treatments-xml/data/03/E8/87/03E887C9FFB95527FF3BE263829EFA49.xml

<document id="73E891BB3DD94D229605CE7A754744EF" ID-DOI="10.11646/phytotaxa.675.2.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="14521820" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734561100275" checkinUser="felipe" docAuthor="Lu, Liang-Ting, Sugita, Ryosuke, Sato, Misaki, Tanaka, Kazuaki &amp; Kirschner, Roland" docDate="2024" docId="03E887C9FFB95527FF3BE263829EFA49" docLanguage="en" docName="phytotaxa.675.2.167-176.pdf" docOrigin="Phytotaxa 675 (2)" docSource="https://doi.org/10.11646/phytotaxa.675.2.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Spegazzinia cryptica Liang T. Lu, R. Sugita, Mis. Sato, Kaz. Tanaka &amp; R. Kirschner 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="173" masterDocId="FFD1FFB1FFBD5521FFB3E65F8621FFED" masterDocTitle="Spegazzinia cryptica sp. nov. (Didymosphaeriaceae) from the western Pacific monsoon region" masterLastPageNumber="176" masterPageNumber="167" pageNumber="171" updateTime="1734565901300" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9C76B13C2BB93FF65B4AAE4AF6C9400F">Spegazzinia cryptica sp. nov. (Didymosphaeriaceae) from the western Pacific monsoon region</mods:title>
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<mods:namePart id="34CCA605522AA01AB4B1A09B2C766FA8">Lu, Liang-Ting</mods:namePart>
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<mods:affiliation id="9A40BDD3D3D593C80922840E704813E3">School of Forestry and Resource Conservation, National Taiwan University, Taipei 10617, Taiwan</mods:affiliation>
<mods:nameIdentifier id="D79A39191FF364EEAD4CC41AE50E857E" type="email">r09625006@ntu.edu.tw</mods:nameIdentifier>
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<mods:namePart id="D486E000150B8B5970A114DE4E40110E">Sugita, Ryosuke</mods:namePart>
<mods:affiliation id="5033699F9597DAF5ABE3D46A91E90158">Faculty of Agriculture and Life Science, Hirosaki University, 3 Bunkyo-cho, Hirosaki, Aomori 036 - 8561, Japan</mods:affiliation>
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<mods:namePart id="0FB5FE9BDF0E7B944875445E5225A6E2">Sato, Misaki</mods:namePart>
<mods:affiliation id="F80CC79E9044E7594ECB86C68FEE32EB">Faculty of Agriculture and Life Science, Hirosaki University, 3 Bunkyo-cho, Hirosaki, Aomori 036 - 8561, Japan</mods:affiliation>
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<mods:namePart id="C0B2599F07908100E947A956F005AEA9">Tanaka, Kazuaki</mods:namePart>
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<mods:affiliation id="E452085752532588DA5684D4AFD5F2D6">Faculty of Agriculture and Life Science, Hirosaki University, 3 Bunkyo-cho, Hirosaki, Aomori 036 - 8561, Japan</mods:affiliation>
<mods:nameIdentifier id="3F76DC514D47532870D26044FB537361" type="email">k-tanaka@hirosaki-u.ac.jp</mods:nameIdentifier>
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<mods:namePart id="8863019511501B3DF7D2BF1C4AD491FE">Kirschner, Roland</mods:namePart>
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<mods:affiliation id="764EDC231408236045517451BF384265">School of Forestry and Resource Conservation, National Taiwan University, Taipei 10617, Taiwan</mods:affiliation>
<mods:nameIdentifier id="E94189AD36632D2C529FE1A3A0CB6972" type="email">kirschner@ntu.edu.tw</mods:nameIdentifier>
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<mods:title id="692E1CA82CE55BC45705D7057AAEC93C">Phytotaxa</mods:title>
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<mods:date id="E160C5E9A8214A0487A35456C1920C8A">2024</mods:date>
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<treatment id="03E887C9FFB95527FF3BE263829EFA49" LSID="urn:lsid:plazi:treatment:03E887C9FFB95527FF3BE263829EFA49" httpUri="http://treatment.plazi.org/id/03E887C9FFB95527FF3BE263829EFA49" lastPageId="6" lastPageNumber="173" pageId="4" pageNumber="171">
<subSubSection id="C35B6554FFB95525FF3BE263843DFB97" pageId="4" pageNumber="171" type="nomenclature">
<paragraph id="8BFE36DFFFB95525FF3BE2638337FBBB" blockId="4.[136,1302,1084,1111]" box="[136,1302,1084,1111]" pageId="4" pageNumber="171">
<heading id="D0B681B3FFB95525FF3BE2638337FBBB" box="[136,1302,1084,1111]" fontSize="11" level="2" pageId="4" pageNumber="171" reason="3">
<taxonomicName id="4C414D5CFFB95525FF3BE263821DFBBB" authority="Liang T. Lu, R. Sugita, Mis. Sato, Kaz. Tanaka &amp; R. Kirschner" authorityName="Liang T. Lu, R. Sugita, Mis. Sato, Kaz. Tanaka &amp; R. Kirschner" authorityYear="2024" box="[136,1084,1084,1111]" class="Dothideomycetes" family="Microthyriaceae" genus="Spegazzinia" kingdom="Fungi" order="Microthyriales" pageId="4" pageNumber="171" phylum="Ascomycota" rank="species" species="cryptica" status="sp. nov.">
<emphasis id="B935EACDFFB95525FF3BE2638755FBBB" bold="true" box="[136,372,1084,1110]" italics="true" pageId="4" pageNumber="171">Spegazzinia cryptica</emphasis>
Liang T. Lu, R. Sugita, Mis. Sato, Kaz. Tanaka &amp; R. Kirschner
</taxonomicName>
, 
<emphasis id="B935EACDFFB95525FBF4E26282BDFBBB" box="[1095,1180,1085,1110]" italics="true" pageId="4" pageNumber="171">
<taxonomicNameLabel id="A20657B6FFB95525FBF4E26282BDFBBB" box="[1095,1180,1085,1110]" pageId="4" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="137A2A5AFFB95525FB19E263832FFBBB" box="[1194,1294,1084,1110]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[136,229,2024,2046]" captionStartId-1="5.[136,229,1863,1885]" captionTargetBox-0="[191,1403,1292,1979]" captionTargetBox-1="[300,1286,190,1839]" captionTargetId-0="figure-74@4.[183,1403,1273,1996]" captionTargetId-1="figure-18@5.[300,1286,190,1839]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 3. Spegazzinia cryptica (LV0167). A. An α conidium arising from conidiogenous cell. B. Detached α conidium. C. Detached β conidium. D. Side view of β conidium. Scale bars: A, B = 10 μm, C, D = 5 μm." captionText-1="FIGURE 4. Spegazzinia cryptica. A. Colony on dead leaf. B. α and β conidia. C, D. Conidia arising from conidiogenous cells (arrows). E, F. Detached α conidia. G, H. Bleached α conidia. I. β conidium arising from conidiogenous cell (arrow). J–L. Detached β conidia. M. Side view of β conidium. N. Bleached β conidium. O. Upper view of PDA culture (3 weeks at 25 °C in the dark). P. Reverse view of culture. Material examined:A, C–E, H–J, M–P LV0167 (TNM). B, L. MAFF 243878. F, G, K. MAFF 244519. Scale bars:A = 0.5 mm, B–H = 10 µm, I–N = 5 µm." figureDoi-0="http://doi.org/10.5281/zenodo.14521826" figureDoi-1="http://doi.org/10.5281/zenodo.14521828" httpUri-0="https://zenodo.org/record/14521826/files/figure.png" httpUri-1="https://zenodo.org/record/14521828/files/figure.png" pageId="4" pageNumber="171">Figs 3–4</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BFE36DFFFB95525FF3BE23F843DFB97" blockId="4.[136,540,1120,1147]" box="[136,540,1120,1147]" pageId="4" pageNumber="171">Index Fungorum number: IF902147</paragraph>
</subSubSection>
<subSubSection id="C35B6554FFB95525FF3BE2F78710FB0B" pageId="4" pageNumber="171" type="etymology">
<paragraph id="8BFE36DFFFB95525FF3BE2F78710FB0B" blockId="4.[136,1451,1192,1254]" pageId="4" pageNumber="171">
<emphasis id="B935EACDFFB95525FF3BE2F7870FFB2F" bold="true" box="[136,302,1192,1218]" pageId="4" pageNumber="171">Etymology:—</emphasis>
Epithet refers to the cryptic species concept, because the species is morphologically indistinguishable from 
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<taxonomicName id="4C414D5CFFB95525FF76E293870DFB0B" box="[197,300,1228,1254]" class="Dothideomycetes" family="Microthyriaceae" genus="Spegazzinia" kingdom="Fungi" order="Microthyriales" pageId="4" pageNumber="171" phylum="Ascomycota" rank="species" species="musae">S. musae</taxonomicName>
.
</emphasis>
</paragraph>
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<paragraph id="8BFE36DFFFB95525FF3BE1B785A0F7CF" blockId="4.[136,1451,2024,2082]" pageId="4" pageNumber="171">
<emphasis id="B935EACDFFB95525FF3BE1B786DFF810" bold="true" box="[136,254,2024,2046]" pageId="4" pageNumber="171">FIGURE 3.</emphasis>
<taxonomicName id="4C414D5CFFB95525FEB7E1B687EDF813" authorityName="Liang T. Lu, R. Sugita, Mis. Sato, Kaz. Tanaka &amp; R. Kirschner" authorityYear="2024" box="[260,460,2025,2046]" class="Dothideomycetes" family="Microthyriaceae" genus="Spegazzinia" kingdom="Fungi" order="Microthyriales" pageId="4" pageNumber="171" phylum="Ascomycota" rank="species" species="cryptica">
<emphasis id="B935EACDFFB95525FEB7E1B687EDF813" box="[260,460,2025,2046]" italics="true" pageId="4" pageNumber="171">Spegazzinia cryptica</emphasis>
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(LV0167). A. An α conidium arising from conidiogenous cell. B. Detached α conidium. C. Detached β conidium. D. Side view of β conidium. Scale bars: A, B = 10 μm, C, D = 5 μm.
</paragraph>
</caption>
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<paragraph id="8BFE36DFFFB85524FF3BE118870CF800" blockId="5.[136,1452,1863,2029]" pageId="5" pageNumber="172">
<emphasis id="B935EACDFFB85524FF3BE11886DCF8B1" bold="true" box="[136,253,1863,1885]" pageId="5" pageNumber="172">FIGURE 4.</emphasis>
<taxonomicName id="4C414D5CFFB85524FEB1E11787E8F8B0" authorityName="Liang T. Lu, R. Sugita, Mis. Sato, Kaz. Tanaka &amp; R. Kirschner" authorityYear="2024" box="[258,457,1864,1885]" class="Dothideomycetes" family="Microthyriaceae" genus="Spegazzinia" kingdom="Fungi" order="Microthyriales" pageId="5" pageNumber="172" phylum="Ascomycota" rank="species" species="cryptica">
<emphasis id="B935EACDFFB85524FEB1E11787E8F8B0" box="[258,457,1864,1885]" italics="true" pageId="5" pageNumber="172">Spegazzinia cryptica</emphasis>
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. A. Colony on dead leaf. B. α and β conidia. C, D. Conidia arising from conidiogenous cells (arrows). E, F. Detached α conidia. G, H. Bleached α conidia. I. β conidium arising from conidiogenous cell (arrow). J–L. Detached β conidia. M. Side view of β conidium. N. Bleached β conidium. O. Upper view of PDA culture (3 weeks at 25 °C in the dark). P. Reverse view of culture. Material examined: A, C–E, H–J, M–P LV0167 (TNM). B, L. MAFF 243878. F, G, K. MAFF 244519. Scale bars: A = 0.5 mm, B–H = 10 µm, I–N = 5 µm.
</paragraph>
</caption>
<subSubSection id="C35B6554FFBB5527FF0EE6C1829EFA49" pageId="6" pageNumber="173" type="materials_examined">
<paragraph id="8BFE36DFFFBB5527FF0EE6C1820CFF55" blockId="6.[136,1452,158,1445]" box="[189,1069,158,185]" pageId="6" pageNumber="173">
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<emphasis id="B935EACDFFBB5527FF0EE6C1876EFF55" bold="true" box="[189,335,158,184]" pageId="6" pageNumber="173">
<typeStatus id="54FA887DFFBB5527FF0EE6C1870CFF55" box="[189,301,158,184]" pageId="6" pageNumber="173" type="holotype">Holotype</typeStatus>
:—
</emphasis>
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.- 
<collectionCode id="ED50AE1AFFBB5527FEC3E6C087A5FF54" box="[368,388,159,185]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="173">T</collectionCode>
. Lu, 
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.- 
<collectionCode id="ED50AE1AFFBB5527FE6DE6C187D5FF55" box="[478,500,158,184]" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="173" type="Herbarium">C</collectionCode>
. Chien, 
<collectionCode id="ED50AE1AFFBB5527FDE3E6C08447FF54" box="[592,614,159,185]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="173">R</collectionCode>
. Kirschner &amp; 
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.- 
<collectionCode id="ED50AE1AFFBB5527FC92E6C0851CFF54" box="[801,829,159,185]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="6" pageNumber="173" type="Herbarium">W</collectionCode>
. Yeh 
<specimenCode id="DBE79EA4FFBB5527FCC9E6C185F5FF55" box="[890,980,158,184]" collectionCode="LV" country="Belgium" lsid="urn:lsid:biocol.org:col:15324" name="Catholic University of Leuven" pageId="6" pageNumber="173" type="Herbarium">LV0167</specimenCode>
(
<collectionCode id="ED50AE1AFFBB5527FC57E6C18207FF55" box="[996,1062,158,184]" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13722" lsid="urn:lsid:biocol.org:col:13722" name="National Museum of Natural Science" pageId="6" pageNumber="173" type="Herbarium">TNM</collectionCode>
)
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</paragraph>
<paragraph id="8BFE36DFFFBB5527FF0EE69D8226FDA9" blockId="6.[136,1452,158,1445]" pageId="6" pageNumber="173">
Description (based on 
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): 
<emphasis id="B935EACDFFBB5527FD85E69D84BBFF31" box="[566,666,194,220]" italics="true" pageId="6" pageNumber="173">Saprobic</emphasis>
on leaf of 
<taxonomicName id="4C414D5CFFBB5527FCA5E69C858EFF31" box="[790,943,194,221]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="6" pageNumber="173" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="B935EACDFFBB5527FCA5E69C858EFF31" box="[790,943,194,221]" italics="true" pageId="6" pageNumber="173">Carex pumila</emphasis>
</taxonomicName>
(
<taxonomicName id="4C414D5CFFBB5527FC0DE69D8269FF31" authorityName="de Jussieu" authorityYear="1789" box="[958,1096,194,220]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="173" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
). 
<emphasis id="B935EACDFFBB5527FBE9E69D8327FF30" bold="true" box="[1114,1286,194,221]" italics="true" pageId="6" pageNumber="173">Sexual morph:</emphasis>
Not observed. 
<emphasis id="B935EACDFFBB5527FF3BE6B98769FEEC" bold="true" box="[136,328,230,257]" italics="true" pageId="6" pageNumber="173">Asexual morph:</emphasis>
Colonies superficial, black, oblong to circular, 
<emphasis id="B935EACDFFBB5527FC36E6B885BEFEED" box="[901,927,231,256]" italics="true" pageId="6" pageNumber="173">ca</emphasis>
. 
<quantity id="4CB99B3AFFBB5527FC07E6B9820DFEEC" box="[948,1068,230,257]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="6" pageNumber="173" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
long. 
<emphasis id="B935EACDFFBB5527FBCDE6B98302FEED" box="[1150,1315,230,256]" italics="true" pageId="6" pageNumber="173">Conidiophores</emphasis>
reduced to conidiogenous cells, densely aggregated, brown to dark brown (those giving rise to the stalks of α conidia darker), subglobose, smooth-walled, 
<emphasis id="B935EACDFFBB5527FE75E77087C1FEA5" box="[454,480,303,328]" italics="true" pageId="6" pageNumber="173">ca</emphasis>
. 4.5–7 × 4–5 µm. 
<emphasis id="B935EACDFFBB5527FD18E7718524FEA5" box="[683,773,302,328]" italics="true" pageId="6" pageNumber="173">Conidia</emphasis>
of 
<specimenCount id="9D47FD56FFBB5527FC9BE77085B6FEA4" box="[808,919,303,329]" count="2" pageId="6" pageNumber="173" type="generic">two types</specimenCount>
: α conidia composed of 4–6 spinose dark brown cells (up to 8 spinose cells visible after bleaching), conidial body (11–)13.5–18(–20) µm (n = 30) diam. excluding spines, with dark brown spines up to 6 µm long, conidial stalk dark brown, sometimes pale brown at base, slightly verruculose to verrucose, up to 155 µm long, ca. 2.5 µm wide, widening up to 7.5 µm before terminating into spinose conidial body; β conidia disc-shaped, dark brown, cross-septate, 4-celled, each cell with slightly concave outermost cell wall in front view, smooth-walled, with 1–3 greenish major guttules per cell after bleaching, conidial body (10.5–)12–13(–13.5) × (10–)11–13(–13.5) µm (n = 30), 6–7.5(–8.5) × (10–)11.5–14(–15) µm (n = 19) when seen from the side, conidial stalk subhyaline to pale brown, up to 6 µm long, 
<emphasis id="B935EACDFFBB5527FCF9E4748545FDA9" box="[842,868,555,580]" italics="true" pageId="6" pageNumber="173">ca</emphasis>
. 2.5 µm wide.
</paragraph>
<paragraph id="8BFE36DFFFBB5527FF0EE411827BFD39" blockId="6.[136,1452,158,1445]" pageId="6" pageNumber="173">
<emphasis id="B935EACDFFBB5527FF0EE41187D4FD85" bold="true" box="[189,501,590,616]" pageId="6" pageNumber="173">Culture characteristics:—</emphasis>
Colonies on 
<collectionCode id="ED50AE1AFFBB5527FD30E4118498FD85" box="[643,697,590,616]" country="Sri Lanka" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13944" lsid="urn:lsid:biocol.org:col:13944" name="Royal Botanic Gardens" pageId="6" pageNumber="173" type="Herbarium">PDA</collectionCode>
almost filling 
<quantity id="4CB99B3AFFBB5527FCEDE411858CFD84" box="[862,941,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="173" unit="mm" value="90.0">90 mm</quantity>
diam. Petri dish after 3 weeks at 25 ° 
<collectionCode id="ED50AE1AFFBB5527FAE2E4118344FD85" box="[1361,1381,590,616]" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="173" type="Herbarium">C</collectionCode>
in the dark. Colony in upper view circular with entire edge, flat, surface slightly rough, with concentric rings near margin, center olive brown (oac832) to greenish brown (oac866), outer most margin creamy white (oac816); in reverse view center dark brown (oac733), outer most margin creamy white (oac816); no sporulation.
</paragraph>
<paragraph id="8BFE36DFFFBB5527FF0EE48186CCFC19" blockId="6.[136,1452,158,1445]" pageId="6" pageNumber="173">
<emphasis id="B935EACDFFBB5527FF0EE48187E9FD15" bold="true" box="[189,456,734,760]" pageId="6" pageNumber="173">Material examined:—</emphasis>
<collectingCountry id="F356764FFFBB5527FE7BE4818419FD15" box="[456,568,734,760]" name="Taiwan" pageId="6" pageNumber="173">TAIWAN</collectingCountry>
. 
<collectingRegion id="4985F83DFFBB5527FDF6E4808526FD15" box="[581,775,734,761]" country="Taiwan" name="New Taipei" pageId="6" pageNumber="173">New Taipei City</collectingRegion>
: Gongliao District, Longmen Beach (
<geoCoordinate id="EE755018FFBB5527FB04E4818315FD14" box="[1207,1332,734,761]" degrees="25.0270" direction="north" orientation="latitude" pageId="6" pageNumber="173" precision="5" value="25.027">25.0270 N</geoCoordinate>
, 
<geoCoordinate id="EE755018FFBB5527FAF1E48186BDFCF1" degrees="121.9378" direction="east" orientation="longitude" pageId="6" pageNumber="173" precision="5" value="121.9378">121.9378 E</geoCoordinate>
), on dead leaves of 
<taxonomicName id="4C414D5CFFBB5527FECEE55C844CFCF1" authority="Thunb." authorityName="Thunb." box="[381,621,770,797]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="6" pageNumber="173" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="B935EACDFFBB5527FECEE55C8437FCF1" box="[381,534,770,797]" italics="true" pageId="6" pageNumber="173">Carex pumila</emphasis>
Thunb.
</taxonomicName>
(
<taxonomicName id="4C414D5CFFBB5527FDCFE55D8524FCF1" authorityName="de Jussieu" authorityYear="1789" box="[636,773,770,796]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="173" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
), 
<date id="FFFF101FFFBB5527FCABE55D85D0FCF1" box="[792,1009,770,796]" pageId="6" pageNumber="173" value="2022-12-26">26 December 2022</date>
, 
<collectionCode id="ED50AE1AFFBB5527FC4FE55C8231FCF0" box="[1020,1040,771,797]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="6" pageNumber="173" type="Herbarium">L</collectionCode>
.- 
<collectionCode id="ED50AE1AFFBB5527FBAEE55C8210FCF0" box="[1053,1073,771,797]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="173">T</collectionCode>
. Lu, 
<collectionCode id="ED50AE1AFFBB5527FBDAE55D825EFCF1" box="[1129,1151,770,796]" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="173" type="Herbarium">C</collectionCode>
.- 
<collectionCode id="ED50AE1AFFBB5527FB3FE55D8283FCF1" box="[1164,1186,770,796]" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="173" type="Herbarium">C</collectionCode>
. Chien, 
<collectionCode id="ED50AE1AFFBB5527FB4DE55C8335FCF0" box="[1278,1300,771,797]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="173">R</collectionCode>
. Kirschner &amp; 
<collectionCode id="ED50AE1AFFBB5527FF3BE57886BCFCAC" box="[136,157,807,833]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="6" pageNumber="173" type="Herbarium">Y</collectionCode>
.- 
<collectionCode id="ED50AE1AFFBB5527FF1BE57886E5FCAC" box="[168,196,807,833]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="6" pageNumber="173" type="Herbarium">W</collectionCode>
. Yeh, 
<collectionCode id="ED50AE1AFFBB5527FEB8E579870EFCAD" box="[267,303,806,832]" country="Belgium" lsid="urn:lsid:biocol.org:col:15324" name="Catholic University of Leuven" pageId="6" pageNumber="173" type="Herbarium">LV</collectionCode>
0167 (
<collectionCode id="ED50AE1AFFBB5527FEC4E579879BFCAD" box="[375,442,806,832]" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13722" lsid="urn:lsid:biocol.org:col:13722" name="National Museum of Natural Science" pageId="6" pageNumber="173" type="Herbarium">TNM</collectionCode>
, 
<typeStatus id="54FA887DFFBB5527FE74E579840CFCAD" box="[455,557,806,832]" pageId="6" pageNumber="173" type="holotype">holotype</typeStatus>
; ex-type strain 
<collectionCode id="ED50AE1AFFBB5527FD6EE5798509FCAD" box="[733,808,806,832]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="6" pageNumber="173">BCRC</collectionCode>
<collectionCode id="ED50AE1AFFBB5527FC82E5798576FCAD" box="[817,855,806,832]" country="China" name="Fudan University, Department of Biology" pageId="6" pageNumber="173">FU</collectionCode>
31884). 
<collectingCountry id="F356764FFFBB5527FC07E579822DFCAD" box="[948,1036,806,832]" name="Japan" pageId="6" pageNumber="173">JAPAN</collectingCountry>
. 
<collectingRegion id="4985F83DFFBB5527FBABE5798245FCAD" box="[1048,1124,806,832]" country="Japan" name="Tokyo" pageId="6" pageNumber="173">Tokyo</collectingRegion>
: Ogasawara, Isl. Chichijima, Ogasawara Subtropical Agriculture Center, on leaves of 
<taxonomicName id="4C414D5CFFBB5527FCA2E51582B5FC89" authority="(DC.) Stapf" authorityName="Stapf" baseAuthorityName="DC." box="[785,1172,842,868]" class="Liliopsida" family="Poaceae" genus="Cymbopogon" kingdom="Plantae" order="Poales" pageId="6" pageNumber="173" phylum="Tracheophyta" rank="species" species="citratus">
<emphasis id="B935EACDFFBB5527FCA2E5158227FC89" box="[785,1030,842,868]" italics="true" pageId="6" pageNumber="173">Cymbopogon citratus</emphasis>
(DC.) Stapf
</taxonomicName>
(
<taxonomicName id="4C414D5CFFBB5527FB16E5158326FC89" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[1189,1287,842,868]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="173" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
), 
<date id="FFFF101FFFBB5527FAAFE51586E2FC65" pageId="6" pageNumber="173" value="2013-09-09">9 September 2013</date>
, 
<collectionCode id="ED50AE1AFFBB5527FF7EE53086C5FC64" box="[205,228,879,905]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="6" pageNumber="173" type="Herbarium">K</collectionCode>
. Tanaka, 
<collectionCode id="ED50AE1AFFBB5527FEFFE5308749FC64" box="[332,360,879,905]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="6" pageNumber="173" type="Herbarium">M</collectionCode>
. Sato, 
<collectionCode id="ED50AE1AFFBB5527FE03E53187E6FC65" box="[432,455,878,904]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="6" pageNumber="173" type="Herbarium">A</collectionCode>
. Hashimoto, and 
<collectionCode id="ED50AE1AFFBB5527FD3AE53084BAFC64" box="[649,667,879,905]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="173">T</collectionCode>
. Sato, KT 3323 (
<collectionCode id="ED50AE1AFFBB5527FCE6E5318582FC65" box="[853,931,878,904]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15135" lsid="urn:lsid:biocol.org:col:15135" name="Hirosaki University" pageId="6" pageNumber="173" type="Herbarium">HHUF</collectionCode>
30271, living strain 
<collectionCode id="ED50AE1AFFBB5527FB39E53182F7FC65" box="[1162,1238,878,904]" country="Fiji" httpUri="http://grbio.org/cool/9fa9-21xc" name="Colo-i-Suva Silvicultural Station" pageId="6" pageNumber="173">MAFF</collectionCode>
244519; originally identified as 
<taxonomicName id="4C414D5CFFBB5527FEAEE5CC8529FC41" authority=", Tanaka et al. 2017" authorityName="Tanaka" authorityYear="2017" box="[285,776,914,941]" class="Dothideomycetes" family="Microthyriaceae" genus="Spegazzinia" kingdom="Fungi" order="Microthyriales" pageId="6" pageNumber="173" phylum="Ascomycota" rank="species" species="tessarthra">
<emphasis id="B935EACDFFBB5527FEAEE5CC843FFC41" box="[285,542,914,941]" italics="true" pageId="6" pageNumber="173">Spegazzinia tessarthra</emphasis>
, 
<bibRefCitation id="EFD04B2EFFBB5527FD9DE5CD8529FC41" author="Tanaka, K. &amp; Hashimoto, A. &amp; Sato, M. &amp; Kaminaga, R. &amp; Ono, T. &amp; Sato, T." box="[558,776,914,940]" pageId="6" pageNumber="173" pagination="1 - 22" refId="ref7841" refString="Tanaka, K., Hashimoto, A., Sato, M., Kaminaga, R., Ono, T. &amp; Sato, T. (2017) Identification and species diversity of microfungi (Ascomycota) in the Bonin Islands. MAFF Microorganism Genetic Resources Manual 39: 1 - 22. [in Japanese]" type="journal article" year="2017">
Tanaka 
<emphasis id="B935EACDFFBB5527FD34E5CC84E3FC41" box="[647,706,914,940]" italics="true" pageId="6" pageNumber="173">et al.</emphasis>
2017
</bibRefCitation>
</taxonomicName>
). 
<collectingCountry id="F356764FFFBB5527FCAFE5CD8555FC41" box="[796,884,914,940]" name="Japan" pageId="6" pageNumber="173">JAPAN</collectingCountry>
. 
<collectingRegion id="4985F83DFFBB5527FC32E5CD85CAFC41" box="[897,1003,914,940]" country="Japan" name="Okinawa" pageId="6" pageNumber="173">Okinawa</collectingRegion>
: Isl. Yonaguni, near Mt. Inbi, on stems of bamboo (
<taxonomicName id="4C414D5CFFBB5527FEA1E5E98755FC3D" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[274,372,950,976]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="173" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
), 
<date id="FFFF101FFFBB5527FE34E5E98444FC3D" box="[391,613,950,976]" pageId="6" pageNumber="173" value="2008-11-24">24 November 2008</date>
, 
<collectionCode id="ED50AE1AFFBB5527FDC2E5E884A9FC3C" box="[625,648,951,977]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="6" pageNumber="173" type="Herbarium">K</collectionCode>
. Tanaka &amp; 
<collectionCode id="ED50AE1AFFBB5527FCB8E5E88503FC3C" box="[779,802,951,977]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="6" pageNumber="173" type="Herbarium">K</collectionCode>
. Hirayama, yone 279 (
<collectionCode id="ED50AE1AFFBB5527FB97E5E98253FC3D" box="[1060,1138,950,976]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15135" lsid="urn:lsid:biocol.org:col:15135" name="Hirosaki University" pageId="6" pageNumber="173" type="Herbarium">HHUF</collectionCode>
30152, living strain 
<collectionCode id="ED50AE1AFFBB5527FAECE5E9838AFC3D" box="[1375,1451,950,976]" country="Fiji" httpUri="http://grbio.org/cool/9fa9-21xc" name="Colo-i-Suva Silvicultural Station" pageId="6" pageNumber="173">MAFF</collectionCode>
243878).
</paragraph>
<paragraph id="8BFE36DFFFBB5527FF0EE5A182B5FB1D" blockId="6.[136,1452,158,1445]" pageId="6" pageNumber="173">
<emphasis id="B935EACDFFBB5527FF0EE5A1878BFBF5" bold="true" box="[189,426,1022,1048]" pageId="6" pageNumber="173">Diagnostic bases:—</emphasis>
Within the sequences of 
<taxonomicName id="4C414D5CFFBB5527FD74E5A08563FBF5" authorityName="Liang T. Lu, R. Sugita, Mis. Sato, Kaz. Tanaka &amp; R. Kirschner" authorityYear="2024" box="[711,834,1023,1049]" class="Dothideomycetes" family="Microthyriaceae" genus="Spegazzinia" kingdom="Fungi" order="Microthyriales" pageId="6" pageNumber="173" phylum="Ascomycota" rank="species" species="cryptica">
<emphasis id="B935EACDFFBB5527FD74E5A08563FBF5" box="[711,834,1023,1049]" italics="true" pageId="6" pageNumber="173">S. cryptica</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D5CFFBB5527FCCEE5A085C2FBF5" box="[893,995,1023,1049]" class="Dothideomycetes" family="Microthyriaceae" genus="Spegazzinia" kingdom="Fungi" order="Microthyriales" pageId="6" pageNumber="173" phylum="Ascomycota" rank="species" species="musae">
<emphasis id="B935EACDFFBB5527FCCEE5A085C2FBF5" box="[893,995,1023,1049]" italics="true" pageId="6" pageNumber="173">S. musae</emphasis>
</taxonomicName>
there are eight and 14 diagnostic bases distributed in the 
<collectionCode id="ED50AE1AFFBB5527FEE2E27D875AFBD1" box="[337,379,1058,1084]" pageId="6" pageNumber="173">ITS</collectionCode>
and 
<emphasis id="B935EACDFFBB5527FE07E27C87C7FBD1" box="[436,486,1059,1084]" italics="true" pageId="6" pageNumber="173">
<collectionCode id="ED50AE1AFFBB5527FE07E27C87C7FBD1" box="[436,486,1059,1084]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="6" pageNumber="173" type="Herbarium">TEF</collectionCode>
</emphasis>
alignment, respectively (Tables 2, 3 and Electronic Supplementary Material 3, 4), in the 
<collectionCode id="ED50AE1AFFBB5527FF01E21986FDFB8D" box="[178,220,1094,1120]" pageId="6" pageNumber="173">ITS</collectionCode>
alignment having transitions 
<collectionCode id="ED50AE1AFFBB5527FD98E219841FFB8D" box="[555,574,1094,1120]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="6" pageNumber="173" type="Herbarium">A</collectionCode>
↔ 
<collectionCode id="ED50AE1AFFBB5527FDDAE219845DFB8D" box="[617,636,1094,1120]" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="173" type="Herbarium">C</collectionCode>
at position 58; 
<collectionCode id="ED50AE1AFFBB5527FC9FE219851EFB8D" box="[812,831,1094,1120]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="6" pageNumber="173" type="Herbarium">A</collectionCode>
↔ 
<collectionCode id="ED50AE1AFFBB5527FCD9E219855FFB8D" box="[874,894,1094,1120]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="6" pageNumber="173" type="Herbarium">G</collectionCode>
at position 296; 
<collectionCode id="ED50AE1AFFBB5527FB8FE219826EFB8D" box="[1084,1103,1094,1120]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="6" pageNumber="173" type="Herbarium">A</collectionCode>
↔ 
<collectionCode id="ED50AE1AFFBB5527FBC9E21882AAFB8C" box="[1146,1163,1095,1121]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="173">T</collectionCode>
at position 344,; 
<collectionCode id="ED50AE1AFFBB5527FAEAE219834DFB8D" box="[1369,1388,1094,1120]" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="173" type="Herbarium">C</collectionCode>
↔ 
<collectionCode id="ED50AE1AFFBB5527FA24E219838AFB8D" box="[1431,1451,1094,1120]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="6" pageNumber="173" type="Herbarium">G</collectionCode>
at position 89; 
<collectionCode id="ED50AE1AFFBB5527FE83E2358762FB69" box="[304,323,1130,1156]" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="173" type="Herbarium">C</collectionCode>
↔ 
<collectionCode id="ED50AE1AFFBB5527FEDFE234875CFB68" box="[364,381,1131,1157]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="173">T</collectionCode>
at positions 33, 75 and 88; and 
<collectionCode id="ED50AE1AFFBB5527FD51E23584D7FB69" box="[738,758,1130,1156]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="6" pageNumber="173" type="Herbarium">G</collectionCode>
↔ 
<collectionCode id="ED50AE1AFFBB5527FC93E2348510FB68" box="[800,817,1131,1157]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="173">T</collectionCode>
at position 360; in the 
<emphasis id="B935EACDFFBB5527FB80E2348244FB69" box="[1075,1125,1131,1156]" italics="true" pageId="6" pageNumber="173">
<collectionCode id="ED50AE1AFFBB5527FB80E2348244FB69" box="[1075,1125,1131,1156]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="6" pageNumber="173" type="Herbarium">TEF</collectionCode>
</emphasis>
alignment 
<collectionCode id="ED50AE1AFFBB5527FB51E23582D4FB69" box="[1250,1269,1130,1156]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="6" pageNumber="173" type="Herbarium">A</collectionCode>
↔ 
<collectionCode id="ED50AE1AFFBB5527FAADE234830EFB68" box="[1310,1327,1131,1157]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="173">T</collectionCode>
at position 299; 
<collectionCode id="ED50AE1AFFBB5527FF72E2D186F5FB45" box="[193,212,1166,1192]" country="Denmark" name="University of Copenhagen" pageId="6" pageNumber="173" type="Herbarium">C</collectionCode>
↔ 
<collectionCode id="ED50AE1AFFBB5527FF4DE2D0872EFB44" box="[254,271,1167,1193]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="173">T</collectionCode>
at positions 205, 226, 247, 280, 310, 463, 484, 514, 616, 688, 697 and 796; 
<collectionCode id="ED50AE1AFFBB5527FBDBE2D1825DFB45" box="[1128,1148,1166,1192]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="6" pageNumber="173" type="Herbarium">G</collectionCode>
↔ 
<collectionCode id="ED50AE1AFFBB5527FB15E2D08296FB44" box="[1190,1207,1167,1193]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="173">T</collectionCode>
at position 
<quantity id="4CB99B3AFFBB5527FA88E2D183A3FB45" box="[1339,1410,1166,1193]" metricMagnitude="0" metricUnit="m" metricValue="6.35" pageId="6" pageNumber="173" unit="in" value="250.0">250 in</quantity>
the 
<emphasis id="B935EACDFFBB5527FF3BE2EC869BFB21" box="[136,186,1203,1228]" italics="true" pageId="6" pageNumber="173">
<collectionCode id="ED50AE1AFFBB5527FF3BE2EC869BFB21" box="[136,186,1203,1228]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="6" pageNumber="173" type="Herbarium">TEF</collectionCode>
</emphasis>
alignment. The alignment of the 
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sequences is not shown because there are more intraspecific differences (one base in 
<taxonomicName id="4C414D5CFFBB5527FEA5E28887AFFB1D" authorityName="Liang T. Lu, R. Sugita, Mis. Sato, Kaz. Tanaka &amp; R. Kirschner" authorityYear="2024" box="[278,398,1239,1265]" class="Dothideomycetes" family="Microthyriaceae" genus="Spegazzinia" kingdom="Fungi" order="Microthyriales" pageId="6" pageNumber="173" phylum="Ascomycota" rank="species" species="cryptica">
<emphasis id="B935EACDFFBB5527FEA5E28887AFFB1D" box="[278,398,1239,1265]" italics="true" pageId="6" pageNumber="173">S. cryptica</emphasis>
</taxonomicName>
and three bases in 
<taxonomicName id="4C414D5CFFBB5527FDD7E28884E9FB1D" box="[612,712,1239,1265]" class="Dothideomycetes" family="Microthyriaceae" genus="Spegazzinia" kingdom="Fungi" order="Microthyriales" pageId="6" pageNumber="173" phylum="Ascomycota" rank="species" species="musae">
<emphasis id="B935EACDFFBB5527FDD7E28884E9FB1D" box="[612,712,1239,1265]" italics="true" pageId="6" pageNumber="173">S. musae</emphasis>
</taxonomicName>
) than interspecific difference (one base).
</paragraph>
<paragraph id="8BFE36DFFFBB5527FF0EE2A584B2FAD5" blockId="6.[136,1452,158,1445]" pageId="6" pageNumber="173">
<emphasis id="B935EACDFFBB5527FF0EE2A58508FAF9" bold="true" box="[189,809,1274,1300]" pageId="6" pageNumber="173">Known substrates and geographical distribution:—</emphasis>
Dead leaves and stems of 
<taxonomicName id="4C414D5CFFBB5527FBE1E2A582BAFAF9" box="[1106,1179,1274,1300]" class="Liliopsida" kingdom="Plantae" order="Poales" pageId="6" pageNumber="173" phylum="Tracheophyta" rank="order">Poales</taxonomicName>
(
<taxonomicName id="4C414D5CFFBB5527FB18E2A58313FAF9" authorityName="de Jussieu" authorityYear="1789" box="[1195,1330,1274,1300]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="173" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
, 
<taxonomicName id="4C414D5CFFBB5527FA8DE2A583BEFAF9" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[1342,1439,1274,1300]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="6" pageNumber="173" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
); islands in the western Pacific monsoon region.
</paragraph>
<paragraph id="8BFE36DFFFBB5527FF0EE31D829EFA49" blockId="6.[136,1452,158,1445]" pageId="6" pageNumber="173">
<emphasis id="B935EACDFFBB5527FF0EE31D8470FAB1" bold="true" box="[189,593,1346,1372]" pageId="6" pageNumber="173">Additional material examined:—</emphasis>
<taxonomicName id="4C414D5CFFBB5527FDE2E31C8277FAB1" authority="(S. Hughes) Subram." authorityName="Subram." baseAuthorityName="S. Hughes" box="[593,1110,1346,1373]" class="Dothideomycetes" family="Microthyriaceae" genus="Spegazzinia" kingdom="Fungi" order="Microthyriales" pageId="6" pageNumber="173" phylum="Ascomycota" rank="species" species="deightonii">
<emphasis id="B935EACDFFBB5527FDE2E31C8575FAB1" box="[593,852,1346,1373]" italics="true" pageId="6" pageNumber="173">Spegazzinia deightonii</emphasis>
(S. Hughes) Subram.
</taxonomicName>
, 
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. 
<collectingRegion id="4985F83DFFBB5527FB50E31C8389FAB1" box="[1251,1448,1346,1373]" country="Taiwan" name="New Taipei" pageId="6" pageNumber="173">New Taipei City</collectingRegion>
: Gongliao District, Longmen Beach (
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, 
<geoCoordinate id="EE755018FFBB5527FD10E3398506FA6D" box="[675,807,1382,1408]" degrees="121.9378" direction="east" orientation="longitude" pageId="6" pageNumber="173" precision="5" value="121.9378">121.9378 E</geoCoordinate>
), on dead leaves of 
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<emphasis id="B935EACDFFBB5527FBB0E33882BAFA6D" box="[1027,1179,1382,1409]" italics="true" pageId="6" pageNumber="173">Carex pumila</emphasis>
</taxonomicName>
, 
<date id="FFFF101FFFBB5527FB1BE339835EFA6D" box="[1192,1407,1382,1408]" pageId="6" pageNumber="173" value="2022-12-26">26 December 2022</date>
, 
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.- 
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. Lu, 
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.- 
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. Chien, 
<collectionCode id="ED50AE1AFFBB5527FED4E3D4875CFA48" box="[359,381,1419,1445]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="173">R</collectionCode>
. Kirschner, &amp; 
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.- 
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. Yeh, 
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0169 (
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, living culture 
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<collectionCode id="ED50AE1AFFBB5527FBF7E3D5824BFA49" box="[1092,1130,1418,1444]" country="China" name="Fudan University, Department of Biology" pageId="6" pageNumber="173">FU</collectionCode>
31883).
</paragraph>
</subSubSection>
</treatment>
</document>