
<emphasis id="B968EAA6F208FF9AEE13FF2B0EE9FF42" box="[559,646,226,251]" italics="true" pageId="15" pageNumber="466">primum</emphasis>
</taxonomicName>
were collected in a very small stream running from the forest through tea plantation. In the river Seethagangula, larvae of 
<taxonomicName id="4C1C4D37F208FF9AEEE0FECF0FE1FEA6" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[732,910,261,287]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F208FF9AEEE0FECF0E82FEA6" box="[732,749,262,287]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
(
<emphasis id="B968EAA6F208FF9AEEC1FECF0F62FEA6" box="[765,781,262,287]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
) 
<emphasis id="B968EAA6F208FF9AEF22FECC0FE1FEA6" box="[798,910,261,287]" italics="true" pageId="15" pageNumber="466">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F208FF9AEFA9FECC0FBBFEA6" bold="true" box="[917,980,261,287]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="A25B57DDF208FF9AEFA9FECC0FBBFEA6" box="[917,980,261,287]" pageId="15" pageNumber="466" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are found on sandy substrate in the widened places with stagnant water.
</paragraph>
<caption id="DF63663CF208FF9AECABF8C40CD7F8F3" ID-DOI="http://doi.org/10.5281/zenodo.3839349" ID-Zenodo-Dep="3839349" httpUri="https://zenodo.org/record/3839349/files/figure.png" pageId="15" pageNumber="466" startId="15.[151,264,1805,1830]" targetBox="[151,1436,345,1780]" targetPageId="15">
<paragraph id="8BA336B4F208FF9AECABF8C40CD7F8F3" blockId="15.[151,1437,1805,1866]" pageId="15" pageNumber="466">
<emphasis id="B968EAA6F208FF9AECABF8C40D3BF89F" bold="true" box="[151,340,1805,1830]" pageId="15" pageNumber="466">FIGURES 50–52.</emphasis>
Egg of 
<taxonomicName id="4C1C4D37F208FF9AED98F8C70E9BF89F" authorityName=": Kluge" authorityYear="2012" box="[420,756,1806,1830]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="primum" subGenus="Indocloeon">
<emphasis id="B968EAA6F208FF9AED98F8C70E79F89F" box="[420,534,1806,1830]" italics="true" pageId="15" pageNumber="466">Indocloeon</emphasis>
(
<emphasis id="B968EAA6F208FF9AEE18F8C70EF9F89F" box="[548,662,1806,1830]" italics="true" pageId="15" pageNumber="466">Indocloeon</emphasis>
) 
<emphasis id="B968EAA6F208FF9AEE99F8C60E9BF89F" box="[677,756,1807,1830]" italics="true" pageId="15" pageNumber="466">primum</emphasis>
</taxonomicName>
. 50, general view; 51–52, enlarged areas marked by frames on Fig. 50.
</paragraph>
</caption>
<paragraph id="8BA336B4F208FF85ECFBF8BE0880FE32" blockId="15.[151,1437,1911,2011]" lastBlockId="16.[151,1437,153,431]" lastPageId="16" lastPageNumber="467" pageId="15" pageNumber="466">
<emphasis id="B968EAA6F208FF9AECFBF8BE0D0DF82B" bold="true" box="[199,354,1911,1938]" pageId="15" pageNumber="466">Comparison.</emphasis>
In contrast to 
<taxonomicName id="4C1C4D37F208FF9AEE35F8B00ECBF82B" authorityName=": Kluge" authorityYear="2012" box="[521,676,1912,1939]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="primum" subGenus="Indocloeon">
<emphasis id="B968EAA6F208FF9AEE35F8B00E75F82B" box="[521,538,1913,1938]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
(
<emphasis id="B968EAA6F208FF9AEE17F8B00E53F82B" box="[555,572,1913,1938]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
) 
<emphasis id="B968EAA6F208FF9AEE71F8B00ECBF82B" box="[589,676,1913,1938]" italics="true" pageId="15" pageNumber="466">primum</emphasis>
</taxonomicName>
, the larva of 
<taxonomicName id="4C1C4D37F208FF9AEF02F8B00F9DF82B" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[830,1010,1912,1939]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F208FF9AEF02F8B00F20F82B" box="[830,847,1913,1938]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
(
<emphasis id="B968EAA6F208FF9AEF5CF8B00F1EF82B" box="[864,881,1913,1938]" italics="true" pageId="15" pageNumber="466">I.</emphasis>
) 
<emphasis id="B968EAA6F208FF9AEFBEF8B10F9DF82B" box="[898,1010,1912,1938]" italics="true" pageId="15" pageNumber="466">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F208FF9AEFC7F8B00853F82A" bold="true" box="[1019,1084,1913,1939]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="A25B57DDF208FF9AEFC7F8B00853F82A" box="[1019,1084,1913,1939]" pageId="15" pageNumber="466" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has less elongated glossae and paraglossae; its 2nd segment of labial palp is less elongated and more widened from base to apex (
<figureCitation id="13272A31F208FF9AE8E3F855094DF80F" box="[1247,1314,1948,1974]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1859,1884]" captionTargetBox="[151,1436,179,1831]" captionTargetId="figure@6.[151,1436,179,1831]" captionTargetPageId="6" captionText="FIGURES 1–5. Larvae of Indocloeon s. str. 1–3, Indocloeon (Indocloeon) secundum sp. n.: 1–2, left and right mandibles of holotype (arrow shows seta on proximal angle of left mola); 3, labium (focus on ventral side); 4–5, Indocloeon (Indocloeon) primum, labium and maxilla (focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.3839331" httpUri="https://zenodo.org/record/3839331/files/figure.png" pageId="15" pageNumber="466">Fig. 3</figureCitation>
); denticles on hind margins of abdominal terga are shorter and less fused together, being mostly non-fused at least on tergum IX, sometimes also on tergum VIII (
<figureCitation id="13272A31F217FF85EE06FF500ED0FF0D" box="[570,703,153,180]" captionStart="FIGURES 25–34" captionStartId="10.[151,264,1838,1863]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure@10.[151,1436,181,1812]" captionTargetPageId="10" captionText="FIGURES 25–34. Larvae of Indocloeon (Indocloeon) secundum sp. n. 25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839339" httpUri="https://zenodo.org/record/3839339/files/figure.png" pageId="16" pageNumber="467">Figs 26–30</figureCitation>
); scales on abdominal terga and sterna are pointed (
<figureCitation id="13272A31F217FF85E928FF500903FF0A" box="[1300,1388,153,179]" captionStart="FIGURES 25–34" captionStartId="10.[151,264,1838,1863]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure@10.[151,1436,181,1812]" captionTargetPageId="10" captionText="FIGURES 25–34. Larvae of Indocloeon (Indocloeon) secundum sp. n. 25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839339" httpUri="https://zenodo.org/record/3839339/files/figure.png" pageId="16" pageNumber="467">Figs 34</figureCitation>
) (in contrast to all other species of 
<taxonomicName id="4C1C4D37F217FF85EDD2FF740E03FF6E" authorityName="Muller-Liebenau" authorityYear="1982" box="[494,620,189,215]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B968EAA6F217FF85EDD2FF740E03FF6E" box="[494,620,189,215]" italics="true" pageId="16" pageNumber="467">Indocloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57DDF217FF85EE4FFF740EF4FF61" box="[627,667,189,216]" pageId="16" pageNumber="467" sensu="lato">s. l.</taxonomicNameLabel>
); cerci have elongated spine-like denticles (
<figureCitation id="13272A31F217FF85E8B5FF7408B3FF6E" box="[1161,1244,189,215]" captionStart="FIGURES 25–34" captionStartId="10.[151,264,1838,1863]" captionTargetBox="[151,1436,181,1812]" captionTargetId="figure@10.[151,1436,181,1812]" captionTargetPageId="10" captionText="FIGURES 25–34. Larvae of Indocloeon (Indocloeon) secundum sp. n. 25–30, posterior margins of abdominal terga I, II, III, VII, IX and X (black triangle shows median line on tergum IX); 31, abdominal tergum VI; 32, abdominal sternum V; 33, cercus; 34, scales on abdominal sternum, enlarged (25–30, 33, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3839339" httpUri="https://zenodo.org/record/3839339/files/figure.png" pageId="16" pageNumber="467">Fig. 33</figureCitation>
). In contrast to 
<taxonomicName id="4C1C4D37F217FF85E9B0FF770D77FF42" authorityName=": Kluge" authorityYear="2012" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="primum" subGenus="Indocloeon">
<emphasis id="B968EAA6F217FF85E9B0FF7709F2FF6E" box="[1420,1437,190,215]" italics="true" pageId="16" pageNumber="467">I.</emphasis>
(
<emphasis id="B968EAA6F217FF85EC9DFF2B0CDEFF42" box="[161,177,226,251]" italics="true" pageId="16" pageNumber="467">I.</emphasis>
) 
<emphasis id="B968EAA6F217FF85ECFDFF2B0D77FF42" box="[193,280,226,251]" italics="true" pageId="16" pageNumber="467">primum</emphasis>
</taxonomicName>
, male imago of 
<taxonomicName id="4C1C4D37F217FF85EDF3FF2B0EEFFF42" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[463,640,225,251]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F217FF85EDF3FF2B0D8FFF42" box="[463,480,226,251]" italics="true" pageId="16" pageNumber="467">I.</emphasis>
(
<emphasis id="B968EAA6F217FF85EDCCFF2B0E6FFF42" box="[496,512,226,251]" italics="true" pageId="16" pageNumber="467">I.</emphasis>
) 
<emphasis id="B968EAA6F217FF85EE2CFF280EEFFF42" box="[528,640,225,251]" italics="true" pageId="16" pageNumber="467">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F217FF85EEBBFF280EA8FF42" bold="true" box="[647,711,225,251]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="A25B57DDF217FF85EEBBFF280EA8FF42" box="[647,711,225,251]" pageId="16" pageNumber="467" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has cross veins contrastingly pigmented (
<figureCitation id="13272A31F217FF85E8A7FF280881FF42" box="[1179,1262,225,251]" captionStart="FIGURES 40–41" captionStartId="12.[151,264,1186,1211]" captionTargetBox="[151,1437,181,1161]" captionTargetId="figure@12.[151,1437,181,1161]" captionTargetPageId="12" captionText="FIGURES 40–41. Indocloeon (Indocloeon) secundum sp. n. (holotype). 40, larval paraprocts; 41, male imago." figureDoi="http://doi.org/10.5281/zenodo.3839343" httpUri="https://zenodo.org/record/3839343/files/figure.png" pageId="16" pageNumber="467">Fig. 41</figureCitation>
); female imago and subimago each have a median reddish stripe on abdominal terga III–VI (
<figureCitation id="13272A31F217FF85EFC5FECC0812FEA6" box="[1017,1149,261,287]" captionStart="FIGURES 42–45" captionStartId="13.[151,264,1819,1844]" captionTargetBox="[151,1436,181,1794]" captionTargetId="figure@13.[151,1436,181,1794]" captionTargetPageId="13" captionText="FIGURES 42–45. Indocloeon (Indocloeon) secundum sp. n., female imagines." figureDoi="http://doi.org/10.5281/zenodo.3839345" httpUri="https://zenodo.org/record/3839345/files/figure.png" pageId="16" pageNumber="467">Figs 42–44</figureCitation>
). The proximal (1st+2nd) segment of the gonostylus is more widened proximally, with its inner margin evenly convex (
<figureCitation id="13272A31F217FF85E88CFEE00940FEFD" box="[1200,1327,297,324]" captionStart="FIGURES 46–49" captionStartId="14.[151,264,1772,1797]" captionTargetBox="[151,1431,460,1743]" captionTargetId="figure@14.[151,1436,453,1747]" captionTargetPageId="14" captionText="FIGURES 46–49. Male genitals of Indocloeon s. str. 46, imaginal genitals of Indocloeon (Indocloeon) secundum sp. n. (holotype) (1+2, proximal segment of gonostylus; 3, distal segment of gonostylus; m.d, dorsal muscle of penis; m.gs, gonostylar muscle; m.gv, gonovectal muscle; m.s., median sterno-styligeral muscle); 47, their subimaginal exuviae; 48, gonostylus of Indocloeon (Indocloeon) primum; 49, its subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.3839347" httpUri="https://zenodo.org/record/3839347/files/figure.png" pageId="16" pageNumber="467">Fig. 46–47</figureCitation>
). The egg of 
<taxonomicName id="4C1C4D37F217FF85EC89FE870D09FEDE" authority="Kluge &amp; Suttinun, 2020" authorityName="Kluge &amp; Suttinun" authorityYear="2020" box="[181,358,333,359]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="secundum" status="sp. nov." subGenus="Indocloeon">
<emphasis id="B968EAA6F217FF85EC89FE870CA9FEDE" box="[181,198,334,359]" italics="true" pageId="16" pageNumber="467">I.</emphasis>
(
<emphasis id="B968EAA6F217FF85ECEAFE870C89FEDE" box="[214,230,334,359]" italics="true" pageId="16" pageNumber="467">I.</emphasis>
) 
<emphasis id="B968EAA6F217FF85ECCAFE840D09FEDE" box="[246,358,333,359]" italics="true" pageId="16" pageNumber="467">secundum</emphasis>
</taxonomicName>
<emphasis id="B968EAA6F217FF85ED51FE840DC3FEDE" bold="true" box="[365,428,333,359]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="A25B57DDF217FF85ED51FE840DC3FEDE" box="[365,428,333,359]" pageId="16" pageNumber="467" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs in the structure of the cell of the reticulate area (cells are wide, with convexities on bottom) (
<figureCitation id="13272A31F217FF85ECC3FEB80D3EFE32" box="[255,337,368,395]" captionStart="FIGURES 53–55" captionStartId="16.[151,264,1913,1938]" captionTargetBox="[151,1436,453,1888]" captionTargetId="figure@16.[151,1436,453,1888]" captionTargetPageId="16" captionText="FIGURES 53–55. Egg of Indocloeon (Indocloeon) secundum sp. n. 53, general view; 54–55, enlarged areas marked by frames on Fig. 53." figureDoi="http://doi.org/10.5281/zenodo.3839351" httpUri="https://zenodo.org/record/3839351/files/figure.png" pageId="16" pageNumber="467">Fig. 55</figureCitation>
) and by having more regular concavities on the remainder of the surface (
<figureCitation id="13272A31F217FF85E8B2FEB8088EFE32" box="[1166,1249,368,395]" captionStart="FIGURES 53–55" captionStartId="16.[151,264,1913,1938]" captionTargetBox="[151,1436,453,1888]" captionTargetId="figure@16.[151,1436,453,1888]" captionTargetPageId="16" captionText="FIGURES 53–55. Egg of Indocloeon (Indocloeon) secundum sp. n. 53, general view; 54–55, enlarged areas marked by frames on Fig. 53." figureDoi="http://doi.org/10.5281/zenodo.3839351" httpUri="https://zenodo.org/record/3839351/files/figure.png" pageId="16" pageNumber="467">Fig. 53</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C306653FF217FF85ECFBFE5D0DB0FE16" box="[199,479,404,431]" pageId="16" pageNumber="467" type="distribution">
<paragraph id="8BA336B4F217FF85ECFBFE5D0DB0FE16" blockId="16.[151,1437,153,431]" box="[199,479,404,431]" pageId="16" pageNumber="467">
<emphasis id="B968EAA6F217FF85ECFBFE5D0D0CFE16" bold="true" box="[199,355,404,431]" pageId="16" pageNumber="467">Distribution:</emphasis>
<collectingCountry id="F30B7624F217FF85ED56FE5C0DB4FE16" box="[362,475,405,431]" name="Sri Lanka" pageId="16" pageNumber="467">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>