322 lines
41 KiB
XML
322 lines
41 KiB
XML
<document id="506F3159BDE85050FBFFEF353FDA9B17" ID-DOI="10.11646/zootaxa.5501.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13628197" ID-ZooBank="F6A55A77-C4C8-4BDD-8D36-482395A01311" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725278975848" checkinUser="plazi" docAuthor="Nagel, Peter" docDate="2024" docId="03F01B1CFFF26D6DA0EE5BD81455FB56" docLanguage="en" docName="zootaxa.5501.3.3.pdf" docOrigin="Zootaxa 5501 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5501.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paussus agatae Nagel, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="451" masterDocId="FFC96364FFF36D69A0795F731654FFC5" masterDocTitle="The Paussus (Scaphipaussus) lanxangensis species group of Indomalayan ant nest beetles (Coleoptera, Carabidae, Paussinae, Paussini) with description of a new species" masterLastPageNumber="457" masterPageNumber="447" pageNumber="448" updateTime="1725279448654" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9C435AFCB8716A7B609016EE265446A6">The Paussus (Scaphipaussus) lanxangensis species group of Indomalayan ant nest beetles (Coleoptera, Carabidae, Paussinae, Paussini) with description of a new species</mods:title>
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<mods:namePart id="3749DA76C5FB2A29933240941A9DAD5B">Nagel, Peter</mods:namePart>
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<mods:title id="01EC8817525E9B47D4DDCB1531C245A9">Zootaxa</mods:title>
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<mods:date id="3E86AE3F32505A6502D79B07D117B58E">2024</mods:date>
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<mods:number id="BED32B5AC2BD5CA3282CEF9A3C23074F">2024-09-02</mods:number>
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<mods:identifier id="147443569CD0F171EDEA9D0134B93609" type="DOI">10.11646/zootaxa.5501.3.3</mods:identifier>
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<treatment id="03F01B1CFFF26D6DA0EE5BD81455FB56" LSID="urn:lsid:plazi:treatment:03F01B1CFFF26D6DA0EE5BD81455FB56" httpUri="http://treatment.plazi.org/id/03F01B1CFFF26D6DA0EE5BD81455FB56" lastPageId="4" lastPageNumber="451" pageId="1" pageNumber="448">
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<subSubSection id="C343F981FFF26D68A0EE5BD817E3FB00" box="[151,439,1195,1221]" pageId="1" pageNumber="448" type="nomenclature">
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<paragraph id="8BE6AA0AFFF26D68A0EE5BD817E3FB00" blockId="1.[151,439,1195,1221]" box="[151,439,1195,1221]" pageId="1" pageNumber="448">
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<heading id="D0AE1D66FFF26D68A0EE5BD817E3FB00" bold="true" box="[151,439,1195,1221]" fontSize="11" level="1" pageId="1" pageNumber="448" reason="1">
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<emphasis id="B92D7618FFF26D68A0EE5BD817E3FB00" bold="true" box="[151,439,1195,1221]" pageId="1" pageNumber="448">
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<taxonomicName id="4C59D189FFF26D68A0EE5BD81706FB00" authorityName="Nagel" authorityYear="2024" box="[151,338,1195,1221]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="agatae" status="sp. nov.">
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<emphasis id="B92D7618FFF26D68A0EE5BD81706FB00" bold="true" box="[151,338,1195,1221]" italics="true" pageId="1" pageNumber="448">Paussus agatae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21ECB63FFF26D68A1205BD817E3FB00" box="[345,439,1195,1221]" pageId="1" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C343F981FFF26D68A0EE5B9C135AF877" pageId="1" pageNumber="448" type="materials_examined">
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<paragraph id="8BE6AA0AFFF26D68A0EE5B9C14E9FAE0" blockId="1.[151,1436,1263,1502]" pageId="1" pageNumber="448">
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<emphasis id="B92D7618FFF26D68A0EE5B9C1754FAC2" bold="true" box="[151,256,1263,1287]" inLineHeading="true" pageId="1" pageNumber="448" reason="1">
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<typeStatus id="54E214A8FFF26D68A0EE5B9C16AFFAC2" box="[151,251,1263,1287]" pageId="1" pageNumber="448" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Female and single specimen, here designated: Dry mounted, glued to pinned card. Head and antennae with scapes detached and glued to the same card. BL
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<quantity id="4CA107EFFFF26D68A2105A7D14EEFAE0" box="[617,698,1294,1318]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="448" unit="mm" value="3.8">3.8 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BE6AA0AFFF26D68A0EE5A5F14F4FA81" blockId="1.[151,1436,1263,1502]" box="[151,672,1324,1348]" pageId="1" pageNumber="448">The following labels are pinned with the specimen:</paragraph>
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<paragraph id="8BE6AA0AFFF26D68A0EE5A3814C1FAA6" blockId="1.[151,1436,1263,1502]" box="[151,661,1355,1379]" pageId="1" pageNumber="448">
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Small, vertical rectangular label, black printing:
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<specimenCount id="9D5F6183FFF26D68A2FB5A3F14C1FAA6" box="[642,661,1356,1379]" count="1" pageId="1" pageNumber="448" type="female">♀</specimenCount>
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</paragraph>
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<paragraph id="8BE6AA0AFFF26D68A0EE5A191501FA44" blockId="1.[151,1436,1263,1502]" box="[151,853,1386,1410]" pageId="1" pageNumber="448">
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White rectangular label (black handwriting):
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<taxonomicName id="4C59D189FFF26D68A2185A1814ABFA47" authorityName="Nagel" authorityYear="2024" box="[609,767,1387,1410]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="agatae" status="sp. nov.">
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<emphasis id="B92D7618FFF26D68A2185A1814ABFA47" box="[609,767,1387,1410]" italics="true" pageId="1" pageNumber="448">Paussus agatae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21ECB63FFF26D68A37F5A191501FA44" box="[774,853,1386,1409]" pageId="1" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BE6AA0AFFF26D68A0EE5AFB148FFA65" blockId="1.[151,1436,1263,1502]" box="[151,731,1416,1440]" pageId="1" pageNumber="448">White rectangular label (black handwriting): NHM Basel</paragraph>
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<paragraph id="8BE6AA0AFFF26D68A0EE5AD41204FA7A" blockId="1.[151,1436,1263,1502]" box="[151,1104,1447,1471]" pageId="1" pageNumber="448">
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Red quadrat label (black handwriting):
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<taxonomicName id="4C59D189FFF26D68A25F5ADB1490FA7A" authorityName="Nagel" authorityYear="2024" box="[550,708,1448,1471]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="agatae" status="sp. nov.">
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<emphasis id="B92D7618FFF26D68A25F5ADB1490FA7A" box="[550,708,1448,1471]" italics="true" pageId="1" pageNumber="448">Paussus agatae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21ECB63FFF26D68A2B25ADB1548FA7A" box="[715,796,1448,1471]" pageId="1" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
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, P. Nagel 2021
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<typeStatus id="54E214A8FFF26D68A3C05AD41204FA7A" box="[953,1104,1447,1471]" pageId="1" pageNumber="448" type="holotype">HOLOTYPUS</typeStatus>
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</paragraph>
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<paragraph id="8BE6AA0AFFF26D68A0EE5AB51576FA1B" blockId="1.[151,1436,1263,1502]" box="[151,802,1478,1502]" pageId="1" pageNumber="448">
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Two white rectangular labels, black printing, as shown in
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<figureCitation id="1362B68FFFF26D68A2985AB51549FA1B" box="[737,797,1478,1502]" captionStart="FIGURE 1" captionStartId="2.[151,250,523,547]" captionTargetBox="[154,1433,185,495]" captionTargetId="figure-18@2.[151,1437,181,498]" captionTargetPageId="2" captionText="FIGURE 1. Original labels (20 X 10 mm each) pinned with holotype specimen of P. agatae sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.13628200" httpUri="https://zenodo.org/record/13628200/files/figure.png" pageId="1" pageNumber="448">Fig. 1</figureCitation>
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.
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</paragraph>
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<paragraph id="8BE6AA0AFFF26D68A0EE597E1730F9E2" blockId="1.[151,1437,1549,1971]" box="[151,356,1549,1575]" pageId="1" pageNumber="448">
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<emphasis id="B92D7618FFF26D68A0EE597E1749F9E3" box="[151,285,1549,1574]" italics="true" pageId="1" pageNumber="448">Depository:</emphasis>
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<collectionCode id="ED4832CFFFF26D68A15D597E1730F9E2" box="[292,356,1549,1575]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="1" pageNumber="448">NMB</collectionCode>
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</paragraph>
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<paragraph id="8BE6AA0AFFF26D68A0BE59421454F973" blockId="1.[151,1437,1549,1971]" pageId="1" pageNumber="448">
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<emphasis id="B92D7618FFF26D68A0BE594214F4F98F" box="[199,672,1584,1610]" italics="true" pageId="1" pageNumber="448">Type locality and additional information:</emphasis>
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<materialsCitation id="3B31A057FFF26D68A2D0594317A8F973" collectingDate="2011-05-04" collectingDateMax="2011-05-18" collectingDateMin="2011-05-04" collectorName="Natural History Museum Basel & Michel Brancucci & Michael Geiser & David Hauck & Zdenek Kraus & Any Phantala & Elawan Vongphachan" country="Laos" county="Northwestern" elevation="600" location="Nature Reserve" longitude="100.75" municipality="Houaixai district" pageId="1" pageNumber="448" specimenCount="1" stateProvince="Bokeo">
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<collectingCounty id="6287D286FFF26D68A2D059431515F98F" box="[681,833,1584,1610]" pageId="1" pageNumber="448">Northwestern</collectingCounty>
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<collectingCountry id="F34EEA9AFFF26D68A332594215D0F98E" box="[843,900,1585,1611]" name="Laos" pageId="1" pageNumber="448">Laos</collectingCountry>
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,
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<collectingRegion id="499D64E8FFF26D68A3EB5943121FF98F" box="[914,1099,1584,1610]" country="Laos" name="Bokeo" pageId="1" pageNumber="448">Bokeo Province</collectingRegion>
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,
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<collectingMunicipality id="6B823070FFF26D68A4205943134EF98F" box="[1113,1306,1584,1610]" pageId="1" pageNumber="448">Houaixai district</collectingMunicipality>
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,
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<locationDeviation id="9A54B328FFF26D68A55E5943175CF9AA" location="Ban Toup" pageId="1" pageNumber="448">
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<quantity id="4CA107EFFFF26D68A55E5943130DF98E" box="[1319,1369,1584,1611]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="448" unit="km" value="5.0">5km</quantity>
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W of Ban Toup
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</locationDeviation>
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,
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<collectingRegion id="499D64E8FFF26D68A16D59271709F9AB" box="[276,349,1620,1646]" country="Laos" name="Bokeo" pageId="1" pageNumber="448">Bokeo</collectingRegion>
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<location id="8E86FCD1FFF26D68A11F59261443F9AA" LSID="urn:lsid:plazi:treatment:03F01B1CFFF26D6DA0EE5BD81455FB56:8E86FCD1FFF26D68A11F59261443F9AA" box="[358,535,1621,1647]" country="Laos" county="Northwestern" longitude="100.75" municipality="Houaixai district" name="Nature Reserve" pageId="1" pageNumber="448" stateProvince="Bokeo">Nature Reserve</location>
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,
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<elevation id="00744D39FFF26D68A25D592714B6F9AB" box="[548,738,1620,1647]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="1" pageNumber="448" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">
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<quantity id="4CA107EFFFF26D68A25D592714F0F9AA" box="[548,676,1620,1647]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="1" pageNumber="448" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500–700 m</quantity>
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a.s.l.
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</elevation>
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, 20°27-28‘N
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<geoCoordinate id="EE6DCCCDFFF26D68A307592715BFF9AA" box="[894,1003,1620,1647]" degrees="100" direction="east" minutes="45" orientation="longitude" pageId="1" pageNumber="448" precision="925" value="100.75">100°45’E</geoCoordinate>
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,
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<date id="FFE78CCAFFF26D68A380592712FAF9AB" box="[1017,1198,1620,1647]" pageId="1" pageNumber="448" value="2011-05-04" valueMax="2011-05-18" valueMin="2011-05-04">
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<collectingDate id="EFA37522FFF26D68A380592712FAF9AB" box="[1017,1198,1620,1647]" pageId="1" pageNumber="448" value="2011-05-04" valueMax="2011-05-18" valueMin="2011-05-04">4–18 May 2011</collectingDate>
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</date>
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||
, leg.
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<collectorName id="26ACCFDCFFF26D68A49559271713F957" pageId="1" pageNumber="448">Natural History Museum Basel</collectorName>
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,
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<collectingCountry id="F34EEA9AFFF26D68A121590A17DAF956" box="[344,398,1657,1683]" name="Laos" pageId="1" pageNumber="448">Laos</collectingCountry>
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2011 expedition
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<collectorName id="26ACCFDCFFF26D68A21B590B1566F957" box="[610,818,1656,1682]" pageId="1" pageNumber="448">Michel Brancucci</collectorName>
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,
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<collectorName id="26ACCFDCFFF26D68A33D590B15A3F957" box="[836,1015,1656,1682]" pageId="1" pageNumber="448">Michael Geiser</collectorName>
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,
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<collectorName id="26ACCFDCFFF26D68A471590B12F1F957" box="[1032,1189,1656,1682]" pageId="1" pageNumber="448">David Hauck</collectorName>
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||
,
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<collectorName id="26ACCFDCFFF26D68A4CF590B1309F956" box="[1206,1373,1656,1683]" pageId="1" pageNumber="448">Zdeněk Kraus</collectorName>
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||
,
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<collectorName id="26ACCFDCFFF26D68A515590B16AEF973" pageId="1" pageNumber="448">Any Phantala</collectorName>
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||
,
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<collectorName id="26ACCFDCFFF26D68A17F59EF17A8F973" box="[262,508,1692,1718]" pageId="1" pageNumber="448">Elawan Vongphachan</collectorName>
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</materialsCitation>
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||
.
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</paragraph>
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<paragraph id="8BE6AA0AFFF26D68A0BE59B3176CF8E7" blockId="1.[151,1437,1549,1971]" pageId="1" pageNumber="448">
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<emphasis id="B92D7618FFF26D68A0BE59B31749F91F" box="[199,285,1728,1754]" italics="true" pageId="1" pageNumber="448">Habitat</emphasis>
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: Primary lowland evergreen dipterocarp forest, patches of secondary forest and small agricultural areas (
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<bibRefCitation id="EFC8D7FBFFF26D68A0E6599717C2F93B" author="Geiser, M. & Nagel, P." box="[159,406,1764,1790]" pageId="1" pageNumber="448" refId="ref6880" refString="Geiser, M. & Nagel, P. (2013) Coleopterology in Laos - an introduction to the nature of the country and its coleopterological exploration. In: Brancucci, M. †, Nagel, P., Kolibac, J. & Geiser, M. (Eds.), Beetle diversity of Laos. Part I. Entomologica Basiliensia et Collectionis Frey. Vol. 34. Pro Entomologia, Basel, pp. 11 - 46. https: // doi. org / 10.5169 / seals- 980997" type="book" year="2013">Geiser & Nagel 2013</bibRefCitation>
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). Captured in a flight interception trap, i.e., the specimen was probably flying at low height above ground.
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</paragraph>
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<paragraph id="8BE6AA0AFFF26D68A0BE585F135AF877" blockId="1.[151,1437,1549,1971]" pageId="1" pageNumber="448">
|
||
<emphasis id="B92D7618FFF26D68A0BE585F176BF883" box="[199,319,1836,1862]" italics="true" pageId="1" pageNumber="448">Etymology</emphasis>
|
||
: Named for Mrs. Agata Luka Stan, Research Institute of Organic Agriculture (FiBL), Frick,
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<collectingCountry id="F34EEA9AFFF26D68A0EE58231774F8AF" box="[151,288,1872,1898]" name="Switzerland" pageId="1" pageNumber="448">Switzerland</collectingCountry>
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, who, on behalf of the Natural History Museum of Basel,
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<collectingCountry id="F34EEA9AFFF26D68A3EB5823124FF8AF" box="[914,1051,1872,1898]" name="Switzerland" pageId="1" pageNumber="448">Switzerland</collectingCountry>
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, and with support of the foundation Pro Entomologia, Basel, searched for months many jars full of preserved tiny arthropods from flight interception traps. The specific epithet is a proper noun in the genitive case, derived from the given name “Agata”.
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</paragraph>
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</subSubSection>
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<caption id="DF26FA82FFF16D6BA0EE5D7812CFFDE6" ID-DOI="http://doi.org/10.5281/zenodo.13628200" ID-Zenodo-Dep="13628200" box="[151,1179,523,547]" httpUri="https://zenodo.org/record/13628200/files/figure.png" pageId="2" pageNumber="449" startId="2.[151,250,523,547]" targetBox="[154,1433,185,495]" targetPageId="2" targetType="figure">
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||
<paragraph id="8BE6AA0AFFF16D6BA0EE5D7812CFFDE6" blockId="2.[151,1179,523,547]" box="[151,1179,523,547]" pageId="2" pageNumber="449">
|
||
<emphasis id="B92D7618FFF16D6BA0EE5D781747FDE6" bold="true" box="[151,275,523,547]" pageId="2" pageNumber="449">FIGURE 1.</emphasis>
|
||
Original labels (20 X 10 mm each) pinned with holotype specimen of
|
||
<taxonomicName id="4C59D189FFF16D6BA3995D7F126AFDE6" authorityName="Nagel" authorityYear="2024" box="[992,1086,524,547]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="agatae" status="sp. nov.">
|
||
<emphasis id="B92D7618FFF16D6BA3995D7F126AFDE6" box="[992,1086,524,547]" italics="true" pageId="2" pageNumber="449">P. agatae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92D7618FFF16D6BA43D5D7812C1FDE6" bold="true" box="[1092,1173,523,547]" pageId="2" pageNumber="449">
|
||
<taxonomicNameLabel id="A21ECB63FFF16D6BA43D5D7812C1FDE6" box="[1092,1173,523,547]" pageId="2" pageNumber="449" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C343F981FFF16D6DA0EE5D061455FB56" lastPageId="4" lastPageNumber="451" pageId="2" pageNumber="449" type="description">
|
||
<paragraph id="8BE6AA0AFFF16D6BA0EE5D0617FFFD4A" blockId="2.[151,427,629,655]" box="[151,427,629,655]" pageId="2" pageNumber="449">
|
||
<emphasis id="B92D7618FFF16D6BA0EE5D0617FFFD4A" bold="true" box="[151,427,629,655]" pageId="2" pageNumber="449">
|
||
Description (
|
||
<figureCitation id="1362B68FFFF16D6BA14E5D0617D1FD4A" box="[311,389,629,655]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[324,1274,192,1907]" captionTargetId="figure-16@3.[305,1282,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Paussus agatae sp.n., female holotype, habitus, dorsal view with pygidium artificially shown at plan view (originally strongly declined). Femora and tibiae of right and left side are shown at broadest and narrowest view respectively. Scale 1 mm. Illustration: Armin Coray, Basel." figureDoi="http://doi.org/10.5281/zenodo.13628202" httpUri="https://zenodo.org/record/13628202/files/figure.png" pageId="2" pageNumber="449">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="1362B68FFFF16D6BA1EB5D0617F6FD4A" box="[402,418,629,655]" captionStart="FIGURE 3" captionStartId="4.[151,250,715,739]" captionTargetBox="[154,1434,184,687]" captionTargetId="figure-18@4.[151,1436,181,691]" captionTargetPageId="4" captionText="FIGURE 3. Paussus agatae sp. nov., female holotype, head, mouthparts, ventral view. Inset: Left maxillary palpomere II, ventral view, slightly distorted to show broadest view, with emarginate mesal margin and slightly convex outer margin. Scale 0.5 mm. Illustration:Armin Coray, Basel." figureDoi="http://doi.org/10.5281/zenodo.13628204" httpUri="https://zenodo.org/record/13628204/files/figure.png" pageId="2" pageNumber="449">3</figureCitation>
|
||
)
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE6AA0AFFF16D6BA0EE5DCE1580FC86" blockId="2.[151,1437,701,2024]" pageId="2" pageNumber="449">
|
||
<emphasis id="B92D7618FFF16D6BA0EE5DCE1685FD12" box="[151,209,701,727]" italics="true" pageId="2" pageNumber="449">Body</emphasis>
|
||
yellowish-brown with the following areas infuscate (dark brown to blackish, not sharply delimited):disc of elytra, lateral flanks of pronotum, frontal margin of head, dorsal and ventral disc of antennal clubs, external antennomeres 1 (scapes), proximal two thirds of all femora. Glossy parts restricted to posterior parts of vertex cavity, central part of transverse pronotal furrow, pygidial disc, and tibiae.
|
||
</paragraph>
|
||
<paragraph id="8BE6AA0AFFF16D6BA0BE5C3E134DFC32" blockId="2.[151,1437,701,2024]" pageId="2" pageNumber="449">
|
||
<emphasis id="B92D7618FFF16D6BA0BE5C3E1705FCA2" box="[199,337,845,871]" italics="true" pageId="2" pageNumber="449">Dorsal head</emphasis>
|
||
1.3 times wider than long, almost as wide as the pronotum, temples large, evenly rounded towards neck constriction, not projecting beyond eyes, frontal margin sinuate. Dorsal head with rough surface, coarsely granulated, and with four deep excavations. Frontal excavation carving the frontal margin. Two laterodorsal excavations with coarse surface. Central posterior excavation indicating a stigmatoid fusion of ancestrally free lateral plate-like structures. Basal surface of the latter excavation smooth, even, impunctate and glossy.
|
||
</paragraph>
|
||
<paragraph id="8BE6AA0AFFF16D6BA0BE5B7212B9FB42" blockId="2.[151,1437,701,2024]" pageId="2" pageNumber="449">
|
||
<emphasis id="B92D7618FFF16D6BA0BE5B721737FBDE" box="[199,355,1025,1051]" italics="true" pageId="2" pageNumber="449">Antennal club</emphasis>
|
||
as shown in
|
||
<figureCitation id="1362B68FFFF16D6BA18F5B72146CFBDE" box="[502,568,1025,1051]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[324,1274,192,1907]" captionTargetId="figure-16@3.[305,1282,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Paussus agatae sp.n., female holotype, habitus, dorsal view with pygidium artificially shown at plan view (originally strongly declined). Femora and tibiae of right and left side are shown at broadest and narrowest view respectively. Scale 1 mm. Illustration: Armin Coray, Basel." figureDoi="http://doi.org/10.5281/zenodo.13628202" httpUri="https://zenodo.org/record/13628202/files/figure.png" pageId="2" pageNumber="449">Fig. 2</figureCitation>
|
||
(female! male unknown). Dorso-ventrally little compressed, 1.9 times longer than wide at dorsal view with anterior (inner) margin slightly and evenly convex and posterior (exterior) margin slightly and evenly emarginate. At hind view, antennal club excavated at little more than the proximal half of its length. At dorsal view, posterior basal angle little pronounced, forming a short, rounded projection.
|
||
</paragraph>
|
||
<paragraph id="8BE6AA0AFFF16D6BA0BE5BE217C0FA46" blockId="2.[151,1437,701,2024]" pageId="2" pageNumber="449">
|
||
<emphasis id="B92D7618FFF16D6BA0BE5BE2171EFB6E" box="[199,330,1169,1195]" italics="true" pageId="2" pageNumber="449">Mouthparts</emphasis>
|
||
(cf.
|
||
<bibRefCitation id="EFC8D7FBFFF16D6BA1025BE214CDFB6E" author="Robertson, J. A. & Moore, W." box="[379,665,1169,1196]" pageId="2" pageNumber="449" pagination="134 - 170" refId="ref7784" refString="Robertson, J. A. & Moore, W. (2016) Phylogeny of Paussus L. (Carabidae: Paussinae): unravelling morphological convergence associated with myrmecophilous life histories. Systematic Entomology, 42 (1), 134 - 170, figs. S 1 - S 2, tabs S 1 - S 5. [2017, DP 28 Sep 2016, version of record online] https: // doi. org / 10.1111 / syen. 12205" type="journal article" year="2016">Robertson & Moore 2016</bibRefCitation>
|
||
) as shown in
|
||
<figureCitation id="1362B68FFFF16D6BA3555BE2153AFB6E" box="[812,878,1169,1195]" captionStart="FIGURE 3" captionStartId="4.[151,250,715,739]" captionTargetBox="[154,1434,184,687]" captionTargetId="figure-18@4.[151,1436,181,691]" captionTargetPageId="4" captionText="FIGURE 3. Paussus agatae sp. nov., female holotype, head, mouthparts, ventral view. Inset: Left maxillary palpomere II, ventral view, slightly distorted to show broadest view, with emarginate mesal margin and slightly convex outer margin. Scale 0.5 mm. Illustration:Armin Coray, Basel." figureDoi="http://doi.org/10.5281/zenodo.13628204" httpUri="https://zenodo.org/record/13628204/files/figure.png" pageId="2" pageNumber="449">Fig. 3</figureCitation>
|
||
. Maxillary palpomere III (penultimate palpomere) not distinctly wider than following (terminal) palpomere IV. Length of maxillary palpomere II (third from last)
|
||
<quantity id="4CA107EFFFF16D6BA5125BC61692FB31" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="449" unit="mm" value="0.26">0.26 mm</quantity>
|
||
. The labial palpomeres are not distinctly wide, flattened and spade-shaped. The terminal labial palpomeres are elliptic, with distal two thirds tapering. Tip diminutive. Lateral lobes of mentum broadest at basis, lateral margins slightly sinuate, not straight. Width/length ratio of gula at narrowest point 0.4 (Character 15, state S
|
||
<quantity id="4CA107EFFFF16D6BA48F5A521376FAFE" box="[1270,1314,1313,1339]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="449" unit="in" value="2.0">2 in</quantity>
|
||
<bibRefCitation id="EFC8D7FBFFF16D6BA5505A521732FA9A" author="Robertson, J. A. & Moore, W." pageId="2" pageNumber="449" pagination="134 - 170" refId="ref7784" refString="Robertson, J. A. & Moore, W. (2016) Phylogeny of Paussus L. (Carabidae: Paussinae): unravelling morphological convergence associated with myrmecophilous life histories. Systematic Entomology, 42 (1), 134 - 170, figs. S 1 - S 2, tabs S 1 - S 5. [2017, DP 28 Sep 2016, version of record online] https: // doi. org / 10.1111 / syen. 12205" type="journal article" year="2016">Robertson and Moore (2016)</bibRefCitation>
|
||
, Table S2. N.B.: here measured at original detached head capsule at plan view of posterior part, not taken from
|
||
<figureCitation id="1362B68FFFF16D6BA13A5A1A17D2FA46" box="[323,390,1385,1411]" captionStart="FIGURE 3" captionStartId="4.[151,250,715,739]" captionTargetBox="[154,1434,184,687]" captionTargetId="figure-18@4.[151,1436,181,691]" captionTargetPageId="4" captionText="FIGURE 3. Paussus agatae sp. nov., female holotype, head, mouthparts, ventral view. Inset: Left maxillary palpomere II, ventral view, slightly distorted to show broadest view, with emarginate mesal margin and slightly convex outer margin. Scale 0.5 mm. Illustration:Armin Coray, Basel." figureDoi="http://doi.org/10.5281/zenodo.13628204" httpUri="https://zenodo.org/record/13628204/files/figure.png" pageId="2" pageNumber="449">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE6AA0AFFF16D6BA0BE5AFD17D8F9D6" blockId="2.[151,1437,701,2024]" pageId="2" pageNumber="449">
|
||
<emphasis id="B92D7618FFF16D6BA0BE5AFD1762FA62" box="[199,310,1422,1447]" italics="true" pageId="2" pageNumber="449">Pronotum</emphasis>
|
||
slightly wider than long (1.13 times), divided by a deep transverse furrow, equipped with one small trichome brush each at both lateral ends. Frontal portion laterally projecting with angles broadly rounded. Hind portion with two large, blunt, central tubercles set left and right of the middle. Medial oblong depressions at fore and hind part shining.
|
||
</paragraph>
|
||
<paragraph id="8BE6AA0AFFF16D6BA0BE596D1285F902" blockId="2.[151,1437,701,2024]" pageId="2" pageNumber="449">
|
||
<emphasis id="B92D7618FFF16D6BA0BE596D16A8F9F2" box="[199,252,1566,1591]" italics="true" pageId="2" pageNumber="449">Legs</emphasis>
|
||
with tibiae moderately compressed, widened towards distal margin, especially at hind tibiae. Exterior margin of all tibiae rounded, outer apical angles rounded, not produced. Tibiae without terminal spurs. Tarsi with tarsomeres 1 to 4 subequal, ventrally with scattered bristles (adhesive pads absent). Infuscate section of dorsal hind femora with transverse, slightly curved to straight, densely arranged, narrow, low and blunt ridges (see
|
||
<figureCitation id="1362B68FFFF16D6BA56A59FA1303F966" box="[1299,1367,1673,1699]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[324,1274,192,1907]" captionTargetId="figure-16@3.[305,1282,181,1920]" captionTargetPageId="3" captionText="FIGURE 2. Paussus agatae sp.n., female holotype, habitus, dorsal view with pygidium artificially shown at plan view (originally strongly declined). Femora and tibiae of right and left side are shown at broadest and narrowest view respectively. Scale 1 mm. Illustration: Armin Coray, Basel." figureDoi="http://doi.org/10.5281/zenodo.13628202" httpUri="https://zenodo.org/record/13628202/files/figure.png" pageId="2" pageNumber="449">Fig. 2</figureCitation>
|
||
). This structure is larger and independent of the proximally adjoining small area of the stridulatory file.
|
||
</paragraph>
|
||
<paragraph id="8BE6AA0AFFF16D6BA0BE59A212B6F806" blockId="2.[151,1437,701,2024]" pageId="2" pageNumber="449">
|
||
<emphasis id="B92D7618FFF16D6BA0BE59A217CBF92E" box="[199,415,1745,1771]" italics="true" pageId="2" pageNumber="449">Stridulatory organ:</emphasis>
|
||
Basically as shown for Gen.
|
||
<taxonomicName id="4C59D189FFF16D6BA29A59A1156BF92E" box="[739,831,1746,1771]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D7618FFF16D6BA29A59A1156BF92E" box="[739,831,1746,1771]" italics="true" pageId="2" pageNumber="449">Paussus</emphasis>
|
||
</taxonomicName>
|
||
in general (
|
||
<bibRefCitation id="EFC8D7FBFFF16D6BA3BB59A212F9F92E" author="Di Giulio, A. & Fattorini, S. & Moore, W. & Robertson, J. & Maurizi, E." box="[962,1197,1745,1771]" pageId="2" pageNumber="449" pagination="692 - 702" refId="ref6710" refString="Di Giulio, A., Fattorini, S., Moore, W., Robertson, J. & Maurizi, E. (2014) Form, function and evolutionary significance of stridulatory organs in ant nest beetles (Coleoptera: Carabidae: Paussini). European Journal of Entomology, 111 (5), 692 - 702. https: // doi. org / 10.14411 / eje. 2014.083" type="journal article" year="2014">
|
||
Di Giulio
|
||
<emphasis id="B92D7618FFF16D6BA44A59A1123EF92E" box="[1075,1130,1745,1771]" italics="true" pageId="2" pageNumber="449">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
). The stridulatory file is a well demarcated, finely and densely ridged small area at the basal part of the dorsal femur. The micro-ridges of the file are orientated longitudinally along the axis of the femur, as usual. The scraper appears to be different from the exemplary
|
||
<taxonomicName id="4C59D189FFF16D6BA100584D1489F89D" authority="Fairmaire, 1851" authorityName="Fairmaire" authorityYear="1851" box="[377,733,1853,1880]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="favieri">
|
||
<emphasis id="B92D7618FFF16D6BA100584D1470F892" box="[377,548,1853,1879]" italics="true" pageId="2" pageNumber="449">Paussus favieri</emphasis>
|
||
<bibRefCitation id="EFC8D7FBFFF16D6BA252584E1489F89D" author="Fairmaire, L." box="[555,733,1853,1880]" pageId="2" pageNumber="449" pagination="527 - 528" refId="ref6785" refString="Fairmaire, L. (1851) Description de sept coleopteres nouveaux provenant du midi de l'Europe et du Maroc. Revue et Magasin de Zoologie Pure et Appliquee, 2 e Serie, Tome 3, 527 - 528." type="journal article" year="1851">Fairmaire, 1851</bibRefCitation>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="EFC8D7FBFFF16D6BA369584E15ACF892" author="Di Giulio, A. & Fattorini, S. & Moore, W. & Robertson, J. & Maurizi, E." box="[784,1016,1853,1879]" pageId="2" pageNumber="449" pagination="692 - 702" refId="ref6710" refString="Di Giulio, A., Fattorini, S., Moore, W., Robertson, J. & Maurizi, E. (2014) Form, function and evolutionary significance of stridulatory organs in ant nest beetles (Coleoptera: Carabidae: Paussini). European Journal of Entomology, 111 (5), 692 - 702. https: // doi. org / 10.14411 / eje. 2014.083" type="journal article" year="2014">
|
||
Di Giulio
|
||
<emphasis id="B92D7618FFF16D6BA3F9584D15ECF892" box="[896,952,1853,1879]" italics="true" pageId="2" pageNumber="449">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
) with regard to the low number of the spines and their wider spacing (less than
|
||
<quantity id="4CA107EFFFF16D6BA224581214C1F8BE" box="[605,661,1889,1915]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="449" unit="in" value="10.0">10 in</quantity>
|
||
<taxonomicName id="4C59D189FFF16D6BA2E258111555F8BE" authorityName="Nagel" authorityYear="2024" box="[667,769,1890,1915]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="agatae" status="sp. nov.">
|
||
<emphasis id="B92D7618FFF16D6BA2E258111555F8BE" box="[667,769,1890,1915]" italics="true" pageId="2" pageNumber="449">P. agatae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92D7618FFF16D6BA37158121534F8BE" bold="true" box="[776,864,1889,1915]" pageId="2" pageNumber="449">
|
||
<taxonomicNameLabel id="A21ECB63FFF16D6BA37158121534F8BE" box="[776,864,1889,1915]" pageId="2" pageNumber="449" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
versus
|
||
<quantity id="4CA107EFFFF16D6BA3CD58121242F8BE" box="[948,1046,1889,1916]" metricMagnitude="-1" metricUnit="m" metricValue="6.984999999999999" metricValueMax="7.62" metricValueMin="6.35" pageId="2" pageNumber="449" unit="in" value="27.5" valueMax="30.0" valueMin="25.0">25–30 in</quantity>
|
||
<taxonomicName id="4C59D189FFF16D6BA465581112D4F8BE" authorityName="Fairmaire" authorityYear="1851" box="[1052,1152,1889,1915]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="favieri">
|
||
<emphasis id="B92D7618FFF16D6BA465581112D4F8BE" box="[1052,1152,1889,1915]" italics="true" pageId="2" pageNumber="449">P. favieri</emphasis>
|
||
</taxonomicName>
|
||
, loc.cit., p. 695). One has to consider, though, that these characters are highly variable in
|
||
<taxonomicName id="4C59D189FFF16D6BA32958F512A9F865" authority="(Di Giulio et al. 2014, p. 695)" baseAuthorityName="Di Giulio" baseAuthorityPageNumber="695" baseAuthorityYear="2014" box="[848,1277,1925,1952]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D7618FFF16D6BA32958F515F8F85A" box="[848,940,1926,1951]" italics="true" pageId="2" pageNumber="449">Paussus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC8D7FBFFF16D6BA3C358F612F6F85A" author="Di Giulio, A. & Fattorini, S. & Moore, W. & Robertson, J. & Maurizi, E." box="[954,1186,1925,1951]" pageId="2" pageNumber="449" pagination="692 - 702" refId="ref6710" refString="Di Giulio, A., Fattorini, S., Moore, W., Robertson, J. & Maurizi, E. (2014) Form, function and evolutionary significance of stridulatory organs in ant nest beetles (Coleoptera: Carabidae: Paussini). European Journal of Entomology, 111 (5), 692 - 702. https: // doi. org / 10.14411 / eje. 2014.083" type="journal article" year="2014">
|
||
Di Giulio
|
||
<emphasis id="B92D7618FFF16D6BA45258F51236F85A" box="[1067,1122,1925,1951]" italics="true" pageId="2" pageNumber="449">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
, p. 695)
|
||
</taxonomicName>
|
||
, that the study of the organ was technically difficult and the details given here are, hence, of limited significance.
|
||
</paragraph>
|
||
<paragraph id="8BE6AA0AFFF16D6BA0BE58BE14BDF822" blockId="2.[151,1437,701,2024]" box="[199,745,1997,2024]" pageId="2" pageNumber="449">
|
||
<taxonomicName id="4C59D189FFF16D6BA0BE58BE1761F822" box="[199,309,1997,2023]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92D7618FFF16D6BA0BE58BE1761F822" box="[199,309,1997,2023]" italics="true" pageId="2" pageNumber="449">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
with apex rounded, finely shagreened.
|
||
</paragraph>
|
||
<caption id="DF26FA82FFF06D6AA0EE58E417B7F83D" ID-DOI="http://doi.org/10.5281/zenodo.13628202" ID-Zenodo-Dep="13628202" httpUri="https://zenodo.org/record/13628202/files/figure.png" pageId="3" pageNumber="450" startId="3.[151,250,1943,1967]" targetBox="[324,1274,192,1907]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BE6AA0AFFF06D6AA0EE58E417B7F83D" blockId="3.[151,1437,1943,2040]" pageId="3" pageNumber="450">
|
||
<emphasis id="B92D7618FFF06D6AA0EE58E41745F86A" bold="true" box="[151,273,1943,1967]" pageId="3" pageNumber="450">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C59D189FFF06D6AA16F58EB17E6F86A" authorityName="Nagel" authorityYear="2024" box="[278,434,1944,1967]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="450" phylum="Arthropoda" rank="species" species="agatae" status="sp. nov.">
|
||
<emphasis id="B92D7618FFF06D6AA16F58EB17E6F86A" box="[278,434,1944,1967]" italics="true" pageId="3" pageNumber="450">Paussus agatae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21ECB63FFF06D6AA1CF58EB17BCF86A" box="[438,488,1944,1967]" pageId="3" pageNumber="450" rank="species">sp.n.</taxonomicNameLabel>
|
||
, female holotype, habitus, dorsal view with pygidium artificially shown at plan view (originally strongly declined). Femora and tibiae of right and left side are shown at broadest and narrowest view respectively. Scale 1 mm. Illustration: Armin Coray, Basel.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF26FA82FFF76D6DA0EE5DB81469FCEE" ID-DOI="http://doi.org/10.5281/zenodo.13628204" ID-Zenodo-Dep="13628204" httpUri="https://zenodo.org/record/13628204/files/figure.png" pageId="4" pageNumber="451" startId="4.[151,250,715,739]" targetBox="[154,1434,184,687]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BE6AA0AFFF76D6DA0EE5DB81469FCEE" blockId="4.[151,1436,715,811]" pageId="4" pageNumber="451">
|
||
<emphasis id="B92D7618FFF76D6DA0EE5DB81742FD26" bold="true" box="[151,278,715,739]" pageId="4" pageNumber="451">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C59D189FFF76D6DA1665DBF1794FD26" authorityName="Nagel" authorityYear="2024" box="[287,448,716,739]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="species" species="agatae" status="sp. nov.">
|
||
<emphasis id="B92D7618FFF76D6DA1665DBF1794FD26" box="[287,448,716,739]" italics="true" pageId="4" pageNumber="451">Paussus agatae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92D7618FFF76D6DA1B05DB81449FD26" bold="true" box="[457,541,715,739]" pageId="4" pageNumber="451">
|
||
<taxonomicNameLabel id="A21ECB63FFF76D6DA1B05DB81449FD26" box="[457,541,715,739]" pageId="4" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype, head, mouthparts, ventral view. Inset: Left maxillary palpomere II, ventral view, slightly distorted to show broadest view, with emarginate mesal margin and slightly convex outer margin. Scale 0.5 mm. Illustration: Armin Coray, Basel.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE6AA0AFFF76D6DA0BE5C0E158DFC79" blockId="4.[151,1437,893,1172]" pageId="4" pageNumber="451">
|
||
<emphasis id="B92D7618FFF76D6DA0BE5C0E1742FC52" box="[199,278,893,919]" italics="true" pageId="4" pageNumber="451">Elytra:</emphasis>
|
||
Dorsal disc with dense punctuation, distance between neighboring punctures ca. 1 puncture at minimum. Pubescence of medium length, proximally upright to distally curved setae.
|
||
</paragraph>
|
||
<paragraph id="8BE6AA0AFFF76D6DA0BE5CB615A0FC1A" blockId="4.[151,1437,893,1172]" box="[199,1012,965,991]" pageId="4" pageNumber="451">
|
||
<emphasis id="B92D7618FFF76D6DA0BE5CB61713FC1A" box="[199,327,965,991]" italics="true" pageId="4" pageNumber="451">Hind wings</emphasis>
|
||
visible through cleft of the dehiscent elytra, fully developed.
|
||
</paragraph>
|
||
<paragraph id="8BE6AA0AFFF76D6DA0BE5C9A1438FBE2" blockId="4.[151,1437,893,1172]" pageId="4" pageNumber="451">
|
||
<taxonomicName id="4C59D189FFF76D6DA0BE5C9A1766FBC6" box="[199,306,1001,1027]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="451" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92D7618FFF76D6DA0BE5C9A1766FBC6" box="[199,306,1001,1027]" italics="true" pageId="4" pageNumber="451">Pygidium</emphasis>
|
||
</taxonomicName>
|
||
excavated, bottom glossy, dorsolateral margins at both sides explanate, upturned, and bent towards the disc. Marginal trichome fringe absent.
|
||
</paragraph>
|
||
<paragraph id="8BE6AA0AFFF76D6DA0BE5B411455FB56" blockId="4.[151,1437,893,1172]" pageId="4" pageNumber="451">
|
||
<emphasis id="B92D7618FFF76D6DA0BE5B411727FB8E" box="[199,371,1074,1099]" italics="true" pageId="4" pageNumber="451">Measurements:</emphasis>
|
||
Body length from anterior margin of head to apex of elytra
|
||
<quantity id="4CA107EFFFF76D6DA46B5B42123FFB89" box="[1042,1131,1073,1100]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="451" unit="mm" value="3.8">3.8 mm</quantity>
|
||
. Elytra 1.7 times as long as wide (= length from anterior margin of shoulder to apex divided by total width of both elytra). Antennal club
|
||
<quantity id="4CA107EFFFF76D6DA5125B261697FB51" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="4" pageNumber="451" unit="mm" value="0.84">0.84 mm</quantity>
|
||
long, twice as long as wide.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |