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<document id="EAC77BAA5AC17C867D292523DFD6C568" ID-CLB-Dataset="6572" ID-DOI="10.5281/zenodo.4502634" ID-GBIF-Dataset="fde5f083-514d-400b-b886-e792390afd86" ID-ISSN="2345-7600" ID-Zenodo-Dep="4502634" ID-ZooBank="E9197A2F-EE46-4E5E-93C5-37E4CEC1A240" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612459456307" checkinUser="carolina" docAuthor="A. D., Tran &amp; Polhemus, D. A." docDate="2017" docId="03E887F5377A3F1BFC4AF97197B53141" docLanguage="en" docName="RafflesBZool.65.109-149.pdf" docOrigin="Raffles Bulletin of Zoology 65" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Metrocoris sapa A.D. &amp; Polhemus 2017, new species" docType="treatment" docVersion="9" lastPageNumber="139" masterDocId="FFD1FF8D37663F05FF95FF8C92763474" masterDocTitle="The genus Metrocoris Mayr, 1865 (Gerromorpha: Gerridae) in Vietnam, with descriptions of five new species" masterLastPageNumber="149" masterPageNumber="109" pageNumber="137" updateTime="1698926865763" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="28121A99F5CD4F45006F7701F16449A2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2DFB5EB115B7682C1701BC7E1B5B1219">The genus Metrocoris Mayr, 1865 (Gerromorpha: Gerridae) in Vietnam, with descriptions of five new species</mods:title>
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<mods:name id="0967E68180E112375751B71088837CE4" type="personal">
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<mods:namePart id="9DE1EF571CD22013B75F5EFD5934558D">A. D., Tran</mods:namePart>
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<mods:namePart id="05516535AA9FBB39F942FF5564EF9D77">Polhemus, D. A.</mods:namePart>
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<mods:title id="A52068E5CF512C1749F85A5F57A7DFC8">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="7F7737F1B6559128A9C44255D9955713">2017</mods:date>
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<mods:number id="F1900EE2938C021FFCCBB9A3EE8808E0">2017-05-24</mods:number>
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<mods:number id="050CE427CCAD4D0EAB49E93049FC4358">65</mods:number>
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<mods:identifier id="D1C6F3ADEF0CC1AEFAD4EB9E87239127" type="DOI">10.5281/zenodo.4502634</mods:identifier>
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<treatment id="03E887F5377A3F1BFC4AF97197B53141" ID-DOI="http://doi.org/10.5281/zenodo.4776495" ID-GBIF-Taxon="182247954" ID-Zenodo-Dep="4776495" LSID="urn:lsid:plazi:treatment:03E887F5377A3F1BFC4AF97197B53141" httpUri="http://treatment.plazi.org/id/03E887F5377A3F1BFC4AF97197B53141" lastPageId="30" lastPageNumber="139" pageId="28" pageNumber="137">
<subSubSection id="C35B6568377A3F19FC4AF97197513361" box="[991,1319,1789,1813]" pageId="28" pageNumber="137" type="nomenclature">
<paragraph id="8BFE36E3377A3F19FC4AF97197513361" blockId="28.[991,1319,1789,1845]" box="[991,1319,1789,1813]" pageId="28" pageNumber="137">
<heading id="D0B6818F377A3F19FC4AF97197513361" bold="true" box="[991,1319,1789,1813]" centered="true" fontSize="10" level="2" pageId="28" pageNumber="137" reason="2">
<emphasis id="B935EAF1377A3F19FC4AF97197513361" bold="true" box="[991,1319,1789,1813]" pageId="28" pageNumber="137">
<taxonomicName id="4C414D60377A3F19FC4AF97196E43361" ID-CoL="42JNV" authority="A.D. &amp; Polhemus, 2017" authorityName="A.D. &amp; Polhemus" authorityYear="2017" box="[991,1170,1789,1813]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="137" phylum="Arthropoda" rank="species" species="sapa" status="sp. nov.">
<emphasis id="B935EAF1377A3F19FC4AF97196E43361" bold="true" box="[991,1170,1789,1813]" italics="true" pageId="28" pageNumber="137">Metrocoris sapa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A206578A377A3F19FB34F97197513361" box="[1185,1319,1789,1813]" pageId="28" pageNumber="137" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35B6568377A3F19FB97F89197753341" box="[1026,1283,1821,1845]" pageId="28" pageNumber="137" type="description">
<paragraph id="8BFE36E3377A3F19FB97F89197753341" blockId="28.[991,1319,1789,1845]" box="[1026,1283,1821,1845]" pageId="28" pageNumber="137">
(
<figureCitation id="137A2A66377A3F19FB9FF89196E63341" box="[1034,1168,1821,1845]" captionStart="Figs" captionStartId="6.[135,179,1936,1958]" captionTargetBox="[242,1379,246,1880]" captionTargetId="figure-7@6.[135,1473,189,1921]" captionTargetPageId="6" captionText="Figs. 18–31. Dorsal colour patterns of Metrocoris species. 18, 19: M. inthanon, specimens from Vietnam, Nghe An Prov., Pu Mat Nature Reserve (CL 4385). 20, 21: M. sicilis, new species, paratypes, from Vietnam, Hanoi, Ba Vi N’Park (CL 4393). 22, 23: M. triangulatus, specimens from Vietnam, Kontum Prov., 60 km NW. of Kontum (CL 4280). 24, 25: M. nigriventris, new species, paratypes, from Vietnam, Lai Chau Prov., 12 km N. of Lai Chau (CL 4410). 26, 27: M. monticola, new species, paratypes, from Vietnam, Lao Cai Prov., Sa Pa, Sin Chai (TAD1363). 28, 29: M. sapa, new species, paratypes, from Vietnam, Lao Cai Prov., Sa Pa, Nui Xe (TAD0416). 30, 31: M. tenuicornis, specimens from Vietnam, Gia Lai Prov., 60 km N. of An Khe (CL 4309)." figureDoi="http://doi.org/10.5281/zenodo.4502642" httpUri="https://zenodo.org/record/4502642/files/figure.png" pageId="28" pageNumber="137">Figs. 28, 29</figureCitation>
,
<figureCitation id="137A2A66377A3F19FB08F891968D3341" box="[1181,1275,1821,1845]" captionStart="Figs" captionStartId="29.[113,157,1216,1238]" captionTargetBox="[113,1452,191,1202]" captionTargetId="figure-9@29.[100,1462,178,1212]" captionTargetPageId="29" captionText="Figs. 117–122. Morphological features of Metrocoris sapa, new species. 117: right fore leg of male. 118: left paramere. 119, 120: endosomal sclerites, lateral and dorsal views. 121: hind coxa and trochanter of female. 122: sternum 7 of female, ventral view (121, 122 same scale)." figureDoi="http://doi.org/10.5281/zenodo.4502666" httpUri="https://zenodo.org/record/4502666/files/figure.png" pageId="28" pageNumber="137">117–122</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35B6568377A3F18FCD6F8D190873116" lastPageId="29" lastPageNumber="138" pageId="28" pageNumber="137" type="materials_examined">
<paragraph id="8BFE36E3377A3F19FCD6F8D1968533A1" blockId="28.[835,1475,1885,2006]" pageId="28" pageNumber="137">
<emphasis id="B935EAF1377A3F19FCD6F8D196553301" bold="true" box="[835,1059,1885,1909]" pageId="28" pageNumber="137">Material examined.</emphasis>
<materialsCitation id="3B293CBE377A3F19FBBEF8D1968533A1" ID-GBIF-Occurrence="3114485701" collectingDate="2004-07-04" collectionCode="ZMHU" collectorName="Tran A. D." country="Vietnam" location="Lao Cai Prov." pageId="28" pageNumber="137" specimenCount="2" stateProvince="Lao Cai" typeStatus="holotype">
<emphasis id="B935EAF1377A3F19FBBEF8D196E53301" bold="true" box="[1067,1171,1885,1909]" pageId="28" pageNumber="137">
<typeStatus id="54FA8841377A3F19FBBEF8D196E53301" box="[1067,1171,1885,1909]" pageId="28" pageNumber="137" type="holotype">Holotype</typeStatus>
</emphasis>
(apterous male) &amp;
<emphasis id="B935EAF1377A3F19FAFCF8D197B53301" bold="true" box="[1385,1475,1885,1909]" pageId="28" pageNumber="137">
<typeStatus id="54FA8841377A3F19FAFCF8D197B53301" box="[1385,1475,1885,1909]" pageId="28" pageNumber="137" type="allotype">allotype</typeStatus>
</emphasis>
(apterous female):
<collectingCountry id="F3567673377A3F19FB9EF8F296FF33E2" box="[1035,1161,1918,1942]" name="Vietnam" pageId="28" pageNumber="137">VIETNAM</collectingCountry>
,
<collectingRegion id="4985F801377A3F19FB06F8F2975F33E1" box="[1171,1321,1917,1941]" country="Vietnam" name="Lao Cai" pageId="28" pageNumber="137">Lao Cai Prov.</collectingRegion>
, Sa Pa, Hoang Lien N’Park, Nui Xe, upstream of Suoi Vang, coll.
<collectorName id="26B45335377A3F19FA1AF812910F33A2" pageId="28" pageNumber="137">Tran A.D.</collectorName>
,
<date id="FFFF1023377A3F19FC13F831967C33A1" box="[902,1034,1981,2005]" pageId="28" pageNumber="137" value="2004-07-04">
<collectingDate id="EFBBE9CB377A3F19FC13F831967C33A1" box="[902,1034,1981,2005]" pageId="28" pageNumber="137" value="2004-07-04">4 July 2004</collectingDate>
</date>
, TAD0416 (
<collectionCode id="ED50AE26377A3F19FB06F831969E33A1" box="[1171,1256,1981,2005]" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="28" pageNumber="137">ZMHU</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BFE36E3377A3F18FCD6F87190873116" blockId="28.[835,1474,2045,2101]" lastBlockId="29.[113,753,1322,1379]" lastPageId="29" lastPageNumber="138" pageId="28" pageNumber="137">
<materialsCitation id="3B293CBE377A3F19FCD6F871910E3C41" ID-GBIF-Occurrence="3114485412" country="Vietnam" location="Lao Cai Prov." pageId="28" pageNumber="137" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Lao Cai" typeStatus="paratype">
<emphasis id="B935EAF1377A3F19FCD6F87191C33C61" bold="true" box="[835,949,2045,2069]" pageId="28" pageNumber="137">
<typeStatus id="54FA8841377A3F19FCD6F87191C33C61" box="[835,949,2045,2069]" pageId="28" pageNumber="137" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="F3567673377A3F19FC53F87296323C62" box="[966,1092,2046,2070]" name="Vietnam" pageId="28" pageNumber="137">VIETNAM</collectingCountry>
,
<emphasis id="B935EAF1377A3F19FBC4F87196893C61" bold="true" box="[1105,1279,2045,2069]" pageId="28" pageNumber="137">
<collectingRegion id="4985F801377A3F19FBC4F871968C3C61" box="[1105,1274,2045,2069]" country="Vietnam" name="Lao Cai" pageId="28" pageNumber="137">Lao Cai Prov.</collectingRegion>
:
</emphasis>
<specimenCount id="9D47FD6A377A3F19FA9DF87197233C61" box="[1288,1365,2045,2069]" pageId="28" pageNumber="137" type="male">1 male</specimenCount>
,
<specimenCount id="9D47FD6A377A3F19FAF6F87197B43C61" box="[1379,1474,2045,2069]" pageId="28" pageNumber="137" type="female">1 female</specimenCount>
(apt)
</materialsCitation>
,
<materialsCitation id="3B293CBE377A3F19FC17F79197B43C41" ID-GBIF-Occurrence="3114485492" box="[898,1474,2077,2101]" country="Vietnam" location="Lao Cai Prov." pageId="28" pageNumber="137" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Lao Cai" typeStatus="holotype">
<specimenCount id="9D47FD6A377A3F19FC17F79191BC3C41" box="[898,970,2077,2101]" pageId="28" pageNumber="137" type="male">1 male</specimenCount>
,
<specimenCount id="9D47FD6A377A3F19FC41F791964C3C41" box="[980,1082,2077,2101]" pageId="28" pageNumber="137" type="female">4 females</specimenCount>
(mpt), same locality data as
<typeStatus id="54FA8841377A3F19FAF0F79197B43C41" box="[1381,1474,2077,2101]" pageId="28" pageNumber="137" type="holotype">holotype</typeStatus>
</materialsCitation>
(
<collectionCode id="ED50AE26377B3F18FFECFAA692C43136" box="[121,178,1322,1346]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="29" pageNumber="138">ZRC</collectionCode>
);
<materialsCitation id="3B293CBE377B3F18FF52FAA6909B3116" ID-GBIF-Occurrence="3114485337" collectingDate="2004-07-04" collectionCode="ZRC" collectorName="Tran A. D." pageId="29" pageNumber="138" specimenCount="1">
<specimenCount id="9D47FD6A377B3F18FF52FAA693643136" box="[199,274,1322,1346]" pageId="29" pageNumber="138" type="male">1 male</specimenCount>
(mpt),
<collectorName id="26B45335377B3F18FEF0FAA693D03136" box="[357,422,1322,1346]" pageId="29" pageNumber="138">Sa Pa</collectorName>
,
<collectorName id="26B45335377B3F18FE26FAA790FB3136" box="[435,653,1322,1347]" pageId="29" pageNumber="138">Hoang Lien N’Park</collectorName>
,
<collectorName id="26B45335377B3F18FD0CFAA6909A3136" box="[665,748,1322,1346]" pageId="29" pageNumber="138">Nui Xe</collectorName>
,
<collectorName id="26B45335377B3F18FFE4FAC692903117" box="[113,230,1354,1379]" pageId="29" pageNumber="138">Suoi Vang</collectorName>
, coll.
<collectorName id="26B45335377B3F18FEBDFAC793EE3117" box="[296,408,1355,1379]" pageId="29" pageNumber="138">Tran A.D.</collectorName>
,
<date id="FFFF1023377B3F18FE31FAC690533116" box="[420,549,1354,1378]" pageId="29" pageNumber="138" value="2004-07-04">
<collectingDate id="EFBBE9CB377B3F18FE31FAC690533116" box="[420,549,1354,1378]" pageId="29" pageNumber="138" value="2004-07-04">4 July 2004</collectingDate>
</date>
, TAD0417 (
<collectionCode id="ED50AE26377B3F18FD39FAC690903116" box="[684,742,1354,1378]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="29" pageNumber="138">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF3E666B377B3F18FFE4FB4C97DA3087" ID-DOI="http://doi.org/10.5281/zenodo.4502666" ID-Zenodo-Dep="4502666" httpUri="https://zenodo.org/record/4502666/files/figure.png" pageId="29" pageNumber="138" startId="29.[113,157,1216,1238]" targetBox="[113,1452,191,1202]" targetPageId="29">
<paragraph id="8BFE36E3377B3F18FFE4FB4C97DA3087" blockId="29.[112,1452,1216,1267]" pageId="29" pageNumber="138">
Figs. 117–122. Morphological features of
<taxonomicName id="4C414D60377B3F18FD97FB4D90E930A2" authority="A.D. &amp; Polhemus, 2017" authorityName="A.D. &amp; Polhemus" authorityYear="2017" box="[514,671,1217,1238]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="138" phylum="Arthropoda" rank="species" species="sapa" status="sp. nov.">
<emphasis id="B935EAF1377B3F18FD97FB4D90E930A2" box="[514,671,1217,1238]" italics="true" pageId="29" pageNumber="138">Metrocoris sapa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A206578A377B3F18FD3EFB4D915630A2" box="[683,800,1216,1238]" pageId="29" pageNumber="138" rank="species">new species</taxonomicNameLabel>
. 117: right fore leg of male. 118: left paramere. 119, 120: endosomal sclerites, lateral and dorsal views. 121: hind coxa and trochanter of female. 122: sternum 7 of female, ventral view (121, 122 same scale).
</paragraph>
</caption>
<subSubSection id="C35B6568377B3F18FFE4FA06901D3C36" pageId="29" pageNumber="138" type="diagnosis">
<paragraph id="8BFE36E3377B3F18FFE4FA06901D3C36" blockId="29.[112,755,1418,2114]" pageId="29" pageNumber="138">
<emphasis id="B935EAF1377B3F18FFE4FA06929D31D6" bold="true" box="[113,235,1418,1442]" pageId="29" pageNumber="138">Diagnosis.</emphasis>
Both sexes: body sub-ovate, ventral surface light coloured ventral surface, without any dark mark; fore femur clearly longer than mesosternum. Male: fore femur moderately slender (ratio length/width: 6.60), proximal half, broader than distal half, constricted near near the middle, pilosity denser and longer at proximal half (
<figureCitation id="137A2A66377B3F18FDE9F9A690943236" box="[636,738,1578,1602]" captionStart="Figs" captionStartId="29.[113,157,1216,1238]" captionTargetBox="[113,1452,191,1202]" captionTargetId="figure-9@29.[100,1462,178,1212]" captionTargetPageId="29" captionText="Figs. 117–122. Morphological features of Metrocoris sapa, new species. 117: right fore leg of male. 118: left paramere. 119, 120: endosomal sclerites, lateral and dorsal views. 121: hind coxa and trochanter of female. 122: sternum 7 of female, ventral view (121, 122 same scale)." figureDoi="http://doi.org/10.5281/zenodo.4502666" httpUri="https://zenodo.org/record/4502666/files/figure.png" pageId="29" pageNumber="138">Fig. 117</figureCitation>
); genitalia small: abdominal segment 8 clearly longer than sternum 7; proctiger simple, elongated and slender; pygophore ovate on ventral view; paramere falciform, simple, curved at middle part, apex rounded and not modified (
<figureCitation id="137A2A66377B3F18FD16F926909532B6" box="[643,739,1706,1730]" captionStart="Figs" captionStartId="29.[113,157,1216,1238]" captionTargetBox="[113,1452,191,1202]" captionTargetId="figure-9@29.[100,1462,178,1212]" captionTargetPageId="29" captionText="Figs. 117–122. Morphological features of Metrocoris sapa, new species. 117: right fore leg of male. 118: left paramere. 119, 120: endosomal sclerites, lateral and dorsal views. 121: hind coxa and trochanter of female. 122: sternum 7 of female, ventral view (121, 122 same scale)." figureDoi="http://doi.org/10.5281/zenodo.4502666" httpUri="https://zenodo.org/record/4502666/files/figure.png" pageId="29" pageNumber="138">Fig. 118</figureCitation>
); endosoma (
<figureCitation id="137A2A66377B3F18FF7EF94693F03296" box="[235,390,1738,1762]" captionStart="Figs" captionStartId="29.[113,157,1216,1238]" captionTargetBox="[113,1452,191,1202]" captionTargetId="figure-9@29.[100,1462,178,1212]" captionTargetPageId="29" captionText="Figs. 117–122. Morphological features of Metrocoris sapa, new species. 117: right fore leg of male. 118: left paramere. 119, 120: endosomal sclerites, lateral and dorsal views. 121: hind coxa and trochanter of female. 122: sternum 7 of female, ventral view (121, 122 same scale)." figureDoi="http://doi.org/10.5281/zenodo.4502666" httpUri="https://zenodo.org/record/4502666/files/figure.png" pageId="29" pageNumber="138">Figs. 119, 120</figureCitation>
): dorsal sclerite long and recurved on apical, apical accessory sclerite distinct, ventral sclerite shorter than dorsal sclerite, lateral sclerite straight, rather broad, partially sclerotised. Female: fore femur simple, not modified as in male, and slender (ratio length/width: 7.94); hind coxa and trochanter enlongated, without apical projection, interior side of hind coxa and trochanter with long hairs (
<figureCitation id="137A2A66377B3F18FF66F826932033B6" box="[243,342,1962,1986]" captionStart="Figs" captionStartId="29.[113,157,1216,1238]" captionTargetBox="[113,1452,191,1202]" captionTargetId="figure-9@29.[100,1462,178,1212]" captionTargetPageId="29" captionText="Figs. 117–122. Morphological features of Metrocoris sapa, new species. 117: right fore leg of male. 118: left paramere. 119, 120: endosomal sclerites, lateral and dorsal views. 121: hind coxa and trochanter of female. 122: sternum 7 of female, ventral view (121, 122 same scale)." figureDoi="http://doi.org/10.5281/zenodo.4502666" httpUri="https://zenodo.org/record/4502666/files/figure.png" pageId="29" pageNumber="138">Fig. 121</figureCitation>
); abdominal sternum 7 broader than long, clearly longer than previous sterna together, constricted laterally, posterior part producing a median lobe with nearly straight apical margin, directed dorsad, and completely enclosing genitalia on ventral view (
<figureCitation id="137A2A66377B3F18FE6AF7A690293C36" box="[511,607,2090,2114]" captionStart="Figs" captionStartId="29.[113,157,1216,1238]" captionTargetBox="[113,1452,191,1202]" captionTargetId="figure-9@29.[100,1462,178,1212]" captionTargetPageId="29" captionText="Figs. 117–122. Morphological features of Metrocoris sapa, new species. 117: right fore leg of male. 118: left paramere. 119, 120: endosomal sclerites, lateral and dorsal views. 121: hind coxa and trochanter of female. 122: sternum 7 of female, ventral view (121, 122 same scale)." figureDoi="http://doi.org/10.5281/zenodo.4502666" httpUri="https://zenodo.org/record/4502666/files/figure.png" pageId="29" pageNumber="138">Fig. 122</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B6568377B3F1BFCB8FAA691713361" lastPageId="30" lastPageNumber="139" pageId="29" pageNumber="138" type="description">
<paragraph id="8BFE36E3377B3F18FCB8FAA697DE31B6" blockId="29.[811,1452,1322,1474]" pageId="29" pageNumber="138">
<emphasis id="B935EAF1377B3F18FCB8FAA691C53136" bold="true" box="[813,947,1322,1346]" pageId="29" pageNumber="138">Description.</emphasis>
Size: apterous males: length 4.80–5.60 (
<typeStatus id="54FA8841377B3F18FADAFAA697DA3136" box="[1359,1452,1322,1346]" pageId="29" pageNumber="138" type="holotype">holotype</typeStatus>
5.60), width 2.48–2.97 (
<typeStatus id="54FA8841377B3F18FBB2FAC696F03116" box="[1063,1158,1354,1378]" pageId="29" pageNumber="138" type="holotype">holotype</typeStatus>
2.97), macropterous males: length 8.00–8.10, width 2.87; apterous females: length 4.80–5.10 (
<typeStatus id="54FA8841377B3F18FC32FA06918931D6" box="[935,1023,1418,1442]" pageId="29" pageNumber="138" type="allotype">allotype</typeStatus>
5.10), width 2.90–2.94 (
<typeStatus id="54FA8841377B3F18FA98FA06971331D6" box="[1293,1381,1418,1442]" pageId="29" pageNumber="138" type="allotype">allotype</typeStatus>
2.94), macropterous females: length 8.00–8.30, width 3.00–3.04.
</paragraph>
<paragraph id="8BFE36E3377B3F1BFCB8FA6693DA35C1" blockId="29.[811,1454,1514,2114]" lastBlockId="30.[135,775,189,437]" lastPageId="30" lastPageNumber="139" pageId="29" pageNumber="138">
Colour (
<figureCitation id="137A2A66377B3F18FC10FA66967E3276" box="[901,1032,1514,1538]" captionStart="Figs" captionStartId="6.[135,179,1936,1958]" captionTargetBox="[242,1379,246,1880]" captionTargetId="figure-7@6.[135,1473,189,1921]" captionTargetPageId="6" captionText="Figs. 18–31. Dorsal colour patterns of Metrocoris species. 18, 19: M. inthanon, specimens from Vietnam, Nghe An Prov., Pu Mat Nature Reserve (CL 4385). 20, 21: M. sicilis, new species, paratypes, from Vietnam, Hanoi, Ba Vi N’Park (CL 4393). 22, 23: M. triangulatus, specimens from Vietnam, Kontum Prov., 60 km NW. of Kontum (CL 4280). 24, 25: M. nigriventris, new species, paratypes, from Vietnam, Lai Chau Prov., 12 km N. of Lai Chau (CL 4410). 26, 27: M. monticola, new species, paratypes, from Vietnam, Lao Cai Prov., Sa Pa, Sin Chai (TAD1363). 28, 29: M. sapa, new species, paratypes, from Vietnam, Lao Cai Prov., Sa Pa, Nui Xe (TAD0416). 30, 31: M. tenuicornis, specimens from Vietnam, Gia Lai Prov., 60 km N. of An Khe (CL 4309)." figureDoi="http://doi.org/10.5281/zenodo.4502642" httpUri="https://zenodo.org/record/4502642/files/figure.png" pageId="29" pageNumber="138">Figs. 28, 29</figureCitation>
). Dorsal body mainly pale, light brown markings expanded and yellow markings reduced. Antennae dark brown or black, proximal part of antennal segment 1 yellowish brown. Interocular region of head mainly dark, with yellow mark on posterior margin of head. Pronotum with black anterior margin confluent with a slender median black marking, forming a T-shaped marking; antero-lateral corners with a large light brown marking, leaving the yellow markings slender along the median black marking and on posterior margin. Mesonotum with a slender median black marking, two large light brown markings along each side of median black marking, and a light brown marking on each antero-lateral corner. Metanotum with a slender median black marking, two yellow markings on each side of the median marking running along posterior margin. Abdominal terga 1–7: each tergum dark brown or black on anterior part, and yellow on posterior part. Metacetabula, inner half yellow, outer half brown to dark brown. All coxae and trochanters mainly yellow with small brown markings. Fore femur yellow with 1 brown stripe on outer side and 1 on inner side, confluent with brown apical part. Middle and hind femora mainly yellowish brown. Tibia and tarsus dark brown or black. Venter of body mainly light yellow. In macropterous form, pronotum with median T-shaped black marking on anterior margin, pronotal lobe mainly yellowish brown with lateral margin light yellow from humeri to tip of pronotal lobe, wings mainly brown.
</paragraph>
<paragraph id="8BFE36E337783F1BFF1DFE5291733701" blockId="30.[134,776,477,885]" pageId="30" pageNumber="139">
Apterous male (
<typeStatus id="54FA884137783F1BFEB9FE5193F83581" box="[300,398,477,501]" pageId="30" pageNumber="139" type="holotype">holotype</typeStatus>
): Head width 1.59, interocular width 0.67, eye size 0.75. Lengths of antennal segments 1–4: 2.54: 1.17: 1.03: 0.71, antennal segment 2 with scattered spines. Pronotum length 0.49, mesonotum length 1.58. Lengths of leg segments (femur: tibia: tarsus 1: tarsus 2): fore leg: 2.64: 2.51: 0.08: 0.86; middle leg: 7.30: 5.45: 2.51: 0.35; hind leg: 7.80: 4.61: 0.35: 0.44. Fore femur: width 0.40, other characteristics as in Diagnosis above. Abdomen length on ventral view: 1.71, pregenital length 0.83; sternum 7 slightly longer than sterna 5 and 6 combined, length 0.27. Genital segments small: segment 8 sub-rectangular on dorsal view, length 0.44, width 0.56; characteristics of pygophore, proctiger, paramere, and endosoma as in Diagnosis above.
</paragraph>
<paragraph id="8BFE36E337783F1BFF1DFC1291713161" blockId="30.[134,776,925,1301]" pageId="30" pageNumber="139">
Apterous female (
<typeStatus id="54FA884137783F1BFEC4FC1193D937C1" box="[337,431,925,949]" pageId="30" pageNumber="139" type="allotype">allotype</typeStatus>
): Head width 1.64, interocular width 0.65, eye size 0.78. Lengths of antennal segments 1–4: 2.23: 1.06: 1.03: 0.71, antennal segment 1 with scattered spines. Pronotum length 0.44, mesonotum length 1.70. Lengths of leg segments: fore leg: 2.54: 2.38: 0.08: 0.83; middle leg: 7.00: 5.10: 2.35: 0.33; hind leg: 7.30: 4.56: 0.32: 0.42. Fore femur: width 0.32, other characteristics as in Diagnosis above. Hind coxae without apical projection, length of hind trochanter 0.76, interior side of hind coxa and trochanter with long hairs (
<figureCitation id="137A2A6637783F1BFE0EFB31938230A1" box="[411,500,1213,1237]" captionStart="Figs" captionStartId="29.[113,157,1216,1238]" captionTargetBox="[113,1452,191,1202]" captionTargetId="figure-9@29.[100,1462,178,1212]" captionTargetPageId="29" captionText="Figs. 117–122. Morphological features of Metrocoris sapa, new species. 117: right fore leg of male. 118: left paramere. 119, 120: endosomal sclerites, lateral and dorsal views. 121: hind coxa and trochanter of female. 122: sternum 7 of female, ventral view (121, 122 same scale)." figureDoi="http://doi.org/10.5281/zenodo.4502666" httpUri="https://zenodo.org/record/4502666/files/figure.png" pageId="30" pageNumber="139">Fig. 121</figureCitation>
). Abdominal venter length 1.08. Abdominal sternum 7 broader than long, length ca. 0.98 (caudal view), shape of sternum 7: see Diagnosis above.
</paragraph>
<paragraph id="8BFE36E337783F1BFF1DFAB290813261" blockId="30.[135,775,1341,1557]" pageId="30" pageNumber="139">Macropterous male: head width 1.56, interocular width 0.63, eye size 0.70. Lengths of antennal segments: 2.48: 1.17: 0.98: 0.71. Pronotum length 3.30, humeral width 2.48. Fore wing length 6.40. Lengths of leg segments: fore leg: 2.48: 2.44: 0.10: 0.79; middle leg: 7.10: 5.15: 2.33: 0.35; hind leg: 7.30: 4.26: 0.32: 0.40. Margin of connexivum with long hairs. Other characteristics similar to apterous male.</paragraph>
<paragraph id="8BFE36E337783F1BFF1DF9B291713361" blockId="30.[135,775,1597,1813]" pageId="30" pageNumber="139">Macropterous female: head with 1.52, interocular width 0.63, eye size 0.71. Lengths of antennal segments: 2.08: 0.90: 0.95: 0.68. Pronotum length 3.45, humeral width 2.48. Fore wing length 6.30. Lengths of leg segments: fore leg: 2.48: 2.31: 0.08: 0.76; middle leg: 6.30: 4.70: 2.38: 0.35; hind leg: 6.80: 4.17: 0.35: 0.44. Margin of connexivum with long hairs. Other characteristics similar to apterous female.</paragraph>
</subSubSection>
<subSubSection id="C35B656837783F1BFF1DF8B197B737E1" pageId="30" pageNumber="139" type="discussion">
<paragraph id="8BFE36E337783F1BFF1DF8B1935A3C61" blockId="30.[136,777,1853,2069]" pageId="30" pageNumber="139">
<emphasis id="B935EAF137783F1BFF1DF8B1928E3321" bold="true" box="[136,248,1853,1877]" pageId="30" pageNumber="139">Remarks.</emphasis>
Based on the definition of
<emphasis id="B935EAF137783F1BFDA6F8B290D83321" box="[563,686,1854,1877]" italics="true" pageId="30" pageNumber="139">M. compar</emphasis>
species group (sensu
<bibRefCitation id="EFD04B1237783F1BFE8EF8D190703301" author="Chen PP &amp; Nieser N" box="[283,518,1885,1909]" pageId="30" pageNumber="139" pagination="45 - 82" refId="ref41516" refString="Chen PP &amp; Nieser N (1993) A taxonomic revision of the Oriental water strider genus Metrocoris Mayr (Hemiptera, Gerridae). Part I and Part II. Steenstrupia, 19 (1 - 2): 1 - 43, 45 - 82." type="journal article" year="1993">Chen &amp; Nieser, 1993</bibRefCitation>
, see Remarks under
<taxonomicName id="4C414D6037783F1BFD7EF8D2928533E1" authority="A.D. &amp; Polhemus, 2017" authorityName="A.D. &amp; Polhemus" authorityYear="2017" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="monticola" status="sp. nov.">
<emphasis id="B935EAF137783F1BFD7EF8D2928533E1" italics="true" pageId="30" pageNumber="139">M. monticola</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A206578A37783F1BFE97F8F293FE33E1" box="[258,392,1917,1941]" pageId="30" pageNumber="139" rank="species">new species</taxonomicNameLabel>
), we can assign also
<taxonomicName id="4C414D6037783F1BFDE7F8F290BC33E1" authority="A.D. &amp; Polhemus, 2017" authorityName="A.D. &amp; Polhemus" authorityYear="2017" box="[626,714,1918,1941]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="sapa" status="sp. nov.">
<emphasis id="B935EAF137783F1BFDE7F8F290BC33E1" box="[626,714,1918,1941]" italics="true" pageId="30" pageNumber="139">M. sapa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A206578A37783F1BFD4FF8F292A333C1" pageId="30" pageNumber="139" rank="species">new species</taxonomicNameLabel>
to the
<emphasis id="B935EAF137783F1BFEB7F81293E133C1" box="[290,407,1950,1973]" italics="true" pageId="30" pageNumber="139">M. compar</emphasis>
species group. However,
<taxonomicName id="4C414D6037783F1BFD24F812917133C1" authorityName="A.D. &amp; Polhemus" authorityYear="2017" box="[689,775,1950,1973]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="sapa">
<emphasis id="B935EAF137783F1BFD24F812917133C1" box="[689,775,1950,1973]" italics="true" pageId="30" pageNumber="139">M. sapa</emphasis>
</taxonomicName>
does not totally fit the diagnostic characteristics of this group because it has a light coloured ventral surface, without any dark markings.
</paragraph>
<paragraph id="8BFE36E337783F1BFCD6FF3296D036C1" blockId="30.[835,1477,189,693]" pageId="30" pageNumber="139">
<taxonomicName id="4C414D6037783F1BFCD6FF32918334A1" authority="A.D. &amp; Polhemus, 2017" authorityName="A.D. &amp; Polhemus" authorityYear="2017" box="[835,1013,190,213]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="sapa" status="sp. nov.">
<emphasis id="B935EAF137783F1BFCD6FF32918334A1" box="[835,1013,190,213]" italics="true" pageId="30" pageNumber="139">Metrocoris sapa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A206578A37783F1BFB93FF3296FF34A1" box="[1030,1161,189,213]" pageId="30" pageNumber="139" rank="species">new species</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C414D6037783F1BFAC8FF3296423481" authority="Chen &amp; Nieser, 1993" authorityName="Chen &amp; Nieser" authorityYear="1993" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B935EAF137783F1BFAC8FF3297B434A1" box="[1373,1474,189,213]" italics="true" pageId="30" pageNumber="139">M. hirtus</emphasis>
<bibRefCitation id="EFD04B1237783F1BFCD6FF5196423481" author="Chen PP &amp; Nieser N" box="[835,1076,221,245]" pageId="30" pageNumber="139" pagination="45 - 82" refId="ref41516" refString="Chen PP &amp; Nieser N (1993) A taxonomic revision of the Oriental water strider genus Metrocoris Mayr (Hemiptera, Gerridae). Part I and Part II. Steenstrupia, 19 (1 - 2): 1 - 43, 45 - 82." type="journal article" year="1993">Chen &amp; Nieser, 1993</bibRefCitation>
</taxonomicName>
(from
<collectingRegion id="4985F80137783F1BFB13FF5196963481" box="[1158,1248,221,245]" country="China" name="Sichuan" pageId="30" pageNumber="139">Sichuan</collectingRegion>
,
<collectingCountry id="F356767337783F1BFB7AFF5197473481" box="[1263,1329,221,245]" name="China" pageId="30" pageNumber="139">China</collectingCountry>
) in having a similar general shape of the male parameres, endosomal structures and female hind coxa. The dorsal colour pattern of
<taxonomicName id="4C414D6037783F1BFCF0FEB291B73521" authorityName="A.D. &amp; Polhemus" authorityYear="2017" box="[869,961,318,341]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="sapa">
<emphasis id="B935EAF137783F1BFCF0FEB291B73521" box="[869,961,318,341]" italics="true" pageId="30" pageNumber="139">M. sapa</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C414D6037783F1BFB2DFEB297B73521" authority="Zettel, 2011" authorityName="Zettel" authorityYear="2011" box="[1208,1473,317,341]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="pardus">
<emphasis id="B935EAF137783F1BFB2DFEB297583521" box="[1208,1326,317,341]" italics="true" pageId="30" pageNumber="139">M. pardus</emphasis>
<bibRefCitation id="EFD04B1237783F1BFAADFEB197B73521" author="Zettel H" box="[1336,1473,317,341]" pageId="30" pageNumber="139" pagination="109 - 114" refId="ref42232" refString="Zettel H (2011) Metrocoris pardus sp. n. (Heteroptera: Gerridae) aus Westmalaysien. Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 63 (2): 109 - 114." type="journal article" year="2011">Zettel, 2011</bibRefCitation>
</taxonomicName>
(from Peninsular
<collectingCountry id="F356767337783F1BFB87FED196093501" box="[1042,1151,349,373]" name="Malaysia" pageId="30" pageNumber="139">Malaysia</collectingCountry>
). However, the apex of the paramere in
<taxonomicName id="4C414D6037783F1BFC5AFEF2965E35E1" authorityName="A.D. &amp; Polhemus" authorityYear="2017" box="[975,1064,382,405]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="sapa">
<emphasis id="B935EAF137783F1BFC5AFEF2965E35E1" box="[975,1064,382,405]" italics="true" pageId="30" pageNumber="139">M. sapa</emphasis>
</taxonomicName>
is rounded, simple and not modified like those of
<taxonomicName id="4C414D6037783F1BFC42FE12964A35C1" authorityName="Chen &amp; Nieser" authorityYear="1993" box="[983,1084,413,437]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B935EAF137783F1BFC42FE12964A35C1" box="[983,1084,413,437]" italics="true" pageId="30" pageNumber="139">M. hirtus</emphasis>
</taxonomicName>
or
<taxonomicName id="4C414D6037783F1BFBF3FE1296A135C1" authorityName="Zettel" authorityYear="2011" box="[1126,1239,413,437]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="pardus">
<emphasis id="B935EAF137783F1BFBF3FE1296A135C1" box="[1126,1239,413,437]" italics="true" pageId="30" pageNumber="139">M. pardus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFD04B1237783F1BFA80FE11910A35A1" author="Chen PP &amp; Nieser N" pageId="30" pageNumber="139" pagination="45 - 82" refId="ref41516" refString="Chen PP &amp; Nieser N (1993) A taxonomic revision of the Oriental water strider genus Metrocoris Mayr (Hemiptera, Gerridae). Part I and Part II. Steenstrupia, 19 (1 - 2): 1 - 43, 45 - 82." type="journal article" year="1993">Chen &amp; Nieser, 1993</bibRefCitation>
: Fig. 195;
<bibRefCitation id="EFD04B1237783F1BFC66FE31960F35A1" author="Zettel H" box="[1011,1145,445,469]" pageId="30" pageNumber="139" pagination="109 - 114" refId="ref42232" refString="Zettel H (2011) Metrocoris pardus sp. n. (Heteroptera: Gerridae) aus Westmalaysien. Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 63 (2): 109 - 114." type="journal article" year="2011">Zettel, 2011</bibRefCitation>
:
<figureCitation id="137A2A6637783F1BFB10FE3196B035A1" box="[1157,1222,445,469]" captionStart="Figs" captionStartId="4.[135,179,1891,1913]" captionTargetBox="[264,1325,243,1822]" captionTargetId="figure-8@4.[152,1455,189,1875]" captionTargetPageId="4" captionText="Figs. 3–17. Dorsal colour patterns of Metrocoris species, all apterous morph. 3, 4: M. stranguloides, specimens from Vietnam, Gia Lai Prov., 60 km N. of An Khe (CL 4309). 5, 6: M. bilobatoides, specimens from Vietnam, Lao Cai Prov., 12 km NW. of Sa Pa (CL 4404). 7, 8: M. johnpolhemi, new species, specimens from Vietnam, Lai Chau Prov.; male from 12 km N. of Lai Chau (CL 4410), female from Nam Ceung stream (CL 4409). 9: M. vietnamensis, paratype, from Vietnam, Da Nang Prov., Ba Na - Nui Chua (TAD0334) (9 reproduced and modified from Tran &amp; Zettel, 2005). 10, 11: M. quynhi, specimens from Vietnam, Lao Cai Prov., 18 km NW. of Sa Pa (CL 4400). 12, 13: M. obscurus, specimens from Vietnam, Lao Cai Prov., 4 km NE. of Sa Pa (CL 4407). 14, 15: M. acutus, specimens from Vietnam, Hai Hung Prov., 22 km N. of Chi Linh (CL 4375). 16, 17: M. nigrofascioides, specimens from Vietnam, Binh Dinh Prov., An Khe Pass (CL 4291)." figureDoi="http://doi.org/10.5281/zenodo.4502640" httpUri="https://zenodo.org/record/4502640/files/figure.png" pageId="30" pageNumber="139">Fig. 8</figureCitation>
). This species can also be distinguished from all other species of the
<emphasis id="B935EAF137783F1BFADCFE5297B43581" box="[1353,1474,478,501]" italics="true" pageId="30" pageNumber="139">M. compar</emphasis>
group by the combination of following characters: apex of paramere rounded and not modified; endosoma with the apical sclerite more developed, lateral sclerite indistinct, ventral sclerite shorter; median lobe of sternum 7 of female with rounded apex; hind coxa of female with less dense pilosity and with the apex not produced.
</paragraph>
<paragraph id="8BFE36E337783F1BFCD6FD5197B737E1" blockId="30.[834,1475,733,917]" pageId="30" pageNumber="139">
The discoveries of
<taxonomicName id="4C414D6037783F1BFB8EFD5296B63681" authorityName="A.D. &amp; Polhemus" authorityYear="2017" box="[1051,1216,734,757]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="B935EAF137783F1BFB8EFD5296B63681" box="[1051,1216,734,757]" italics="true" pageId="30" pageNumber="139">M. nigriventris</emphasis>
</taxonomicName>
,
<taxonomicName id="4C414D6037783F1BFB44FD5297123681" authorityName="A.D. &amp; Polhemus" authorityYear="2017" box="[1233,1380,733,757]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B935EAF137783F1BFB44FD5297123681" box="[1233,1380,733,757]" italics="true" pageId="30" pageNumber="139">M. monticola</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C414D6037783F1BFA30FD5291013761" authorityName="A.D. &amp; Polhemus" authorityYear="2017" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="sapa">
<emphasis id="B935EAF137783F1BFA30FD5291013761" italics="true" pageId="30" pageNumber="139">M. sapa</emphasis>
</taxonomicName>
have raised the total number of species in the
<emphasis id="B935EAF137783F1BFA30FD7291E23741" italics="true" pageId="30" pageNumber="139">M. compar</emphasis>
group to nine, and they are also the first records of this group in
<collectingCountry id="F356767337783F1BFC5BFCB1965B3721" box="[974,1069,829,853]" name="Vietnam" pageId="30" pageNumber="139">Vietnam</collectingCountry>
. Other previously known species have been found in
<collectingCountry id="F356767337783F1BFC42FCD196663701" box="[983,1040,861,885]" name="India" pageId="30" pageNumber="139">India</collectingCountry>
,
<collectingCountry id="F356767337783F1BFB8FFCD1962A3701" box="[1050,1116,861,885]" name="Nepal" pageId="30" pageNumber="139">Nepal</collectingCountry>
,
<collectingCountry id="F356767337783F1BFBF3FCD296A63702" box="[1126,1232,862,886]" name="Myanmar" pageId="30" pageNumber="139">Myanmar</collectingCountry>
,
<collectingCountry id="F356767337783F1BFB4CFCD1976D3701" box="[1241,1307,861,885]" name="China" pageId="30" pageNumber="139">China</collectingCountry>
, and Peninsular
<collectingCountry id="F356767337783F1BFCD7FCF191D237E1" box="[834,932,893,917]" name="Malaysia" pageId="30" pageNumber="139">Malaysia</collectingCountry>
(
<bibRefCitation id="EFD04B1237783F1BFC27FCF196EC37E1" author="Chen PP &amp; Nieser N" box="[946,1178,893,917]" pageId="30" pageNumber="139" pagination="45 - 82" refId="ref41516" refString="Chen PP &amp; Nieser N (1993) A taxonomic revision of the Oriental water strider genus Metrocoris Mayr (Hemiptera, Gerridae). Part I and Part II. Steenstrupia, 19 (1 - 2): 1 - 43, 45 - 82." type="journal article" year="1993">Chen &amp; Nieser, 1993</bibRefCitation>
;
<bibRefCitation id="EFD04B1237783F1BFB33FCF1975337E1" author="Chen PP" box="[1190,1317,893,917]" pageId="30" pageNumber="139" pagination="124 - 134" refId="ref41564" refString="Chen PP (1994) An overview of Chinese Metrocoris Mayr with description of three new species (Hemiptera: Gerridae). Entomologia Sinica, 1 (2): 124 - 134." type="journal article" year="1994">Chen, 1994</bibRefCitation>
;
<bibRefCitation id="EFD04B1237783F1BFAA5FCF197C037E1" author="Zettel H" box="[1328,1462,893,917]" pageId="30" pageNumber="139" pagination="109 - 114" refId="ref42232" refString="Zettel H (2011) Metrocoris pardus sp. n. (Heteroptera: Gerridae) aus Westmalaysien. Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 63 (2): 109 - 114." type="journal article" year="2011">Zettel, 2011</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B656837783F1BFCD6FC3191A13061" pageId="30" pageNumber="139" type="etymology">
<paragraph id="8BFE36E337783F1BFCD6FC3191A13061" blockId="30.[835,1474,957,1045]" pageId="30" pageNumber="139">
<emphasis id="B935EAF137783F1BFCD6FC3191B137A1" bold="true" box="[835,967,957,981]" pageId="30" pageNumber="139">Etymology.</emphasis>
This species is named after its
<typeStatus id="54FA884137783F1BFAA4FC32972937A2" box="[1329,1375,958,982]" pageId="30" pageNumber="139">type</typeStatus>
locality, Sa Pa. The word “
<taxonomicName id="4C414D6037783F1BFB9EFC52964B3781" authorityName="A.D. &amp; Polhemus" authorityYear="2017" box="[1035,1085,990,1013]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="sapa">
<emphasis id="B935EAF137783F1BFB9EFC52964B3781" box="[1035,1085,990,1013]" italics="true" pageId="30" pageNumber="139">sapa</emphasis>
</taxonomicName>
” is used here specifically as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C35B656837783F1BFCD6FBB197B53141" pageId="30" pageNumber="139" type="distribution">
<paragraph id="8BFE36E337783F1BFCD6FBB1976B3021" blockId="30.[835,1309,1085,1109]" box="[835,1309,1085,1109]" pageId="30" pageNumber="139">
<emphasis id="B935EAF137783F1BFCD6FBB191A03021" bold="true" box="[835,982,1085,1109]" pageId="30" pageNumber="139">Distribution.</emphasis>
<collectingCountry id="F356767337783F1BFC4BFBB196483021" box="[990,1086,1085,1109]" name="Vietnam" pageId="30" pageNumber="139">Vietnam</collectingCountry>
:
<collectingRegion id="4985F80137783F1BFBDFFBB296D73021" box="[1098,1185,1085,1109]" country="Vietnam" name="Lao Cai" pageId="30" pageNumber="139">Lao Cai</collectingRegion>
(
<figureCitation id="137A2A6637783F1BFB27FBB197663021" box="[1202,1296,1085,1109]" captionStart="Fig" captionStartId="39.[113,148,2051,2073]" captionTargetBox="[113,1451,199,2036]" captionTargetId="figure-8@39.[113,1451,196,2036]" captionTargetPageId="39" captionText="Fig. 141. Distribution records in Vietnam of Metrocoris nigriventris, new species, M. monticola, new species, M. sapa, new species, and M. tenuicornis." figureDoi="http://doi.org/10.5281/zenodo.4502686" httpUri="https://zenodo.org/record/4502686/files/figure.png" pageId="30" pageNumber="139">Fig. 141</figureCitation>
).
</paragraph>
<paragraph id="8BFE36E337783F1BFCD6FBF197B53141" blockId="30.[834,1475,1149,1333]" pageId="30" pageNumber="139">
<emphasis id="B935EAF137783F1BFCD6FBF191DA30E1" bold="true" box="[835,940,1149,1173]" pageId="30" pageNumber="139">Habitats.</emphasis>
<taxonomicName id="4C414D6037783F1BFC23FBF2961C30E1" authorityName="A.D. &amp; Polhemus" authorityYear="2017" box="[950,1130,1150,1173]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="sapa">
<emphasis id="B935EAF137783F1BFC23FBF2961C30E1" box="[950,1130,1150,1173]" italics="true" pageId="30" pageNumber="139">Metrocoris sapa</emphasis>
</taxonomicName>
was collected from a montane stream, at elevations between 1800 and
<quantity id="4CB99B0637783F1BFB71FB11974130C1" box="[1252,1335,1181,1205]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="30" pageNumber="139" unit="m" value="1900.0">1900 m</quantity>
. It was found in association with
<taxonomicName id="4C414D6037783F1BFB81FB3296F730A1" authorityName="Tran &amp; Zettel" authorityYear="2005" box="[1044,1153,1213,1237]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="quynhi">
<emphasis id="B935EAF137783F1BFB81FB3296F730A1" box="[1044,1153,1213,1237]" italics="true" pageId="30" pageNumber="139">M. quynhi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C414D6037783F1BFB23FB32976930A1" authorityName="Chen &amp; Nieser" authorityYear="1993" box="[1206,1311,1214,1237]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="species" species="acutus">
<emphasis id="B935EAF137783F1BFB23FB3296BA30A1" box="[1206,1228,1214,1237]" italics="true" pageId="30" pageNumber="139">M</emphasis>
.
<emphasis id="B935EAF137783F1BFB4CFB32976930A1" box="[1241,1311,1214,1237]" italics="true" pageId="30" pageNumber="139">acutus</emphasis>
</taxonomicName>
, both of which are much larger species. Due to the small size and pale dorsal colour of living specimens in this new species, it may superficially resemble nymphs of other
<taxonomicName id="4C414D6037783F1BFB65FA9297113141" box="[1264,1383,1310,1333]" class="Insecta" family="Gerridae" genus="Metrocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAF137783F1BFB65FA9297113141" box="[1264,1383,1310,1333]" italics="true" pageId="30" pageNumber="139">Metrocoris</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>
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