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<mods:title id="65FF777BFD25F264DEE88E502AA98664">Two new generic synonyms in Palearctic Ciinae (Coleoptera: Ciidae)</mods:title>
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<paragraph id="D20D36CCFFC0FFA6FF0FC6C3FCCA85E7" blockId="0.[151,804,1687,1785]" box="[151,804,1687,1714]" pageId="0" pageNumber="297">
<heading id="894581A0FFC0FFA6FF0FC6C3FCCA85E7" bold="true" box="[151,804,1687,1714]" fontSize="11" level="1" pageId="0" pageNumber="297" reason="1">
<emphasis id="E0C6EADEFFC0FFA6FF0FC6C3FCCA85E7" bold="true" box="[151,804,1687,1714]" pageId="0" pageNumber="297">
<taxonomicName id="15B24D4FFFC0FFA6FF0FC6C3FE1785E7" authority="Miyatake, 1954" authorityName="Miyatake" authorityYear="1954" box="[151,505,1687,1713]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC0FFA6FF0FC6C3FEDE85E7" bold="true" box="[151,304,1687,1713]" italics="true" pageId="0" pageNumber="297">Paraxestocis</emphasis>
<bibRefCitation id="B6234B3DFFC0FFA6FEA0C6C3FE1785E7" author="Miyatake, M." box="[312,505,1687,1713]" pageId="0" pageNumber="297" pagination="40 - 67" refId="ref1942" refString="Miyatake, M. (1954) Studies on the Japanese Ciidae, I (Coleoptera). The Scientific Reports of the Matsuyama Agricultural College, 14, 40 - 67, pls. 1 - 11." type="journal article" year="1954">Miyatake, 1954</bibRefCitation>
</taxonomicName>
,
<emphasis id="E0C6EADEFFC0FFA6FD9FC6CCFD6385E4" bold="true" box="[519,653,1688,1714]" italics="true" pageId="0" pageNumber="297">
<taxonomicNameLabel id="FBF557A5FFC0FFA6FD9FC6CCFD6385E4" box="[519,653,1688,1714]" pageId="0" pageNumber="297">sensu novo</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="4A892A49FFC0FFA6FD05C6C3FCF285E7" box="[669,796,1687,1713]" captionStart="FIGURE 1" captionStartId="1.[151,250,1785,1809]" captionTargetBox="[151,1436,487,1760]" captionTargetId="figure-133@1.[151,1436,487,1760]" captionTargetPageId="1" captionText="FIGURE 1. Species of Paraxestocis Miyatake, 1954, sensu novo. AD. Paraxestocis unicornis Miyatake, 1954, male holotype. Dorsal view, with arrow showing the pronotum pointed anteriorly (A), lateral view (B), ventral view (C) and labels (D). The type locality &quot;Naose&quot; was a small village on the foot of Mt. Ishizumi, about 500700 m of altitude (33º42N, 132º57E), &quot;Kuma&quot; is now the city of &quot;Kumakogen-cho&quot; and “IYO” is now Ehime Prefecture, Shikoku. E. Paraxestocis denticollis (Nakane &amp; Nobuchi, 1955), new combination, male from Chitose, Hokkaido, Japan. F. Paraxestocis wagae (Wankowiez, 1869), new combination, male from Bialowieska Forest, Poland. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189902" httpUri="https://zenodo.org/record/12189902/files/figure.png" pageId="0" pageNumber="297">Fig. 1AF</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9AA86547FFC0FFA6FF5FC6E9FD1B8583" box="[199,757,1725,1749]" pageId="0" pageNumber="297" type="synonymic_list">
<paragraph id="D20D36CCFFC0FFA6FF5FC6E9FD1B8583" blockId="0.[151,804,1687,1785]" box="[199,757,1725,1749]" pageId="0" pageNumber="297">
<taxonomicName id="15B24D4FFFC0FFA6FF5FC6E9FDBE8583" authority="Nakane &amp; Nobuchi, 1955" authorityName="Nakane &amp; Nobuchi" authorityYear="1955" box="[199,592,1725,1749]" class="Insecta" family="Ciidae" genus="Odontocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="297" phylum="Arthropoda" rank="genus" status="new synonym">
<emphasis id="E0C6EADEFFC0FFA6FF5FC6E9FEDB8583" box="[199,309,1725,1749]" italics="true" pageId="0" pageNumber="297">Odontocis</emphasis>
<bibRefCitation id="B6234B3DFFC0FFA6FEA3C6E9FDBE8583" author="Nakane, T. &amp; Nobuchi, A." box="[315,592,1725,1749]" pageId="0" pageNumber="297" pagination="47 - 52" refId="ref1981" refString="Nakane, T. &amp; Nobuchi, A. (1955) On a new Genus and six new Species of Ciid-Beetles from Japan (Ciidae, Coleoptera). Scientific Reports of the Saikyo University (Natural Science and Living Science), A Series, 2 (1), 47 - 52." type="journal article" year="1955">Nakane &amp; Nobuchi, 1955</bibRefCitation>
</taxonomicName>
,
<emphasis id="E0C6EADEFFC0FFA6FDC3C6E9FD1B8583" bold="true" box="[603,757,1725,1749]" pageId="0" pageNumber="297">
<taxonomicNameLabel id="FBF557A5FFC0FFA6FDC3C6E9FD1B8583" box="[603,757,1725,1749]" pageId="0" pageNumber="297">new synonym</taxonomicNameLabel>
</emphasis>
</paragraph>
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<paragraph id="D20D36CCFFC0FFA6FF5FC6B6FDB485AF" blockId="0.[151,804,1687,1785]" box="[199,602,1761,1785]" pageId="0" pageNumber="297">
<taxonomicName id="15B24D4FFFC0FFA6FF5FC6B6FE5B85AF" authority="Lohse, 1964" authorityName="Lohse" authorityYear="1964" box="[199,437,1761,1785]" class="Insecta" family="Ciidae" genus="Wagaicis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="297" phylum="Arthropoda" rank="genus" status="new synonym">
<emphasis id="E0C6EADEFFC0FFA6FF5FC6B6FEC685AF" box="[199,296,1762,1785]" italics="true" pageId="0" pageNumber="297">Wagaicis</emphasis>
<bibRefCitation id="B6234B3DFFC0FFA6FEB7C6B5FE5B85AF" author="Lohse, G. A." box="[303,437,1761,1785]" pageId="0" pageNumber="297" pagination="116 - 122" refId="ref1827" refString="Lohse, G. A. (1964) Die in Mitteleuropa vertretenen Gattungen der Cisidae. (1. Beitrag zur Kenntnis der mitteleuropaischen Cisidae). Entomologische Blatter, 60 (2), 116 - 122." type="journal article" year="1964">Lohse, 1964</bibRefCitation>
</taxonomicName>
,
<emphasis id="E0C6EADEFFC0FFA6FE58C6B5FDB485AF" bold="true" box="[448,602,1761,1785]" pageId="0" pageNumber="297">
<taxonomicNameLabel id="FBF557A5FFC0FFA6FE58C6B5FDB485AF" box="[448,602,1761,1785]" pageId="0" pageNumber="297">new synonym</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="9AA86547FFC0FFA6FF0FC77DFC808417" box="[151,878,1833,1857]" pageId="0" pageNumber="297" type="type_taxon">
<paragraph id="D20D36CCFFC0FFA6FF0FC77DFC808417" blockId="0.[151,1436,1833,2038]" box="[151,878,1833,1857]" pageId="0" pageNumber="297">
<emphasis id="E0C6EADEFFC0FFA6FF0FC77DFE2B8417" bold="true" box="[151,453,1833,1857]" inLineHeading="true" pageId="0" pageNumber="297" reason="1">
<typeStatus id="0D09886EFFC0FFA6FF0FC77DFF218417" box="[151,207,1833,1857]" pageId="0" pageNumber="297">Type</typeStatus>
species by monotypy.
</emphasis>
<taxonomicName id="15B24D4FFFC0FFA6FE54C77EFC808417" authority="Miyatake, 1954" authorityName="Miyatake" authorityYear="1954" box="[460,878,1833,1857]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="297" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="E0C6EADEFFC0FFA6FE54C77EFD2F8417" box="[460,705,1834,1857]" italics="true" pageId="0" pageNumber="297">Paraxestocis unicornis</emphasis>
<bibRefCitation id="B6234B3DFFC0FFA6FD5FC77DFC808417" author="Miyatake, M." box="[711,878,1833,1857]" pageId="0" pageNumber="297" pagination="40 - 67" refId="ref1942" refString="Miyatake, M. (1954) Studies on the Japanese Ciidae, I (Coleoptera). The Scientific Reports of the Matsuyama Agricultural College, 14, 40 - 67, pls. 1 - 11." type="journal article" year="1954">Miyatake, 1954</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="9AA86547FFC0FFA4FF5FC719FB5B8249" lastPageId="2" lastPageNumber="299" pageId="0" pageNumber="297" type="diagnosis">
<paragraph id="D20D36CCFFC0FFA6FF5FC719FC1184DF" blockId="0.[151,1436,1833,2038]" pageId="0" pageNumber="297">
<emphasis id="E0C6EADEFFC0FFA6FF5FC719FED28433" bold="true" box="[199,316,1869,1893]" inLineHeading="true" pageId="0" pageNumber="297" reason="1">Diagnosis.</emphasis>
Body convex, apparently glabrous dorsally (minute setae seen under magnification of 100x or higher), with a total length of
<quantity id="154A9B29FFC0FFA6FEE4C725FE5784DF" box="[380,441,1905,1929]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="297" unit="mm" value="2.0">2 mm</quantity>
or less; dorsal punctation coarse, single and confused.
</paragraph>
<paragraph id="D20D36CCFFC0FFA6FF5FC7C1FC2684A0" blockId="0.[151,1436,1833,2038]" pageId="0" pageNumber="297">Antennae 9- or 10- segmented; third antennomere conspicuously longer than the fourth; apical three antennomeres forming a loose club, with each bearing four multi-pronged sensilla (=sensillifers).Anterocephalic edge of males produced as a lamina and modified to form a short triangular projection near each eye.</paragraph>
<paragraph id="D20D36CCFFC1FFA7FF5FC0CCFCB683A1" blockId="1.[151,1437,151,427]" pageId="1" pageNumber="298">
Pronotum with lateral edges smooth, not explanate and not visible from above; in males, the anterior edge is projected forward at the middle forming a subtle horn-like process (
<figureCitation id="4A892A49FFC1FFA7FC9CC0EFFCB68382" box="[772,856,187,212]" captionStart="FIGURE 1" captionStartId="1.[151,250,1785,1809]" captionTargetBox="[151,1436,487,1760]" captionTargetId="figure-133@1.[151,1436,487,1760]" captionTargetPageId="1" captionText="FIGURE 1. Species of Paraxestocis Miyatake, 1954, sensu novo. AD. Paraxestocis unicornis Miyatake, 1954, male holotype. Dorsal view, with arrow showing the pronotum pointed anteriorly (A), lateral view (B), ventral view (C) and labels (D). The type locality &quot;Naose&quot; was a small village on the foot of Mt. Ishizumi, about 500700 m of altitude (33º42N, 132º57E), &quot;Kuma&quot; is now the city of &quot;Kumakogen-cho&quot; and “IYO” is now Ehime Prefecture, Shikoku. E. Paraxestocis denticollis (Nakane &amp; Nobuchi, 1955), new combination, male from Chitose, Hokkaido, Japan. F. Paraxestocis wagae (Wankowiez, 1869), new combination, male from Bialowieska Forest, Poland. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189902" httpUri="https://zenodo.org/record/12189902/files/figure.png" pageId="1" pageNumber="298">Fig. 1A</figureCitation>
, arrow). Prosternum short, slightly tumid and carinate at the longitudinal midline; prosternal process very thin, laminate.
</paragraph>
<paragraph id="D20D36CCFFC1FFA7FF5FC150FE558235" blockId="1.[151,1437,151,427]" pageId="1" pageNumber="298">Procoxae are somewhat subconical, but not projecting beyond the plane of the prosternal process. Protibiae gradually expanding from base to apex; outer apical angle broadly rounded, with a row of spines extending from the apex to at least one-third of the outer edge.</paragraph>
<paragraph id="D20D36CCFFC1FFA7FF5FC124FD1582FD" blockId="1.[151,1437,151,427]" pageId="1" pageNumber="298">Meso- and metatibiae moderately expanded, with a row of spines extending from apex to at least the outer apical angle. First abdominal ventrite of males with a sex patch.</paragraph>
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<paragraph id="D20D36CCFFC1FFA7FF0FC6ADFC908492" blockId="1.[151,1437,1785,1989]" pageId="1" pageNumber="298">
<emphasis id="E0C6EADEFFC1FFA7FF0FC6ADFEFC8447" bold="true" box="[151,274,1785,1809]" pageId="1" pageNumber="298">FIGURE 1.</emphasis>
Species of
<taxonomicName id="15B24D4FFFC1FFA7FE1DC6AEFD408447" authority="Miyatake, 1954" authorityName="Miyatake" authorityYear="1954" box="[389,686,1785,1809]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="298" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC1FFA7FE1DC6AEFDE68447" box="[389,520,1786,1809]" italics="true" pageId="1" pageNumber="298">Paraxestocis</emphasis>
<bibRefCitation id="B6234B3DFFC1FFA7FD95C6ADFD408447" author="Miyatake, M." box="[525,686,1785,1809]" pageId="1" pageNumber="298" pagination="40 - 67" refId="ref1942" refString="Miyatake, M. (1954) Studies on the Japanese Ciidae, I (Coleoptera). The Scientific Reports of the Matsuyama Agricultural College, 14, 40 - 67, pls. 1 - 11." type="journal article" year="1954">Miyatake, 1954</bibRefCitation>
</taxonomicName>
,
<emphasis id="E0C6EADEFFC1FFA7FD2FC6AEFCC88447" box="[695,806,1786,1809]" italics="true" pageId="1" pageNumber="298">
<taxonomicName id="15B24D4FFFC1FFA7FD2FC6AEFD018447" box="[695,751,1786,1809]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="298" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
novo
</emphasis>
.
<emphasis id="E0C6EADEFFC1FFA7FCAAC6ADFC848447" bold="true" box="[818,874,1785,1809]" pageId="1" pageNumber="298">AD.</emphasis>
<taxonomicName id="15B24D4FFFC1FFA7FCF7C6AEFB138447" authority="Miyatake, 1954" authorityName="Miyatake" authorityYear="1954" box="[879,1277,1785,1809]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="298" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="E0C6EADEFFC1FFA7FCF7C6AEFBB98447" box="[879,1111,1786,1809]" italics="true" pageId="1" pageNumber="298">Paraxestocis unicornis</emphasis>
<bibRefCitation id="B6234B3DFFC1FFA7FBC5C6ADFB138447" author="Miyatake, M." box="[1117,1277,1785,1809]" pageId="1" pageNumber="298" pagination="40 - 67" refId="ref1942" refString="Miyatake, M. (1954) Studies on the Japanese Ciidae, I (Coleoptera). The Scientific Reports of the Matsuyama Agricultural College, 14, 40 - 67, pls. 1 - 11." type="journal article" year="1954">Miyatake, 1954</bibRefCitation>
</taxonomicName>
, male holotype. Dorsal view, with arrow showing the pronotum pointed anteriorly (
<emphasis id="E0C6EADEFFC1FFA7FCD4C749FCB08463" bold="true" box="[844,862,1821,1845]" pageId="1" pageNumber="298">A</emphasis>
), lateral view (
<emphasis id="E0C6EADEFFC1FFA7FC64C749FBE38463" bold="true" box="[1020,1037,1821,1845]" pageId="1" pageNumber="298">B</emphasis>
), ventral view (
<emphasis id="E0C6EADEFFC1FFA7FB2BC749FB2B8463" bold="true" box="[1203,1221,1821,1845]" pageId="1" pageNumber="298">C</emphasis>
) and labels (
<emphasis id="E0C6EADEFFC1FFA7FAD5C749FAB18463" bold="true" box="[1357,1375,1821,1845]" pageId="1" pageNumber="298">D</emphasis>
). The type locality &quot;Naose&quot; was a small village on the foot of Mt. Ishizumi, about 500700 m of altitude (33º42N, 132º57E), &quot;Kuma&quot; is now the city of &quot;Kumakogen-cho&quot; and “IYO” is now Ehime Prefecture, Shikoku.
<emphasis id="E0C6EADEFFC1FFA7FB94C731FBCA842B" bold="true" box="[1036,1060,1893,1917]" pageId="1" pageNumber="298">E.</emphasis>
<taxonomicName id="15B24D4FFFC1FFA7FBB4C732FED384F7" authority="(Nakane &amp; Nobuchi, 1955)" authorityName="Lopes-Andrade" authorityYear="2024" baseAuthorityName="Nakane &amp; Nobuchi" baseAuthorityYear="1955" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="298" phylum="Arthropoda" rank="species" species="denticollis" status="comb. nov.">
<emphasis id="E0C6EADEFFC1FFA7FBB4C732FACC842B" box="[1068,1314,1893,1917]" italics="true" pageId="1" pageNumber="298">Paraxestocis denticollis</emphasis>
(
<bibRefCitation id="B6234B3DFFC1FFA7FAA9C731FEDB84F7" author="Nakane, T. &amp; Nobuchi, A." pageId="1" pageNumber="298" pagination="47 - 52" refId="ref1981" refString="Nakane, T. &amp; Nobuchi, A. (1955) On a new Genus and six new Species of Ciid-Beetles from Japan (Ciidae, Coleoptera). Scientific Reports of the Saikyo University (Natural Science and Living Science), A Series, 2 (1), 47 - 52." type="journal article" year="1955">Nakane &amp; Nobuchi, 1955</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="FBF557A5FFC1FFA7FED2C7DDFE1184F7" box="[330,511,1929,1953]" pageId="1" pageNumber="298" rank="species">new combination</taxonomicNameLabel>
, male from Chitose, Hokkaido, Japan.
<emphasis id="E0C6EADEFFC1FFA7FC38C7DDFC5D84F7" bold="true" box="[928,947,1929,1953]" pageId="1" pageNumber="298">F.</emphasis>
<taxonomicName id="15B24D4FFFC1FFA7FC26C7DEFA8D84F7" authority="(Wankowiez, 1869)" authorityName="Lopes-Andrade" authorityYear="2024" baseAuthorityName="Wankowiez" baseAuthorityYear="1869" box="[958,1379,1929,1953]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="298" phylum="Arthropoda" rank="species" species="wagae" status="comb. nov.">
<emphasis id="E0C6EADEFFC1FFA7FC26C7DEFB6384F7" box="[958,1165,1930,1953]" italics="true" pageId="1" pageNumber="298">Paraxestocis wagae</emphasis>
(
<bibRefCitation id="B6234B3DFFC1FFA7FB06C7DDFAB384F7" author="Wankowiez, J." box="[1182,1373,1929,1953]" pageId="1" pageNumber="298" pagination="411 - 422" refId="ref2361" refString="Wankowiez, J. (1869) Notices entomologiques. Annales de la Societe entomologique de France, 4 (9), 411 - 422." type="journal article" year="1869">Wankowiez, 1869</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="FBF557A5FFC1FFA7FAEAC7DDFEF68493" pageId="1" pageNumber="298" rank="species">new combination</taxonomicNameLabel>
, male from Bialowieska Forest, Poland. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="D20D36CCFFC2FFA4FF5FC0C3FB5B8249" blockId="2.[151,1437,151,661]" pageId="2" pageNumber="299">
<emphasis id="E0C6EADEFFC2FFA4FF5FC0C3FE4983E6" bold="true" box="[199,423,151,176]" inLineHeading="true" pageId="2" pageNumber="299" reason="1">Comparative notes.</emphasis>
Comparisons were based upon the examination of species of all the genera mentioned below, but also on
<bibRefCitation id="B6234B3DFFC2FFA4FE8DC0E9FD7B8383" author="Lopes-Andrade, C. &amp; Lawrence, J. F." box="[277,661,189,213]" pageId="2" pageNumber="299" pagination="43 - 60" refId="ref1867" refString="Lopes-Andrade, C. &amp; Lawrence, J. F. (2005) Phellinocis, a new genus of Neotropical Ciidae (Coleoptera: Tenebrionoidea). Zootaxa, 1034 (1), 43 - 60. https: // doi. org / 10.11646 / zootaxa. 1034.1.3" type="journal article" year="2005">Lopes-Andrade &amp; Lawrence (2005)</bibRefCitation>
and
<bibRefCitation id="B6234B3DFFC2FFA4FD52C0E9FC948383" author="Lawrence, J. F." box="[714,890,189,213]" pageId="2" pageNumber="299" pagination="419 - 522" refId="ref1749" refString="Lawrence, J. F. (1971) Revision of the North American Ciidae (Coleoptera). Bulletin of the Museum of Comparative Zoology, 142, 419 - 522." type="journal article" year="1971">Lawrence (1971</bibRefCitation>
,
<bibRefCitation id="B6234B3DFFC2FFA4FC1EC0E9FC2E8383" author="Lawrence, J. F." box="[902,960,189,213]" pageId="2" pageNumber="299" pagination="1" refId="ref1782" refString="Lawrence, J. F. (2016) The Australian Ciidae (Coleoptera: Tenebrionoidea): a preliminary revision. Zootaxa, 4198 (1), 001 - 208. https: // doi. org / 10.11646 / zootaxa. 4198.1.1" type="journal article" year="2016">2016</bibRefCitation>
).
<taxonomicName id="15B24D4FFFC2FFA4FC4AC0EAFB348383" box="[978,1242,190,213]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="subSpecies" species="sensu" subSpecies="novo">
<emphasis id="E0C6EADEFFC2FFA4FC4AC0EAFB348383" box="[978,1242,190,213]" italics="true" pageId="2" pageNumber="299">Paraxestocis sensu novo</emphasis>
</taxonomicName>
differs from most
<taxonomicName id="15B24D4FFFC2FFA4FF0FC0B6FF2283AC" authorityName="Leach" authorityYear="1819" box="[151,204,226,250]" class="Insecta" family="Ciidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="tribe" tribe="Ciini">Ciini</taxonomicName>
genera in the combination of a short prosternum with a laminate prosternal process, and tibiae expanded and bearing a row of spines from apex to at least the outer apical angle, sometimes extending to the outer edge.
</paragraph>
</subSubSection>
<subSubSection id="9AA86547FFC2FFA4FF5FC179FD0081C2" pageId="2" pageNumber="299" type="description">
<paragraph id="D20D36CCFFC2FFA4FF5FC179FDAE82D9" blockId="2.[151,1437,151,661]" pageId="2" pageNumber="299">
Within
<taxonomicName id="15B24D4FFFC2FFA4FE86C179FEB88213" authorityName="Leach" authorityYear="1819" box="[286,342,301,325]" class="Insecta" family="Ciidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="tribe" tribe="Ciini">Ciini</taxonomicName>
,
<taxonomicName id="15B24D4FFFC2FFA4FEF0C179FDBA8213" authority="Miyatake" authorityName="Miyatake" authorityYear="1955" box="[360,596,301,325]" class="Insecta" family="Ciidae" genus="Acanthocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FEF0C179FE318213" box="[360,479,301,325]" italics="true" pageId="2" pageNumber="299">Acanthocis</emphasis>
Miyatake
</taxonomicName>
,
<taxonomicName id="15B24D4FFFC2FFA4FDFDC179FCE08213" authority="Mellie" authorityName="Mellie" box="[613,782,301,325]" class="Insecta" family="Ciidae" genus="Ceracis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FDFDC179FD568212" box="[613,696,301,324]" italics="true" pageId="2" pageNumber="299">Ceracis</emphasis>
Mellié
</taxonomicName>
,
<taxonomicName id="15B24D4FFFC2FFA4FCB8C179FBD38213" authority="Kawanabe" authorityName="Kawanabe" authorityYear="1994" box="[800,1085,301,325]" class="Insecta" family="Ciidae" genus="Dichodontocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FCB8C179FC528213" box="[800,956,301,325]" italics="true" pageId="2" pageNumber="299">Dichodontocis</emphasis>
Kawanabe
</taxonomicName>
,
<taxonomicName id="15B24D4FFFC2FFA4FBD7C179FB328213" authority="Pic" authorityName="Pic" authorityYear="1916" box="[1103,1244,301,325]" class="Insecta" family="Ciidae" genus="Falsocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FBD7C179FB478213" box="[1103,1193,301,325]" italics="true" pageId="2" pageNumber="299">Falsocis</emphasis>
Pic
</taxonomicName>
,
<taxonomicName id="15B24D4FFFC2FFA4FB75C179FEED823C" authority="Lawrence" authorityName="Lawrence" authorityYear="2016" genus="Pseudeuxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="299" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FB75C179FA728213" box="[1261,1436,301,325]" italics="true" pageId="2" pageNumber="299">Pseudeuxestocis</emphasis>
Lawrence
</taxonomicName>
,
<taxonomicName id="15B24D4FFFC2FFA4FE95C106FD51823C" authority="Lopes-Andrade &amp; Lawrence" authorityName="Lopes-Andrade &amp; Lawrence" authorityYear="2005" box="[269,703,338,362]" class="Insecta" family="Ciidae" genus="Phellinocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FE95C106FE6B823C" box="[269,389,338,362]" italics="true" pageId="2" pageNumber="299">Phellinocis</emphasis>
Lopes-Andrade &amp; Lawrence
</taxonomicName>
,
<taxonomicName id="15B24D4FFFC2FFA4FD52C106FC9F823C" authority="Casey" authorityName="Casey" authorityYear="1898" box="[714,881,338,362]" class="Insecta" family="Ciidae" genus="Plesiocis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FD52C106FCC4823C" box="[714,810,338,362]" italics="true" pageId="2" pageNumber="299">Plesiocis</emphasis>
Casey
</taxonomicName>
and
<taxonomicName id="15B24D4FFFC2FFA4FC3DC106FB9A823C" authority="Lawrence" authorityName="Lawrence" authorityYear="1987" box="[933,1140,338,362]" class="Insecta" family="Ciidae" genus="Porculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FC3DC106FBEA823C" box="[933,1028,338,362]" italics="true" pageId="2" pageNumber="299">Porculus</emphasis>
Lawrence
</taxonomicName>
are similar in having a sublaminate to laminate prosternal process.
</paragraph>
<paragraph id="D20D36CCFFC2FFA4FF5FC1C9FB38811C" blockId="2.[151,1437,151,661]" pageId="2" pageNumber="299">
However, in
<taxonomicName id="15B24D4FFFC2FFA4FEC8C1C9FE4D82E2" box="[336,419,413,436]" class="Insecta" family="Ciidae" genus="Ceracis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FEC8C1C9FE4D82E2" box="[336,419,413,436]" italics="true" pageId="2" pageNumber="299">Ceracis</emphasis>
</taxonomicName>
the spines of meso- and metatibiae are mostly confined to their apices. In
<taxonomicName id="15B24D4FFFC2FFA4FB20C1C9FAC082E3" authorityName="Miyatake" authorityYear="1955" box="[1208,1326,413,437]" class="Insecta" family="Ciidae" genus="Acanthocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FB20C1C9FAC082E3" box="[1208,1326,413,437]" italics="true" pageId="2" pageNumber="299">Acanthocis</emphasis>
</taxonomicName>
,
<taxonomicName id="15B24D4FFFC2FFA4FAA3C1C9FA7B82E3" authorityName="Pic" authorityYear="1916" box="[1339,1429,413,437]" class="Insecta" family="Ciidae" genus="Falsocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FAA3C1C9FA7B82E3" box="[1339,1429,413,437]" italics="true" pageId="2" pageNumber="299">Falsocis</emphasis>
</taxonomicName>
,
<taxonomicName id="15B24D4FFFC2FFA4FF0FC196FF19828C" authorityName="Casey" authorityYear="1898" box="[151,247,450,474]" class="Insecta" family="Ciidae" genus="Plesiocis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FF0FC196FF19828C" box="[151,247,450,474]" italics="true" pageId="2" pageNumber="299">Plesiocis</emphasis>
</taxonomicName>
,
<taxonomicName id="15B24D4FFFC2FFA4FE9CC196FE5D828C" authorityName="Lawrence" authorityYear="2016" box="[260,435,450,474]" genus="Pseudeuxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="299" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FE9CC196FE5D828C" box="[260,435,450,474]" italics="true" pageId="2" pageNumber="299">Pseudeuxestocis</emphasis>
</taxonomicName>
and
<taxonomicName id="15B24D4FFFC2FFA4FE7CC196FDAD828C" authorityName="Lawrence" authorityYear="1987" box="[484,579,450,474]" class="Insecta" family="Ciidae" genus="Porculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FE7CC196FDAD828C" box="[484,579,450,474]" italics="true" pageId="2" pageNumber="299">Porculus</emphasis>
</taxonomicName>
, the outer apical angle of protibiae is angulate and produced to form a stout tooth. Additionally,
<taxonomicName id="15B24D4FFFC2FFA4FEBFC1B3FE3882A9" authorityName="Lawrence" authorityYear="2016" box="[295,470,487,511]" genus="Pseudeuxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="299" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FEBFC1B3FE3882A9" box="[295,470,487,511]" italics="true" pageId="2" pageNumber="299">Pseudeuxestocis</emphasis>
</taxonomicName>
differs in the serrate outer edge of protibiae and the 8-segmented antennae. In
<taxonomicName id="15B24D4FFFC2FFA4FA99C1B3FA7382A9" authorityName="Kawanabe" authorityYear="1994" box="[1281,1437,487,511]" class="Insecta" family="Ciidae" genus="Dichodontocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FA99C1B3FA7382A9" box="[1281,1437,487,511]" italics="true" pageId="2" pageNumber="299">Dichodontocis</emphasis>
</taxonomicName>
and
<taxonomicName id="15B24D4FFFC2FFA4FF5CC259FED28173" authorityName="Lopes-Andrade &amp; Lawrence" authorityYear="2005" box="[196,316,525,549]" class="Insecta" family="Ciidae" genus="Phellinocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FF5CC259FED28173" box="[196,316,525,549]" italics="true" pageId="2" pageNumber="299">Phellinocis</emphasis>
</taxonomicName>
, the outer apical angle of protibiae has two teeth; in the former, the protibiae are serrate at the outer edge while in the latter the elytral apices are strongly declined, convex to slightly concave and often fluted.
</paragraph>
<paragraph id="D20D36CCFFC2FFA4FF5FC20CFD0081C2" blockId="2.[151,1437,151,661]" pageId="2" pageNumber="299">
<taxonomicName id="15B24D4FFFC2FFA4FF5FC20CFEBF8139" authorityName="Miyatake" authorityYear="1954" box="[199,337,600,623]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="E0C6EADEFFC2FFA4FF5FC20CFEBF8139" box="[199,337,600,623]" italics="true" pageId="2" pageNumber="299">Paraxestocis</emphasis>
</taxonomicName>
resembles Orophiini genera in the configuration of the prosternum and tibiae but differs in the procoxae not projected beyond the level of the prosternal process.
</paragraph>
</subSubSection>
<subSubSection id="9AA86547FFC2FFA4FF0FC2BEFCFB80C5" pageId="2" pageNumber="299" type="taxon_list">
<paragraph id="D20D36CCFFC2FFA4FF0FC2BEFE8D8055" blockId="2.[151,913,745,916]" box="[151,355,745,772]" pageId="2" pageNumber="299">
<heading id="894581A0FFC2FFA4FF0FC2BEFE8D8055" bold="true" box="[151,355,745,772]" fontSize="11" level="1" pageId="2" pageNumber="299" reason="1">
<emphasis id="E0C6EADEFFC2FFA4FF0FC2BEFE8D8055" bold="true" box="[151,355,745,772]" pageId="2" pageNumber="299">Included species</emphasis>
</heading>
</paragraph>
<paragraph id="D20D36CCFFC2FFA4FF0FC344FC7F8071" blockId="2.[151,913,745,916]" box="[151,913,783,808]" pageId="2" pageNumber="299">
<taxonomicName id="15B24D4FFFC2FFA4FF0FC344FD2A8071" authority="(Nakane &amp; Nobuchi, 1955)" authorityName="Lopes-Andrade" authorityYear="2024" baseAuthorityName="Nakane &amp; Nobuchi" baseAuthorityYear="1955" box="[151,708,783,808]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="denticollis" status="comb. nov.">
<emphasis id="E0C6EADEFFC2FFA4FF0FC344FE798071" box="[151,407,783,807]" italics="true" pageId="2" pageNumber="299">Paraxestocis denticollis</emphasis>
(
<bibRefCitation id="B6234B3DFFC2FFA4FE3CC35BFD528071" author="Nakane, T. &amp; Nobuchi, A." box="[420,700,783,808]" pageId="2" pageNumber="299" pagination="47 - 52" refId="ref1981" refString="Nakane, T. &amp; Nobuchi, A. (1955) On a new Genus and six new Species of Ciid-Beetles from Japan (Ciidae, Coleoptera). Scientific Reports of the Saikyo University (Natural Science and Living Science), A Series, 2 (1), 47 - 52." type="journal article" year="1955">Nakane &amp; Nobuchi, 1955</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="E0C6EADEFFC2FFA4FD56C344FC7F8071" bold="true" box="[718,913,783,808]" pageId="2" pageNumber="299">
<taxonomicNameLabel id="FBF557A5FFC2FFA4FD56C344FC7F8071" box="[718,913,783,808]" pageId="2" pageNumber="299" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="D20D36CCFFC2FFA4FF0FC360FDD4801D" blockId="2.[151,913,745,916]" box="[151,570,819,843]" pageId="2" pageNumber="299">
<taxonomicName id="15B24D4FFFC2FFA4FF0FC360FDD4801D" authority="Miyatake, 1954" authorityName="Miyatake" authorityYear="1954" box="[151,570,819,843]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="E0C6EADEFFC2FFA4FF0FC360FE63801D" box="[151,397,820,843]" italics="true" pageId="2" pageNumber="299">Paraxestocis unicornis</emphasis>
<bibRefCitation id="B6234B3DFFC2FFA4FE0BC367FDD4801D" author="Miyatake, M." box="[403,570,819,843]" pageId="2" pageNumber="299" pagination="40 - 67" refId="ref1942" refString="Miyatake, M. (1954) Studies on the Japanese Ciidae, I (Coleoptera). The Scientific Reports of the Matsuyama Agricultural College, 14, 40 - 67, pls. 1 - 11." type="journal article" year="1954">Miyatake, 1954</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="D20D36CCFFC2FFA4FF5FC303FD578026" blockId="2.[151,913,745,916]" box="[199,697,855,880]" pageId="2" pageNumber="299">
<taxonomicName id="15B24D4FFFC2FFA4FF5FC303FDBC8039" authority="Nobuchi, 1955" authorityName="Nobuchi" authorityYear="1955" box="[199,594,855,879]" class="Insecta" family="Ciidae" genus="Xylographus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="nakanei">
<emphasis id="E0C6EADEFFC2FFA4FF5FC303FE448039" box="[199,426,855,879]" italics="true" pageId="2" pageNumber="299">Xylographus nakanei</emphasis>
<bibRefCitation id="B6234B3DFFC2FFA4FE29C303FDBC8039" author="Nobuchi, A." box="[433,594,855,879]" pageId="2" pageNumber="299" pagination="53 - 58" refId="ref2037" refString="Nobuchi, A. (1955) Studies on the Ciid-Beetles from Japan. (I) With the Descriptions of a New Genus and Some New Species. (Ciidae, Coleoptera). Entomological Review of Japan, 6 (7), 53 - 58, pl. 12." type="journal article" year="1955">Nobuchi, 1955</bibRefCitation>
</taxonomicName>
, jun. syn.
</paragraph>
<paragraph id="D20D36CCFFC2FFA4FF0FC328FCFB80C5" blockId="2.[151,913,745,916]" box="[151,789,891,916]" pageId="2" pageNumber="299">
<taxonomicName id="15B24D4FFFC2FFA4FF0FC328FDA680C5" authority="(Wankowiez, 1869)" authorityName="Lopes-Andrade" authorityYear="2024" baseAuthorityName="Wankowiez" baseAuthorityYear="1869" box="[151,584,891,915]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="wagae" status="comb. nov.">
<emphasis id="E0C6EADEFFC2FFA4FF0FC328FE8080C5" box="[151,366,892,915]" italics="true" pageId="2" pageNumber="299">Paraxestocis wagae</emphasis>
(
<bibRefCitation id="B6234B3DFFC2FFA4FEE3C32FFDAE80C5" author="Wankowiez, J." box="[379,576,891,915]" pageId="2" pageNumber="299" pagination="411 - 422" refId="ref2361" refString="Wankowiez, J. (1869) Notices entomologiques. Annales de la Societe entomologique de France, 4 (9), 411 - 422." type="journal article" year="1869">Wankowiez, 1869</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="E0C6EADEFFC2FFA4FDCBC328FCFB80C5" bold="true" box="[595,789,891,916]" pageId="2" pageNumber="299">
<taxonomicNameLabel id="FBF557A5FFC2FFA4FDCBC328FCFB80C5" box="[595,789,891,916]" pageId="2" pageNumber="299" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="9AA86547FFC2FFA4FF0FC392FF338725" pageId="2" pageNumber="299" type="discussion">
<paragraph id="D20D36CCFFC2FFA4FF0FC392FF338725" blockId="2.[151,1437,966,1139]" pageId="2" pageNumber="299">
<emphasis id="E0C6EADEFFC2FFA4FF0FC392FEF78088" bold="true" box="[151,281,966,990]" inLineHeading="true" pageId="2" pageNumber="299" reason="1">Comments.</emphasis>
The female terminalia of
<taxonomicName id="15B24D4FFFC2FFA4FDD9C393FD4B8088" authorityName="Lopes-Andrade" authorityYear="2024" baseAuthorityName="Wankowiez" baseAuthorityYear="1869" box="[577,677,967,990]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="wagae">
<emphasis id="E0C6EADEFFC2FFA4FDD9C393FD4B8088" box="[577,677,967,990]" italics="true" pageId="2" pageNumber="299">P. wagae</emphasis>
</taxonomicName>
have reduced paraprocts, shorter than the gonocoxites, with distinct baculi. The gonocoxites are transversely divided into only two lobes ventrally, and basal baculi are distinct. The baculi of proctiger are present, but less sclerotized than the other baculi. The gonostyli are absent. It would be important to examine the female terminalia of
<taxonomicName id="15B24D4FFFC2FFA4FE01C463FDCF8718" authorityName="Lopes-Andrade" authorityYear="2024" baseAuthorityName="Nakane &amp; Nobuchi" baseAuthorityYear="1955" box="[409,545,1078,1102]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="E0C6EADEFFC2FFA4FE01C463FDCF8718" box="[409,545,1078,1102]" italics="true" pageId="2" pageNumber="299">P. denticollis</emphasis>
</taxonomicName>
and
<taxonomicName id="15B24D4FFFC2FFA4FDCBC463FD3F8718" authorityName="Miyatake" authorityYear="1954" box="[595,721,1079,1102]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="E0C6EADEFFC2FFA4FDCBC463FD3F8718" box="[595,721,1079,1102]" italics="true" pageId="2" pageNumber="299">P. unicornis</emphasis>
</taxonomicName>
to evaluate whether this is a pattern within
<taxonomicName id="15B24D4FFFC2FFA4FB00C463FA728718" box="[1176,1436,1079,1102]" class="Insecta" family="Ciidae" genus="Paraxestocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="subSpecies" species="sensu" subSpecies="novo">
<emphasis id="E0C6EADEFFC2FFA4FB00C463FA728718" box="[1176,1436,1079,1102]" italics="true" pageId="2" pageNumber="299">Paraxestocis sensu novo</emphasis>
</taxonomicName>
or not.
</paragraph>
</subSubSection>
</treatment>
</document>