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<document id="02DB0A5CCA4158683C1A5629DB63EB27" ID-CLB-Dataset="27634" ID-DOI="10.11646/zootaxa.4545.2.8" ID-GBIF-Dataset="9e5c003e-59a5-4d2e-abc6-664d9450042a" ID-ISSN="1175-5326" ID-Zenodo-Dep="2618856" ID-ZooBank="A486724D-1981-4A38-ADD6-48834837D1FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553930735751" checkinUser="plazi" docAuthor="Garrison, Rosser W. &amp; Ramón Cabrera, Giovanni M." docDate="2019" docId="03F947798B69BF76FF20FD9BFC26FAB6" docLanguage="en" docName="zootaxa.4545.2.8.pdf" docOrigin="Zootaxa 4545 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Argia mauffrayi Garrison &amp; Ramón Cabrera 2019, n. sp." docType="treatment" docVersion="8" lastPageNumber="291" masterDocId="FFC03F018B68BF73FFB7FFF3FFA7FFB9" masterDocTitle="Argia mauffrayi n. sp. from Ecuador (Odonata: Coenagrionidae)" masterLastPageNumber="292" masterPageNumber="286" pageNumber="287" updateTime="1698715938196" updateUser="ExternalLinkService">
<mods:mods id="C258DF79DFB695C31623258AA583C290" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="825A1F56A4893414F6FDF3DE4CA21F33">
<mods:title id="29F4DB8EB5FEC6C5AAA1382EBB1B83AC">Argia mauffrayi n. sp. from Ecuador (Odonata: Coenagrionidae)</mods:title>
</mods:titleInfo>
<mods:name id="517E1EE85698ADF5D3C421CF1A741110" type="personal">
<mods:role id="77B36DBD40A0B310176F847AC7437D19">
<mods:roleTerm id="2A2B54B832BF463E3C6BEF296DD06A3F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9BF0FC522A643ECE79CFA84884D6B4FE">Garrison, Rosser W.</mods:namePart>
</mods:name>
<mods:name id="0FFD61BD3CEC551F8F2B57E9ED02A1E4" type="personal">
<mods:role id="90EB880D65C728AD4821BF0BF501F83B">
<mods:roleTerm id="AA20AEC76D1A893ABC7EF9326340FD60">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3A3B5D443F8FE641F5126BCE6FADF50E">Ramón Cabrera, Giovanni M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="019CDC71AD553C289CCEB53BC9214343">text</mods:typeOfResource>
<mods:relatedItem id="43368AE88B7F92E30D09A357DB3D8C7E" type="host">
<mods:titleInfo id="A9155175A05025F852D677E98AE03974">
<mods:title id="A146606A98149A18C2CA876F88949389">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="92FD48697D9538BB4FBC8D01297447A1">
<mods:date id="9F3A7385D65D067A203FE45170837D18">2019</mods:date>
<mods:detail id="1B3CFDD8903FAD0CCDE13911937CD6F7" type="pubDate">
<mods:number id="9687EEF062E59E2650862136CF76A5B5">2019-01-17</mods:number>
</mods:detail>
<mods:detail id="094CDFC3255588A1D75B85E8D3024817" type="volume">
<mods:number id="BB7A2AF0DCB135ACB82AB50C5C863A03">4545</mods:number>
</mods:detail>
<mods:detail id="8FC5DF8BF6A927DF2D2D73C2ED0E9853" type="issue">
<mods:number id="A164D0CC3750E286D08D704B98D2F100">2</mods:number>
</mods:detail>
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<mods:start id="5BBA0A2D7503F614FB902CF6F4361309">286</mods:start>
<mods:end id="9B83B317E6EAAAE3CCADB2EE6E238D31">292</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="611F7D5A75F4EA14C464B5B6C4508699">journal article</mods:classification>
<mods:identifier id="9A0010EDEFCD6D43F9AEA41F600A54CC" type="CLB-Dataset">27634</mods:identifier>
<mods:identifier id="956F175C4988A7A6429AF5214ADDDBDC" type="DOI">10.11646/zootaxa.4545.2.8</mods:identifier>
<mods:identifier id="E09FC54C96B6E8C3FD0428DDAE2C66CD" type="GBIF-Dataset">9e5c003e-59a5-4d2e-abc6-664d9450042a</mods:identifier>
<mods:identifier id="1FB9E504AEBDD1D5945DBE2148628FF8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="37DE58790F5B9A612CFE9B231C40CAD9" type="Zenodo-Dep">2618856</mods:identifier>
<mods:identifier id="F1287BD8B0A1FC9BA4E4BE80D8370125" type="ZooBank">A486724D-1981-4A38-ADD6-48834837D1FE</mods:identifier>
</mods:mods>
<treatment id="03F947798B69BF76FF20FD9BFC26FAB6" ID-DOI="http://doi.org/10.5281/zenodo.5934139" ID-GBIF-Taxon="154531098" ID-Zenodo-Dep="5934139" LSID="urn:lsid:plazi:treatment:03F947798B69BF76FF20FD9BFC26FAB6" httpUri="http://treatment.plazi.org/id/03F947798B69BF76FF20FD9BFC26FAB6" lastPageId="5" lastPageNumber="291" pageId="1" pageNumber="287">
<subSubSection id="C34AA5E48B69BF72FF20FD9BFCBDFD3A" box="[151,794,616,643]" pageId="1" pageNumber="287" type="nomenclature">
<paragraph id="8BEFF66F8B69BF72FF20FD9BFCBDFD3A" blockId="1.[151,794,616,643]" box="[151,794,616,643]" pageId="1" pageNumber="287">
<heading id="D0A741038B69BF72FF20FD9BFCBDFD3A" bold="true" box="[151,794,616,643]" fontSize="11" level="1" pageId="1" pageNumber="287" reason="1">
<emphasis id="B9242A7D8B69BF72FF20FD9BFCBDFD3A" bold="true" box="[151,794,616,643]" pageId="1" pageNumber="287">
<taxonomicName id="4C508DEC8B69BF72FF20FD9BFD6CFD3A" ID-CoL="9X22T" authority="Garrison &amp; Ramon-Cabrera" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[151,715,616,643]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="mauffrayi" status="sp. nov.">
<emphasis id="B9242A7D8B69BF72FF20FD9BFEFCFD3B" bold="true" box="[151,347,616,642]" italics="true" pageId="1" pageNumber="287">Argia mauffrayi</emphasis>
Garrison &amp; Ramón-Cabrera
</taxonomicName>
,
<taxonomicNameLabel id="A21797068B69BF72FD60FD9AFCBDFD3A" box="[727,794,617,643]" pageId="1" pageNumber="287" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34AA5E48B69BF72FF20FD5DFEA3FCB6" pageId="1" pageNumber="287" type="description">
<paragraph id="8BEFF66F8B69BF72FF20FD5DFEA3FCB6" blockId="1.[151,1437,686,2010]" pageId="1" pageNumber="287">
<figureCitation id="136BEAEA8B69BF72FF20FD5DFF40FD7E" box="[151,231,686,711]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="1" pageNumber="287">Figs. 1</figureCitation>
(head, thorax, S1–4), 2 (S7–10 lateral), 3 (S5–10 dorsal); 4 (appendages mediodorsal); 5 (appendages lateral), 6 (cerci dorsal), 7 (genital ligula lateral), 8 (genital ligula entolateral); 9 (genital ligula ectal); 10 (thorax), 18 (map).
</paragraph>
</subSubSection>
<subSubSection id="C34AA5E48B69BF72FF70FCE9FEDFFB97" pageId="1" pageNumber="287" type="etymology">
<paragraph id="8BEFF66F8B69BF72FF70FCE9FC13FCEE" blockId="1.[151,1437,686,2010]" pageId="1" pageNumber="287">
<emphasis id="B9242A7D8B69BF72FF70FCE9FEE1FC8A" bold="true" box="[199,326,794,819]" pageId="1" pageNumber="287">Etymology</emphasis>
. Named
<taxonomicName id="4C508DEC8B69BF72FE19FCE9FD86FC8A" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[430,545,794,819]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="287" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="B9242A7D8B69BF72FE19FCE9FD86FC8A" bold="true" box="[430,545,794,819]" italics="true" pageId="1" pageNumber="287">mauffrayi</emphasis>
</taxonomicName>
(Latinized name) in honor of the collector William F. Mauffray in recognition to his valuable contributions to the knowledge of Neotropical
<taxonomicName id="4C508DEC8B69BF72FCFAFCCDFC17FCEE" authorityName="Fabricius" authorityYear="1793" box="[845,944,830,855]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="287" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
.
</paragraph>
<paragraph id="8BEFF66F8B69BF72FF70FC91FD56FC5E" blockId="1.[151,1437,686,2010]" pageId="1" pageNumber="287">
<emphasis id="B9242A7D8B69BF72FF70FC91FE63FCC2" bold="true" box="[199,452,866,891]" pageId="1" pageNumber="287">Specimens examined</emphasis>
.
<specimenCount id="9D563DE68B69BF72FE6FFC90FDB8FCC3" box="[472,543,867,891]" pageId="1" pageNumber="287" type="male">2 ♂♂</specimenCount>
.
<emphasis id="B9242A7D8B69BF72FD98FC91FDD4FCC2" bold="true" box="[559,627,866,891]" pageId="1" pageNumber="287">Types</emphasis>
.
<emphasis id="B9242A7D8B69BF72FD3FFC91FD52FCC2" bold="true" box="[648,757,866,891]" pageId="1" pageNumber="287">
<typeStatus id="54EB48CD8B69BF72FD3FFC91FD52FCC2" box="[648,757,866,891]" pageId="1" pageNumber="287" type="holotype">Holotype</typeStatus>
</emphasis>
<materialsCitation id="3B38FC328B69BF72FCB5FC90FD8FFC7A" ID-GBIF-Occurrence="2147526480" collectingDate="2018-09-04" collectionCode="ZSFQ" country="Ecuador" county="San Miguel de Los Bancos Canton" elevation="1101" latitude="0.0333" location="Milpe Bird Sanctuary" longLatPrecision="7" longitude="-78.8661" municipality="Recinto Milpe" pageId="1" pageNumber="287" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha" typeStatus="paratype">
<specimenCount id="9D563DE68B69BF72FCB5FC90FCBDFCC3" box="[770,794,867,890]" pageId="1" pageNumber="287" type="male">♂</specimenCount>
:
<collectingCountry id="F347B6FF8B69BF72FC9CFC90FC12FCC2" box="[811,949,867,891]" name="Ecuador" pageId="1" pageNumber="287">ECUADOR</collectingCountry>
,
<collectingRegion id="4994388D8B69BF72FC71FC91FB01FCC2" box="[966,1190,866,891]" country="Ecuador" name="Pichincha" pageId="1" pageNumber="287">Pichincha Province</collectingRegion>
,
<collectingCounty id="628E8EE38B69BF72FB00FC90FEE0FC26" pageId="1" pageNumber="287">San Miguel de Los Bancos Cantón</collectingCounty>
,
<collectingMunicipality id="6B8B6C158B69BF72FEE2FC75FE5BFC26" box="[341,508,902,927]" pageId="1" pageNumber="287">Recinto Milpe</collectingMunicipality>
,
<location id="8E8FA0B48B69BF72FDBEFC75FCA5FC26" LSID="urn:lsid:plazi:treatment:03F947798B69BF76FF20FD9BFC26FAB6:8E8FA0B48B69BF72FDBEFC75FCA5FC26" box="[521,770,902,927]" country="Ecuador" county="San Miguel de Los Bancos Canton" latitude="0.0333" longLatPrecision="7" longitude="-78.8661" municipality="Recinto Milpe" name="Milpe Bird Sanctuary" pageId="1" pageNumber="287" stateProvince="Pichincha">Milpe Bird Sanctuary</location>
,
<location id="8E8FA0B48B69BF72FCA7FC75FC4FFC26" LSID="urn:lsid:plazi:treatment:03F947798B69BF76FF20FD9BFC26FAB6:8E8FA0B48B69BF72FCA7FC75FC4FFC26" box="[784,1000,902,927]" country="Ecuador" county="San Miguel de Los Bancos Canton" latitude="0.0333" longLatPrecision="7" longitude="-78.8661" municipality="Recinto Milpe" name="Mirador Uno Trail" pageId="1" pageNumber="287" stateProvince="Pichincha">Mirador Uno Trail</location>
,
<geoCoordinate id="EE6490A88B69BF72FC41FC74FBC8FC27" box="[1014,1135,903,927]" direction="north" orientation="latitude" pageId="1" pageNumber="287" precision="5" value="0.0333">0.0333° N</geoCoordinate>
,
<geoCoordinate id="EE6490A88B69BF72FBCAFC75FB5AFC27" box="[1149,1277,902,927]" direction="west" orientation="longitude" pageId="1" pageNumber="287" precision="5" value="-78.8661">78.8661 W</geoCoordinate>
,
<quantity id="4CA85B8A8B69BF72FABDFC74FAFAFC26" box="[1290,1373,903,927]" metricMagnitude="3" metricUnit="m" metricValue="1.101" pageId="1" pageNumber="287" unit="m" value="1101.0">
<elevation id="007D115C8B69BF72FABDFC74FAFAFC26" box="[1290,1373,903,927]" metricMagnitude="3" metricUnit="m" metricValue="1.101" pageId="1" pageNumber="287" unit="m" value="1101.0">1101m</elevation>
</quantity>
.,
<date id="FFEED0AF8B69BF72FAD8FC74FF75FC7A" pageId="1" pageNumber="287" value="2018-09-04">
<collectingDate id="EFAA29478B69BF72FAD8FC74FF75FC7A" pageId="1" pageNumber="287" value="2018-09-04">4 ix 2018</collectingDate>
</date>
, WFM leg. (
<collectionCode id="ED416EAA8B69BF72FEE8FC58FE0EFC7A" box="[351,425,939,963]" pageId="1" pageNumber="287">ZSFQ</collectionCode>
);
<emphasis id="B9242A7D8B69BF72FE0BFC59FD8FFC7A" bold="true" box="[444,552,938,963]" pageId="1" pageNumber="287">
<typeStatus id="54EB48CD8B69BF72FE0BFC59FD8FFC7A" box="[444,552,938,963]" pageId="1" pageNumber="287" type="paratype">Paratype</typeStatus>
</emphasis>
</materialsCitation>
<materialsCitation id="3B38FC328B69BF72FD87FC58FD49FC5E" ID-GBIF-Occurrence="2147526533" collectingDate="2012-05-15" collectionCode="RWG" collectorName="Jessica Ware &amp; Melissa Sanchez" country="Ecuador" elevation="470" latitude="0.5203" location="Reserva Canande" longLatPrecision="7" longitude="-79.0866" pageId="1" pageNumber="287" specimenCount="1" specimenCount-male="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCount id="9D563DE68B69BF72FD87FC58FDEFFC7B" box="[560,584,939,962]" pageId="1" pageNumber="287" type="male">♂</specimenCount>
:
<collectingRegion id="4994388D8B69BF72FDE3FC59FCE4FC7A" box="[596,835,938,963]" country="Ecuador" name="Esmeraldas" pageId="1" pageNumber="287">Esmeraldas Province</collectingRegion>
,
<location id="8E8FA0B48B69BF72FCFAFC58FBB3FC7A" LSID="urn:lsid:plazi:treatment:03F947798B69BF76FF20FD9BFC26FAB6:8E8FA0B48B69BF72FCFAFC58FBB3FC7A" box="[845,1044,938,963]" country="Ecuador" latitude="0.5203" longLatPrecision="7" longitude="-79.0866" name="Reserva Canande" pageId="1" pageNumber="287" stateProvince="Esmeraldas">Reserva Canande</location>
,
<geoCoordinate id="EE6490A88B69BF72FBA9FC59FB33FC7B" box="[1054,1172,938,963]" direction="north" orientation="latitude" pageId="1" pageNumber="287" precision="5" value="0.5203">0.5203° N</geoCoordinate>
,
<geoCoordinate id="EE6490A88B69BF72FB17FC59FA8FFC7B" box="[1184,1320,938,963]" direction="west" orientation="longitude" pageId="1" pageNumber="287" precision="5" value="-79.0866">79.0866° W</geoCoordinate>
,
<elevation id="007D115C8B69BF72FA85FC58FAD2FC7A" box="[1330,1397,939,963]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="287" unit="m" value="470.0">
<quantity id="4CA85B8A8B69BF72FA85FC58FAD2FC7A" box="[1330,1397,939,963]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="287" unit="m" value="470.0">470m</quantity>
</elevation>
,
<date id="FFEED0AF8B69BF72FA37FC59FF40FC5E" pageId="1" pageNumber="287" value="2012-05-15">
<collectingDate id="EFAA29478B69BF72FA37FC59FF40FC5E" pageId="1" pageNumber="287" value="2012-05-15">15 v 2012</collectingDate>
</date>
,
<collectorName id="26A593B98B69BF72FF45FC3DFE26FC5F" box="[242,385,974,999]" pageId="1" pageNumber="287">Jessica Ware</collectorName>
&amp;
<collectorName id="26A593B98B69BF72FE12FC3DFDC7FC5E" box="[421,608,974,999]" pageId="1" pageNumber="287">Melissa Sanchez</collectorName>
leg. (
<collectionCode id="ED416EAA8B69BF72FD28FC3CFD42FC5E" box="[671,741,975,999]" pageId="1" pageNumber="287">RWG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEFF66F8B69BF72FF70FC00FEDFFB97" blockId="1.[151,1437,686,2010]" pageId="1" pageNumber="287">A largely orange species with forcipate cercus armed terminally with a decumbent tooth and with a small unilobate paraproct.</paragraph>
</subSubSection>
<subSubSection id="C34AA5E48B69BF76FF70FBC9FA83FF41" lastPageId="5" lastPageNumber="291" pageId="1" pageNumber="287" type="description">
<paragraph id="8BEFF66F8B69BF72FF70FBC9FE52FB06" blockId="1.[151,1437,686,2010]" pageId="1" pageNumber="287">
<emphasis id="B9242A7D8B69BF72FF70FBC9FE7AFBEA" bold="true" box="[199,477,1082,1107]" pageId="1" pageNumber="287">
Description of
<typeStatus id="54EB48CD8B69BF72FEC0FBC9FE7AFBEA" box="[375,477,1082,1107]" pageId="1" pageNumber="287" type="holotype">holotype</typeStatus>
</emphasis>
.
<emphasis id="B9242A7D8B69BF72FE5BFBC9FD8FFBEA" box="[492,552,1082,1107]" italics="true" pageId="1" pageNumber="287">Head</emphasis>
(
<figureCitation id="136BEAEA8B69BF72FD8EFBC9FDDDFBEA" box="[569,634,1082,1107]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="1" pageNumber="287">Fig. 1</figureCitation>
). Labrum, genae, clypeus and antefrons orange, epicranium black with small orange spot anterolateral to lateral ocellus, large orange postocular spots confluent laterally with margin of compound eye; scape orange, pedicel black; rear of head black above, pale orange below with an ill-defined isolated black spot above gena.
</paragraph>
<paragraph id="8BEFF66F8B69BF72FF70FB39FC5BFA02" blockId="1.[151,1437,686,2010]" pageId="1" pageNumber="287">
<emphasis id="B9242A7D8B69BF72FF70FB39FEBFFB5A" box="[199,280,1226,1251]" italics="true" pageId="1" pageNumber="287">Thorax</emphasis>
. Prothorax orange except for anterior margin of anterior lobe black and dorsal portion of middle and hind lobes black with narrow middorsal stripe of orange on middle and posterior lobes. Mesothorax (
<figureCitation id="136BEAEA8B69BF72FAA2FB1DFAF0FABE" box="[1301,1367,1262,1287]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="1" pageNumber="287">Fig. 1</figureCitation>
) with mesepisternum orange, paler orange laterally, black parallel middorsal stripe as wide as pale antehumeral stripe; black humeral stripe divided at upper 0.60, the anterior branch a narrow line along mesopleural suture with mesopleural fossa black, the posterior branch gradually narrowing and ending before subalar carina; metapleural stripe black, slightly narrower than humeral stripe, metapleural carina edged with black; anterior half of mesepisternum and most of metepimeron black; venter of thorax pale orange.
</paragraph>
<paragraph id="8BEFF66F8B69BF72FF70FA34FF4BF9F7" blockId="1.[151,1437,686,2010]" pageId="1" pageNumber="287">
<emphasis id="B9242A7D8B69BF72FF70FA34FEADFA67" box="[199,266,1479,1502]" italics="true" pageId="1" pageNumber="287">Wings</emphasis>
. Slightly flavescent with venation black; pterostigma trapezoidal, dark brown, surmounting one cell in all wings; postnodals Fw 13/13, Hw 12/11; postquadrangular cells Fw 3/3, Hw 3/3; RP
<subScript id="17D4F42A8B69BF72FBDDFA0AFBD5F9BE" attach="left" box="[1130,1138,1529,1543]" fontSize="6" pageId="1" pageNumber="287">2</subScript>
at Fw 6/7, Hw 5/5. Coxae and trochanters orange, femora orange with tip becoming black, tarsi and armature black, tibiae orange, black basally.
</paragraph>
<paragraph id="8BEFF66F8B69BF72FF70F9AAFF7CF8F3" blockId="1.[151,1437,686,2010]" pageId="1" pageNumber="287">
<emphasis id="B9242A7D8B69BF72FF70F9AAFE96F9CB" box="[199,305,1625,1650]" italics="true" pageId="1" pageNumber="287">Abdomen</emphasis>
(
<figureCitation id="136BEAEA8B69BF72FE88F9AAFE0CF9CB" box="[319,427,1625,1650]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="1" pageNumber="287">Figs. 1–3</figureCitation>
). Mostly orange; S1 orange with a wash of black at basal 0.30; S2 orange laterally with an ill-defined black stripe constricted medially, apical annulus black; S3 orange with a poorly defined narrow dark lateral stripe interrupted medially, annulus black; S4–6 similar to S3 but posterior darker portion of succeeding segments becoming more extensive and defined; S7 with orange basal annulus followed by dorsolateral black interrupted dorsally with orange terminating with black at apical fifth; S8 black dorsally, orange laterally (
<figureCitation id="136BEAEA8B69BF72FAFFF91AFA2AF8BB" box="[1352,1421,1769,1794]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="1" pageNumber="287">Fig. 2</figureCitation>
); S9 black with a small longitudinal tear-shaped, middorsal spot (
<figureCitation id="136BEAEA8B69BF72FCE9F8FEFC07F89F" box="[862,928,1805,1830]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="1" pageNumber="287">Fig. 3</figureCitation>
); 10 entirely black; torus orange, appendages black.
</paragraph>
<paragraph id="8BEFF66F8B69BF72FF70F8A6FD32F863" blockId="1.[151,1437,686,2010]" pageId="1" pageNumber="287">
<emphasis id="B9242A7D8B69BF72FF70F8A6FEC0F8D7" box="[199,359,1877,1902]" italics="true" pageId="1" pageNumber="287">Genital ligula</emphasis>
(as in
<figureCitation id="136BEAEA8B69BF72FE0CF8A6FD8EF8D7" box="[443,553,1877,1902]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="1" pageNumber="287">
Figs. 7
<emphasis id="B9242A7D8B69BF72FDBCF8A5FDBEF8D4" box="[523,537,1878,1901]" italics="true" pageId="1" pageNumber="287">‒</emphasis>
9
</figureCitation>
). With a long stiff flagellum, and a shorter medial trilobed process, the two lateral lobes of which are triangular in ventral (ectal) view (as in
<figureCitation id="136BEAEA8B69BF72FC11F88AFC49F82B" box="[934,1006,1913,1938]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="1" pageNumber="287">Fig. 9</figureCitation>
), and the decumbent median lobe is quadrate apically (as in
<figureCitation id="136BEAEA8B69BF72FE19F86EFE52F80F" box="[430,501,1949,1974]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="1" pageNumber="287">Fig. 8</figureCitation>
); the latter process with an evenly curved arcuate ridge; more basally at base of flagellum a pair of decumbent rounded lobes.
</paragraph>
<caption id="DF2FA6E78B6ABF71FF20F887FC9FF871" httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="2" pageNumber="288" startId="2.[151,264,1908,1931]" targetBox="[151,1436,195,1884]" targetPageId="2">
<paragraph id="8BEFF66F8B6ABF71FF20F887FC9FF871" blockId="2.[151,1436,1908,1992]" pageId="2" pageNumber="288">
<emphasis id="B9242A7D8B6ABF71FF20F887FEECF830" bold="true" box="[151,331,1908,1931]" pageId="2" pageNumber="288">FIGURES 1–10.</emphasis>
Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10).
</paragraph>
</caption>
<caption id="DF2FA6E78B6BBF70FF20F855FAACF862" httpUri="https://zenodo.org/record/2618860/files/figure.png" pageId="3" pageNumber="289" startId="3.[151,264,1958,1981]" targetBox="[178,1369,266,1771]" targetPageId="3">
<paragraph id="8BEFF66F8B6BBF70FF20F855FAACF862" blockId="3.[151,1435,1958,2011]" pageId="3" pageNumber="289">
<emphasis id="B9242A7D8B6BBF70FF20F855FEF3F805" bold="true" box="[151,340,1958,1981]" pageId="3" pageNumber="289">FIGURES 11–17.</emphasis>
Color pattern of head, thorax, S1–4, lateral view (11); S7–10, lateral view (12); appendages, mediodorsal view (13); same, lateral view (14); cerci, dorsal view (15); genital ligula, lateral view (16); same ectal view (17).
</paragraph>
</caption>
<caption id="DF2FA6E78B6CBF77FF20F9CAFCAAF9F6" box="[151,781,1593,1615]" httpUri="https://zenodo.org/record/2618862/files/figure.png" pageId="4" pageNumber="290" startId="4.[151,250,1593,1615]" targetBox="[228,1335,239,1493]" targetPageId="4">
<paragraph id="8BEFF66F8B6CBF77FF20F9CAFCAAF9F6" blockId="4.[151,781,1593,1615]" box="[151,781,1593,1615]" pageId="4" pageNumber="290">
<emphasis id="B9242A7D8B6CBF77FF20F9CAFEB8F9F7" bold="true" box="[151,287,1593,1615]" pageId="4" pageNumber="290">FIGURE 18.</emphasis>
Distribution of
<taxonomicName id="4C508DEC8B6CBF77FE75F9C9FDFFF9F6" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[450,600,1593,1615]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="290" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9242A7D8B6CBF77FE75F9C9FDFFF9F6" box="[450,600,1593,1615]" italics="true" pageId="4" pageNumber="290">Argia acridens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C508DEC8B6CBF77FD3EF9C9FCA0F9F6" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[649,775,1593,1615]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="290" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="B9242A7D8B6CBF77FD3EF9C9FCA0F9F6" box="[649,775,1593,1615]" italics="true" pageId="4" pageNumber="290">A. mauffrayi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BEFF66F8B6CBF77FF70F98FFE8BF8D4" blockId="4.[151,1437,1660,2009]" pageId="4" pageNumber="290">
<emphasis id="B9242A7D8B6CBF77FF70F98FFE0DF92C" box="[199,426,1660,1685]" italics="true" pageId="4" pageNumber="290">Caudal appendages</emphasis>
. Torifer orange, torus large, planar, roundly triangular (as in
<figureCitation id="136BEAEA8B6CBF77FBCBF98FFB64F92C" box="[1148,1219,1660,1685]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="4" pageNumber="290">Fig. 4</figureCitation>
), occupying entire ventral margin of torifer and not overlapping bilobed epiproct; epiproct black, small, slightly bilobed (as in
<figureCitation id="136BEAEA8B6CBF77FAFDF953FA2AF900" box="[1354,1421,1696,1721]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="4" pageNumber="290">Fig. 6</figureCitation>
); cercus (as in
<figureCitation id="136BEAEA8B6CBF77FE9CF937FE33F964" box="[299,404,1732,1757]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="4" pageNumber="290">Figs. 4–6</figureCitation>
) slightly longer than paraproct, about three times longer than wide, slightly swollen basally with medial basal margin slightly convex and slightly narrowing distally with a strongly acuminate mediolaterally directed tooth, dorsal surface of cercus with long cluster of thick hairs; mediodorsal surface of cercus slightly depressed along medial margin; paraproct unilobate, almost quadrate in lateral view with a narrow tuberculate tip (as in
<figureCitation id="136BEAEA8B6CBF77FF6CF8A7FEB9F8D4" box="[219,286,1876,1901]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="4" pageNumber="290">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BEFF66F8B6CBF77FF70F88BFE3BF828" blockId="4.[151,1437,1660,2009]" box="[199,412,1912,1937]" pageId="4" pageNumber="290">
<emphasis id="B9242A7D8B6CBF77FF70F88BFEB9F828" bold="true" box="[199,286,1912,1937]" pageId="4" pageNumber="290">Female</emphasis>
. unknown.
</paragraph>
<paragraph id="8BEFF66F8B6CBF77FF70F86FFCFAF80C" blockId="4.[151,1437,1660,2009]" box="[199,861,1948,1973]" pageId="4" pageNumber="290">
<emphasis id="B9242A7D8B6CBF77FF70F86FFED1F80C" bold="true" box="[199,374,1948,1973]" pageId="4" pageNumber="290">Measurements</emphasis>
. Hw 19.0, abdomen 26.5, total length 33.0.
</paragraph>
<paragraph id="8BEFF66F8B6CBF76FF70F833FA83FF41" blockId="4.[151,1437,1660,2009]" lastBlockId="5.[151,1436,151,1295]" lastPageId="5" lastPageNumber="291" pageId="4" pageNumber="290">
<emphasis id="B9242A7D8B6CBF77FF70F833FE6FF860" bold="true" box="[199,456,1984,2009]" pageId="4" pageNumber="290">
Variation in
<typeStatus id="54EB48CD8B6CBF77FEEAF833FE6FF860" box="[349,456,1984,2009]" pageId="4" pageNumber="290" type="paratype">paratype</typeStatus>
</emphasis>
. Similar to
<typeStatus id="54EB48CD8B6CBF77FDFAF833FD08F860" box="[589,687,1984,2009]" pageId="4" pageNumber="290" type="holotype">holotype</typeStatus>
but with black humeral stripe divided at upper 0.70 (e. g.
<figureCitation id="136BEAEA8B6CBF77FA8BF833FA29F860" box="[1340,1422,1984,2009]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="4" pageNumber="290">Fig. 10</figureCitation>
), wings slightly more flavescent and with small middorsal orange stripe confined to hind lobe of prothorax and S9 entirely black. Pterostigma surmounting one cell in all wings; postnodals: Fw 12–12, Hw 11–10; postquadrangular cells Fw 3–3, Hw 3–3; RP
<subScript id="17D4F42A8B6DBF76FE09FF1DFE61FF45" attach="left" box="[446,454,238,252]" fontSize="6" pageId="5" pageNumber="291">2</subScript>
at Fw 6–5, Hw 4–4.
<emphasis id="B9242A7D8B6DBF76FD05FF2CFC9BFF41" bold="true" box="[690,828,223,248]" pageId="5" pageNumber="291">Dimensions</emphasis>
. Hw 18.0, abdomen 24.5, total length 31.0.
</paragraph>
</subSubSection>
<subSubSection id="C34AA5E48B6DBF76FF70FEF5FE3BFD76" pageId="5" pageNumber="291" type="diagnosis">
<paragraph id="8BEFF66F8B6DBF76FF70FEF5FE3BFD76" blockId="5.[151,1436,151,1295]" pageId="5" pageNumber="291">
<emphasis id="B9242A7D8B6DBF76FF70FEF5FE9DFEA6" bold="true" box="[199,314,262,287]" pageId="5" pageNumber="291">Diagnosis</emphasis>
. This species is unique by overall orange coloration, mostly black S9 and 10, and genital ligula and caudal appendage morphology. The only other South American species of
<taxonomicName id="4C508DEC8B6DBF76FC5FFED8FB80FEFB" box="[1000,1063,299,322]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="B9242A7D8B6DBF76FC5FFED8FB80FEFB" box="[1000,1063,299,322]" italics="true" pageId="5" pageNumber="291">Argia</emphasis>
</taxonomicName>
with a largely orange body is
<taxonomicName id="4C508DEC8B6DBF76FA33FED8FEE6FEDE" authority="Selys" authorityName="Selys" authorityYear="1865" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="infumata">
<emphasis id="B9242A7D8B6DBF76FA33FED8FF5CFEDE" italics="true" pageId="5" pageNumber="291">A. infumata</emphasis>
Selys
</taxonomicName>
but it differs greatly in genital and appendage morphology (Garrison &amp; von
<bibRefCitation id="EFC18B9E8B6DBF76FB7AFEBDFA36FEDE" author="Garrison, R. W. &amp; von Ellenrieder, N." box="[1229,1425,334,359]" pageId="5" pageNumber="291" pagination="1 - 134" refId="ref2508" refString="Garrison, R. W. &amp; von Ellenrieder, N. (2015) Damselflies of the genus Argia of the Guiana Shield (Odonata: Coenagrionidae). Zootaxa, 4042 (1), 1 - 134. https: // doi. org / 10.11646 / zootaxa. 4042.1.1" type="journal article" year="2015">Ellenrieder 2015</bibRefCitation>
). Genital ligula and appendage morphology of
<taxonomicName id="4C508DEC8B6DBF76FD0BFE80FCE8FE32" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[700,847,371,395]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="B9242A7D8B6DBF76FD0BFE80FCE8FE32" box="[700,847,371,395]" italics="true" pageId="5" pageNumber="291">A. mauffrayi</emphasis>
</taxonomicName>
resemble those of
<taxonomicName id="4C508DEC8B6DBF76FB89FE80FEFCFE16" authority="Garrison &amp; von Ellenrieder, 2018" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B9242A7D8B6DBF76FB89FE80FB76FE32" box="[1086,1233,370,395]" italics="true" pageId="5" pageNumber="291">A. cuspidata</emphasis>
Garrison &amp; von
<bibRefCitation id="EFC18B9E8B6DBF76FF20FE65FEFCFE16" author="Garrison, R. W. &amp; von Ellenrieder, N." box="[151,347,406,431]" pageId="5" pageNumber="291" pagination="1 - 69" refId="ref2561" refString="Garrison, R. W. &amp; von Ellenrieder, N. (2018) Damselflies of the genus Argia (Odonata: Coenagrionidae) from Ecuador with descriptions of five new species. Zootaxa, 4470 (1), 1 - 69. https: // doi. org / 10.11646 / zootaxa. 4470.1.1" type="journal article" year="2018">Ellenrieder, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C508DEC8B6DBF76FE2FFE64FDBCFE16" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[408,539,406,431]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9242A7D8B6DBF76FE2FFE64FDBCFE16" box="[408,539,406,431]" italics="true" pageId="5" pageNumber="291">A. acridens</emphasis>
</taxonomicName>
, although appendage morphology is most similar to that of
<taxonomicName id="4C508DEC8B6DBF76FB6EFE64FAFCFE16" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[1241,1371,406,431]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9242A7D8B6DBF76FB6EFE64FAFCFE16" box="[1241,1371,406,431]" italics="true" pageId="5" pageNumber="291">A. acridens</emphasis>
</taxonomicName>
(
<figureCitation id="136BEAEA8B6DBF76FAD9FE65FF47FE6A" captionStart="FIGURES 11–17" captionStartId="3.[151,264,1958,1981]" captionTargetBox="[178,1369,266,1771]" captionTargetId="figure@3.[151,1436,225,1914]" captionTargetPageId="3" captionText="FIGURES 11–17. Color pattern of head, thorax, S1–4, lateral view (11); S7–10, lateral view (12); appendages, mediodorsal view (13); same, lateral view (14); cerci, dorsal view (15); genital ligula, lateral view (16); same ectal view (17)." httpUri="https://zenodo.org/record/2618860/files/figure.png" pageId="5" pageNumber="291">Fig. 11–17</figureCitation>
). However, they differ by: 1) dorsal surface of cercus covered with thick hairs in
<taxonomicName id="4C508DEC8B6DBF76FB30FE48FAB2FE6A" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[1159,1301,443,467]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="B9242A7D8B6DBF76FB30FE48FAB2FE6A" box="[1159,1301,443,467]" italics="true" pageId="5" pageNumber="291">A. mauffrayi</emphasis>
</taxonomicName>
(
<figureCitation id="136BEAEA8B6DBF76FA91FE49FA34FE6A" box="[1318,1427,442,467]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="5" pageNumber="291">Figs. 4, 5</figureCitation>
) compared to being bare in
<taxonomicName id="4C508DEC8B6DBF76FE75FE2CFDE6FE4E" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[450,577,478,503]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9242A7D8B6DBF76FE75FE2CFDE6FE4E" box="[450,577,478,503]" italics="true" pageId="5" pageNumber="291">A. acridens</emphasis>
</taxonomicName>
(
<figureCitation id="136BEAEA8B6DBF76FDE6FE2DFD7FFE4E" box="[593,728,478,503]" captionStart="FIGURES 11–17" captionStartId="3.[151,264,1958,1981]" captionTargetBox="[178,1369,266,1771]" captionTargetId="figure@3.[151,1436,225,1914]" captionTargetPageId="3" captionText="FIGURES 11–17. Color pattern of head, thorax, S1–4, lateral view (11); S7–10, lateral view (12); appendages, mediodorsal view (13); same, lateral view (14); cerci, dorsal view (15); genital ligula, lateral view (16); same ectal view (17)." httpUri="https://zenodo.org/record/2618860/files/figure.png" pageId="5" pageNumber="291">Figs. 13, 14</figureCitation>
), and 2) the cercus in dorsal view being slightly forcipate, with basomedial margin only slightly convex in
<taxonomicName id="4C508DEC8B6DBF76FD32FDF0FCB4FDA2" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[645,787,515,539]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="B9242A7D8B6DBF76FD32FDF0FCB4FDA2" box="[645,787,515,539]" italics="true" pageId="5" pageNumber="291">A. mauffrayi</emphasis>
</taxonomicName>
(
<figureCitation id="136BEAEA8B6DBF76FC93FDF1FCCEFDA2" box="[804,873,514,539]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="5" pageNumber="291">Fig. 6</figureCitation>
) versus greatly expanded into a basomesal round lobe as in
<taxonomicName id="4C508DEC8B6DBF76FEBCFDD4FE2CFD86" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[267,395,550,575]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9242A7D8B6DBF76FEBCFDD4FE2CFD86" box="[267,395,550,575]" italics="true" pageId="5" pageNumber="291">A. acridens</emphasis>
</taxonomicName>
(
<figureCitation id="136BEAEA8B6DBF76FE2CFDD5FE48FD86" box="[411,495,550,575]" captionStart="FIGURES 11–17" captionStartId="3.[151,264,1958,1981]" captionTargetBox="[178,1369,266,1771]" captionTargetId="figure@3.[151,1436,225,1914]" captionTargetPageId="3" captionText="FIGURES 11–17. Color pattern of head, thorax, S1–4, lateral view (11); S7–10, lateral view (12); appendages, mediodorsal view (13); same, lateral view (14); cerci, dorsal view (15); genital ligula, lateral view (16); same ectal view (17)." httpUri="https://zenodo.org/record/2618860/files/figure.png" pageId="5" pageNumber="291">Fig. 15</figureCitation>
). Additionally, the torus in
<taxonomicName id="4C508DEC8B6DBF76FC92FDD4FC15FD86" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[805,946,551,575]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="B9242A7D8B6DBF76FC92FDD4FC91FD87" box="[805,822,551,574]" italics="true" pageId="5" pageNumber="291">A</emphasis>
.
<emphasis id="B9242A7D8B6DBF76FCF2FDD4FC15FD86" box="[837,946,551,575]" italics="true" pageId="5" pageNumber="291">mauffrayi</emphasis>
</taxonomicName>
(
<figureCitation id="136BEAEA8B6DBF76FC75FDD5FBA1FD86" box="[962,1030,550,575]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="5" pageNumber="291">Fig. 4</figureCitation>
) is roughly triangular but is smaller and more oval in
<taxonomicName id="4C508DEC8B6DBF76FEEAFDB8FE7AFDDA" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[349,477,586,611]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9242A7D8B6DBF76FEEAFDB8FE7AFDDA" box="[349,477,586,611]" italics="true" pageId="5" pageNumber="291">A. acridens</emphasis>
</taxonomicName>
(
<figureCitation id="136BEAEA8B6DBF76FE5AFDB9FDE6FDDA" box="[493,577,586,611]" captionStart="FIGURES 11–17" captionStartId="3.[151,264,1958,1981]" captionTargetBox="[178,1369,266,1771]" captionTargetId="figure@3.[151,1436,225,1914]" captionTargetPageId="3" captionText="FIGURES 11–17. Color pattern of head, thorax, S1–4, lateral view (11); S7–10, lateral view (12); appendages, mediodorsal view (13); same, lateral view (14); cerci, dorsal view (15); genital ligula, lateral view (16); same ectal view (17)." httpUri="https://zenodo.org/record/2618860/files/figure.png" pageId="5" pageNumber="291">Fig. 13</figureCitation>
). The apical process of the paraproct of
<taxonomicName id="4C508DEC8B6DBF76FBBFFDB8FB33FDDA" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[1032,1172,587,611]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="B9242A7D8B6DBF76FBBFFDB8FB33FDDA" box="[1032,1172,587,611]" italics="true" pageId="5" pageNumber="291">A. mauffrayi</emphasis>
</taxonomicName>
in lateral view is small and minutely bilobed (
<figureCitation id="136BEAEA8B6DBF76FE21FD9DFE7EFD3E" box="[406,473,622,647]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="5" pageNumber="291">Fig. 5</figureCitation>
) whereas the apical process in
<taxonomicName id="4C508DEC8B6DBF76FC8BFD9CFC1BFD3E" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[828,956,622,647]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9242A7D8B6DBF76FC8BFD9CFC1BFD3E" box="[828,956,622,647]" italics="true" pageId="5" pageNumber="291">A. acridens</emphasis>
</taxonomicName>
is broader and roughly quadrate (
<figureCitation id="136BEAEA8B6DBF76FA8CFD9DFA28FD3E" box="[1339,1423,622,647]" captionStart="FIGURES 11–17" captionStartId="3.[151,264,1958,1981]" captionTargetBox="[178,1369,266,1771]" captionTargetId="figure@3.[151,1436,225,1914]" captionTargetPageId="3" captionText="FIGURES 11–17. Color pattern of head, thorax, S1–4, lateral view (11); S7–10, lateral view (12); appendages, mediodorsal view (13); same, lateral view (14); cerci, dorsal view (15); genital ligula, lateral view (16); same ectal view (17)." httpUri="https://zenodo.org/record/2618860/files/figure.png" pageId="5" pageNumber="291">Fig. 14</figureCitation>
). The basal trilobate process and paired lateral lobes in the genital ligula in
<taxonomicName id="4C508DEC8B6DBF76FC67FD60FBFBFD12" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[976,1116,659,683]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="B9242A7D8B6DBF76FC67FD60FBFBFD12" box="[976,1116,659,683]" italics="true" pageId="5" pageNumber="291">A. mauffrayi</emphasis>
</taxonomicName>
(
<figureCitation id="136BEAEA8B6DBF76FBDCFD61FB71FD12" box="[1131,1238,658,683]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="5" pageNumber="291">Figs. 7–9</figureCitation>
) are lacking in
<taxonomicName id="4C508DEC8B6DBF76FA33FD60FF50FD76" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9242A7D8B6DBF76FA33FD60FF50FD76" italics="true" pageId="5" pageNumber="291">A. acridens</emphasis>
</taxonomicName>
(
<figureCitation id="136BEAEA8B6DBF76FEB1FD45FE29FD76" box="[262,398,694,719]" captionStart="FIGURES 11–17" captionStartId="3.[151,264,1958,1981]" captionTargetBox="[178,1369,266,1771]" captionTargetId="figure@3.[151,1436,225,1914]" captionTargetPageId="3" captionText="FIGURES 11–17. Color pattern of head, thorax, S1–4, lateral view (11); S7–10, lateral view (12); appendages, mediodorsal view (13); same, lateral view (14); cerci, dorsal view (15); genital ligula, lateral view (16); same ectal view (17)." httpUri="https://zenodo.org/record/2618860/files/figure.png" pageId="5" pageNumber="291">Figs. 16, 17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34AA5E48B6DBF76FF70FD29FC26FAB6" pageId="5" pageNumber="291" type="discussion">
<paragraph id="8BEFF66F8B6DBF76FF70FD29FAE6FBAA" blockId="5.[151,1436,151,1295]" pageId="5" pageNumber="291">
<emphasis id="B9242A7D8B6DBF76FF70FD29FE9AFD4A" bold="true" box="[199,317,730,755]" pageId="5" pageNumber="291">Remarks:</emphasis>
<taxonomicName id="4C508DEC8B6DBF76FEF4FD28FE52FD4A" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[323,501,731,755]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="B9242A7D8B6DBF76FEF4FD28FE52FD4A" box="[323,501,731,755]" italics="true" pageId="5" pageNumber="291">Argia mauffrayi</emphasis>
</taxonomicName>
is unusual in its overall orange coloration and lack of any pale color on the terminal abdominal segments. Specimens are noticeable in the field as Sanchez, one of the collectors of the
<typeStatus id="54EB48CD8B6DBF76FB58FCF3FAF5FCA1" box="[1263,1362,768,792]" pageId="5" pageNumber="291" type="paratype">paratype</typeStatus>
, noted its orange color when reporting its capture to Garrison. Mauffray and Tennessen noted the following: &quot;[The
<typeStatus id="54EB48CD8B6DBF76FF20FCB5FF5BFCE6" box="[151,252,838,863]" pageId="5" pageNumber="291" type="holotype">holotype</typeStatus>
] was collected at approx 9:30am along forested trail on a very steep hill with a few small sun-lit spots. It was collected as it flew upward and perched on a leaf in one of these sunny spots along with
<taxonomicName id="4C508DEC8B6DBF76FB50FC98FA3BFC3A" box="[1255,1436,875,899]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="variegata">
<emphasis id="B9242A7D8B6DBF76FB50FC98FA3BFC3A" box="[1255,1436,875,899]" italics="true" pageId="5" pageNumber="291">Argia variegata</emphasis>
</taxonomicName>
[Förster],
<taxonomicName id="4C508DEC8B6DBF76FEBEFC7CFE02FC1E" authorityName="Hagen in Selys" authorityYear="1865" box="[265,421,910,935]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="B9242A7D8B6DBF76FEBEFC7CFE02FC1E" box="[265,421,910,935]" italics="true" pageId="5" pageNumber="291">Argia oculata</emphasis>
</taxonomicName>
[Hagen
<emphasis id="B9242A7D8B6DBF76FDBDFC63FD87FC1E" box="[522,544,912,935]" italics="true" pageId="5" pageNumber="291">in</emphasis>
Selys] and
<taxonomicName id="4C508DEC8B6DBF76FD1DFC7DFC24FC1E" authorityName="Selys" authorityYear="1853" box="[682,899,910,935]" class="Insecta" family="Polythoridae" genus="Polythore" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="B9242A7D8B6DBF76FD1DFC7DFC24FC1E" box="[682,899,910,935]" italics="true" pageId="5" pageNumber="291">Polythore gigantea</emphasis>
</taxonomicName>
[(Selys)]. There was no seepage nor stream at that location; however, there was a small seepage stream about 50 meters from the location. Ken and I continued to look for other specimens and even returned to the site about an hour later, but the weather had turned cloudy&quot; and &quot;..when I first saw the specimen flying–it appeared mostly orange–[and thought] it's a small
<taxonomicName id="4C508DEC8B6DBF76FB15FC0FFA8BFBAA" authorityName="Selys" authorityYear="1862" box="[1186,1324,1020,1043]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="B9242A7D8B6DBF76FB15FC0FFA8BFBAA" box="[1186,1324,1020,1043]" italics="true" pageId="5" pageNumber="291">Heteragrion</emphasis>
</taxonomicName>
!&quot;
</paragraph>
<paragraph id="8BEFF66F8B6DBF76FF70FBEDFCDBFB7E" blockId="5.[151,1436,151,1295]" pageId="5" pageNumber="291">
Although the female of
<taxonomicName id="4C508DEC8B6DBF76FE62FBECFDC6FB8E" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[469,609,1055,1079]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="B9242A7D8B6DBF76FE62FBECFDC6FB8E" box="[469,609,1055,1079]" italics="true" pageId="5" pageNumber="291">A. mauffrayi</emphasis>
</taxonomicName>
is unknown, we suspect that the mesostigmal plates will likely be poorly developed similar to the female sex of
<taxonomicName id="4C508DEC8B6DBF76FDF8FBB0FD68FBE2" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[591,719,1090,1115]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9242A7D8B6DBF76FDF8FBB0FD68FBE2" box="[591,719,1090,1115]" italics="true" pageId="5" pageNumber="291">A. acridens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C508DEC8B6DBF76FD69FBB0FCCCFBE2" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[734,875,1090,1115]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B9242A7D8B6DBF76FD69FBB0FCCCFBE2" box="[734,875,1090,1115]" italics="true" pageId="5" pageNumber="291">A. cuspidata</emphasis>
</taxonomicName>
(Garrison &amp; von
<bibRefCitation id="EFC18B9E8B6DBF76FB82FBB1FB54FBE2" author="Garrison, R. W. &amp; von Ellenrieder, N." box="[1077,1267,1090,1115]" pageId="5" pageNumber="291" pagination="1 - 69" refId="ref2561" refString="Garrison, R. W. &amp; von Ellenrieder, N. (2018) Damselflies of the genus Argia (Odonata: Coenagrionidae) from Ecuador with descriptions of five new species. Zootaxa, 4470 (1), 1 - 69. https: // doi. org / 10.11646 / zootaxa. 4470.1.1" type="journal article" year="2018">Ellenrieder 2018</bibRefCitation>
, Figs. 69 &amp; 72 respectively) and
<taxonomicName id="4C508DEC8B6DBF76FEE8FB94FD2EFBC6" authority="Fraser, 1946" authorityName="Fraser" authorityYear="1946" box="[351,649,1126,1151]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="mishuyaca">
<emphasis id="B9242A7D8B6DBF76FEE8FB94FE51FBC6" box="[351,502,1126,1151]" italics="true" pageId="5" pageNumber="291">A. mishuyaca</emphasis>
Fraser, 1946
</taxonomicName>
(Garrison &amp; von
<bibRefCitation id="EFC18B9E8B6DBF76FCE4FB95FBB6FBC6" author="Garrison, R. W. &amp; von Ellenrieder, N." box="[851,1041,1126,1151]" pageId="5" pageNumber="291" pagination="1 - 134" refId="ref2508" refString="Garrison, R. W. &amp; von Ellenrieder, N. (2015) Damselflies of the genus Argia of the Guiana Shield (Odonata: Coenagrionidae). Zootaxa, 4042 (1), 1 - 134. https: // doi. org / 10.11646 / zootaxa. 4042.1.1" type="journal article" year="2015">Ellenrieder 2015</bibRefCitation>
,
<figureCitation id="136BEAEA8B6DBF76FBAAFB95FBC9FBC6" box="[1053,1134,1126,1151]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1436,195,1884]" captionTargetId="figure@2.[151,1436,195,1884]" captionTargetPageId="2" captionText="FIGURES 1–10. Color pattern of head, thorax, S1–4, lateral view (1); S7–10, lateral view (2); S5–10, dorsal view (3); appendages, mediodorsal view (4); same, lateral view (5); cerci, dorsal view (6); genital ligula, lateral view (7); same entolateral view (8); same ectal view (9); thorax, lateral view (10)." httpUri="https://zenodo.org/record/2618858/files/figure.png" pageId="5" pageNumber="291">Fig. 7c</figureCitation>
). The curvilinear cercus in the males of these three species are longer than the paraproct and are armed terminally with a apical decumbent tooth—a condition similar to that illustrated here for
<taxonomicName id="4C508DEC8B6DBF76FD5EFB5CFCD2FB7E" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[745,885,1199,1223]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="B9242A7D8B6DBF76FD5EFB5CFCD2FB7E" box="[745,885,1199,1223]" italics="true" pageId="5" pageNumber="291">A. mauffrayi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BEFF66F8B6DBF76FF70FB20FC26FAB6" blockId="5.[151,1436,151,1295]" pageId="5" pageNumber="291">
<taxonomicName id="4C508DEC8B6DBF76FF70FB20FE26FB52" authorityName="Garrison &amp; Ramón Cabrera" authorityYear="2019" box="[199,385,1235,1259]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="mauffrayi">
<emphasis id="B9242A7D8B6DBF76FF70FB20FE26FB52" box="[199,385,1235,1259]" italics="true" pageId="5" pageNumber="291">Argia mauffrayi</emphasis>
</taxonomicName>
, which is sympatric with
<taxonomicName id="4C508DEC8B6DBF76FD7FFB20FCE8FB52" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[712,847,1234,1259]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="acridens">
<emphasis id="B9242A7D8B6DBF76FD7FFB20FCE8FB52" box="[712,847,1234,1259]" italics="true" pageId="5" pageNumber="291">A. acridens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C508DEC8B6DBF76FC22FB20FB8EFB52" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[917,1065,1234,1259]" class="Insecta" family="Coenagrionidae" genus="Argia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="cuspidata">
<emphasis id="B9242A7D8B6DBF76FC22FB20FB8EFB52" box="[917,1065,1234,1259]" italics="true" pageId="5" pageNumber="291">A. cuspidata</emphasis>
</taxonomicName>
, is thus far known only from
<collectingRegion id="4994388D8B6DBF76FF20FB05FEBFFAB6" box="[151,280,1270,1295]" country="Ecuador" name="Esmeraldas" pageId="5" pageNumber="291">Esmeraldas</collectingRegion>
and
<collectingRegion id="4994388D8B6DBF76FEF9FB05FE1BFAB6" box="[334,444,1270,1295]" country="Ecuador" name="Pichincha" pageId="5" pageNumber="291">Pichincha</collectingRegion>
Provinces in western
<collectingCountry id="F347B6FF8B6DBF76FD03FB05FCB6FAB6" box="[692,785,1270,1295]" name="Ecuador" pageId="5" pageNumber="291">Ecuador</collectingCountry>
(
<figureCitation id="136BEAEA8B6DBF76FC96FB05FCD3FAB6" box="[801,884,1270,1295]" captionStart="FIGURE 18" captionStartId="4.[151,250,1593,1615]" captionTargetBox="[228,1335,239,1493]" captionTargetId="figure@4.[151,1436,193,1572]" captionTargetPageId="4" captionText="FIGURE 18. Distribution of Argia acridens and A. mauffrayi." httpUri="https://zenodo.org/record/2618862/files/figure.png" pageId="5" pageNumber="291">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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