221 lines
28 KiB
XML
221 lines
28 KiB
XML
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<mods:title id="A6698EAD904E154B6D6210CCB8D6A01E">Additions to the taxonomy of New World Pheidole (Hymenoptera: Formicidae).</mods:title>
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<mods:namePart id="7405B2777D9240C80F79A0B928CC45D0">Longino, J. T.</mods:namePart>
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<mods:date id="9E505B2BB488FC5A1FA8E8829F8A81C6">2009</mods:date>
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<mods:identifier id="504D748719D96FF2CA645621340E5CE7" type="HNS-PUB">22820</mods:identifier>
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<treatment id="0C75634EA3D793B882F33AC21E1C5D25" ID-DOI="http://doi.org/10.5281/zenodo.6222821" ID-GBIF-Taxon="100121379" ID-Zenodo-Dep="6222821" LSID="urn:lsid:zoobank.org:act:7BB67F46-FD47-43E5-98A2-129C3311E4F4" httpUri="http://treatment.plazi.org/id/0C75634EA3D793B882F33AC21E1C5D25" lastPageNumber="25" pageNumber="23">
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<subSubSection id="AEA339C9733F92052EEBE99C83AA7F6A" type="nomenclature">
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<paragraph id="B8550B7428EC881AFEC9DBC2A203AE2E" pageNumber="23">
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<taxonomicName id="043AFDE663A3CA73ECB0BECDA55665CA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253256" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole debilis Longino" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="debilis">
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<pageBreakToken id="F57BE0BBE69D5D804DB134876848DC57" pageNumber="23">Pheidole</pageBreakToken>
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debilis
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</taxonomicName>
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<taxonomicNameLabel id="4F5A49DED336A42364517DB225A7EE85" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="ABA091A8C6FBDA355D4A12455DCA3712" type="description">
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<paragraph id="C729303E5E3C56B50665F44E11769B05" pageNumber="23">Figure 4</paragraph>
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</subSubSection>
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<subSubSection id="0C9D6C3816CE90384C73CE83D33FFEE8" type="materials_examined">
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<paragraph id="0A09C538FC26A9B95C0FDE10A06B2035" pageNumber="23">
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Holotype major worker. Costa Rica, Heredia: 16km SSE La Virgen,
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<geoCoordinate id="D63E7230BCDA8AFDE9719D5968C511CB" direction="north" orientation="latitude" value="10.26667">10.26667°N</geoCoordinate>
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<geoCoordinate id="492BD6E7495E9F533FE4792F9CA95B12" direction="west" orientation="longitude" value="-84.08333">84.08333°W</geoCoordinate>
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, ±2000m, 1100m, 15 Feb 2001 (R. Vargas#01-RVC-015) [
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<collectionCode id="6DCDDBA8FFECD0984543764A2A36AF01" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBC</collectionCode>
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, unique specimen identifier INB0003622662].
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</paragraph>
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<paragraph id="6DDB29AD6113A8A7137A8D9A43D5EC91" pageNumber="23">
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Paratypes: major and minor workers. Same data as holotype; same locality as holotype but 17 Nov 2000 (J. Longino#4361) [
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<collectionCode id="AD473A2E8CA58AD6390DCDCCB6023D24" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]">BMNH</collectionCode>
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,
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<collectionCode id="7861B4C6CDF753BAF4AA7056C99EE515" collectionName="USA, California, San Francisco, California Academy of Sciences">CAS</collectionCode>
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,
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<collectionCode id="E4809050CD6317ECE44F54CF6D025706" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)">FMNH</collectionCode>
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,
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<collectionCode id="E7EC7D7D9F2D48BAC3812A10163EC5CC" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBC</collectionCode>
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,
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<collectionCode id="624FC1F0ABA501B7422AF311D00E949A" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
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,
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<collectionCode id="12B0FA757E641D55679A92F2042FC08C" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZ</collectionCode>
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,
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<collectionCode id="45B319190DFCDFA1F246AD69CABA00D6" collectionName="USA, California, Davis, University of California, R.M. Bohart Museum of Entomology">UCD</collectionCode>
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,
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<collectionCode id="D51D602BA54108256D53BC9407838E92" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="9D26662F6A01459AF925910D74897C11" type="distribution">
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<paragraph id="1A902D2E36321C32928B5847E2D8435E" pageNumber="23">Geographic Range</paragraph>
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<paragraph id="AE0715515B7464927F87BA95993EDB62" pageNumber="23">Costa Rica.</paragraph>
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</subSubSection>
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<subSubSection id="B2D656E81CB054F5BC1556C5F6E0228B" type="diagnosis">
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<paragraph id="35D2D6F51CBED78BE6B3BE93C393DA33" pageNumber="23">Diagnosis</paragraph>
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<paragraph id="68E26C10412EFF43384C0B4D85DF84CA" pageNumber="23">
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With the morphometric profile of
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<taxonomicName id="ACD0987F35DE6CEE80CC193C6D355752" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182001" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole gnomus Wilson" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="gnomus">P. gnomus,</taxonomicName>
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<taxonomicName id="BE3EBBD57938B003A61E682BB5F80911" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181900" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="0" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole mera Wilson" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="mera">P. mera</taxonomicName>
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,
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<taxonomicName id="A010653A09B5E5F2558C311634200CF1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253273" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="36" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole sparsisculpta Longino" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="sparsisculpta">P. sparsisculpta</taxonomicName>
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, and
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<taxonomicName id="685FDCDDA85AA0F728BCCD79C467A848" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181943" class="Insecta" family="Formicidae" genus="Pheidole" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole sospes Forel" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="sospes">P. sospes</taxonomicName>
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. Minor worker: face foveolate with overlying rugulae versus face uniformly foveolate, without overlying rugulae (
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<taxonomicName id="51D2C96BE91C14582BB20CE21B62336C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181900" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing" genus.completionRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole mera Wilson" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="mera">mera</taxonomicName>
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,
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<taxonomicName id="7353F43E7F6CC57369ACDAB828B18CA7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181943" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing" genus.completionRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole sospes Forel" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="sospes">sospes</taxonomicName>
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) or with extensive smooth areas (
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<taxonomicName id="DD0BA51C31280189CBF3AB8F99911BAC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253273" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole sparsisculpta Longino" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="sparsisculpta">sparsisculpta</taxonomicName>
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); lateral pronotum foveolate versus with smooth shiny patch (
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<taxonomicName id="993DCFAC8607974D48A85663FC2ABD04" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181900" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole mera Wilson" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="mera">mera</taxonomicName>
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); minor worker of
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<taxonomicName id="139ADE7C41B62C1FFA03ADE47A98EF3A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182001" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole gnomus Wilson" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="gnomus">gnomus</taxonomicName>
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unknown. Major worker: vertex lobes weakly sculptured with faint foveolate sculpture versus vertex lobes with pronounced transversely arcuate rugae (
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<taxonomicName id="F48D7F4B47C66CB0D0DC50DF1D48D86C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182001" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole gnomus Wilson" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="gnomus">gnomus</taxonomicName>
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), or coarsely reticulate rugose (
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<taxonomicName id="1C6A638011BBE1AC33C1C63235E17A19" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181943" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole sospes Forel" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="sospes">sospes</taxonomicName>
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), or completely smooth and shining (
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<taxonomicName id="BB663B8B0F768D618BA470484A557903" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253273" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole sparsisculpta Longino" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="sparsisculpta">sparsisculpta</taxonomicName>
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); head in lateral view normally convex versus with pronounced medial bulge on face (
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<taxonomicName id="900E27F4E7EC06CCAA544E1C6BDCE86A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182001" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole gnomus Wilson" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="gnomus">gnomus</taxonomicName>
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); lacking antennal scrobe versus with shallow but distinct antennal scrobe (
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<taxonomicName id="BC16CF92B6C5F75A5AA7887926DAF1B5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181900" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole mera Wilson" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="mera">mera</taxonomicName>
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,
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<taxonomicName id="B1C87D771CA5FBD1C10CDCCC9BE2B106" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181943" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole sospes Forel" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="sospes">sospes</taxonomicName>
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); promesonotal groove present versus promesonotum evenly arched, with no trace of promesonotal groove (
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<taxonomicName id="29CC6BD855A64A82A024C181397022D4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181900" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole mera Wilson" order="Hymenoptera" pageNumber="23" phylum="Arthropoda" rank="species" species="mera">mera</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="67DDA8063053F6B776FC89ABDD55F867" type="description">
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<paragraph id="E2146EF8C58E72DAFFEBE637BE0E7E5D" pageNumber="23">Description of minor worker</paragraph>
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<paragraph id="5CAA643A52E741C0764AAFA292F2AE7C" pageNumber="23">Measurements (paratype): HL 0.52, HW 0.49, HLA 0.17, SL 0.42, EL 0.09, ML 0.60, PSL 0.06, PMG 0.00, SPL 0.03, PTW 0.09, PPW 0.14, CI 93, SI 86, PSLI 11, PMGI 0, SPLI 6, PPI 152.</paragraph>
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<paragraph id="339C5BE8C23A2AD84E69A33810ED2771" pageNumber="23">Measurements (n=13): HL 0.46-0.55, HW 0.42-0.49, SL 0.36-0.46, CI 88-93, SI 84-95.</paragraph>
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<paragraph id="A4D4D37A3E5A185A5F21E5718541A5E6" pageNumber="23">Clypeus smooth with 1-3 longitudinal rugulae; eye small, with fewer than 10 ommatidia; face densely foveolate, overlain with reticulate rugulae; margin of vertex flattened with weak median impression; occipital carina narrow, not visible in full face view; scape with abundant subdecumbent to suberect setae, mostly shorter than maximum width of scape; promesonotal groove weakly impressed; propodeal spines present; entire mesosoma foveolate, overlain with faint reticulate rugulae on promesonotal dorsum; abundant setae on promesonotal dorsum; dorsal (outer) margin of hind tibia with short decumbent pilosity, no long erect setae; first gastral tergum smooth and shining; gastral dorsum with moderately abundant suberect setae; color variable, light to dark red brown.</paragraph>
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<paragraph id="994F0127D4F199750A122718890182C9" pageNumber="23">Description of major worker</paragraph>
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<paragraph id="F322F1C9879C52BF80F767CAECA7307E" pageNumber="23">Measurements (holotype): HL 1.13, HW 0.94, HLA 0.30, SL 0.48, EL 0.14, ML 0.81, PSL 0.06, PMG 0.02, SPL 0.06, PTW 0.19, PPW 0.38, IHT 0.18, OHT 0.41, CI 83, SI 51, PSLI 5, PMGI 2, SPLI 6, PPI 204, HTI 43.</paragraph>
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<paragraph id="A25922402E3D9E634A61A081F54F9C4E" pageNumber="23">Measurements (n=11): HL 0.92-1.15, HW 0.80-1.04, SL 0.42-0.52, CI 81-88, SI 50-56.</paragraph>
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<paragraph id="DC9EC3FB834785DDB0DC09EDC3FA1CD7" lastPageNumber="24" pageNumber="23">
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Mandibles smooth and shiny; clypeus smooth and shallowly concave, with shallow anterior notch, and very weak posteromedian gibbosity or carina; face with longitudinal rugulae on cheeks and between frontal carinae, grading to faint foveolate sculpture overlain with reticulate rugulae on scrobe, fading to very faint foveolate sculpture on posterior vertex lobes; scrobe a faint, unbounded impression; head with abundant suberect setae projecting from sides of head in face view; scape smooth and shining, terete at base, with subdecumbent pubescence and scattered longer suberect setae; hypostomal margin flat; median tooth small; inner hypostomal teeth pointed, stout, closer to midline than to outer hypostomal teeth; promesonotal groove impressed; propodeal spines present; mesosoma with a mix of foveolate, smooth, and rugulose sculpture; dorsal (outer) margin of hind tibia with short decumbent pilosity, no long erect hairs; pilosity abundant on
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<pageBreakToken id="F7D2F910F7E4B31221D1F0BE3D32EC8F" pageNumber="24">mesosomal</pageBreakToken>
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dorsum; first gastral tergite smooth and shining, with abundant flexuous erect setae; color orange brown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C8CE1BF8FFBC4F065967CB37F1FFD2A1" type="multiple">
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<caption id="BEA6BFA3EA2AB9E98609527B8D8A8635">
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<paragraph id="ED390291D00C1153530CA59F48B53E56" pageNumber="24">
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FIGURE 4.
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<taxonomicName id="E3FAA2C318F9B086302FF25673FB0945" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253256" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole debilis Longino" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="debilis">Pheidole debilis</taxonomicName>
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. Major worker: A, face view; B, lateral view; C, dorsal view; D, hypostomal margin. Minor worker: E, face view; F, lateral view; G, dorsal view; H, hind tibia. A-C, Holotype major worker; D, non-type major worker; E-H, Paratype minor worker. Scale bar 0.5mm for E, F, 1mm for others.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="4EAE846A3FAA05D0B6454718981EBC9F" type="biology_ecology">
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<paragraph id="AF35F323ECADE580C49099ADA7A69555" pageNumber="25">
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<pageBreakToken id="863A907DC801B802845D7F3162B79F48" pageNumber="25">Biology</pageBreakToken>
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</paragraph>
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<paragraph id="7F67769E4B3676C1DD35ABC32AE414F0" pageNumber="25">
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<taxonomicName id="DDDDF03144DA02DB1C877C0F3AE79E67" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253256" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole debilis Longino" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="debilis">Pheidole debilis</taxonomicName>
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occurs in wet forest leaf litter. It is most often collected in Winkler samples of sifted litter but is rarely collected at baits. Major workers may occur together with minor workers in Winkler samples. One nest was collected in soil, and one nest was collected from rotten wood.
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</paragraph>
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</subSubSection>
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<subSubSection id="3E4A299074CB39F8A3426F4F40547B53" type="etymology">
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<paragraph id="0DDC8A81F0F489AED86ED081F61D174D" pageNumber="25">Etymology</paragraph>
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<paragraph id="AFE27E12ACC8897B4E5FA2B07E81CE8A" pageNumber="25">
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The name is in reference to the faint sculpture on the face of the major worker, compared to
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<taxonomicName id="F295AE74C7BD1C375F1DA5C43262AFBE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181882" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="5852" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole harrisonfordi Wilson" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="harrisonfordi">P. harrisonfordi</taxonomicName>
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and other common litter ants that have more heavily sculptured major workers.
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</paragraph>
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</subSubSection>
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<subSubSection id="C5CEE35A4859E0AC5ADF42BD87D15D01" type="discussion">
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<paragraph id="1FFBD864D0AD1B034015CAECC3BF9192" pageNumber="25">Comments</paragraph>
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<paragraph id="4EA413E4A66CD9A4ED28E95604AA4739" pageNumber="25">This species is mainly known from the Barva Transect in Braulio Carrillo National Park, on the Atlantic slope of Costa Rica. It is moderately abundant across a range of elevations, from La Selva Biological Station to 1500m. Minor workers tentatively assigned to this species have been collected elsewhere in Costa Rica: Hitoy Cerere, the Monteverde area, and the southern Pacific lowlands. A collection of minor workers in a Winkler sample from the Osa Peninsula also contained a major worker that matches the typical form in most respects, differing in the lack of any clypeal gibbosity or carina, and the presence of shiny patches on the antennal scrobes.</paragraph>
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<paragraph id="4510986FB06E36D893C58A44207A0893" pageNumber="25">
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In Costa Rica, minor workers of this species can be distinguished from the common
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<taxonomicName id="14CBC6F4B01D88ED7C52BCD0CF9389C7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181882" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="5701" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole harrisonfordi Wilson" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="harrisonfordi">P. harrisonfordi</taxonomicName>
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by more pronounced rugulae on the face and mesosomal dorsum, the slightly impressed promesonotal groove, and the lack of any longer setae above the short, subdecumbent pilosity on the hind tibia. However, these distinctions do not hold over the broad concept of
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<taxonomicName id="CD2B99164957047BE143F9AA50A599FD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181882" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="5651" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole harrisonfordi Wilson" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="harrisonfordi">P. harrisonfordi</taxonomicName>
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as a whole (see Comments under
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<taxonomicName id="67E8941FC97EDCA8C1038B1DBD904647" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181882" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="5641" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole harrisonfordi Wilson" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="harrisonfordi">P. harrisonfordi</taxonomicName>
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). Minor workers are distinguished from the similar
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<taxonomicName id="5728672B655B88E29D1E617B6B4D9A08" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181843" class="Insecta" family="Formicidae" genus="Pheidole" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole beloceps Wilson" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="beloceps">P. beloceps</taxonomicName>
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by the foveolate side of the pronotum (smooth and shiny on
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<taxonomicName id="562513A13FC8719D6850F299AE18E04F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181843" class="Insecta" family="Formicidae" genus="Pheidole" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole beloceps Wilson" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="beloceps">P. beloceps</taxonomicName>
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). Major workers are distinguished from
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<taxonomicName id="1405FC90A49D7DBDCCEEA3A158DBD2F7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181843" class="Insecta" family="Formicidae" genus="Pheidole" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole beloceps Wilson" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="beloceps">P. beloceps</taxonomicName>
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by having a relatively broader head and having the scrobe foveolate (small shiny patches on
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<taxonomicName id="206C730691CECB9B94E03419BC86D2FB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181843" class="Insecta" family="Formicidae" genus="Pheidole" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole beloceps Wilson" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="beloceps">P. beloceps</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="F1A3258B6427E1DD9A527F02CB8FC6AA" type="materials_examined">
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<paragraph id="5AB89CDF07B2B5927C0FEEFB15BC5397" pageNumber="25">Additional material examined</paragraph>
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<paragraph id="3EDAD92E67EFFA63EFF2B36B4FDA6704" pageNumber="25">
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COSTA RICA: Heredia, La Selva Biological Station,
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<geoCoordinate id="B983F701CCC9C311F8B5750E50D89538" direction="north" orientation="latitude" value="10.416667">10°25'N</geoCoordinate>
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,
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<geoCoordinate id="296F0AED0AFB15E4F7A3DCC89AD9C8CE" direction="west" orientation="longitude" value="-84.01667">84°01'W</geoCoordinate>
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, 50m (numerous collectors and collections); 16km SSW Pto. Viejo,
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<geoCoordinate id="77950BF9B55838ADFA027F78FA4BAFB9" direction="north" orientation="latitude" value="10.3175">10°19'03"N</geoCoordinate>
|
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,
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<geoCoordinate id="82AB21C4853BAE9CE1A5692E7989C5CD" direction="west" orientation="longitude" value="-84.04889">84°02'56"W</geoCoordinate>
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, 500m (numerous collectors and collections); Cantarrana, 11km ESE La Virgen,
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|
<geoCoordinate id="C776DA06E38600679DC7F1E735CB9572" direction="north" orientation="latitude" value="10.345278">10°20'43"N</geoCoordinate>
|
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,
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|
<geoCoordinate id="B645A5E93DB16C21884C34E30DABB771" direction="west" orientation="longitude" value="-84.05778">84°03'28"W</geoCoordinate>
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, 300m (numerous collectors and collections); Vara Blanca, 10km NE Vara Blanca,
|
|
<geoCoordinate id="BD6F7E3AED7D1095C89DB664FA5E2041" direction="north" orientation="latitude" value="10.233334">10°14'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="66661A16601FEF8A66F75E67E8A4A745" direction="west" orientation="longitude" value="-84.083336">84°05'W</geoCoordinate>
|
|
, 1500m (numerous collectors and collections); Limon, Hitoy Cerere Biol. Reserve,
|
|
<geoCoordinate id="B8797E4D8E7277BA2FA3CE0637628EC8" direction="north" orientation="latitude" value="9.666667">9°40'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="9213DF64FB1FF0087CECAA4B75FF4570" direction="west" orientation="longitude" value="-83.03333">83°02'W</geoCoordinate>
|
|
, 200m (J. Longino); Puntarenas, Monteverde,
|
|
<geoCoordinate id="D08FC4FB5B255B18E05097F03312DD15" direction="north" orientation="latitude" value="10.3">10°18'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="C38A6170F5F11AAECB7F8D0EFB5C063E" direction="west" orientation="longitude" value="-84.8">84°48'W</geoCoordinate>
|
|
, 1500m (L. A. Schonberg); Bijagual, Carara Biol. Reserve,
|
|
<geoCoordinate id="4FCF5BC88B676592B983197E371C2AA5" direction="north" orientation="latitude" value="9.783334">9°47'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="F2F324DD00C938F12905DF1046C36FC4" direction="west" orientation="longitude" value="-84.6">84°36'W</geoCoordinate>
|
|
, 500m (J. Longino); Est. Biol. Los Llanos, near Santa Elena,
|
|
<geoCoordinate id="84012775CE4F49E357F13792E138E63E" direction="north" orientation="latitude" value="10.305">10°18'18"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="647181A54320BE937E3DD1A9A4DA86E5" direction="west" orientation="longitude" value="-84.83723">84°50'14"W</geoCoordinate>
|
|
, 1150m (J. Longino); 13km SSW Pto. Jimenez,
|
|
<geoCoordinate id="101E16BC4504FD3A219408002E9A8CFC" direction="north" orientation="latitude" value="8.406667">8°24'24"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="C6D1CDE722D0BFD9046F132D1EC5829F" direction="west" orientation="longitude" value="-83.32833">83°19'42"W</geoCoordinate>
|
|
, 130m (J. Longino).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |