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<document id="2E068A1A83AD8CADDCA5EB63E95A0301" ID-DOI="10.5852/ejt.2018.410" ID-GBIF-Dataset="6ec71c9e-1dcc-4135-a1e3-7389872abdcd" ID-ISSN="2118-9773" ID-Zenodo-Dep="1211198" ID-ZooBank="80659A6D-A9F2-4C90-95AF-C87C0127137C" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1522682334120" checkinUser="plazi" docAuthor="Skelton, Paul H., Swartz, Ernst R. &amp; Vreven, Emmanuel J." docDate="2018" docId="038DB87EFFD7FFC27A1BFA69FEDB27CC" docLanguage="en" docName="ejt-410_skelton_swartz_vreven.pdf.imf" docOrigin="European Journal of Taxonomy 410" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cheilobarbus Smith 1841" docType="treatment" docVersion="15" lastPageNumber="19" masterDocId="FFB4C006FFC7FFD07807FFB6FFF8227E" masterDocTitle="The identity of Barbus capensis Smith, 1841 and the generic status of southern African tetraploid cyprinids (Teleostei, Cyprinidae)" masterLastPageNumber="29" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547732810861" originalUpdateUser="yanell" pageNumber="17" updateTime="1735821858870" updateUser="ExternalLinkService">
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<mods:title id="D468B90AD4C4EC8B6B817E69BFF4F60F">The identity of Barbus capensis Smith, 1841 and the generic status of southern African tetraploid cyprinids (Teleostei, Cyprinidae)</mods:title>
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<mods:namePart id="902602681138D3D4948DA3D933367C6F">Skelton, Paul H.</mods:namePart>
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<mods:namePart id="A5C70CA9651610EE4971590E1E404827">Swartz, Ernst R.</mods:namePart>
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<mods:namePart id="044C975ABE37077A5DBA9CCC3EB7A86E">Vreven, Emmanuel J.</mods:namePart>
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<mods:title id="ABB10C1F522A24D1B3DE10B42406F211">European Journal of Taxonomy</mods:title>
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<mods:date id="7518770E27D332B818EE202333B3C912">2018</mods:date>
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<treatment id="038DB87EFFD7FFC27A1BFA69FEDB27CC" ID-DOI="http://doi.org/10.5281/zenodo.5687607" ID-GBIF-Taxon="143448358" ID-Zenodo-Dep="5687607" LSID="urn:lsid:plazi:treatment:038DB87EFFD7FFC27A1BFA69FEDB27CC" httpUri="http://treatment.plazi.org/id/038DB87EFFD7FFC27A1BFA69FEDB27CC" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" scope_family="Cyprinidae" scope_subFamily="Cyprininae" scope_tribe="Smiliogastrini">
<subSubSection id="C33E5AE3FFD7FFC07A1BFA69FBE02787" box="[540,1048,1502,1529]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B9B0968FFD7FFC07A1BFA69FBE02787" blockId="16.[540,1048,1502,1529]" box="[540,1048,1502,1529]" pageId="16" pageNumber="17">
<heading id="D0D3BE04FFD7FFC07A1BFA69FBE02787" box="[540,1048,1502,1529]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
Genus
<taxonomicName id="4C2472EBFFD7FFC07A69FA68FC502787" authority="Smith, 1841" authorityName="Smith" authorityYear="1841" box="[622,936,1502,1529]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD7FFC07A69FA68FCE92787" bold="true" box="[622,785,1502,1529]" italics="true" pageId="16" pageNumber="17">Cheilobarbus</emphasis>
<bibRefCitation id="EFB57499FFD7FFC07B1FFA68FC502787" author="Smith A." box="[792,936,1502,1529]" journalOrPublisher="London" pageId="16" pageNumber="17" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith, 1841</bibRefCitation>
</taxonomicName>
stat. nov.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33E5AE3FFD7FFC078BAF996FD7F24BD" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8B9B0968FFD7FFC078BAF996FAE82444" blockId="16.[189,1296,1568,1631]" box="[189,1296,1568,1595]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB342E46FFD7FFC078BAF996FAE82444" box="[189,1296,1568,1595]" pageId="16" pageNumber="17">
<taxonomicName id="4C2472EBFFD7FFC078BAF996FE1D2445" ID-CoL="382L" authority="Daudin, 1805: 58" authorityName="Daudin" authorityPageNumber="58" authorityYear="1805" box="[189,485,1568,1595]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD7FFC078BAF996FEEA2444" box="[189,274,1568,1594]" italics="true" pageId="16" pageNumber="17">Barbus</emphasis>
<treatmentCitation id="0A852F79FFD7FFC0791EF996FE1D2445" author="" box="[281,485,1568,1595]" page="58" pageId="16" pageNumber="17">
<bibRefCitation id="EFB57499FFD7FFC0791EF996FE452445" author="Daudin F. M." box="[281,445,1568,1595]" editor="Cuvier F." journalOrPublisher="Serie" pageId="16" pageNumber="17" pagination="58" part="1" refId="ref12911" refString="Daudin F. M. 1805. Barbus. In: Cuvier F. (ed.) Dictionnaire des Sciences naturelles. BAN - BLU. Serie 1. Tome quatrieme: 58. Levrault, Schoell &amp; Cie, Paris." title="Barbus" type="journal article" volumeTitle="Dictionnaire des Sciences naturelles. BAN - BLU" year="1805">Daudin, 1805</bibRefCitation>
: 58
</treatmentCitation>
</taxonomicName>
(in part, non-Daudin:
<taxonomicName id="4C2472EBFFD7FFC07AEAF996FAFB2444" authority="Smith, 1841" authorityName="Smith" authorityYear="1841" box="[749,1283,1568,1594]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="capensis" subGenus="Cheilobarbus">
<emphasis id="B950D57AFFD7FFC07AEAF996FB902444" box="[749,1128,1568,1594]" italics="true" pageId="16" pageNumber="17">Barbus (Cheilobarbus) capensis</emphasis>
<bibRefCitation id="EFB57499FFD7FFC07C77F996FAFB2444" author="Smith A." box="[1136,1283,1568,1594]" journalOrPublisher="London" pageId="16" pageNumber="17" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith, 1841</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9B0968FFD7FFC078BAF9F2FB3B2421" blockId="16.[189,1296,1568,1631]" box="[189,1219,1604,1631]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB342E46FFD7FFC078BAF9F2FB3B2421" box="[189,1219,1604,1631]" pageId="16" pageNumber="17">
<taxonomicName id="4C2472EBFFD7FFC078BAF9F2FD9B2420" authority="Smith, 1841" authorityName="Smith" authorityYear="1841" box="[189,611,1604,1630]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="subGenus" subGenus="Cheilobarbus">
<emphasis id="B950D57AFFD7FFC078BAF9F2FE352420" box="[189,461,1604,1630]" italics="true" pageId="16" pageNumber="17">Barbus (Cheilobarbus)</emphasis>
<bibRefCitation id="EFB57499FFD7FFC079D3F9F2FD9B2420" author="Smith A." box="[468,611,1604,1630]" journalOrPublisher="London" pageId="16" pageNumber="17" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith, 1841</bibRefCitation>
</taxonomicName>
: description of pl. X., fig.1 (applied as a subgenus).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9B0968FFD7FFC078BAF933FDFA24E1" blockId="16.[189,647,1669,1732]" box="[189,514,1669,1696]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB342E46FFD7FFC078BAF933FDFA24E1" box="[189,514,1669,1696]" pageId="16" pageNumber="17">
<taxonomicName id="4C2472EBFFD7FFC078BAF933FEEA24E1" ID-CoL="382L" authorityName="Daudin" authorityYear="1805" box="[189,274,1669,1695]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD7FFC078BAF933FEEA24E1" box="[189,274,1669,1695]" italics="true" pageId="16" pageNumber="17">Barbus</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A852F79FFD7FFC07928F933FE0724E1" author="" box="[303,511,1669,1696]" page="244" pageId="16" pageNumber="17">
<bibRefCitation id="EFB57499FFD7FFC07928F933FE3824E1" author="Jordan D. S." box="[303,448,1669,1695]" journalOrPublisher="Leland Stanford Jr. University Publications, University Series" pageId="16" pageNumber="17" publicationUrl="http://gallica.bnf.fr/ark:/12148/bpt6k97097q" refId="ref13519" refString="Jordan D. S. 1919. The Genera of Fishes, Part II, from Agassiz to Bleeker, 1833 - 1858, Twenty-Six Years, with the Accepted Type of Each. A Contribution to the Stability of Scientific Nomenclature. Leland Stanford Jr. University Publications, University Series. Available from http: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 97097 q [accessed 5 Feb. 2018]." title="The Genera of Fishes, Part II, from Agassiz to Bleeker, 1833 - 1858, Twenty-Six Years, with the Accepted Type of Each. A Contribution to the Stability of Scientific Nomenclature" type="book chapter" year="1919">Jordan 1919</bibRefCitation>
: 244
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9B0968FFD7FFC078BAF91FFD7F24BD" blockId="16.[189,647,1669,1732]" box="[189,647,1705,1732]" pageId="16" pageNumber="17">
‘
<taxonomicName id="4C2472EBFFD7FFC078C0F91FFE9724BD" authorityName="Smith" authorityYear="1841" box="[199,367,1705,1731]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD7FFC078C0F91FFE9724BD" box="[199,367,1705,1731]" italics="true" pageId="16" pageNumber="17">Pseudobarbus</emphasis>
</taxonomicName>
’ –
<bibRefCitation id="EFB57499FFD7FFC07995F91FFDAB24BA" author="Yang L. &amp; Tetsuya S. &amp; Hirt M. &amp; Pasco-Viel E. &amp; Arunachalam M. &amp; Junbing L. &amp; Wang X. &amp; Freyhof J. &amp; Saitoh K. &amp; Simons A. M. &amp; Masaki M. &amp; Shunping H. S. &amp; Mayden R. L." box="[402,595,1705,1732]" journalOrPublisher="Molecular Phylogeny and Evolution" pageId="16" pageNumber="17" pagination="97 - 116" part="85" publicationUrl="https://doi.org/10.1016/j.ympev.2015.01.014" refId="ref14973" refString="Yang L., Tetsuya S., Hirt M. V., Pasco-Viel E., Arunachalam M., Junbing L., Wang X., Freyhof J., Saitoh K., Simons A. M., Masaki M., Shunping H. S. &amp; Mayden R. L. 2015. Phylogeny and polyploidy: resolving the classification of cyprinine fishes (Teleostei: Cypriniformes). Molecular Phylogeny and Evolution 85: 97 - 116. https: // doi. org / 10.1016 / j. ympev. 2015.01.014" title="Phylogeny and polyploidy: resolving the classification of cyprinine fishes (Teleostei: Cypriniformes)" type="journal article" year="2015">
Yang
<emphasis id="B950D57AFFD7FFC079D1F91CFDE924BD" box="[470,529,1705,1731]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2015
</bibRefCitation>
: 99.
</paragraph>
</subSubSection>
<subSubSection id="C33E5AE3FFD7FFC078BAF95DFD1A254D" pageId="16" pageNumber="17" type="type_taxon">
<paragraph id="8B9B0968FFD7FFC078BAF95DFEA0257A" blockId="16.[189,344,1770,1797]" box="[189,344,1770,1797]" pageId="16" pageNumber="17">
<heading id="D0D3BE04FFD7FFC078BAF95DFEA0257A" bold="true" box="[189,344,1770,1797]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B950D57AFFD7FFC078BAF95DFEA0257A" bold="true" box="[189,344,1770,1797]" pageId="16" pageNumber="17">Type species</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B0968FFD7FFC078BAF8AFFD1A254D" blockId="16.[189,738,1817,1843]" box="[189,738,1817,1843]" pageId="16" pageNumber="17">
<taxonomicName id="4C2472EBFFD7FFC078BAF8AFFD98254D" ID-CoL="7TS8Q" authority="Smith, 1841" authorityName="Smith" authorityYear="1841" box="[189,608,1817,1843]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="capensis">
<emphasis id="B950D57AFFD7FFC078BAF8AFFE31254D" box="[189,457,1817,1843]" italics="true" pageId="16" pageNumber="17">Cheilobarbus capensis</emphasis>
<bibRefCitation id="EFB57499FFD7FFC079D7F8AFFD98254D" author="Smith A." box="[464,608,1817,1843]" journalOrPublisher="London" pageId="16" pageNumber="17" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith, 1841</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131F15EDFFD7FFC07A77F8AFFD21254D" box="[624,729,1817,1843]" captionStart="Fig. 10" captionStartId="17.[189,232,1860,1886]" captionTargetBox="[189,1399,970,1817]" captionTargetId="figure@17.[189,1399,967,1818]" captionTargetPageId="17" captionText="Fig. 10. Species representative of southernAfrican tetraploid cyprinine genera. A. Pseudobarbus afer (Peters, 1864). B. Cheilobarbus capensis Smith, 1841. C. Amatolacypris trevelyani (Günther, 1877) gen.et comb.nov. D. Sedercypris calidus(Barnard, 1938) gen.et comb.nov. E. Namaquacyprishospes (Barnard, 1938) gen. et comb. nov. Photograph credits:A–D by P.H. Skelton (SAIAB); E by R.I. Bills (SAIAB)." figureDoi="http://doi.org/10.5281/zenodo.1211218" httpUri="https://zenodo.org/record/1211218/files/figure.png" pageId="16" pageNumber="17">Fig. 10B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33E5AE3FFD7FFC078BAF8EDFA8F2592" pageId="16" pageNumber="17" type="taxon_list">
<paragraph id="8B9B0968FFD7FFC078BAF8EDFE71250B" blockId="16.[189,393,1883,1909]" box="[189,393,1883,1909]" pageId="16" pageNumber="17">
<heading id="D0D3BE04FFD7FFC078BAF8EDFE71250B" bold="true" box="[189,393,1883,1909]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B950D57AFFD7FFC078BAF8EDFE71250B" bold="true" box="[189,393,1883,1909]" pageId="16" pageNumber="17">Included species</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B0968FFD7FFC078BAF83CFA8F2592" blockId="16.[189,1399,1930,2028]" pageId="16" pageNumber="17">
<taxonomicName id="4C2472EBFFD7FFC078BAF83CFD9F25DA" ID-CoL="7TS8Q" authority="Smith, 1841" authorityName="Smith" authorityYear="1841" box="[189,615,1930,1956]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="capensis">
<emphasis id="B950D57AFFD7FFC078BAF83CFE3325DA" box="[189,459,1930,1956]" italics="true" pageId="16" pageNumber="17">Cheilobarbus capensis</emphasis>
<bibRefCitation id="EFB57499FFD7FFC079D2F83CFD9F25DA" author="Smith A." box="[469,615,1930,1956]" journalOrPublisher="London" pageId="16" pageNumber="17" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith, 1841</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131F15EDFFD7FFC07A7EF83CFD1E25DA" box="[633,742,1930,1956]" captionStart="Fig. 10" captionStartId="17.[189,232,1860,1886]" captionTargetBox="[189,1399,970,1817]" captionTargetId="figure@17.[189,1399,967,1818]" captionTargetPageId="17" captionText="Fig. 10. Species representative of southernAfrican tetraploid cyprinine genera. A. Pseudobarbus afer (Peters, 1864). B. Cheilobarbus capensis Smith, 1841. C. Amatolacypris trevelyani (Günther, 1877) gen.et comb.nov. D. Sedercypris calidus(Barnard, 1938) gen.et comb.nov. E. Namaquacyprishospes (Barnard, 1938) gen. et comb. nov. Photograph credits:A–D by P.H. Skelton (SAIAB); E by R.I. Bills (SAIAB)." figureDoi="http://doi.org/10.5281/zenodo.1211218" httpUri="https://zenodo.org/record/1211218/files/figure.png" pageId="16" pageNumber="17">Fig. 10B</figureCitation>
), known as the witvis or Berg-Breede River whitefish, from the Berg (Atlantic drainage) and Breede Rivers (Indian Ocean drainage);
<taxonomicName id="4C2472EBFFD7FFC07C8AF81BFE992592" authority="(Peters, 1864)" authorityName="Skelton &amp; Swartz &amp; Vreven" authorityYear="2018" baseAuthorityName="Peters" baseAuthorityYear="1864" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="serra">
<emphasis id="B950D57AFFD7FFC07C8AF81BFA8E25B9" box="[1165,1398,1965,1991]" italics="true" pageId="16" pageNumber="17">Cheilobarbus serra</emphasis>
(Peters, 1864)
</taxonomicName>
known as the sawfin or saagvin, from the Clanwilliam Olifants River (Atlantic drainage).
</paragraph>
</subSubSection>
<subSubSection id="C33E5AE3FFD6FFC278BAFEBDFBEA2335" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8B9B0968FFD6FFC178BAFEBDFECE235B" blockId="17.[189,310,267,293]" box="[189,310,267,293]" pageId="17" pageNumber="18">
<heading id="D0D3BE04FFD6FFC178BAFEBDFECE235B" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B950D57AFFD6FFC178BAFEBDFECE235B" bold="true" box="[189,310,267,293]" pageId="17" pageNumber="18">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B0968FFD6FFC278BAFE8FFBEA2335" blockId="17.[189,1399,313,868]" lastBlockId="18.[189,1399,267,331]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<taxonomicName id="4C2472EBFFD6FFC178BAFE8FFEA5232D" authorityName="Smith" authorityYear="1841" box="[189,349,313,339]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD6FFC178BAFE8FFEA5232D" box="[189,349,313,339]" italics="true" pageId="17" pageNumber="18">Cheilobarbus</emphasis>
</taxonomicName>
is distinguished from all other southern African tetraploid cyprinine genera by attaining a relatively large size (adult&gt;
<quantity id="4CDCA48DFFD6FFC17A0EFEEAFD912309" box="[521,617,348,375]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="18" unit="mm" value="150.0">150 mm</quantity>
SL vs &lt;
<quantity id="4CDCA48DFFD6FFC17AC8FEEAFCD72309" box="[719,815,348,375]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="18" unit="mm" value="150.0">150 mm</quantity>
SL), and by having an extended snout as reflected in an elongated lachrymal bone (length 3 × depth vs rectangular-shaped lachrymal length 2 × depth).
<taxonomicName id="4C2472EBFFD6FFC178BAFE15FEA523C3" authorityName="Smith" authorityYear="1841" box="[189,349,419,445]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD6FFC178BAFE15FEA523C3" box="[189,349,419,445]" italics="true" pageId="17" pageNumber="18">Cheilobarbus</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="4C2472EBFFD6FFC17A2AFE15FD4E23C3" authority="Skelton &amp; Swartz &amp; Vreven, 2018" authorityName="Skelton &amp; Swartz &amp; Vreven" authorityYear="2018" box="[557,694,419,445]" class="Actinopterygii" family="Cyprinidae" genus="Sedercypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B950D57AFFD6FFC17A2AFE15FD4E23C3" box="[557,694,419,445]" italics="true" pageId="17" pageNumber="18">Sedercypris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2636801FFD6FFC17AB9FE12FCD023C0" box="[702,808,420,446]" pageId="17" pageNumber="18" rank="genus">gen. nov.</taxonomicNameLabel>
by an absence of red colour at the base of the fins, in the position of the mouth (subterminal vs terminal) and in the length of the gut (&gt; 1.5 × SL vs &lt;1.5 × SL).
<taxonomicName id="4C2472EBFFD6FFC178FFFE5FFE60207D" authorityName="Smith" authorityYear="1841" box="[248,408,489,515]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD6FFC178FFFE5FFE60207D" box="[248,408,489,515]" italics="true" pageId="17" pageNumber="18">Cheilobarbus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C2472EBFFD6FFC17A32FE5FFD18207D" authority="Skelton &amp; Swartz &amp; Vreven, 2018" authorityName="Skelton &amp; Swartz &amp; Vreven" authorityYear="2018" box="[565,736,489,515]" class="Actinopterygii" family="Cyprinidae" genus="Amatolacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B950D57AFFD6FFC17A32FE5FFD18207D" box="[565,736,489,515]" italics="true" pageId="17" pageNumber="18">Amatolacypris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2636801FFD6FFC17AEFFE5CFCAB207A" box="[744,851,490,516]" pageId="17" pageNumber="18" rank="genus">gen. nov.</taxonomicNameLabel>
in overall size, colour (silvery to olive-bronze vs grey with double black lateral stripe), number of barbels (two pairs vs one pair), number (five vs four) and form of the 3rd and 4th infraorbital bones (narrow and slender vs broad).
<taxonomicName id="4C2472EBFFD6FFC17C45FD86FB1A2034" authorityName="Smith" authorityYear="1841" box="[1090,1250,560,586]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD6FFC17C45FD86FB1A2034" box="[1090,1250,560,586]" italics="true" pageId="17" pageNumber="18">Cheilobarbus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C2472EBFFD6FFC178BAFDE2FE8F2013" authority="Skelton &amp; Swartz &amp; Vreven, 2018" authorityName="Skelton &amp; Swartz &amp; Vreven" authorityYear="2018" box="[189,375,596,621]" class="Actinopterygii" family="Cyprinidae" genus="Namaquacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B950D57AFFD6FFC178BAFDE2FE8F2013" box="[189,375,596,621]" italics="true" pageId="17" pageNumber="18">Namaquacypris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2636801FFD6FFC1797AFDE2FE1F2010" box="[381,487,596,622]" pageId="17" pageNumber="18" rank="genus">gen. nov.</taxonomicNameLabel>
in the position of the dorsal fin (entirely before anal fin base vs reaching above anal fin base), form of the scales (regular vs thin), and the lack of a membrane between the innermost pelvic rays and the body (vs present).
<taxonomicName id="4C2472EBFFD6FFC17A96FD2FFCC920CD" authorityName="Smith" authorityYear="1841" box="[657,817,665,691]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD6FFC17A96FD2FFCC920CD" box="[657,817,665,691]" italics="true" pageId="17" pageNumber="18">Cheilobarbus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C2472EBFFD6FFC17BD2FD2FFB8520CD" authorityName="Smith" authorityYear="1841" box="[981,1149,665,691]" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD6FFC17BD2FD2FFB8520CD" box="[981,1149,665,691]" italics="true" pageId="17" pageNumber="18">Pseudobarbus</emphasis>
</taxonomicName>
in having a serrated dorsal-fin unbranched ray (vs simple dorsal-fin unbranched ray), an absence of red patches at the base of the fins (vs presence), no sexual dimorphism in fin size and shape and in the form and expression of nuptial tubercles as adults (vs clear sexual dimorphism in fin size and shape [males with larger fins] and nuptial tubercles [males with large conical tubercles on snout and head, small tubercles on scales and fins]).
<taxonomicName id="4C2472EBFFD6FFC1790EFCFFFE51211D" authorityName="Smith" authorityYear="1841" box="[265,425,841,867]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD6FFC1790EFCFFFE51211D" box="[265,425,841,867]" italics="true" pageId="17" pageNumber="18">Cheilobarbus</emphasis>
</taxonomicName>
can be separated from all other large sized southern African cyprinids by karyology (tetraploid vs diploid or hexaploid) the form of the scales (radiate striae vs parallel striae) and a serrated unbranched dorsal-fin ray (vs simple spinous unbranched dorsal-fin ray).
</paragraph>
</subSubSection>
<caption id="DF5B59E0FFD6FFC178BAF8F2FEC82595" ID-DOI="http://doi.org/10.5281/zenodo.1211218" httpUri="https://zenodo.org/record/1211218/files/figure.png" pageId="17" pageNumber="18" startId="17.[189,232,1860,1886]" targetBox="[189,1399,970,1817]" targetPageId="17">
<paragraph id="8B9B0968FFD6FFC178BAF8F2FEC82595" blockId="17.[189,1399,1860,2027]" pageId="17" pageNumber="18">
<emphasis id="B950D57AFFD6FFC178BAF8F2FEE22520" bold="true" box="[189,282,1860,1886]" pageId="17" pageNumber="18">Fig. 10.</emphasis>
Species representative of southern African tetraploid cyprinine genera.
<emphasis id="B950D57AFFD6FFC17C74F8F2FB702520" bold="true" box="[1139,1160,1860,1886]" pageId="17" pageNumber="18">A</emphasis>
.
<taxonomicName id="4C2472EBFFD6FFC17C9FF8F2FE9125FC" authority="(Peters, 1864)" authorityName="Peters" authorityYear="1864" baseAuthorityName="Peters" baseAuthorityYear="1864" class="Actinopterygii" family="Cyprinidae" genus="Pseudobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="afer">
<emphasis id="B950D57AFFD6FFC17C9FF8F2FA8E2520" box="[1176,1398,1860,1886]" italics="true" pageId="17" pageNumber="18">Pseudobarbus afer</emphasis>
(Peters, 1864)
</taxonomicName>
.
<emphasis id="B950D57AFFD6FFC17970F8D1FE7325FF" bold="true" box="[375,395,1895,1921]" pageId="17" pageNumber="18">B</emphasis>
.
<taxonomicName id="4C2472EBFFD6FFC1799AF8D1FCB225FF" authority="Smith, 1841" authorityName="Smith" authorityYear="1841" box="[413,842,1895,1921]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="capensis">
<emphasis id="B950D57AFFD6FFC1799AF8D1FD5425FF" box="[413,684,1895,1921]" italics="true" pageId="17" pageNumber="18">Cheilobarbus capensis</emphasis>
Smith, 1841
</taxonomicName>
.
<taxonomicName id="4C2472EBFFD6FFC17B5BF8D1FA8E25FF" authority="(Gunther, 1877)" authorityName="Skelton &amp; Swartz &amp; Vreven" authorityYear="2018" baseAuthorityName="Gunther" baseAuthorityYear="1877" box="[860,1398,1895,1922]" pageId="17" pageNumber="18" rank="subSpecies" status="gen. et comb. nov." subSpecies="trevelyani">
<emphasis id="B950D57AFFD6FFC17B5BF8D1FC8925FF" bold="true" box="[860,881,1895,1921]" pageId="17" pageNumber="18">C</emphasis>
.
<emphasis id="B950D57AFFD6FFC17B84F8D1FB5525FF" box="[899,1197,1895,1921]" italics="true" pageId="17" pageNumber="18">Amatolacypris trevelyani</emphasis>
(Günther, 1877)
</taxonomicName>
<taxonomicNameLabel id="A2636801FFD6FFC178BAF83DFE6B25DA" box="[189,403,1930,1957]" pageId="17" pageNumber="18">gen. et comb. nov.</taxonomicNameLabel>
<taxonomicName id="4C2472EBFFD6FFC1799DF83CFC9A25DA" authority="(Barnard, 1938)" authorityName="Skelton &amp; Swartz &amp; Vreven" authorityYear="2018" baseAuthorityName="Barnard" baseAuthorityYear="1938" box="[410,866,1930,1957]" pageId="17" pageNumber="18" rank="subSpecies" status="gen. et comb. nov." subSpecies="calidus">
<emphasis id="B950D57AFFD6FFC1799DF83CFE5725DA" bold="true" box="[410,431,1930,1956]" pageId="17" pageNumber="18">D</emphasis>
.
<emphasis id="B950D57AFFD6FFC179BAF83CFD5A25DA" box="[445,674,1930,1956]" italics="true" pageId="17" pageNumber="18">Sedercypris calidus</emphasis>
(Barnard, 1938)
</taxonomicName>
<taxonomicNameLabel id="A2636801FFD6FFC17B6EF83DFBC625DA" box="[873,1086,1930,1957]" pageId="17" pageNumber="18">gen. et comb. nov.</taxonomicNameLabel>
<taxonomicName id="4C2472EBFFD6FFC17C42F83CFE8125B6" authority="(Barnard, 1938)" authorityName="Skelton &amp; Swartz &amp; Vreven" authorityYear="2018" baseAuthorityName="Barnard" baseAuthorityYear="1938" pageId="17" pageNumber="18" rank="subSpecies" status="gen. et comb. nov." subSpecies="hospes">
<emphasis id="B950D57AFFD6FFC17C42F83CFBA125DA" bold="true" box="[1093,1113,1930,1956]" pageId="17" pageNumber="18">E</emphasis>
.
<emphasis id="B950D57AFFD6FFC17C60F83DFA8F25DA" box="[1127,1399,1930,1956]" italics="true" pageId="17" pageNumber="18">Namaquacypris hospes</emphasis>
(Barnard, 1938)
</taxonomicName>
<taxonomicNameLabel id="A2636801FFD6FFC17986F818FDA325B9" box="[385,603,1965,1992]" pageId="17" pageNumber="18">gen. et comb. nov.</taxonomicNameLabel>
Photograph credits: A–D by P.H. Skelton (SAIAB); E by R.I. Bills (SAIAB).
</paragraph>
</caption>
<subSubSection id="C33E5AE3FFD5FFC278BAFEC2FCE12077" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8B9B0968FFD5FFC278BAFEC2FEBC23F0" blockId="18.[189,324,372,398]" box="[189,324,372,398]" pageId="18" pageNumber="19">
<heading id="D0D3BE04FFD5FFC278BAFEC2FEBC23F0" bold="true" box="[189,324,372,398]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B950D57AFFD5FFC278BAFEC2FEBC23F0" bold="true" box="[189,324,372,398]" pageId="18" pageNumber="19">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B0968FFD5FFC278BAFE12FCE12077" blockId="18.[189,1399,420,521]" pageId="18" pageNumber="19">
The name
<taxonomicName id="4C2472EBFFD5FFC27930FE12FE2F23C0" authorityName="Smith" authorityYear="1841" box="[311,471,420,446]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD5FFC27930FE12FE2F23C0" box="[311,471,420,446]" italics="true" pageId="18" pageNumber="19">Cheilobarbus</emphasis>
</taxonomicName>
is derived from the Greek ‘
<emphasis id="B950D57AFFD5FFC27B12FE12FC9023C0" box="[789,872,420,446]" italics="true" pageId="18" pageNumber="19">cheilos</emphasis>
’ meaning a margin, lip or brim, and the Latin ‘
<emphasis id="B950D57AFFD5FFC278C0FE7CFEF5239A" box="[199,269,458,484]" italics="true" pageId="18" pageNumber="19">barba</emphasis>
’ meaning a beard and referring to the oral barbels.
<bibRefCitation id="EFB57499FFD5FFC27B72FE7FFBEA239A" author="Smith A." box="[885,1042,457,484]" journalOrPublisher="London" pageId="18" pageNumber="19" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith (1841)</bibRefCitation>
mentioned the “lips” of these species as being “full and firm”. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C33E5AE3FFD5FFC278BAFD84FEDB27CC" pageId="18" pageNumber="19" type="description">
<paragraph id="8B9B0968FFD5FFC278BAFD84FEB62032" blockId="18.[189,334,562,588]" box="[189,334,562,588]" pageId="18" pageNumber="19">
<heading id="D0D3BE04FFD5FFC278BAFD84FEB62032" bold="true" box="[189,334,562,588]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B950D57AFFD5FFC278BAFD84FEB62032" bold="true" box="[189,334,562,588]" pageId="18" pageNumber="19">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B0968FFD5FFC278BAFDD5FAEC21FF" blockId="18.[189,1399,611,897]" pageId="18" pageNumber="19">
<taxonomicName id="4C2472EBFFD5FFC278BAFDD5FEA52003" authorityName="Smith" authorityYear="1841" box="[189,349,611,637]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD5FFC278BAFDD5FEA52003" box="[189,349,611,637]" italics="true" pageId="18" pageNumber="19">Cheilobarbus</emphasis>
</taxonomicName>
is a genus of tetraploid cyprinine fishes with moderately sized, radiately striated scales; an elongated snout with elongated lacrymals, mouth subterminal with firm, well-developed lips, two pairs of simple oral barbels, pharyngeal bones with three rows of hooked pharyngeal teeth, tooth formula 5,3,2-2,3,5; intestine involuted and longer than the SL; dorsal fin positioned over or slightly behind the origin of the pelvic fins, the last simple ray bony weakly or strongly serrated, and eight branched rays; anal fin with three simple rays and five or six branched rays; mature breeding adults of both sexes develop small erupted nuptial tubercles densely scattered over head dorsum and in single rows over the pectoral fin rays; both species breed in male dominated nuptial shoals over gravel and cobbles.
</paragraph>
<paragraph id="8B9B0968FFD5FFC278BAFC1CFEE421BA" blockId="18.[189,284,938,964]" box="[189,284,938,964]" pageId="18" pageNumber="19">
<heading id="D0D3BE04FFD5FFC278BAFC1CFEE421BA" bold="true" box="[189,284,938,964]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B950D57AFFD5FFC278BAFC1CFEE421BA" bold="true" box="[189,284,938,964]" pageId="18" pageNumber="19">History</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B0968FFD5FFC278BAFC6DFEDB27CC" blockId="18.[189,1399,987,1458]" pageId="18" pageNumber="19">
<bibRefCitation id="EFB57499FFD5FFC278BAFC6DFEAF218B" author="Smith A." box="[189,343,987,1013]" journalOrPublisher="London" pageId="18" pageNumber="19" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith (1841)</bibRefCitation>
attributed both the large
<taxonomicName id="4C2472EBFFD5FFC27A70FC6DFD34218B" authorityName="Daudin" authorityYear="1805" box="[631,716,987,1013]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD5FFC27A70FC6DFD34218B" box="[631,716,987,1013]" italics="true" pageId="18" pageNumber="19">Barbus</emphasis>
</taxonomicName>
species he described to a new subgenus
<taxonomicName id="4C2472EBFFD5FFC27C99FC6DFAC6218B" authorityName="Smith" authorityYear="1841" box="[1182,1342,987,1013]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD5FFC27C99FC6DFAC6218B" box="[1182,1342,987,1013]" italics="true" pageId="18" pageNumber="19">Cheilobarbus</emphasis>
</taxonomicName>
, i.e.,
<taxonomicName id="4C2472EBFFD5FFC278BAFBB6FDC62664" authorityName="Smith" authorityYear="1841" box="[189,574,1024,1050]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="capensis" subGenus="Cheilobarbus">
<emphasis id="B950D57AFFD5FFC278BAFBB6FEEA2664" box="[189,274,1024,1050]" italics="true" pageId="18" pageNumber="19">Barbus</emphasis>
(
<emphasis id="B950D57AFFD5FFC27921FBB6FE3E2664" box="[294,454,1024,1050]" italics="true" pageId="18" pageNumber="19">Cheilobarbus</emphasis>
)
<emphasis id="B950D57AFFD5FFC279DEFBB7FDC62664" box="[473,574,1025,1050]" italics="true" pageId="18" pageNumber="19">capensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2472EBFFD5FFC27A7AFBB6FBC82664" baseAuthorityName="non Smith" baseAuthorityYear="1841" box="[637,1072,1024,1050]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="marequensis" subGenus="Cheilobarbus">
<emphasis id="B950D57AFFD5FFC27A7AFBB6FD2A2664" box="[637,722,1024,1050]" italics="true" pageId="18" pageNumber="19">Barbus</emphasis>
(
<emphasis id="B950D57AFFD5FFC27AE1FBB6FC7E2664" box="[742,902,1024,1050]" italics="true" pageId="18" pageNumber="19">Cheilobarbus</emphasis>
)
<emphasis id="B950D57AFFD5FFC27B9EFBB7FBC82664" box="[921,1072,1025,1050]" italics="true" pageId="18" pageNumber="19">marequensis</emphasis>
</taxonomicName>
<emphasis id="B950D57AFFD5FFC27C37FBB7FBCC2664" box="[1072,1076,1025,1050]" italics="true" pageId="18" pageNumber="19">.</emphasis>
While the latter species is identified as a
<taxonomicName id="4C2472EBFFD5FFC27960FB93FDFA2641" authorityName="Ruppell" authorityYear="1835" box="[359,514,1061,1087]" class="Actinopterygii" family="Cyprinidae" genus="Labeobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD5FFC27960FB93FDFA2641" box="[359,514,1061,1087]" italics="true" pageId="18" pageNumber="19">Labeobarbus</emphasis>
</taxonomicName>
, the former, a tetraploid species, does not belong in
<taxonomicName id="4C2472EBFFD5FFC27C61FB93FAF92641" authorityName="Ruppell" authorityYear="1835" box="[1126,1281,1061,1087]" class="Actinopterygii" family="Cyprinidae" genus="Labeobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD5FFC27C61FB93FAF92641" box="[1126,1281,1061,1087]" italics="true" pageId="18" pageNumber="19">Labeobarbus</emphasis>
</taxonomicName>
. The
<typeStatus id="549FB7CAFFD5FFC27D43FB93FA8F2641" box="[1348,1399,1061,1087]" pageId="18" pageNumber="19">type</typeStatus>
species of the subgenus,
<taxonomicName id="4C2472EBFFD5FFC279DAFBFDFD9A261A" authorityName="Smith" authorityYear="1841" box="[477,610,1099,1124]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="capensis">
<emphasis id="B950D57AFFD5FFC279DAFBFDFD9A261A" box="[477,610,1099,1124]" italics="true" pageId="18" pageNumber="19">B. capensis</emphasis>
</taxonomicName>
, was designated by monotypy by
<bibRefCitation id="EFB57499FFD5FFC27BEAFBFCFB33261A" author="Jordan D. S." box="[1005,1227,1098,1125]" journalOrPublisher="Leland Stanford Jr. University Publications, University Series" pageId="18" pageNumber="19" publicationUrl="http://gallica.bnf.fr/ark:/12148/bpt6k97097q" refId="ref13519" refString="Jordan D. S. 1919. The Genera of Fishes, Part II, from Agassiz to Bleeker, 1833 - 1858, Twenty-Six Years, with the Accepted Type of Each. A Contribution to the Stability of Scientific Nomenclature. Leland Stanford Jr. University Publications, University Series. Available from http: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 97097 q [accessed 5 Feb. 2018]." title="The Genera of Fishes, Part II, from Agassiz to Bleeker, 1833 - 1858, Twenty-Six Years, with the Accepted Type of Each. A Contribution to the Stability of Scientific Nomenclature" type="book chapter" year="1919">
<collectingCountry id="F33349F8FFD5FFC27BEAFBFCFBC3261A" box="[1005,1083,1098,1124]" name="Jordan" pageId="18" pageNumber="19">Jordan</collectingCountry>
(1919: 244)
</bibRefCitation>
as first revisor and, therefore, is not a junior synonym of
<taxonomicName id="4C2472EBFFD5FFC27AA8FBD9FCB226F7" authorityName="Ruppell" authorityYear="1835" box="[687,842,1135,1161]" class="Actinopterygii" family="Cyprinidae" genus="Labeobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD5FFC27AA8FBD9FCB226F7" box="[687,842,1135,1161]" italics="true" pageId="18" pageNumber="19">Labeobarbus</emphasis>
</taxonomicName>
. The subgenus
<taxonomicName id="4C2472EBFFD5FFC27C03FBD9FB5C26F7" authorityName="Smith" authorityYear="1841" box="[1028,1188,1135,1161]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD5FFC27C03FBD9FB5C26F7" box="[1028,1188,1135,1161]" italics="true" pageId="18" pageNumber="19">Cheilobarbus</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFB57499FFD5FFC278BAFB22FEA426D1" author="Smith A." box="[189,348,1172,1199]" journalOrPublisher="London" pageId="18" pageNumber="19" refId="ref14583" refString="Smith A. 1841. Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co., London." title="Illustrations of the Zoology of South Africa, Pisces. Smith, Elder and Co" type="book" year="1841">Smith (1841)</bibRefCitation>
as follows: “Mouth opening forwards; lips full, and firm; intermaxillary bones slightly extensible; nostrils double; four cirri, two from snout, and one from each angle of mouth; lateral line consisting of a series of small tubes; scales large; dorsal fin short, and commencing slightly in front of base of ventral fins; commencement of anal fin about midway between ventral and caudal fins.” However this description is not diagnostic, e.g., it does not mention the nature of the unbranched dorsal-fin ray nor the nature of the scale radii. In order to institute
<taxonomicName id="4C2472EBFFD5FFC27B3BFAF8FC242716" authorityName="Smith" authorityYear="1841" box="[828,988,1358,1384]" class="Actinopterygii" family="Cyprinidae" genus="Cheilobarbus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B950D57AFFD5FFC27B3BFAF8FC242716" box="[828,988,1358,1384]" italics="true" pageId="18" pageNumber="19">Cheilobarbus</emphasis>
</taxonomicName>
as the genus for these species, an expanded diagnostic definition is required and the species to be included identified along phylogenetic grounds.
</paragraph>
</subSubSection>
</treatment>
</document>
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