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<document id="71F65FA6CFB221D2AB57DD0B67F4474C" ID-DOI="10.11646/zootaxa.5414.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10681263" ID-ZooBank="3B1DFADA-C20E-473B-A5E9-843548B54AD3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708377054888" checkinUser="plazi" docAuthor="Paredes-Munguía, Williams, Brescovit, Antonio D. &amp; Teixeira, Renato A." docDate="2024" docId="03E887D2BA35FE16FDDBFDE560D6FDCB" docLanguage="en" docName="zootaxa.5414.1.1.pdf" docOrigin="Zootaxa 5414 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5414.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Hogna andina Paredes-Munguía &amp; Brescovit &amp; Teixeira 2024, comb. nov." docType="treatment" docVersion="1" lastPageNumber="59" masterDocId="FFD1FFAABA0DFE2CFD4CFF8E6559FFD2" masterDocTitle="Revision of Neotropical wolf spider genus Arctosa C. L. Koch, 1847 (Araneae: Lycosidae), with description of seven new species" masterLastPageNumber="83" masterPageNumber="1" pageNumber="57" updateTime="1708377875290" updateUser="ExternalLinkService">
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<mods:title id="60AEBCF8A09283BADDF65BE046D8CE3D">Revision of Neotropical wolf spider genus Arctosa C. L. Koch, 1847 (Araneae: Lycosidae), with description of seven new species</mods:title>
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<mods:namePart id="EABD5C0308192A1958D51C5BFEEFCF60">Paredes-Munguía, Williams</mods:namePart>
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<mods:affiliation id="E324C43721C85719E1CB4401A0A24951">Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Laboratorio de Entomología, Av. Arenales 1256, Lima 14, Peru. &amp; williams. paredes @ acad. pucrs. br; https: // orcid. org / 0000 - 0002 - 3320 - 4611</mods:affiliation>
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<mods:namePart id="CDDBC7591A1522D89852645E5C17EA54">Brescovit, Antonio D.</mods:namePart>
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<mods:affiliation id="7A311961B74BC6D32E1DDB7C9E21D4F5">Laboratório de Coleções Zoológicas, Av. Vital Brasil, 1500, Butantã, 05503 - 900, São Paulo, SP, Brazil. &amp; antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324</mods:affiliation>
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<mods:namePart id="034D54735AFCFF812EF6550E169E157A">Teixeira, Renato A.</mods:namePart>
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<mods:affiliation id="E80933E7389CA37D01BE3FCDCE5A3D92">Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; renatoaug. tx @ gmail. com; https: // orcid. org / 0000 - 0002 - 1756 - 9821</mods:affiliation>
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<mods:date id="B5C1DE6E0A73DA0F61A60F05FC2A63DF">2024</mods:date>
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<heading id="D0B681A8BA35FE14FDDBFDE56789FD56" bold="true" box="[151,720,618,645]" fontSize="11" level="2" pageId="56" pageNumber="57" reason="4">
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<taxonomicName id="4C414D47BA35FE14FDDBFDE5671BFD56" authority="(Chamberlin, 1916)" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" baseAuthorityName="Chamberlin" baseAuthorityYear="1916" box="[151,578,618,645]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="andina" status="comb. nov.">
<emphasis id="B935EAD6BA35FE14FDDBFDE5641EFD57" bold="true" box="[151,327,618,645]" italics="true" pageId="56" pageNumber="57">Hogna andina</emphasis>
(
<bibRefCitation id="EFD04B35BA35FE14FC1AFDE46763FD56" author="Chamberlin, R. V." box="[342,570,618,644]" pageId="56" pageNumber="57" pagination="177 - 299" refId="ref48794" refString="Chamberlin, R. V. (1916) Results of the Yale Peruvian Expedition of 1911. The Arachnida. Bulletin of The Museum of Comparative Zoology, 60 (6), 177 - 299." type="journal article" year="1916">Chamberlin, 1916</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20657ADBA35FE14FF06FDE46789FD56" box="[586,720,618,644]" pageId="56" pageNumber="57" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C35B654FBA35FE14FDDBFD006467FD7A" box="[151,318,654,681]" pageId="56" pageNumber="57" type="description">
<paragraph id="8BFE36C4BA35FE14FDDBFD006467FD7A" blockId="56.[151,720,618,681]" box="[151,318,654,681]" pageId="56" pageNumber="57">
<figureCitation id="137A2A41BA35FE14FDDBFD00644EFD7B" box="[151,279,654,681]" captionStart-0="FIGURE 50" captionStart-1="FIGURE 51" captionStartId-0="56.[151,250,1914,1938]" captionStartId-1="57.[151,250,1560,1584]" captionTargetBox-0="[151,1436,844,1890]" captionTargetBox-1="[151,1436,181,1537]" captionTargetId-0="figure-187@56.[151,1436,832,1890]" captionTargetId-1="figure-21@57.[151,1436,181,1537]" captionTargetPageId-0="56" captionTargetPageId-1="57" captionText-0="FIGURE 50. Hogna andina Chamberlin, 1916 comb. nov. Male pedipalp, holotype ac (a, retrolateral; b, ventral; c, prolateral) Collecting and ID labels de (d, type locality label; e, ID label). Scale bars: ac, 1 mm. E, embolus, DMA depression of MA, MA median apophysis, P palea, SD seminal duct, T tegulum, TA terminal apophysis, TL tegular lobe, VP ventral process." captionText-1="FIGURE 51. Hogna andina Chamberlin, 1916 comb.nov. Female habitus, paratype ab, d (a, ventral; b, prosoma dorsal; d, opisthosoma dorsal) Epigynum c (c dorsal) Collecting and ID labels ef (e, type locality; f, ID label). Scale bars: a, f, 5 mm, b, 1 mm, c, 0.5 mm. CM cardiac mark, H hood, HS head of spermatheca, MS median septum, SS stalk of spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.10681367" figureDoi-1="http://doi.org/10.5281/zenodo.10681369" httpUri-0="https://zenodo.org/record/10681367/files/figure.png" httpUri-1="https://zenodo.org/record/10681369/files/figure.png" pageId="56" pageNumber="57">Figs 5051</figureCitation>
,
<figureCitation id="137A2A41BA35FE14FC6EFD006467FD7A" box="[290,318,654,680]" captionStart="FIGURE 62" captionStartId="72.[151,250,1618,1642]" captionTargetBox="[171,1418,564,1590]" captionTargetId="figure-197@72.[164,1423,561,1595]" captionTargetPageId="72" captionText="FIGURE 62. Geographic distribution records of Arctosa ayaymama sp. nov. (blue dot), Arctosa pacaya sp. nov. (red dot), Hogna andina comb. nov. (orange dot)." figureDoi="http://doi.org/10.5281/zenodo.10681391" httpUri="https://zenodo.org/record/10681391/files/figure.png" pageId="56" pageNumber="57">62</figureCitation>
</paragraph>
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<paragraph id="8BFE36C4BA35FE14FDDBFD5D6630FCDB" blockId="56.[151,1436,723,808]" pageId="56" pageNumber="57">
<treatmentCitationGroup id="AB5111EABA35FE14FDDBFD5D6630FCDB" pageId="56" pageNumber="57">
<treatmentCitation id="0AE010D5BA35FE14FDDBFD5D677EFD39" author="Chamberlin, R. V." box="[151,551,723,747]" page="286" pageId="56" pageNumber="57" year="1916">
<taxonomicName id="4C414D47BA35FE14FDDBFD5D677EFD39" authority="Chamberlin, 1916: 286" authorityName="Chamberlin" authorityPageNumber="286" authorityYear="1916" box="[151,551,723,747]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="B935EAD6BA35FE14FDDBFD5D6474FD39" box="[151,301,723,747]" italics="true" pageId="56" pageNumber="57">Lycosa andina</emphasis>
<materialsCitation id="3B293C99BA35FE14FC7AFD5D677EFD39" box="[310,551,723,747]" country="Peru" location="Tincochaca" pageId="56" pageNumber="57" specimenCount="2" stateProvince="Cusco" typeStatus="holotype">
<bibRefCitation id="EFD04B35BA35FE14FC7AFD5D677EFD39" author="Chamberlin, R. V." box="[310,551,723,747]" pageId="56" pageNumber="57" pagination="177 - 299" refId="ref48794" refString="Chamberlin, R. V. (1916) Results of the Yale Peruvian Expedition of 1911. The Arachnida. Bulletin of The Museum of Comparative Zoology, 60 (6), 177 - 299." type="journal article" year="1916">Chamberlin, 1916: 286</bibRefCitation>
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, pl. 24, figs 23. Male
<typeStatus id="54FA8866BA35FE14FE53FD5D6621FD39" box="[799,888,723,747]" pageId="56" pageNumber="57" type="holotype">holotype</typeStatus>
and female
<typeStatus id="54FA8866BA35FE14FEB7FD5D610BFD39" box="[1019,1106,723,747]" pageId="56" pageNumber="57" type="paratype">paratype</typeStatus>
from
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,
<collectingRegion id="4985F826BA35FE14F854FD5D6002FD39" box="[1304,1371,723,747]" country="Peru" name="Cusco" pageId="56" pageNumber="57">Cuzco</collectingRegion>
,
<collectingCountry id="F3567654BA35FE14F82BFD5D60C1FD38" box="[1383,1432,723,746]" name="Peru" pageId="56" pageNumber="57">Peru</collectingCountry>
, deposited in MCZ 285 and 286, respectively, examined by photos.
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</paragraph>
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<treatmentCitationGroup id="AB5111EABA35FE14FDDBFC9E6758FCFA" box="[151,513,784,808]" pageId="56" pageNumber="57">
<taxonomicName id="4C414D47BA35FE14FDDBFC9E6468FCFA" ID-CoL="GBDM" baseAuthorityName="Chamberlin" baseAuthorityYear="1916" box="[151,305,784,808]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="B935EAD6BA35FE14FDDBFC9E6468FCFA" box="[151,305,784,808]" italics="true" pageId="56" pageNumber="57">Arctosa andina</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AE010D5BA35FE14FC73FC9E64A4FCFA" author="Roewer, V. C. F." box="[319,509,784,808]" page="230" pageId="56" pageNumber="57" year="1955">
<bibRefCitation id="EFD04B35BA35FE14FC73FC9E64A4FCFA" author="Roewer, V. C. F." box="[319,509,784,808]" pageId="56" pageNumber="57" refId="ref51988" refString="Roewer, V. C. F. (1955) Katalog der Araneae von 1758 bis 1940, bzw. 1954. 2. Band, Abt. a (Lycosaeformia, Dionycha [excl. Salticiformia]). 2. Band, Abt. b (Salticiformia, Cribellata) (Synonyma-Verzeichnis, Gesamtindex). Institut Royal des Sciences Naturelles de Belgique, Bruxelles, 1751 pp." type="book" year="1955">Roewer 1955: 230</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BFE36C4BA35FE14FDDBF8F4602EF80B" blockId="56.[151,1437,1914,2010]" pageId="56" pageNumber="57">
<emphasis id="B935EAD6BA35FE14FDDBF8F46446F840" bold="true" box="[151,287,1914,1938]" pageId="56" pageNumber="57">FIGURE 50.</emphasis>
<taxonomicName id="4C414D47BA35FE14FC69F8F56728F840" authority="Chamberlin, 1916" authorityName="Chamberlin" authorityYear="1916" baseAuthorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" baseAuthorityYear="2024" box="[293,625,1914,1938]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="andina" status="comb. nov.">
<emphasis id="B935EAD6BA35FE14FC69F8F564ECF840" box="[293,437,1914,1938]" italics="true" pageId="56" pageNumber="57">Hogna andina</emphasis>
<bibRefCitation id="EFD04B35BA35FE14FCF7F8F46728F840" author="Chamberlin, R. V." box="[443,625,1914,1938]" pageId="56" pageNumber="57" pagination="177 - 299" refId="ref48794" refString="Chamberlin, R. V. (1916) Results of the Yale Peruvian Expedition of 1911. The Arachnida. Bulletin of The Museum of Comparative Zoology, 60 (6), 177 - 299." type="journal article" year="1916">Chamberlin, 1916</bibRefCitation>
</taxonomicName>
<emphasis id="B935EAD6BA35FE14FF3BF8F467B3F840" bold="true" box="[631,746,1914,1938]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="A20657ADBA35FE14FF3BF8F467B3F840" box="[631,746,1914,1938]" pageId="56" pageNumber="57" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
Male pedipalp, holotype
<emphasis id="B935EAD6BA35FE14FEA7F8F46156F840" bold="true" box="[1003,1039,1914,1938]" pageId="56" pageNumber="57">ac</emphasis>
(a, retrolateral; b, ventral; c, prolateral) Collecting and ID labels
<emphasis id="B935EAD6BA35FE14FCDBF81064E4F864" bold="true" box="[407,445,1950,1974]" pageId="56" pageNumber="57">de</emphasis>
(d, type locality label; e, ID label). Scale bars: ac, 1 mm.
<emphasis id="B935EAD6BA35FE14F955F8106173F864" bold="true" box="[1049,1066,1950,1974]" pageId="56" pageNumber="57">E</emphasis>
, embolus,
<emphasis id="B935EAD6BA35FE14F9D7F810618EF864" bold="true" box="[1179,1239,1950,1974]" pageId="56" pageNumber="57">DMA</emphasis>
depression of MA,
<emphasis id="B935EAD6BA35FE14FDDBF84C6598F808" bold="true" box="[151,193,1986,2010]" pageId="56" pageNumber="57">MA</emphasis>
median apophysis,
<emphasis id="B935EAD6BA35FE14FCC7F84C64C3F808" bold="true" box="[395,410,1986,2010]" pageId="56" pageNumber="57">P</emphasis>
palea,
<emphasis id="B935EAD6BA35FE14FCAFF84F675AF80B" bold="true" box="[483,515,1985,2009]" pageId="56" pageNumber="57">SD</emphasis>
seminal duct,
<emphasis id="B935EAD6BA35FE14FFDAF84C67FEF808" bold="true" box="[662,679,1986,2010]" pageId="56" pageNumber="57">T</emphasis>
tegulum,
<emphasis id="B935EAD6BA35FE14FE41F84C6677F808" bold="true" box="[781,814,1986,2010]" pageId="56" pageNumber="57">TA</emphasis>
terminal apophysis,
<emphasis id="B935EAD6BA35FE14F94EF84C617DF808" bold="true" box="[1026,1060,1986,2010]" pageId="56" pageNumber="57">TL</emphasis>
tegular lobe,
<emphasis id="B935EAD6BA35FE14F9E3F84C6188F808" bold="true" box="[1199,1233,1986,2010]" pageId="56" pageNumber="57">VP</emphasis>
ventral process.
</paragraph>
</caption>
<caption id="DF3E664CBA34FE15FDDBF9966064F9AB" ID-DOI="http://doi.org/10.5281/zenodo.10681369" ID-Zenodo-Dep="10681369" httpUri="https://zenodo.org/record/10681369/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,250,1560,1584]" targetBox="[151,1436,181,1537]" targetPageId="57" targetType="figure">
<paragraph id="8BFE36C4BA34FE15FDDBF9966064F9AB" blockId="57.[151,1436,1560,1657]" pageId="57" pageNumber="58">
<emphasis id="B935EAD6BA34FE15FDDBF9966478F9E2" bold="true" box="[151,289,1560,1584]" pageId="57" pageNumber="58">FIGURE 51.</emphasis>
<taxonomicName id="4C414D47BA34FE15FC65F9976725F9E3" authority="Chamberlin, 1916" authorityName="Chamberlin" authorityYear="1916" baseAuthorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" baseAuthorityYear="2024" box="[297,636,1561,1585]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="andina" status="comb. nov.">
<emphasis id="B935EAD6BA34FE15FC65F99764E2F9E3" box="[297,443,1561,1585]" italics="true" pageId="57" pageNumber="58">Hogna andina</emphasis>
<bibRefCitation id="EFD04B35BA34FE15FC8FF9976725F9E3" author="Chamberlin, R. V." box="[451,636,1561,1585]" pageId="57" pageNumber="58" pagination="177 - 299" refId="ref48794" refString="Chamberlin, R. V. (1916) Results of the Yale Peruvian Expedition of 1911. The Arachnida. Bulletin of The Museum of Comparative Zoology, 60 (6), 177 - 299." type="journal article" year="1916">Chamberlin, 1916</bibRefCitation>
</taxonomicName>
<emphasis id="B935EAD6BA34FE15FFCFF99767ABF9E3" bold="true" box="[643,754,1561,1585]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="A20657ADBA34FE15FFCFF99767ABF9E3" box="[643,754,1561,1585]" pageId="57" pageNumber="58" rank="species">comb.nov.</taxonomicNameLabel>
</emphasis>
Female habitus, paratype
<emphasis id="B935EAD6BA34FE15F94DF997611CF9E3" bold="true" box="[1025,1093,1561,1585]" pageId="57" pageNumber="58">ab, d</emphasis>
(a, ventral; b, prosoma dorsal; d, opisthosoma dorsal) Epigynum
<emphasis id="B935EAD6BA34FE15FC96F9B364BCF987" bold="true" box="[474,485,1597,1621]" pageId="57" pageNumber="58">c</emphasis>
(c dorsal) Collecting and ID labels
<emphasis id="B935EAD6BA34FE15FE1CF9B26628F986" bold="true" box="[848,881,1596,1620]" pageId="57" pageNumber="58">ef</emphasis>
(e, type locality; f, ID label). Scale bars: a, f, 5 mm, b, 1 mm, c, 0.5 mm.
<emphasis id="B935EAD6BA34FE15FC1CF9EE6423F9AA" bold="true" box="[336,378,1632,1656]" pageId="57" pageNumber="58">CM</emphasis>
cardiac mark,
<emphasis id="B935EAD6BA34FE15FF5DF9EF677CF9AB" bold="true" box="[529,549,1633,1657]" pageId="57" pageNumber="58">H</emphasis>
hood,
<emphasis id="B935EAD6BA34FE15FF26F9EE67D5F9AA" bold="true" box="[618,652,1632,1656]" pageId="57" pageNumber="58">HS</emphasis>
head of spermatheca,
<emphasis id="B935EAD6BA34FE15FE23F9EE66CCF9AA" bold="true" box="[879,917,1632,1656]" pageId="57" pageNumber="58">MS</emphasis>
median septum,
<emphasis id="B935EAD6BA34FE15F90EF9EE6107F9AA" bold="true" box="[1090,1118,1632,1656]" pageId="57" pageNumber="58">SS</emphasis>
stalk of spermatheca.
</paragraph>
</caption>
<subSubSection id="C35B654FBA34FE16FDDBF92961CAFE5B" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="discussion">
<paragraph id="8BFE36C4BA34FE16FDDBF92961CAFE5B" blockId="57.[151,1437,1703,2018]" lastBlockId="58.[151,1437,151,537]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
<emphasis id="B935EAD6BA34FE15FDDBF9296452F913" bold="true" box="[151,267,1703,1729]" pageId="57" pageNumber="58">Remarks.</emphasis>
The male and female description by
<bibRefCitation id="EFD04B35BA34FE15FFFDF92966D1F913" author="Chamberlin, R. V." box="[689,904,1703,1729]" pageId="57" pageNumber="58" pagination="177 - 299" refId="ref48794" refString="Chamberlin, R. V. (1916) Results of the Yale Peruvian Expedition of 1911. The Arachnida. Bulletin of The Museum of Comparative Zoology, 60 (6), 177 - 299." type="journal article" year="1916">Chamberlin (1916)</bibRefCitation>
presented detailed information on morphology for both sexes.
<typeStatus id="54FA8866BA34FE15FC03F94564EEF937" box="[335,439,1739,1765]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
photos confronted with drawings of both epigynum and pedipalp (
<bibRefCitation id="EFD04B35BA34FE15F987F945659DF8DB" author="Chamberlin, R. V." pageId="57" pageNumber="58" pagination="177 - 299" refId="ref48794" refString="Chamberlin, R. V. (1916) Results of the Yale Peruvian Expedition of 1911. The Arachnida. Bulletin of The Museum of Comparative Zoology, 60 (6), 177 - 299." type="journal article" year="1916">Chamberlin 1916: 286</bibRefCitation>
, pl. 24, figs 23) suggest transferring to
<taxonomicName id="4C414D47BA34FE15FFD5F97E67BCF8DB" authorityName="Simon" authorityYear="1885" box="[665,741,1776,1801]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA34FE15FFD5F97E67BCF8DB" box="[665,741,1776,1801]" italics="true" pageId="57" pageNumber="58">Hogna</emphasis>
</taxonomicName>
. The male pedipalp has a rounded palea and a sickle shaped terminal apophysis (
<figureCitation id="137A2A41BA34FE15FC33F89D64BAF8FC" box="[383,483,1811,1838]" captionStart="FIGURE 50" captionStartId="56.[151,250,1914,1938]" captionTargetBox="[151,1436,844,1890]" captionTargetId="figure-187@56.[151,1436,832,1890]" captionTargetPageId="56" captionText="FIGURE 50. Hogna andina Chamberlin, 1916 comb. nov. Male pedipalp, holotype ac (a, retrolateral; b, ventral; c, prolateral) Collecting and ID labels de (d, type locality label; e, ID label). Scale bars: ac, 1 mm. E, embolus, DMA depression of MA, MA median apophysis, P palea, SD seminal duct, T tegulum, TA terminal apophysis, TL tegular lobe, VP ventral process." figureDoi="http://doi.org/10.5281/zenodo.10681367" httpUri="https://zenodo.org/record/10681367/files/figure.png" pageId="57" pageNumber="58">Fig. 50b</figureCitation>
), which is diagnostic for the majority of
<taxonomicName id="4C414D47BA34FE15FEF7F89A6138F8FF" box="[955,1121,1811,1837]" pageId="57" pageNumber="58">
<emphasis id="B935EAD6BA34FE15FEF7F89A615EF8FF" box="[955,1031,1812,1837]" italics="true" pageId="57" pageNumber="58">Hogna</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="EFD04B35BA34FE15F93EF89D60C9F8FF" author="Dondale, C. D. &amp; Redner, J. H." box="[1138,1424,1811,1837]" pageId="57" pageNumber="58" pagination="1 - 383" refId="ref49112" refString="Dondale, C. D. &amp; Redner, J. H. (1990) The insects and arachnids of Canada, Part 17. The wolf spiders, nurseryweb spiders, and lynx spiders of Canada and Alaska, Araneae: Lycosidae, Pisauridae, and Oxyopidae. Research Branch Agriculture Canada Publication, 1856, 1 - 383." type="journal article" year="1990">Dondale &amp; Redner 1990</bibRefCitation>
). The epigynum presents an inverted T-shaped pattern and a smooth atrium (
<figureCitation id="137A2A41BA34FE15F94EF8B9613FF880" box="[1026,1126,1847,1874]" captionStart="FIGURE 51" captionStartId="57.[151,250,1560,1584]" captionTargetBox="[151,1436,181,1537]" captionTargetId="figure-21@57.[151,1436,181,1537]" captionTargetPageId="57" captionText="FIGURE 51. Hogna andina Chamberlin, 1916 comb.nov. Female habitus, paratype ab, d (a, ventral; b, prosoma dorsal; d, opisthosoma dorsal) Epigynum c (c dorsal) Collecting and ID labels ef (e, type locality; f, ID label). Scale bars: a, f, 5 mm, b, 1 mm, c, 0.5 mm. CM cardiac mark, H hood, HS head of spermatheca, MS median septum, SS stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.10681369" httpUri="https://zenodo.org/record/10681369/files/figure.png" pageId="57" pageNumber="58">Fig. 51c</figureCitation>
). Different from
<taxonomicName id="4C414D47BA34FE15F862F8B960CCF883" authorityName="C. L. Koch" authorityYear="1847" box="[1326,1429,1847,1873]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA34FE15F862F8B960CCF883" box="[1326,1429,1847,1873]" italics="true" pageId="57" pageNumber="58">Trochosa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C414D47BA34FE15FDDBF8D265BAF8A7" authorityName="Simon" authorityYear="1885" box="[151,227,1884,1909]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA34FE15FDDBF8D265BAF8A7" box="[151,227,1884,1909]" italics="true" pageId="57" pageNumber="58">Hogna</emphasis>
</taxonomicName>
median septum is longer than wide, the transversal septum is narrow and about half the length of the median septum (
<bibRefCitation id="EFD04B35BA34FE15FDBAF8F16754F84B" author="Dondale, C. D. &amp; Redner, J. H." box="[246,525,1919,1945]" pageId="57" pageNumber="58" pagination="1 - 383" refId="ref49112" refString="Dondale, C. D. &amp; Redner, J. H. (1990) The insects and arachnids of Canada, Part 17. The wolf spiders, nurseryweb spiders, and lynx spiders of Canada and Alaska, Araneae: Lycosidae, Pisauridae, and Oxyopidae. Research Branch Agriculture Canada Publication, 1856, 1 - 383." type="journal article" year="1990">Dondale &amp; Redner 1990</bibRefCitation>
). Additionally,
<taxonomicName id="4C414D47BA34FE15FFFBF80E667FF84B" authorityName="Chamberlin" authorityYear="1916" baseAuthorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" baseAuthorityYear="2024" box="[695,806,1919,1945]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="B935EAD6BA34FE15FFFBF80E667FF84B" box="[695,806,1919,1945]" italics="true" pageId="57" pageNumber="58">H. andina</emphasis>
</taxonomicName>
hoods are parallel (
<figureCitation id="137A2A41BA34FE15F94DF8F16139F848" box="[1025,1120,1919,1946]" captionStart="FIGURE 51" captionStartId="57.[151,250,1560,1584]" captionTargetBox="[151,1436,181,1537]" captionTargetId="figure-21@57.[151,1436,181,1537]" captionTargetPageId="57" captionText="FIGURE 51. Hogna andina Chamberlin, 1916 comb.nov. Female habitus, paratype ab, d (a, ventral; b, prosoma dorsal; d, opisthosoma dorsal) Epigynum c (c dorsal) Collecting and ID labels ef (e, type locality; f, ID label). Scale bars: a, f, 5 mm, b, 1 mm, c, 0.5 mm. CM cardiac mark, H hood, HS head of spermatheca, MS median septum, SS stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.10681369" httpUri="https://zenodo.org/record/10681369/files/figure.png" pageId="57" pageNumber="58">Fig. 51c</figureCitation>
), different from
<taxonomicName id="4C414D47BA34FE15F85BF8F16027F84B" authorityName="C. L. Koch" authorityYear="1847" box="[1303,1406,1919,1945]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA34FE15F85BF8F16027F84B" box="[1303,1406,1919,1945]" italics="true" pageId="57" pageNumber="58">Trochosa</emphasis>
</taxonomicName>
or
<taxonomicName id="4C414D47BA34FE15FDDBF82A672DF86F" box="[151,628,1955,1981]" pageId="57" pageNumber="58">
<emphasis id="B935EAD6BA34FE15FDDBF82A65A6F86F" box="[151,255,1956,1981]" italics="true" pageId="57" pageNumber="58">Varacosa</emphasis>
<bibRefCitation id="EFD04B35BA34FE15FC49F82D6742F86F" author="Chamberlin, R. V. &amp; Ivie, W." box="[261,539,1955,1981]" pageId="57" pageNumber="58" pagination="1 - 117" refId="ref48869" refString="Chamberlin, R. V. &amp; Ivie, W. (1942) A hundred new species of American spiders. Bulletin of the University of Utah, 32 (13), 1 - 117." type="journal article" year="1942">Chamberlin &amp; Ivie, 1942</bibRefCitation>
species
</taxonomicName>
, which have curved hoods. The two parallel bands inside the longitudinal median band are absent in both sexes (
<figureCitation id="137A2A41BA34FE15FF01F84967E9F830" box="[589,688,1991,2018]" captionStart="FIGURE 51" captionStartId="57.[151,250,1560,1584]" captionTargetBox="[151,1436,181,1537]" captionTargetId="figure-21@57.[151,1436,181,1537]" captionTargetPageId="57" captionText="FIGURE 51. Hogna andina Chamberlin, 1916 comb.nov. Female habitus, paratype ab, d (a, ventral; b, prosoma dorsal; d, opisthosoma dorsal) Epigynum c (c dorsal) Collecting and ID labels ef (e, type locality; f, ID label). Scale bars: a, f, 5 mm, b, 1 mm, c, 0.5 mm. CM cardiac mark, H hood, HS head of spermatheca, MS median septum, SS stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.10681369" httpUri="https://zenodo.org/record/10681369/files/figure.png" pageId="57" pageNumber="58">Fig. 51b</figureCitation>
), which also excludes placing this species in
<taxonomicName id="4C414D47BA34FE15F9FCF849604EF833" authorityName="C. L. Koch" authorityYear="1847" box="[1200,1303,1991,2017]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA34FE15F9FCF849604EF833" box="[1200,1303,1991,2017]" italics="true" pageId="57" pageNumber="58">Trochosa</emphasis>
</taxonomicName>
. The hoods in
<taxonomicName id="4C414D47BA37FE16FDFAFF196754FF63" authority="Mello-Leitao, 1941" authorityName="Mello-Leitao" authorityYear="1941" box="[182,525,151,177]" class="Arachnida" family="Lycosidae" genus="Birabenia" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA37FE16FDFAFF19647FFF63" box="[182,294,151,177]" italics="true" pageId="58" pageNumber="59">Birabenia</emphasis>
Mello-Leitão, 1941
</taxonomicName>
, also called anterior pockets (
<bibRefCitation id="EFD04B35BA37FE16FE2FFF1961C7FF63" author="Piacentini, L. N. &amp; Laborda, A." box="[867,1182,151,177]" pageId="58" pageNumber="59" pagination="39 - 52" refId="ref51621" refString="Piacentini, L. N. &amp; Laborda, A. (2013) The South American wolf spider genus Birabenia Mello-Leitao, 1941 (Araneae: Lycosidae: Lycosinae). Zootaxa, 3716 (1), 39 - 52. https: // doi. org / 10.11646 / zootaxa. 3716.1.3" type="journal article" year="2013">Piacentini &amp; Laborda 2013</bibRefCitation>
) are deep, as opposed to the shallow ones in
<taxonomicName id="4C414D47BA37FE16FCEBFF3564AAFF06" authorityName="Simon" authorityYear="1885" box="[423,499,187,212]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA37FE16FCEBFF3564AAFF06" box="[423,499,187,212]" italics="true" pageId="58" pageNumber="59">Hogna</emphasis>
</taxonomicName>
females. Likewise, the long legs and thick body (
<figureCitation id="137A2A41BA37FE16F908FF35619FFF07" box="[1092,1222,186,213]" captionStart="FIGURE 51" captionStartId="57.[151,250,1560,1584]" captionTargetBox="[151,1436,181,1537]" captionTargetId="figure-21@57.[151,1436,181,1537]" captionTargetPageId="57" captionText="FIGURE 51. Hogna andina Chamberlin, 1916 comb.nov. Female habitus, paratype ab, d (a, ventral; b, prosoma dorsal; d, opisthosoma dorsal) Epigynum c (c dorsal) Collecting and ID labels ef (e, type locality; f, ID label). Scale bars: a, f, 5 mm, b, 1 mm, c, 0.5 mm. CM cardiac mark, H hood, HS head of spermatheca, MS median septum, SS stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.10681369" httpUri="https://zenodo.org/record/10681369/files/figure.png" pageId="58" pageNumber="59">Fig. 51a, d</figureCitation>
) are diagnostic of
<taxonomicName id="4C414D47BA37FE16FDDBFF5164A7FF2B" authority="Dondale &amp; Redner 1990" authorityName="Dondale &amp; Redner" authorityYear="1990" box="[151,510,223,249]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA37FE16FDDBFF5165BAFF2A" box="[151,227,223,248]" italics="true" pageId="58" pageNumber="59">Hogna</emphasis>
<bibRefCitation id="EFD04B35BA37FE16FDA6FF5164A7FF2B" author="Dondale, C. D. &amp; Redner, J. H." box="[234,510,223,249]" pageId="58" pageNumber="59" pagination="1 - 383" refId="ref49112" refString="Dondale, C. D. &amp; Redner, J. H. (1990) The insects and arachnids of Canada, Part 17. The wolf spiders, nurseryweb spiders, and lynx spiders of Canada and Alaska, Araneae: Lycosidae, Pisauridae, and Oxyopidae. Research Branch Agriculture Canada Publication, 1856, 1 - 383." type="journal article" year="1990">Dondale &amp; Redner 1990</bibRefCitation>
</taxonomicName>
), along with the three, instead of four retromarginal teeth of
<taxonomicName id="4C414D47BA37FE16F9D0FF516055FF2B" authorityName="Mello-Leitao" authorityYear="1941" box="[1180,1292,223,249]" class="Arachnida" family="Lycosidae" genus="Birabenia" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA37FE16F9D0FF516055FF2B" box="[1180,1292,223,249]" italics="true" pageId="58" pageNumber="59">Birabenia</emphasis>
</taxonomicName>
. Finally, this species should not be kept in
<taxonomicName id="4C414D47BA37FE16FCA4FE8D6767FECF" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[488,574,259,285]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA37FE16FCA4FE8D6767FECF" box="[488,574,259,285]" italics="true" pageId="58" pageNumber="59">Arctosa</emphasis>
</taxonomicName>
because the males do not have the reduced palea or TA in different degrees of sclerotization (
<bibRefCitation id="EFD04B35BA37FE16FC17FEA96726FE93" author="Dondale, C. D. &amp; Redner, J. H." box="[347,639,295,321]" pageId="58" pageNumber="59" pagination="1 - 30" refId="ref48997" refString="Dondale, C. D. &amp; Redner, J. H. (1983 a) Revision of the wolf spiders of the genus Arctosa C. L. Koch in North and Central America (Araneae: Lycosidae). The Journal of Arachnology, 11 (1), 1 - 30." type="journal article" year="1983">Dondale &amp; Redner 1983a</bibRefCitation>
), and females possess the inverted T-shape of the epigynal plate, which is triangular to trapezoidal in
<taxonomicName id="4C414D47BA37FE16FCABFEC56624FEB7" authority="(Dondale &amp; Redner 1983 a)" baseAuthorityName="Dondale &amp; Redner" baseAuthorityYear="1983" box="[487,893,331,357]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA37FE16FCABFEC56764FEB7" box="[487,573,331,357]" italics="true" pageId="58" pageNumber="59">Arctosa</emphasis>
(
<bibRefCitation id="EFD04B35BA37FE16FF00FEC5662DFEB7" author="Dondale, C. D. &amp; Redner, J. H." box="[588,884,331,357]" pageId="58" pageNumber="59" pagination="1 - 30" refId="ref48997" refString="Dondale, C. D. &amp; Redner, J. H. (1983 a) Revision of the wolf spiders of the genus Arctosa C. L. Koch in North and Central America (Araneae: Lycosidae). The Journal of Arachnology, 11 (1), 1 - 30." type="journal article" year="1983">Dondale &amp; Redner 1983a</bibRefCitation>
)
</taxonomicName>
. The synapomorphies of
<taxonomicName id="4C414D47BA37FE16F9D7FEC561BEFEB6" authorityName="Dondale &amp; Redner" authorityYear="1990" box="[1179,1255,331,356]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA37FE16F9D7FEC561BEFEB6" box="[1179,1255,331,356]" italics="true" pageId="58" pageNumber="59">Hogna</emphasis>
</taxonomicName>
used to remove species from this genus and be transferred to
<taxonomicName id="4C414D47BA37FE16FFDEFEE167BEFE5A" authorityName="Brady" authorityYear="2012" box="[658,743,367,392]" class="Arachnida" family="Lycosidae" genus="Tigrosa" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B935EAD6BA37FE16FFDEFEE167BEFE5A" box="[658,743,367,392]" italics="true" pageId="58" pageNumber="59">Tigrosa</emphasis>
</taxonomicName>
are summarized in
<bibRefCitation id="EFD04B35BA37FE16FE88FEE161D7FE5B" author="Brady, A. R." box="[964,1166,367,393]" pageId="58" pageNumber="59" pagination="182 - 208" refId="ref48506" refString="Brady, A. R. (2012) Nearctic species of the new genus Tigrosa (Araneae: Lycosidae). The Journal of Arachnology, 40 (2), 182 - 208. https: // doi. org / 10.1636 / k 11 - 77.1" type="journal article" year="2012">Brady (2012:182)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B654FBA37FE16FD8BFE1C6454FE27" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="8BFE36C4BA37FE16FD8BFE1C6454FE27" blockId="58.[151,1437,151,537]" pageId="58" pageNumber="59">
<emphasis id="B935EAD6BA37FE16FD8BFE1C641DFE7E" bold="true" box="[199,324,402,428]" pageId="58" pageNumber="59">Diagnosis:</emphasis>
Males of
<taxonomicName id="4C414D47BA37FE16FCF1FE1D6739FE7F" authorityName="Chamberlin" authorityYear="1916" baseAuthorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" baseAuthorityYear="2024" box="[445,608,403,429]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="B935EAD6BA37FE16FCF1FE1D6739FE7F" box="[445,608,403,429]" italics="true" pageId="58" pageNumber="59">Hogna andina</emphasis>
</taxonomicName>
can be recognized by the curved cymbium (
<figureCitation id="137A2A41BA37FE16F92BFE1D6190FE7F" box="[1127,1225,402,429]" captionStart="FIGURE 50" captionStartId="56.[151,250,1914,1938]" captionTargetBox="[151,1436,844,1890]" captionTargetId="figure-187@56.[151,1436,832,1890]" captionTargetPageId="56" captionText="FIGURE 50. Hogna andina Chamberlin, 1916 comb. nov. Male pedipalp, holotype ac (a, retrolateral; b, ventral; c, prolateral) Collecting and ID labels de (d, type locality label; e, ID label). Scale bars: ac, 1 mm. E, embolus, DMA depression of MA, MA median apophysis, P palea, SD seminal duct, T tegulum, TA terminal apophysis, TL tegular lobe, VP ventral process." figureDoi="http://doi.org/10.5281/zenodo.10681367" httpUri="https://zenodo.org/record/10681367/files/figure.png" pageId="58" pageNumber="59">Fig. 50b</figureCitation>
) and short ventral projection of median apophysis (
<figureCitation id="137A2A41BA37FE16FCB0FE396702FE03" box="[508,603,438,465]" captionStart="FIGURE 50" captionStartId="56.[151,250,1914,1938]" captionTargetBox="[151,1436,844,1890]" captionTargetId="figure-187@56.[151,1436,832,1890]" captionTargetPageId="56" captionText="FIGURE 50. Hogna andina Chamberlin, 1916 comb. nov. Male pedipalp, holotype ac (a, retrolateral; b, ventral; c, prolateral) Collecting and ID labels de (d, type locality label; e, ID label). Scale bars: ac, 1 mm. E, embolus, DMA depression of MA, MA median apophysis, P palea, SD seminal duct, T tegulum, TA terminal apophysis, TL tegular lobe, VP ventral process." figureDoi="http://doi.org/10.5281/zenodo.10681367" httpUri="https://zenodo.org/record/10681367/files/figure.png" pageId="58" pageNumber="59">Fig. 50c</figureCitation>
). Females can be recognized by the dilated and rounded head of spermatheca (
<figureCitation id="137A2A41BA37FE16FDD3FE556459FE27" box="[159,256,474,501]" captionStart="FIGURE 51" captionStartId="57.[151,250,1560,1584]" captionTargetBox="[151,1436,181,1537]" captionTargetId="figure-21@57.[151,1436,181,1537]" captionTargetPageId="57" captionText="FIGURE 51. Hogna andina Chamberlin, 1916 comb.nov. Female habitus, paratype ab, d (a, ventral; b, prosoma dorsal; d, opisthosoma dorsal) Epigynum c (c dorsal) Collecting and ID labels ef (e, type locality; f, ID label). Scale bars: a, f, 5 mm, b, 1 mm, c, 0.5 mm. CM cardiac mark, H hood, HS head of spermatheca, MS median septum, SS stalk of spermatheca." figureDoi="http://doi.org/10.5281/zenodo.10681369" httpUri="https://zenodo.org/record/10681369/files/figure.png" pageId="58" pageNumber="59">Fig. 51c</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B654FBA37FE16FD8BFE7060D6FDCB" box="[199,1423,510,537]" pageId="58" pageNumber="59" type="distribution">
<paragraph id="8BFE36C4BA37FE16FD8BFE7060D6FDCB" blockId="58.[151,1437,151,537]" box="[199,1423,510,537]" pageId="58" pageNumber="59">
<emphasis id="B935EAD6BA37FE16FD8BFE706438FDCA" bold="true" box="[199,353,510,536]" pageId="58" pageNumber="59">Distribution.</emphasis>
Known from four localities
<collectingRegion id="4985F826BA37FE16FFECFE7167B3FDCB" box="[672,746,511,537]" country="Peru" name="Cusco" pageId="58" pageNumber="59">Cuzco</collectingRegion>
,
<collectingCountry id="F3567654BA37FE16FFBAFE716670FDCB" box="[758,809,511,537]" name="Peru" pageId="58" pageNumber="59">Peru</collectingCountry>
including the
<typeStatus id="54FA8866BA37FE16FE86FE7166A3FDCB" box="[970,1018,511,537]" pageId="58" pageNumber="59">type</typeStatus>
locality (
<figureCitation id="137A2A41BA37FE16F928FE7161EEFDCB" box="[1124,1207,511,537]" captionStart="FIGURE 62" captionStartId="72.[151,250,1618,1642]" captionTargetBox="[171,1418,564,1590]" captionTargetId="figure-197@72.[164,1423,561,1595]" captionTargetPageId="72" captionText="FIGURE 62. Geographic distribution records of Arctosa ayaymama sp. nov. (blue dot), Arctosa pacaya sp. nov. (red dot), Hogna andina comb. nov. (orange dot)." figureDoi="http://doi.org/10.5281/zenodo.10681391" httpUri="https://zenodo.org/record/10681391/files/figure.png" pageId="58" pageNumber="59">Fig. 62</figureCitation>
,
<bibRefCitation id="EFD04B35BA37FE16F98FFE7160DEFDCB" author="Chamberlin, R. V." box="[1219,1415,511,537]" pageId="58" pageNumber="59" pagination="177 - 299" refId="ref48794" refString="Chamberlin, R. V. (1916) Results of the Yale Peruvian Expedition of 1911. The Arachnida. Bulletin of The Museum of Comparative Zoology, 60 (6), 177 - 299." type="journal article" year="1916">Chamberlin 1916</bibRefCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>