169 lines
20 KiB
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169 lines
20 KiB
XML
<document id="6E84717B747E9266729A6051A522E989" ID-CLB-Dataset="20103" ID-DOI="10.11646/zootaxa.5120.3.5" ID-GBIF-Dataset="e162bab9-fcba-4176-a8f7-84de8be953c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="6389529" ID-ZooBank="7E5C9060-305E-4B52-BCFB-A0EDB3EF5292" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648455646056" checkinUser="plazi" docAuthor="Gaimari, Stephen D. & Soares, Matheus M. M." docDate="2022" docId="03E88794FFE0AD7AFF4BF9EA3CA9FDA2" docLanguage="en" docName="zootaxa.5120.3.5.pdf" docOrigin="Zootaxa 5120 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trivialia Malloch 1923" docType="treatment" docVersion="6" lastPageNumber="393" masterDocId="FFD1FFECFFE1AD78FFDCFFBA3E44FF9F" masterDocTitle="Unusual sexually dimorphic head morphology in Lauxaniidae (Diptera: Lauxanioidea) - a new species of the genus Trivialia Malloch from Peru" masterLastPageNumber="401" masterPageNumber="391" pageNumber="392" updateTime="1699318620531" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="2A4E9D272AA86199AE4F99C7A0DF2F2A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C2A79C9D074493D252EEB3B763BA5A38">
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<mods:title id="D4A9D44BD90D8C78B252A79BA518758A">Unusual sexually dimorphic head morphology in Lauxaniidae (Diptera: Lauxanioidea) - a new species of the genus Trivialia Malloch from Peru</mods:title>
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</mods:titleInfo>
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<mods:name id="EB8A9B7928D683540BC0223B703F9A25" type="personal">
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<mods:roleTerm id="CE0A02F9DFC2590C05E7630072363A18">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FB21D71A0BED3BB8438BDB0DA7B30054">Gaimari, Stephen D.</mods:namePart>
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<mods:nameIdentifier id="F7BCF892E72553447A68C5CB1CFC97CC" type="ORCID">0000-0002-4240-7154</mods:nameIdentifier>
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<mods:affiliation id="F604A5F17EFCA765A52C237EAE234CB1">California State Collection of Arthropods, California Department of Food & Agriculture, Plant Pest Diagnostics Branch, 3294 Meadowview Road, Sacramento, California 95832 - 1448, USA. sgaimari @ gmail. com; https: // orcid. org / 0000 - 0002 - 4240 - 7154 Graduate Program in Entomology, Instituto Nacional de Pesquisas da Amazônia - INPA, Av. Andr Araújo, Petrópolis, 2936, CEP 69067 - 375, Manaus, Amazonas, Brazil. matheusmmsoares @ gmail. com; https: // orcid. org / 0000 - 0002 - 2355 - 1441</mods:affiliation>
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<mods:nameIdentifier id="506E610B91A6ABA240089AFB3FB36F2F" type="email">sgaimari@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="E7FA20E79DAD5D2BAA9BF81B852D4883" type="personal">
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<mods:role id="704E810B9BB2445763C8D69FF4BC6EC9">
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<mods:roleTerm id="A8471790D97FD3C6440B9E75426B660B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0BB22C8645FD32370D39DB9DD9B10465">Soares, Matheus M. M.</mods:namePart>
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<mods:nameIdentifier id="A12E069A1DE73793E0BA492030303E3F" type="ORCID">0000-0002-2355-1441</mods:nameIdentifier>
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<mods:nameIdentifier id="2589E1F8514648CB6FC09BE778E0D6E5" type="email">matheusmmsoares@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="FC0A6FDF2A0DC97702FA374F3320F119">text</mods:typeOfResource>
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<mods:title id="CE53F792257BB5E0B27D9E5F0A356AF8">Zootaxa</mods:title>
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<mods:part id="4B608AA7C3CF1A84E716391E8AA29198">
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<mods:date id="22EF8819CFBD18F8384B9CDD0F213D20">2022</mods:date>
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<mods:number id="E4A97C97CE0EB3C38F19B34A3169ED6C">2022-03-28</mods:number>
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</mods:detail>
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<mods:detail id="8E02D152890E30686EFD2E079577CD03" type="volume">
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<mods:number id="A2476CBF44362ABB5149F202444A8FDB">5120</mods:number>
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</mods:detail>
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<mods:detail id="06710EC9C61CF49C221D2FD4F343C51F" type="issue">
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<mods:number id="EB2A6E9D2F35EA7B933C0C6E087C99CB">3</mods:number>
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<mods:start id="4C40D09F45E8CD7D4AD1B8CD6C4CE43C">391</mods:start>
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<mods:identifier id="8369CCB330C2F9AF59E485A9EC07A7CB" type="DOI">10.11646/zootaxa.5120.3.5</mods:identifier>
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<treatment id="03E88794FFE0AD7AFF4BF9EA3CA9FDA2" ID-DOI="http://doi.org/10.5281/zenodo.6389527" ID-GBIF-Taxon="194285846" ID-Zenodo-Dep="6389527" LSID="urn:lsid:plazi:treatment:03E88794FFE0AD7AFF4BF9EA3CA9FDA2" httpUri="http://treatment.plazi.org/id/03E88794FFE0AD7AFF4BF9EA3CA9FDA2" lastPageId="2" lastPageNumber="393" pageId="1" pageNumber="392">
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<subSubSection id="C35B6509FFE0AD79FF4BF9EA3FDEF9F5" box="[151,410,1616,1642]" pageId="1" pageNumber="392" type="nomenclature">
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<paragraph id="8BFE3682FFE0AD79FF4BF9EA3FDEF9F5" blockId="1.[151,410,1616,1642]" box="[151,410,1616,1642]" pageId="1" pageNumber="392">
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<heading id="D0B681EEFFE0AD79FF4BF9EA3FDEF9F5" bold="true" box="[151,410,1616,1642]" fontSize="11" level="1" pageId="1" pageNumber="392" reason="1">
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<taxonomicName id="4C414D01FFE0AD79FF4BF9EA3FDEF9F5" ID-CoL="82NG" authority="Malloch" authorityName="Malloch" authorityYear="1923" box="[151,410,1616,1642]" class="Insecta" family="Lauxaniidae" genus="Trivialia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="genus">
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<emphasis id="B935EA90FFE0AD79FF4BF9EA3FDEF9F5" bold="true" box="[151,410,1616,1642]" pageId="1" pageNumber="392">
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<emphasis id="B935EA90FFE0AD79FF4BF9EA3F6FF9F5" bold="true" box="[151,299,1616,1642]" italics="true" pageId="1" pageNumber="392">TRIVIALIA</emphasis>
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Malloch
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35B6509FFE0AD79FF4BF92E3D09F932" box="[151,845,1684,1709]" pageId="1" pageNumber="392" type="reference_group">
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<paragraph id="8BFE3682FFE0AD79FF4BF92E3D09F932" blockId="1.[151,845,1684,1709]" box="[151,845,1684,1709]" pageId="1" pageNumber="392">
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<treatmentCitationGroup id="AB5111ACFFE0AD79FF4BF92E3D09F932" box="[151,845,1684,1709]" pageId="1" pageNumber="392">
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<treatmentCitation id="0AE01093FFE0AD79FF4BF92E3FF5F933" author="Malloch, J. R." box="[151,433,1684,1709]" page="53" pageId="1" pageNumber="392" year="1923">
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<taxonomicName id="4C414D01FFE0AD79FF4BF92E3FF5F933" ID-CoL="82NG" authority="Malloch, 1923: 53" authorityName="Malloch" authorityPageNumber="53" authorityYear="1923" box="[151,433,1684,1709]" class="Insecta" family="Lauxaniidae" genus="Trivialia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="genus">
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<emphasis id="B935EA90FFE0AD79FF4BF92E3EB4F933" bold="true" box="[151,240,1684,1708]" italics="true" pageId="1" pageNumber="392">Trivialia</emphasis>
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<bibRefCitation id="EFD04B73FFE0AD79FF2BF92E3FF5F933" author="Malloch, J. R." box="[247,433,1684,1709]" pageId="1" pageNumber="392" pagination="45 - 53" refId="ref5528" refString="Malloch, J. R. (1923) Some new genera and species of Lonchaeidae and Sapromyzidae (Diptera). Proceedings of the Entomological Society of Washington, 25 (2), 45 - 53." type="journal article" year="1923">Malloch, 1923: 53</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(as subgenus of
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<taxonomicName id="4C414D01FFE0AD79FD87F92F3D05F932" ID-CoL="42CM" authority="Malloch, 1923" authorityName="Malloch" authorityYear="1923" box="[603,833,1684,1709]" class="Insecta" family="Lauxaniidae" genus="Deceia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="genus">
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<emphasis id="B935EA90FFE0AD79FD87F92F3CE7F933" box="[603,675,1685,1708]" italics="true" pageId="1" pageNumber="392">Deceia</emphasis>
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<bibRefCitation id="EFD04B73FFE0AD79FD75F92E3D05F932" author="Malloch, J. R." box="[681,833,1684,1709]" pageId="1" pageNumber="392" pagination="45 - 53" refId="ref5528" refString="Malloch, J. R. (1923) Some new genera and species of Lonchaeidae and Sapromyzidae (Diptera). Proceedings of the Entomological Society of Washington, 25 (2), 45 - 53." type="journal article" year="1923">Malloch, 1923</bibRefCitation>
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</taxonomicName>
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).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35B6509FFE0AD79FF4BF9613DB8F969" box="[151,1020,1754,1782]" pageId="1" pageNumber="392" type="type_taxon">
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<paragraph id="8BFE3682FFE0AD79FF4BF9613DB8F969" blockId="1.[151,1437,1754,2033]" box="[151,1020,1754,1782]" pageId="1" pageNumber="392">
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<emphasis id="B935EA90FFE0AD79FF4BF9613F76F96B" bold="true" box="[151,306,1754,1781]" pageId="1" pageNumber="392">
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<typeStatus id="54FA8820FFE0AD79FF4BF9613E96F96A" box="[151,210,1755,1781]" pageId="1" pageNumber="392">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C414D01FFE0AD79FEE5F9613CBFF96A" ID-CoL="8BRFL" authority="Malloch, 1923: 53" authorityName="Malloch" authorityPageNumber="53" authorityYear="1923" box="[313,763,1754,1781]" class="Insecta" family="Lauxaniidae" genus="Deceia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="fuscocapitata">
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<emphasis id="B935EA90FFE0AD79FEE5F9613C63F96A" box="[313,551,1755,1781]" italics="true" pageId="1" pageNumber="392">Deceia fuscocapitata</emphasis>
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<bibRefCitation id="EFD04B73FFE0AD79FDF1F9613CBFF96A" author="Malloch, J. R." box="[557,763,1754,1781]" pageId="1" pageNumber="392" pagination="45 - 53" refId="ref5528" refString="Malloch, J. R. (1923) Some new genera and species of Lonchaeidae and Sapromyzidae (Diptera). Proceedings of the Entomological Society of Washington, 25 (2), 45 - 53." type="journal article" year="1923">Malloch, 1923: 53</bibRefCitation>
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</taxonomicName>
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(original designation).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35B6509FFE0AD7AFF1BF9443C2CFEDE" lastPageId="2" lastPageNumber="393" pageId="1" pageNumber="392" type="diagnosis">
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<paragraph id="8BFE3682FFE0AD7AFF1BF9443A84FE82" blockId="1.[151,1437,1754,2033]" lastBlockId="2.[151,1436,151,574]" lastPageId="2" lastPageNumber="393" pageId="1" pageNumber="392">
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<emphasis id="B935EA90FFE0AD79FF1BF9443F06F887" bold="true" box="[199,322,1790,1816]" pageId="1" pageNumber="392">Diagnosis.</emphasis>
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This genus can be differentiated from other lauxaniids by the following combination of characteristics. On the head, the frons is broader than long, bare of setulae, with the fronto-orbital setae reclinate and strong, with the posterior seta stronger. The ocellar setae are short and divergent, shorter than the postocellar setae. The eyes are almost round and are not narrowed below. The face is flat and dull (not glossy). The antennal first flagellomere is black, and the arista is short plumose with the basal-most dorsal rays usually the longest, and with shorter rays below. The scutum has 2 pairs of postsutural dorsocentral setae (and no presutural dorsocentral setae), with the anterior seta at about the midpoint between the posterior seta and the transverse suture. The postsutural intra-alar seta is absent (i.e., this genus is not in the
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<taxonomicName id="4C414D01FFE3AD7AFD5AFF2D3CA1FF2F" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[646,741,151,176]" class="Insecta" family="Lauxaniidae" genus="Minettia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="genus">
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<emphasis id="B935EA90FFE3AD7AFD5AFF2D3CA1FF2F" box="[646,741,151,176]" italics="true" pageId="2" pageNumber="393">Minettia</emphasis>
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</taxonomicName>
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group of genera). The anterior scutellar setae are parallel to slightly convergent, while the posterior scutellar setae are divergent. On the pleuron, the anepimeron is bare and there is 1 strong katepisternal seta (sometimes with an additional weaker seta anteriorly). On the wing, the costal setulae end well before the apex of R
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<subScript id="17C534C7FFE3AD7AFDEBFEAF3C15FEBC" attach="left" box="[567,593,277,291]" fontSize="6" pageId="2" pageNumber="393">4+5</subScript>
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(as is typical in
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<taxonomicName id="4C414D01FFE3AD7AFCD5FEB93DDCFE81" authorityName="Macquart" authorityYear="1835" box="[777,920,259,286]" class="Insecta" family="Lauxaniidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="subFamily" subFamily="Lauxaniinae">Lauxaniinae</taxonomicName>
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), and the R veins are bare.
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</paragraph>
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<paragraph id="8BFE3682FFE3AD7AFF1BFE9D3C2CFEDE" blockId="2.[151,1436,151,574]" box="[199,616,295,321]" pageId="2" pageNumber="393">
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<emphasis id="B935EA90FFE3AD7AFF1BFE9D3F16FEDE" bold="true" box="[199,338,295,321]" pageId="2" pageNumber="393">Immatures.</emphasis>
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Unknown for the genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35B6509FFE3AD7AFF1BFEF03A3DFE16" pageId="2" pageNumber="393" type="distribution">
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<paragraph id="8BFE3682FFE3AD7AFF1BFEF03A3DFE16" blockId="2.[151,1436,151,574]" pageId="2" pageNumber="393">
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<emphasis id="B935EA90FFE3AD7AFF1BFEF03F25FEFB" bold="true" box="[199,353,330,356]" pageId="2" pageNumber="393">Distribution.</emphasis>
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North America (
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<collectingCountry id="F3567612FFE3AD7AFDF4FEF13C24FEFA" box="[552,608,331,357]" name="United States of America" pageId="2" pageNumber="393">USA</collectingCountry>
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[eastern,
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<collectingRegion id="4985F860FFE3AD7AFD08FEF13D6BFEFA" box="[724,815,331,357]" country="United States of America" name="Virginia" pageId="2" pageNumber="393">Virginia</collectingRegion>
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south to
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<collectingRegion id="4985F860FFE3AD7AFC7DFEF03DBDFEFA" box="[929,1017,330,357]" country="United States of America" name="Florida" pageId="2" pageNumber="393">Florida</collectingRegion>
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]), West Indies (
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<collectingCountry id="F3567612FFE3AD7AFB73FEF13B70FEF9" box="[1199,1332,331,358]" name="Saint Vincent and the Grenadines" pageId="2" pageNumber="393">St. Vincent</collectingCountry>
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), Central America (
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<collectingCountry id="F3567612FFE3AD7AFEDAFED53FC1FE16" box="[262,389,367,393]" name="Costa Rica" pageId="2" pageNumber="393">Costa Rica</collectingCountry>
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,
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<collectingCountry id="F3567612FFE3AD7AFE4CFED53FA9FE16" box="[400,493,367,393]" name="Panama" pageId="2" pageNumber="393">Panama</collectingCountry>
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), South America (
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<collectingCountry id="F3567612FFE3AD7AFD6AFED53D6DFE16" box="[694,809,367,393]" name="Colombia" pageId="2" pageNumber="393">Colombia</collectingCountry>
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,
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<collectingCountry id="F3567612FFE3AD7AFCE8FED53DD7FE16" box="[820,915,367,393]" name="Ecuador" pageId="2" pageNumber="393">Ecuador</collectingCountry>
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,
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<collectingCountry id="F3567612FFE3AD7AFC42FED53D95FE16" box="[926,977,367,393]" name="Peru" pageId="2" pageNumber="393">Peru</collectingCountry>
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[new record]).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35B6509FFE3AD7AFF1BFE293CA9FDA2" pageId="2" pageNumber="393" type="discussion">
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<paragraph id="8BFE3682FFE3AD7AFF1BFE293CA9FDA2" blockId="2.[151,1436,151,574]" pageId="2" pageNumber="393">
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<emphasis id="B935EA90FFE3AD7AFF1BFE293F7FFE32" bold="true" box="[199,315,403,429]" pageId="2" pageNumber="393">Remarks.</emphasis>
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<bibRefCitation id="EFD04B73FFE3AD7AFE9FFE293C65FE32" author="Stuckenberg, B. R." box="[323,545,403,429]" pageId="2" pageNumber="393" pagination="499 - 610" refId="ref6085" refString="Stuckenberg, B. R. (1971) A review of the Old World genera of Lauxaniidae (Diptera). Annals of the Natal Museum, 20, 499 - 610." type="journal article" year="1971">Stuckenberg (1971)</bibRefCitation>
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and
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<bibRefCitation id="EFD04B73FFE3AD7AFD85FE293D42FE32" author="Shewell, G. C." box="[601,774,403,429]" pageId="2" pageNumber="393" pagination="951 - 964" refId="ref5985" refString="Shewell, G. C. (1987) Lauxaniidae. In: McAlpine, J. F. (Ed.), Manual of Nearctic Diptera. Vol. 2. Monograph 28. Research Branch, Agriculture Canada, Ottawa, pp. 951 - 964." type="book chapter" year="1987">Shewell (1987)</bibRefCitation>
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considered this an endemic Nearctic genus. Besides being included in the key to Nearctic genera,
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<bibRefCitation id="EFD04B73FFE3AD7AFD83FE0D3D49FE4E" author="Shewell, G. C." box="[607,781,439,465]" pageId="2" pageNumber="393" pagination="951 - 964" refId="ref5985" refString="Shewell, G. C. (1987) Lauxaniidae. In: McAlpine, J. F. (Ed.), Manual of Nearctic Diptera. Vol. 2. Monograph 28. Research Branch, Agriculture Canada, Ottawa, pp. 951 - 964." type="book chapter" year="1987">Shewell (1987)</bibRefCitation>
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provided a figure of the lateral head of the type species,
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<taxonomicName id="4C414D01FFE3AD7AFF4BFE613C02FE6A" authority="Malloch, 1923" authorityName="Malloch" authorityYear="1923" box="[151,582,475,501]" class="Insecta" family="Lauxaniidae" genus="Trivialia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="species" species="fuscocapitata">
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<emphasis id="B935EA90FFE3AD7AFF4BFE613FD3FE6A" box="[151,407,475,501]" italics="true" pageId="2" pageNumber="393">Trivialia fuscocapitata</emphasis>
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<bibRefCitation id="EFD04B73FFE3AD7AFE43FE613C02FE6A" author="Malloch, J. R." box="[415,582,475,501]" pageId="2" pageNumber="393" pagination="45 - 53" refId="ref5528" refString="Malloch, J. R. (1923) Some new genera and species of Lonchaeidae and Sapromyzidae (Diptera). Proceedings of the Entomological Society of Washington, 25 (2), 45 - 53." type="journal article" year="1923">Malloch, 1923</bibRefCitation>
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</taxonomicName>
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, which was also included in
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<bibRefCitation id="EFD04B73FFE3AD7AFC53FE613AE9FE6A" author="Gaimari, S. D. & Silva, V. C." box="[911,1197,475,501]" pageId="2" pageNumber="393" refId="ref5051" refString="Gaimari, S. D. & Silva, V. C. (2010 a) Lauxaniidae. In: Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, N. E. & Zumbado, M. (Eds.), Manual of Central American Diptera. Vol. 2. National Research Council Press, Ottawa, pp. 971 - 995." type="book" year="2010">Gaimari & Silva (2010a)</bibRefCitation>
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. Photographs (lateral and dorsal habitus and oblique head) of the
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<typeStatus id="54FA8820FFE3AD7AFD50FE453CB7FD86" box="[652,755,511,537]" pageId="2" pageNumber="393" type="lectotype">lectotype</typeStatus>
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of one of the included species,
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<taxonomicName id="4C414D01FFE3AD7AFB82FE453E9FFDA1" authority="(Williston, 1896)" baseAuthorityName="Williston" baseAuthorityYear="1896" class="Insecta" family="Lauxaniidae" genus="Trivialia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="species" species="venusta">
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<emphasis id="B935EA90FFE3AD7AFB82FE453B59FD87" box="[1118,1309,511,537]" italics="true" pageId="2" pageNumber="393">Trivialia venusta</emphasis>
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(
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<bibRefCitation id="EFD04B73FFE3AD7AFAF0FE453E97FDA1" author="Williston, S. W." pageId="2" pageNumber="393" pagination="253 - 446" refId="ref6173" refString="Williston, S. W. (1896) On the Diptera of St. Vincent (West Indies). Transactions of the Entomological Society of London, 1896 (3), 253 - 446. https: // doi. org / 10.1111 / j. 1365 - 2311.1896. tb 00965. x" type="journal article" year="1896">Williston, 1896</bibRefCitation>
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)
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</taxonomicName>
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, was also provided by
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<bibRefCitation id="EFD04B73FFE3AD7AFE07FD993CADFDA2" author="Gaimari, S. D. & Silva, V. C." box="[475,745,547,573]" pageId="2" pageNumber="393" pagination="1 - 217" refId="ref5189" refString="Gaimari, S. D. & Silva, V. C. (2020) A conspectus of Neotropical Lauxaniidae (Diptera: Lauxanioidea). Zootaxa, 4862 (1), 1 - 217. https: // doi. org / 10.11646 / zootaxa. 4862.1.1" type="journal article" year="2020">Gaimari & Silva (2020)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |