treatments-xml/data/03/E8/87/03E88783FF81FFDAFF5C30B0FF68D741.xml

<document id="774061D2A33AFB47E603480AFADE1CF0" ID-CLB-Dataset="132754" ID-DOI="10.11646/zootaxa.5178.5.6" ID-GBIF-Dataset="934298b6-eddd-469d-8a95-c734cedb584a" ID-ISSN="1175-5326" ID-Zenodo-Dep="7034050" ID-ZooBank="ABC5B768-A3CE-4965-BFD4-90FA8271B799" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661844100065" checkinUser="plazi" docAuthor="Vilela, Diogo Silva, Rodrigues, Marciel Elio &amp; Lencioni, Frederico A. A." docDate="2022" docId="03E88783FF81FFDAFF5C30B0FF68D741" docLanguage="en" docName="zootaxa.5178.5.6.pdf" docOrigin="Zootaxa 5178 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heteragrion roquei Vilela, Rodrigues &amp; Lencioni 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="499" masterDocId="FFD1FFFBFF83FFDCFFCB306EFF98D56A" masterDocTitle="Revealing the Odonatofauna of Northeastern Brazil: new Heteragrion Selys, 1862 (Odonata: Heteragrionidae) species from Bahia state" masterLastPageNumber="500" masterPageNumber="493" pageNumber="495" updateTime="1699374471167" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B55A74D601E6C55E8ACF2B14DB423EFC">Revealing the Odonatofauna of Northeastern Brazil: new Heteragrion Selys, 1862 (Odonata: Heteragrionidae) species from Bahia state</mods:title>
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<mods:namePart id="BE479C9704D5B9BC188A389750DD2BDD">Vilela, Diogo Silva</mods:namePart>
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<mods:affiliation id="1163C68485E6A7368CC7EC7C5F040D4B">Laboratório de Biologia Aquática, Departamento de Ciências Biológicas, Faculdade de Ciências e Letras de Assis, Universidade Estadual Paulista, Assis, SP, Brazil. deeogoo @ gmail. com; https: // orcid. org / 0000 - 0001 - 6510 - 7018</mods:affiliation>
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<mods:namePart id="835843E5B7B8A84EB189832B1CB943EB">Rodrigues, Marciel Elio</mods:namePart>
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<mods:affiliation id="EA71C93822AC6BAEFB831774F7EEBAB9">Laborátorio de Organismos Aquáticos, Departamento de Ciências Biológicas, Universidade Estadual de Santa Cruz (UESC), Ilhéus, BA, Brazil. merodrigues @ uesc. br; https: // orcid. org / 0000 - 0001 - 8161 - 6234</mods:affiliation>
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<mods:namePart id="CFB815DECA67817E6ADE9143D4BCD17D">Lencioni, Frederico A. A.</mods:namePart>
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<taxonomicName id="4C414D16FF81FFDEFF5C30B0FD6CD592" authority="Vilela, Rodrigues &amp; Lencioni" authorityName="Vilela, Rodrigues &amp; Lencioni" authorityYear="2022" box="[151,756,222,248]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="roquei" status="sp. nov.">
<emphasis id="B935EA87FF81FFDEFF5C30B0FE19D592" bold="true" box="[151,385,222,248]" italics="true" pageId="2" pageNumber="495">Heteragrion roquei</emphasis>
Vilela, Rodrigues &amp; Lencioni
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(
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a−b, 2, 3a−b, 4a−b, 5d)
</paragraph>
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<paragraph id="8BFE3695FF81FFDEFF5C312BFD18D4E9" blockId="2.[151,1437,325,1036]" pageId="2" pageNumber="495">
<materialsCitation id="3B293CC8FF81FFDEFF5C312BFD18D4E9" ID-GBIF-Occurrence="3897088301" collectingDate="2019-09-19" collectionCode="CZYG" collectorName="M. E. Rodrigues" country="Brazil" elevation="40" latitude="-15.3146" location="Fazenda Araruna" longLatPrecision="7" longitude="-39.1621" municipality="Municipality of Una" pageId="2" pageNumber="495" specimenCode="CZYG3085" specimenCount="1" specimenCount-male="1" stateProvince="Bahia" typeStatus="holotype">
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<typeStatus id="54FA8837FF81FFDEFF5C312BFE9ED435" box="[151,262,325,351]" pageId="2" pageNumber="495" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D47FD1CFF81FFDEFEDA312BFEBED435" box="[273,294,325,351]" pageId="2" pageNumber="495" type="male">♂</specimenCount>
(
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BA), 
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, 
<collectingRegion id="4985F877FF81FFDEFD82312BFD5ED435" box="[585,710,325,351]" country="Brazil" name="Bahia" pageId="2" pageNumber="495">Bahia state</collectingRegion>
, 
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, 
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, (
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, 
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, 
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<collectingDate id="EFBBE9BDFF81FFDEFF703107FEADD4E9" box="[187,309,361,387]" pageId="2" pageNumber="495" value="2019-09-19">19.ix.2019</collectingDate>
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, 
<collectorName id="26B45343FF81FFDEFE8A3107FE6BD4E9" box="[321,499,361,387]" pageId="2" pageNumber="495">M.E. Rodrigues</collectorName>
leg., UESC.
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<materialsCitation id="3B293CC8FF81FFDEFF0C31E3FB57D4CD" ID-GBIF-Occurrence="3897088302" box="[199,1231,397,424]" collectingDate="2019-09-19" collectionCode="FAAL" collectorName="M. E. Rodrigues" country="Brazil" elevation="40" latitude="-15.3101" location="Fazenda Araruna" longLatPrecision="7" longitude="-39.1621" municipality="Municipality of Una" pageId="2" pageNumber="495" specimenCode="CZYG3065" specimenCount="1" specimenCount-male="1" stateProvince="Bahia" typeStatus="paratype">
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<typeStatus id="54FA8837FF81FFDEFF0C31E3FEAED4CD" box="[199,310,397,423]" pageId="2" pageNumber="495" type="paratype">Paratype</typeStatus>
.
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<specimenCount id="9D47FD1CFF81FFDEFE8931E3FEFDD4CD" box="[322,357,397,423]" pageId="2" pageNumber="495" type="male">1♂</specimenCount>
(
<specimenCode id="DBE79EEEFF81FFDEFEBF31E3FE65D4C2" box="[372,509,397,424]" collectionCode="CZYG" pageId="2" pageNumber="495">CZYG3065</specimenCode>
BA), same data as holotype, except (
<geoCoordinate id="EE755052FF81FFDEFC5E31E3FC64D4C2" box="[917,1020,397,424]" degrees="15.3101" direction="south" orientation="latitude" pageId="2" pageNumber="495" precision="5" value="-15.3101">-15.3101</geoCoordinate>
, 
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), 
<collectionCode id="ED50AE50FF81FFDEFB4F31E3FB57D4CD" box="[1156,1231,397,423]" pageId="2" pageNumber="495">FAAL</collectionCode>
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.
</paragraph>
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<subSubSection id="C35B651EFF81FFD8FF0C31DFFE3AD210" lastPageId="4" lastPageNumber="497" pageId="2" pageNumber="495" type="etymology">
<paragraph id="8BFE3695FF81FFDEFF0C31DFFE04D731" blockId="2.[151,1437,325,1036]" pageId="2" pageNumber="495">
<emphasis id="B935EA87FF81FFDEFF0C31DFFED6D4A1" bold="true" box="[199,334,433,459]" pageId="2" pageNumber="495">Etymology.</emphasis>
Named 
<taxonomicName id="4C414D16FF81FFDEFE6631DCFE6DD4A1" authorityName="Vilela, Rodrigues &amp; Lencioni" authorityYear="2022" box="[429,501,434,459]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="roquei">
<emphasis id="B935EA87FF81FFDEFE6631DCFE6DD4A1" box="[429,501,434,459]" italics="true" pageId="2" pageNumber="495">roquei</emphasis>
</taxonomicName>
(noun in genitive case) after Dr. Fábio de Oliveira Roque, professor and researcher at the Federal University of 
<collectingRegion id="4985F877FF81FFDEFE1831BBFD2ED485" box="[467,694,469,495]" country="Brazil" name="Mato Grosso do Sul" pageId="2" pageNumber="495">Mato Grosso do Sul</collectingRegion>
, 
<collectingCountry id="F3567605FF81FFDEFD0931BBFC91D485" box="[706,777,469,495]" name="Brazil" pageId="2" pageNumber="495">Brazil</collectingCountry>
, for his dedication and contribution to science with studies focused on ecology and conservation of continental aquatic ecosystems, especially with the aquatic insects. And also, for the friendship, partnership in research and orientation to studies with 
<taxonomicName id="4C414D16FF81FFDEFBC03273FBF2D75D" authorityName="Fabricius" authorityYear="1793" box="[1035,1130,541,567]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="495" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
during the second author’s master’s and doctorate.
</paragraph>
<paragraph id="8BFE3695FF81FFDEFF0C320BFDD3D781" blockId="2.[151,1437,325,1036]" pageId="2" pageNumber="495">
<emphasis id="B935EA87FF81FFDEFF0C320BFE9CD715" box="[199,260,613,639]" italics="true" pageId="2" pageNumber="495">Head</emphasis>
(
<figureCitation id="137A2A10FF81FFDEFED9320BFEC2D715" box="[274,346,613,639]" captionStart="FIGURE 1" captionStartId="1.[151,250,1911,1935]" captionTargetBox="[151,1416,611,1860]" captionTargetId="figure-181@1.[151,1435,594,1885]" captionTargetPageId="1" captionText="FIGURE 1. Heteragrion roquei sp. nov. male holotype: (a) lateral view of habitus, (b) dorsal view of head and thorax; Heteragrion gracile Machado, 2006 male holotype: (c) lateral view of habitus." figureDoi="http://doi.org/10.5281/zenodo.7034052" httpUri="https://zenodo.org/record/7034052/files/figure.png" pageId="2" pageNumber="495">Figs. 1</figureCitation>
a−b, 3a). Labium and labrum brown; base of mandibles and genae pale colored, apex of mandibles dark brown; remainder of the head orange, except for the brown vertex, and following black areas: a black stripe lateral to each antenna, a black stripe ranging from each postocular area to the occipital bar. Rear of the head pale colored, darker on the occipital region.
</paragraph>
<paragraph id="8BFE3695FF81FFDEFF0C329BFECED6A9" blockId="2.[151,1437,325,1036]" pageId="2" pageNumber="495">
<emphasis id="B935EA87FF81FFDEFF0C329BFE80D665" box="[199,280,757,783]" italics="true" pageId="2" pageNumber="495">Thorax</emphasis>
(
<figureCitation id="137A2A10FF81FFDEFEE2329BFEEDD665" box="[297,373,757,783]" captionStart="FIGURE 1" captionStartId="1.[151,250,1911,1935]" captionTargetBox="[151,1416,611,1860]" captionTargetId="figure-181@1.[151,1435,594,1885]" captionTargetPageId="1" captionText="FIGURE 1. Heteragrion roquei sp. nov. male holotype: (a) lateral view of habitus, (b) dorsal view of head and thorax; Heteragrion gracile Machado, 2006 male holotype: (c) lateral view of habitus." figureDoi="http://doi.org/10.5281/zenodo.7034052" httpUri="https://zenodo.org/record/7034052/files/figure.png" pageId="2" pageNumber="495">Figs. 1</figureCitation>
a−b, 3a−b). Prothorax mostly orange; anterior lobe black laterally and posteriorly, remainder orange; medial lobe mostly orange with two black stripes on each side; hind lobe orange with a black medial spot. Pterothorax mostly orange/yellow, with a black middorsal carina; mesepisternum orange with a black basal stripe; mesopleural suture pale; mesepimeron and metepisternum each orange with a medial thin black stripe; remainder of pterothorax yellow. Coxae and trochanter pale colored; legs pale with smoky femoral areas, darker on distal femora and apical tibiae.
</paragraph>
<paragraph id="8BFE3695FF81FFDEFF0C33A0FC2BD161" blockId="2.[151,1437,325,1036]" pageId="2" pageNumber="495">
<emphasis id="B935EA87FF81FFDEFF0C33A0FE93D68D" box="[199,267,974,999]" italics="true" pageId="2" pageNumber="495">Wings</emphasis>
(
<figureCitation id="137A2A10FF81FFDEFED733A3FEF8D68D" box="[284,352,973,999]" captionStart="FIGURE 2" captionStartId="2.[151,250,1547,1571]" captionTargetBox="[235,1349,1065,1519]" captionTargetId="figure-325@2.[231,1357,1059,1523]" captionTargetPageId="2" captionText="FIGURE 2. Wings of Heteragrion roquei sp. nov., male holotype." figureDoi="http://doi.org/10.5281/zenodo.7034054" httpUri="https://zenodo.org/record/7034054/files/figure.png" pageId="2" pageNumber="495">Fig. 2</figureCitation>
). Hyaline, reaching S6; venation and Pt dark brown; Pt overlying 2.5 cells in Fw, 2 cells in Hw, oblique proximally; 2 Pq cells in all wings; 19 Px in FW, 15 Px in HW.
</paragraph>
<caption id="DF3E661DFF81FFDEFF5C3665FCA4D349" ID-DOI="http://doi.org/10.5281/zenodo.7034054" ID-Zenodo-Dep="7034054" box="[151,828,1547,1571]" httpUri="https://zenodo.org/record/7034054/files/figure.png" pageId="2" pageNumber="495" startId="2.[151,250,1547,1571]" targetBox="[235,1349,1065,1519]" targetPageId="2">
<paragraph id="8BFE3695FF81FFDEFF5C3665FCA4D349" blockId="2.[151,828,1547,1571]" box="[151,828,1547,1571]" pageId="2" pageNumber="495">
<emphasis id="B935EA87FF81FFDEFF5C3665FE8BD349" bold="true" box="[151,275,1547,1571]" pageId="2" pageNumber="495">FIGURE 2.</emphasis>
Wings of 
<taxonomicName id="4C414D16FF81FFDEFEB73662FDD8D349" authority="Vilela &amp; Rodrigues &amp; Lencioni, 2022" authorityName="Vilela &amp; Rodrigues &amp; Lencioni" authorityYear="2022" box="[380,576,1548,1571]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="roquei" status="sp. nov.">
<emphasis id="B935EA87FF81FFDEFEB73662FDD8D349" box="[380,576,1548,1571]" italics="true" pageId="2" pageNumber="495">Heteragrion roquei</emphasis>
</taxonomicName>
<emphasis id="B935EA87FF81FFDEFD8C3665FD0FD349" bold="true" box="[583,663,1547,1571]" pageId="2" pageNumber="495">
<taxonomicNameLabel id="A20657FCFF81FFDEFD8C3665FD0FD349" box="[583,663,1547,1571]" pageId="2" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype.
</paragraph>
</caption>
<paragraph id="8BFE3695FF81FFDEFF0C363FFB48D3DE" blockId="2.[151,1437,1617,2040]" pageId="2" pageNumber="495">
<emphasis id="B935EA87FF81FFDEFF0C363FFEA9D301" box="[199,305,1617,1643]" italics="true" pageId="2" pageNumber="495">Abdomen</emphasis>
(
<figureCitation id="137A2A10FF81FFDEFE8A363FFE0AD306" box="[321,402,1617,1644]" captionStart="FIGURE 1" captionStartId="1.[151,250,1911,1935]" captionTargetBox="[151,1416,611,1860]" captionTargetId="figure-181@1.[151,1435,594,1885]" captionTargetPageId="1" captionText="FIGURE 1. Heteragrion roquei sp. nov. male holotype: (a) lateral view of habitus, (b) dorsal view of head and thorax; Heteragrion gracile Machado, 2006 male holotype: (c) lateral view of habitus." figureDoi="http://doi.org/10.5281/zenodo.7034052" httpUri="https://zenodo.org/record/7034052/files/figure.png" pageId="2" pageNumber="495">Fig. 1a</figureCitation>
). S1 mostly yellow, with a brown dorsal spot; S2 pale laterally, gradually becoming brown dorsally, with a yellow dorsal stripe; S3−6 mostly black with pale basal rings and pale spots on their 4/5; S7 black, becoming dark brown posteriorly; S8−9 dark brown; S10 black, becoming dark brown ventrally.
</paragraph>
<paragraph id="8BFE3695FF81FFDEFF0C36D3FE51D20D" blockId="2.[151,1437,1617,2040]" pageId="2" pageNumber="495">
<emphasis id="B935EA87FF81FFDEFF0C36D3FE13D3BD" box="[199,395,1725,1751]" italics="true" pageId="2" pageNumber="495">Anal appendages</emphasis>
(
<figureCitation id="137A2A10FF81FFDEFE5036D3FE73D3BD" box="[411,491,1725,1751]" captionStart="FIGURE 4" captionStartId="4.[151,250,1782,1806]" captionTargetBox="[187,1414,195,1744]" captionTargetId="figure-20@4.[161,1425,181,1758]" captionTargetPageId="4" captionText="FIGURE 4. Mediodorsal and lateral view of cercus of: (a−b) Heteragrion roquei sp. nov. male holotype, (c−d) Heteragrion triangulare Hagen in Selys, 1862 from Goiás state, (e−f) Heteragrion gracile Machado, 2006 from Espírito Santo state." figureDoi="http://doi.org/10.5281/zenodo.7034058" httpUri="https://zenodo.org/record/7034058/files/figure.png" pageId="2" pageNumber="495">Figs. 4</figureCitation>
a−b). Dark brown with darker coloration on the carinas and spines; BP subequal to MP; ventrobasal expansion (VbE) present, ‘C’ shaped in posterior view; dorsobasal expansion (DbE) broad, as large as VbE; MP roughly quadrangular, plate like, squared apex; sclerotized ridge of MP oblique, slightly curved, bearing at least six denticles, reaching MP posterior margin; AP the shortest, ending blunt tip curved inwards. Paraprocts greatly reduced.
</paragraph>
<paragraph id="8BFE3695FF81FFDEFF0C371CFCDFD2E6" blockId="2.[151,1437,1617,2040]" box="[199,839,1905,1932]" pageId="2" pageNumber="495">Measurements. FW: 27.3; HW: 26.5; AL: 41.7; TL: 50.1.</paragraph>
<paragraph id="8BFE3695FF81FFDEFF0C37FBFDF8D292" blockId="2.[151,1437,1617,2040]" pageId="2" pageNumber="495">
<emphasis id="B935EA87FF81FFDEFF0C37FBFE76D2DA" bold="true" box="[199,494,1941,1968]" pageId="2" pageNumber="495">
Variations in 
<typeStatus id="54FA8837FF81FFDEFEBB37F8FE70D2DA" box="[368,488,1942,1968]" pageId="2" pageNumber="495" type="paratype">paratypes</typeStatus>
.
</emphasis>
Male 
<typeStatus id="54FA8837FF81FFDEFDF537F8FD06D2DA" box="[574,670,1942,1968]" pageId="2" pageNumber="495" type="paratype">paratype</typeStatus>
do not vary in cerci morphology. Only slight thoracic coloration differences were spotted, probably related to aging. Male 
<typeStatus id="54FA8837FF81FFDEFCE437D4FC17D2BE" box="[815,911,1978,2004]" pageId="2" pageNumber="495" type="paratype">paratype</typeStatus>
varied in size as follows: FW: 27−27.3, HW: 25.9−26.5, AL: 40−41.7, Total: 48−50.1.
</paragraph>
<caption id="DF3E661DFF80FFDFFF5C37F9FF55D29D" ID-DOI="http://doi.org/10.5281/zenodo.7034056" ID-Zenodo-Dep="7034056" httpUri="https://zenodo.org/record/7034056/files/figure.png" pageId="3" pageNumber="496" startId="3.[151,250,1943,1967]" targetBox="[320,1276,190,1920]" targetPageId="3">
<paragraph id="8BFE3695FF80FFDFFF5C37F9FF55D29D" blockId="3.[151,1437,1943,2039]" pageId="3" pageNumber="496">
<emphasis id="B935EA87FF80FFDFFF5C37F9FE8FD2C5" bold="true" box="[151,279,1943,1967]" pageId="3" pageNumber="496">FIGURE 3.</emphasis>
Dorsal view of head and lateral view of thorax of: (a−b) 
<taxonomicName id="4C414D16FF80FFDFFC4637F6FBCED2C5" authority="Vilela &amp; Rodrigues &amp; Lencioni, 2022" authorityName="Vilela &amp; Rodrigues &amp; Lencioni" authorityYear="2022" box="[909,1110,1944,1967]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="roquei" status="sp. nov.">
<emphasis id="B935EA87FF80FFDFFC4637F6FBCED2C5" box="[909,1110,1944,1967]" italics="true" pageId="3" pageNumber="496">Heteragrion roquei</emphasis>
</taxonomicName>
<emphasis id="B935EA87FF80FFDFFBAB37F6FB2DD2DA" bold="true" box="[1120,1205,1944,1968]" pageId="3" pageNumber="496">
<taxonomicNameLabel id="A20657FCFF80FFDFFBAB37F6FB2DD2DA" box="[1120,1205,1944,1968]" pageId="3" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype, (c−d) 
<taxonomicName id="4C414D16FF80FFDFFF5C37D2FE4DD2BE" authority="Hagen" authorityName="Hagen" box="[151,469,1980,2004]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="triangulare">
<emphasis id="B935EA87FF80FFDFFF5C37D2FE15D2BE" box="[151,397,1980,2004]" italics="true" pageId="3" pageNumber="496">Heteragrion triangulare</emphasis>
Hagen
</taxonomicName>
in Selys, 1862 from Goiás state, (e−f) 
<taxonomicName id="4C414D16FF80FFDFFC9437D2FB57D2BE" authority="Machado, 2006" authorityName="Machado" authorityYear="2006" box="[863,1231,1980,2004]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B935EA87FF80FFDFFC9437D2FBB3D2BE" box="[863,1067,1980,2004]" italics="true" pageId="3" pageNumber="496">Heteragrion gracile</emphasis>
Machado, 2006
</taxonomicName>
from Espírito Santo state.
</paragraph>
</caption>
<caption id="DF3E661DFF87FFD8FF5C3698FACED258" ID-DOI="http://doi.org/10.5281/zenodo.7034058" ID-Zenodo-Dep="7034058" httpUri="https://zenodo.org/record/7034058/files/figure.png" pageId="4" pageNumber="497" startId="4.[151,250,1782,1806]" targetBox="[187,1414,195,1744]" targetPageId="4">
<paragraph id="8BFE3695FF87FFD8FF5C3698FACED258" blockId="4.[151,1436,1782,1843]" pageId="4" pageNumber="497">
<emphasis id="B935EA87FF87FFD8FF5C3698FE8CD264" bold="true" box="[151,276,1782,1806]" pageId="4" pageNumber="497">FIGURE 4.</emphasis>
Mediodorsal and lateral view of cercus of: (a−b) 
<taxonomicName id="4C414D16FF87FFD8FCD33699FC46D264" authority="Vilela &amp; Rodrigues &amp; Lencioni, 2022" authorityName="Vilela &amp; Rodrigues &amp; Lencioni" authorityYear="2022" box="[792,990,1783,1806]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="roquei" status="sp. nov.">
<emphasis id="B935EA87FF87FFD8FCD33699FC46D264" box="[792,990,1783,1806]" italics="true" pageId="4" pageNumber="497">Heteragrion roquei</emphasis>
</taxonomicName>
<emphasis id="B935EA87FF87FFD8FC2D3698FBA0D264" bold="true" box="[998,1080,1782,1806]" pageId="4" pageNumber="497">
<taxonomicNameLabel id="A20657FCFF87FFD8FC2D3698FBA0D264" box="[998,1080,1782,1806]" pageId="4" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype, (c−d) 
<taxonomicName id="4C414D16FF87FFD8FAD43699FECAD259" authority="Hagen" authorityName="Hagen" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="triangulare">
<emphasis id="B935EA87FF87FFD8FAD43699FE91D258" italics="true" pageId="4" pageNumber="497">Heteragrion triangulare</emphasis>
Hagen
</taxonomicName>
in Selys, 1862 from Goiás state, (e−f) 
<taxonomicName id="4C414D16FF87FFD8FD173775FBD4D258" authority="Machado, 2006" authorityName="Machado" authorityYear="2006" box="[732,1100,1818,1842]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B935EA87FF87FFD8FD173775FC30D258" box="[732,936,1818,1842]" italics="true" pageId="4" pageNumber="497">Heteragrion gracile</emphasis>
Machado, 2006
</taxonomicName>
from Espírito Santo state.
</paragraph>
</caption>
<paragraph id="8BFE3695FF87FFD8FF0C370EFE3AD210" blockId="4.[151,1436,1888,2023]" box="[199,418,1888,1914]" pageId="4" pageNumber="497">
<emphasis id="B935EA87FF87FFD8FF0C370EFEBDD210" bold="true" box="[199,293,1888,1914]" pageId="4" pageNumber="497">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35B651EFF87FFDAFF0C37EAFF68D741" lastPageId="6" lastPageNumber="499" pageId="4" pageNumber="497" type="diagnosis">
<paragraph id="8BFE3695FF87FFD9FF0C37EAFBD3D4C7" blockId="4.[151,1436,1888,2023]" lastBlockId="5.[151,1437,151,429]" lastPageId="5" lastPageNumber="498" pageId="4" pageNumber="497">
<emphasis id="B935EA87FF87FFD8FF0C37EAFE4CD2F4" bold="true" box="[199,468,1924,1950]" pageId="4" pageNumber="497">Differential diagnosis.</emphasis>
<taxonomicName id="4C414D16FF87FFD8FE2937EBFD5AD2F4" authorityName="Vilela &amp; Rodrigues &amp; Lencioni" authorityYear="2022" box="[482,706,1925,1950]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="roquei">
<emphasis id="B935EA87FF87FFD8FE2937EBFD5AD2F4" box="[482,706,1925,1950]" italics="true" pageId="4" pageNumber="497">Heteragrion roquei</emphasis>
</taxonomicName>
belongs to Lencioni’s Group A, composed of males lacking developed paraprocts (
<bibRefCitation id="EFD04B64FF87FFD8FE5337C6FDDCD2A8" author="Lencioni, F. A. A." box="[408,580,1960,1986]" pageId="4" pageNumber="497" pagination="1 - 80" refId="ref2713" refString="Lencioni, F. A. A. (2013) Diagnoses and discussion of the group 1 and 2 Brazilian species of Heteragrion, with descriptions of four new species (Odonata: Megapodagrionidae). Zootaxa, 3685 (1), 1 - 80. http: // dx. doi. org / 10.11646 / zootaxa. 3685.1.1" type="journal article" year="2013">Lencioni 2013</bibRefCitation>
). Based mainly on cercus morphology, and some coloration characters, the morphologically closest taxa of the new species are 
<taxonomicName id="4C414D16FF87FFD8FD2137A3FC46D28D" authority="Hagen" authorityName="Hagen" box="[746,990,1996,2023]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="triangulare">
<emphasis id="B935EA87FF87FFD8FD2137A3FC15D28C" box="[746,909,1996,2022]" italics="true" pageId="4" pageNumber="497">H. triangulare</emphasis>
Hagen
</taxonomicName>
<emphasis id="B935EA87FF87FFD8FC2C37A3FC65D28C" box="[999,1021,1997,2022]" italics="true" pageId="4" pageNumber="497">in</emphasis>
Selys (
<figureCitation id="137A2A10FF87FFD8FB9837A2FB07D28C" box="[1107,1183,1996,2022]" captionStart="FIGURE 3" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[320,1276,190,1920]" captionTargetId="figure-17@3.[311,1276,181,1920]" captionTargetPageId="3" captionText="FIGURE 3. Dorsal view of head and lateral view of thorax of: (a−b) Heteragrion roquei sp. nov. male holotype, (c−d) Heteragrion triangulare Hagen in Selys, 1862 from Goiás state, (e−f) Heteragrion gracile Machado, 2006 from Espírito Santo state." figureDoi="http://doi.org/10.5281/zenodo.7034056" httpUri="https://zenodo.org/record/7034056/files/figure.png" pageId="4" pageNumber="497">Figs. 3</figureCitation>
c−d, 4c−d), 1862, and 
<taxonomicName id="4C414D16FF86FFD9FF5C30F9FE52D5DB" authority="Machado, 2006" authorityName="Machado" authorityYear="2006" box="[151,458,151,177]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B935EA87FF86FFD9FF5C30F9FE94D5DB" box="[151,268,151,177]" italics="true" pageId="5" pageNumber="498">H. gracile</emphasis>
Machado, 2006
</taxonomicName>
(
<figureCitation id="137A2A10FF86FFD9FE1630F9FDA4D5DB" box="[477,572,151,177]" captionStart="FIGURE 1" captionStartId="1.[151,250,1911,1935]" captionTargetBox="[151,1416,611,1860]" captionTargetId="figure-181@1.[151,1435,594,1885]" captionTargetPageId="1" captionText="FIGURE 1. Heteragrion roquei sp. nov. male holotype: (a) lateral view of habitus, (b) dorsal view of head and thorax; Heteragrion gracile Machado, 2006 male holotype: (c) lateral view of habitus." figureDoi="http://doi.org/10.5281/zenodo.7034052" httpUri="https://zenodo.org/record/7034052/files/figure.png" pageId="5" pageNumber="498">Figs. 1c</figureCitation>
, 
<figureCitation id="137A2A10FF86FFD9FD8630F9FDC1D5DB" box="[589,601,151,177]" captionStart="FIGURE 3" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[320,1276,190,1920]" captionTargetId="figure-17@3.[311,1276,181,1920]" captionTargetPageId="3" captionText="FIGURE 3. Dorsal view of head and lateral view of thorax of: (a−b) Heteragrion roquei sp. nov. male holotype, (c−d) Heteragrion triangulare Hagen in Selys, 1862 from Goiás state, (e−f) Heteragrion gracile Machado, 2006 from Espírito Santo state." figureDoi="http://doi.org/10.5281/zenodo.7034056" httpUri="https://zenodo.org/record/7034056/files/figure.png" pageId="5" pageNumber="498">3</figureCitation>
e−f, 4e−f). Despite the resemblances, the new species can be separated from these congeners by the following character combination (see 
<tableCitation id="C6C3032EFF86FFD9FC4D30D5FC41D5BF" box="[902,985,187,213]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Comparative morphology of species close to H. roquei. All characters shown in Figure 4." httpUri="http://table.plazi.org/id/DF3E661DFF85FFDAFF5330F6FB55D5DB" pageId="5" pageNumber="498" tableUuid="DF3E661DFF85FFDAFF5330F6FB55D5DB">Table 1</tableCitation>
, and respective figures; other species in parentheses): ventrobasal expansion (VbE) present, ‘C’ shaped in posterior view (absent in 
<taxonomicName id="4C414D16FF86FFD9FB5330B1FAA2D593" box="[1176,1338,223,249]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="triangulare">
<emphasis id="B935EA87FF86FFD9FB5330B1FAA2D593" box="[1176,1338,223,249]" italics="true" pageId="5" pageNumber="498">H. triangulare</emphasis>
</taxonomicName>
, 
<figureCitation id="137A2A10FF86FFD9FA8C30B1FA0FD593" box="[1351,1431,223,249]" captionStart="FIGURE 4" captionStartId="4.[151,250,1782,1806]" captionTargetBox="[187,1414,195,1744]" captionTargetId="figure-20@4.[161,1425,181,1758]" captionTargetPageId="4" captionText="FIGURE 4. Mediodorsal and lateral view of cercus of: (a−b) Heteragrion roquei sp. nov. male holotype, (c−d) Heteragrion triangulare Hagen in Selys, 1862 from Goiás state, (e−f) Heteragrion gracile Machado, 2006 from Espírito Santo state." figureDoi="http://doi.org/10.5281/zenodo.7034058" httpUri="https://zenodo.org/record/7034058/files/figure.png" pageId="5" pageNumber="498">Fig. 4c</figureCitation>
, who has an apophysis; larger in 
<taxonomicName id="4C414D16FF86FFD9FDCC316DFDE2D477" authorityName="Machado" authorityYear="2006" box="[519,634,259,285]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B935EA87FF86FFD9FDCC316DFDE2D477" box="[519,634,259,285]" italics="true" pageId="5" pageNumber="498">H. gracile</emphasis>
</taxonomicName>
, 
<figureCitation id="137A2A10FF86FFD9FD41316DFD44D477" box="[650,732,259,285]" captionStart="FIGURE 4" captionStartId="4.[151,250,1782,1806]" captionTargetBox="[187,1414,195,1744]" captionTargetId="figure-20@4.[161,1425,181,1758]" captionTargetPageId="4" captionText="FIGURE 4. Mediodorsal and lateral view of cercus of: (a−b) Heteragrion roquei sp. nov. male holotype, (c−d) Heteragrion triangulare Hagen in Selys, 1862 from Goiás state, (e−f) Heteragrion gracile Machado, 2006 from Espírito Santo state." figureDoi="http://doi.org/10.5281/zenodo.7034058" httpUri="https://zenodo.org/record/7034058/files/figure.png" pageId="5" pageNumber="498">Fig. 4e</figureCitation>
); dorsobasal expansion (DbE) broad, as large as VbE (narrow in 
<taxonomicName id="4C414D16FF86FFD9FF7E3149FEB0D42B" authorityName="Machado" authorityYear="2006" box="[181,296,295,321]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B935EA87FF86FFD9FF7E3149FEB0D42B" box="[181,296,295,321]" italics="true" pageId="5" pageNumber="498">H. gracile</emphasis>
</taxonomicName>
, 
<figureCitation id="137A2A10FF86FFD9FEF33149FE12D42B" box="[312,394,295,321]" captionStart="FIGURE 4" captionStartId="4.[151,250,1782,1806]" captionTargetBox="[187,1414,195,1744]" captionTargetId="figure-20@4.[161,1425,181,1758]" captionTargetPageId="4" captionText="FIGURE 4. Mediodorsal and lateral view of cercus of: (a−b) Heteragrion roquei sp. nov. male holotype, (c−d) Heteragrion triangulare Hagen in Selys, 1862 from Goiás state, (e−f) Heteragrion gracile Machado, 2006 from Espírito Santo state." figureDoi="http://doi.org/10.5281/zenodo.7034058" httpUri="https://zenodo.org/record/7034058/files/figure.png" pageId="5" pageNumber="498">Fig. 4e</figureCitation>
); single MP ridge (two ridges in 
<taxonomicName id="4C414D16FF86FFD9FCCF3149FCEED42B" authorityName="Machado" authorityYear="2006" box="[772,886,295,321]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B935EA87FF86FFD9FCCF3149FCEED42B" box="[772,886,295,321]" italics="true" pageId="5" pageNumber="498">H. gracile</emphasis>
</taxonomicName>
, 
<figureCitation id="137A2A10FF86FFD9FC4D3149FC41D42B" box="[902,985,295,321]" captionStart="FIGURE 4" captionStartId="4.[151,250,1782,1806]" captionTargetBox="[187,1414,195,1744]" captionTargetId="figure-20@4.[161,1425,181,1758]" captionTargetPageId="4" captionText="FIGURE 4. Mediodorsal and lateral view of cercus of: (a−b) Heteragrion roquei sp. nov. male holotype, (c−d) Heteragrion triangulare Hagen in Selys, 1862 from Goiás state, (e−f) Heteragrion gracile Machado, 2006 from Espírito Santo state." figureDoi="http://doi.org/10.5281/zenodo.7034058" httpUri="https://zenodo.org/record/7034058/files/figure.png" pageId="5" pageNumber="498">Fig. 4e</figureCitation>
); ridge reaching MP apex (not reaching apex in 
<taxonomicName id="4C414D16FF86FFD9FF3A3125FE0BD40F" box="[241,403,331,357]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="triangulare">
<emphasis id="B935EA87FF86FFD9FF3A3125FE0BD40F" box="[241,403,331,357]" italics="true" pageId="5" pageNumber="498">H. triangulare</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FF86FFD9FE013125FDA4D40F" authorityName="Machado" authorityYear="2006" box="[458,572,331,357]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B935EA87FF86FFD9FE013125FDA4D40F" box="[458,572,331,357]" italics="true" pageId="5" pageNumber="498">H. gracile</emphasis>
</taxonomicName>
). The new species can readily be separated from 
<taxonomicName id="4C414D16FF86FFD9FBA03125FA94D40F" box="[1131,1292,331,357]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="triangulare">
<emphasis id="B935EA87FF86FFD9FBA03125FA94D40F" box="[1131,1292,331,357]" italics="true" pageId="5" pageNumber="498">H. triangulare</emphasis>
</taxonomicName>
because this species presents an apophysis on the base of the cercus (
<figureCitation id="137A2A10FF86FFD9FCCC3101FCCFD4E3" box="[775,855,367,393]" captionStart="FIGURE 4" captionStartId="4.[151,250,1782,1806]" captionTargetBox="[187,1414,195,1744]" captionTargetId="figure-20@4.[161,1425,181,1758]" captionTargetPageId="4" captionText="FIGURE 4. Mediodorsal and lateral view of cercus of: (a−b) Heteragrion roquei sp. nov. male holotype, (c−d) Heteragrion triangulare Hagen in Selys, 1862 from Goiás state, (e−f) Heteragrion gracile Machado, 2006 from Espírito Santo state." figureDoi="http://doi.org/10.5281/zenodo.7034058" httpUri="https://zenodo.org/record/7034058/files/figure.png" pageId="5" pageNumber="498">Fig. 4c</figureCitation>
, VbA), instead of an excavated VbE. The remainder combination of characters should be sufficient to separate 
<taxonomicName id="4C414D16FF86FFD9FCEF31FDFC16D4C6" authorityName="Vilela &amp; Rodrigues &amp; Lencioni" authorityYear="2022" box="[804,910,403,428]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="roquei">
<emphasis id="B935EA87FF86FFD9FCEF31FDFC16D4C6" box="[804,910,403,428]" italics="true" pageId="5" pageNumber="498">H. roquei</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C414D16FF86FFD9FC1931FDFBDCD4C7" authorityName="Machado" authorityYear="2006" box="[978,1092,403,429]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B935EA87FF86FFD9FC1931FDFBDCD4C7" box="[978,1092,403,429]" italics="true" pageId="5" pageNumber="498">H. gracile</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF3E661DFF86FFD9FF5C37D4FC63D29C" ID-DOI="http://doi.org/10.5281/zenodo.7034062" ID-Zenodo-Dep="7034062" httpUri="https://zenodo.org/record/7034062/files/figure.png" pageId="5" pageNumber="498" startId="5.[151,250,1978,2002]" targetBox="[227,1361,453,1954]" targetPageId="5">
<paragraph id="8BFE3695FF86FFD9FF5C37D4FC63D29C" blockId="5.[151,1436,1978,2038]" pageId="5" pageNumber="498">
<emphasis id="B935EA87FF86FFD9FF5C37D4FE8FD2B8" bold="true" box="[151,279,1978,2002]" pageId="5" pageNumber="498">FIGURE 5.</emphasis>
(a−c) Habitat of 
<taxonomicName id="4C414D16FF86FFD9FE1D37D5FD06D2B8" authority="Vilela &amp; Rodrigues &amp; Lencioni, 2022" authorityName="Vilela &amp; Rodrigues &amp; Lencioni" authorityYear="2022" box="[470,670,1979,2002]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="roquei" status="sp. nov.">
<emphasis id="B935EA87FF86FFD9FE1D37D5FD06D2B8" box="[470,670,1979,2002]" italics="true" pageId="5" pageNumber="498">Heteragrion roquei</emphasis>
</taxonomicName>
<emphasis id="B935EA87FF86FFD9FD6237D4FD65D2B8" bold="true" box="[681,765,1978,2002]" pageId="5" pageNumber="498">
<taxonomicNameLabel id="A20657FCFF86FFD9FD6237D4FD65D2B8" box="[681,765,1978,2002]" pageId="5" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Fazenda Araruna, Una municipality, Bahia state, Brazil; (d) 
<taxonomicName id="4C414D16FF86FFD9FF5C37B1FEC4D29C" authority="Vilela &amp; Rodrigues &amp; Lencioni, 2022" authorityName="Vilela &amp; Rodrigues &amp; Lencioni" authorityYear="2022" box="[151,348,2015,2038]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="roquei" status="sp. nov.">
<emphasis id="B935EA87FF86FFD9FF5C37B1FEC4D29C" box="[151,348,2015,2038]" italics="true" pageId="5" pageNumber="498">Heteragrion roquei</emphasis>
</taxonomicName>
<emphasis id="B935EA87FF86FFD9FEA937B0FE2BD29C" bold="true" box="[354,435,2014,2038]" pageId="5" pageNumber="498">
<taxonomicNameLabel id="A20657FCFF86FFD9FEA937B0FE2BD29C" box="[354,435,2014,2038]" pageId="5" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype perched on leaves from its natural habitat.
</paragraph>
</caption>
<caption id="DF3E661DFF85FFDAFF5330F6FB55D5DB" ID-Table-UUID="DF3E661DFF85FFDAFF5330F6FB55D5DB" box="[152,1229,152,177]" httpUri="http://table.plazi.org/id/DF3E661DFF85FFDAFF5330F6FB55D5DB" pageId="6" pageNumber="499" startId="6.[152,244,152,176]" targetBox="[166,1326,192,337]" targetIsTable="true" targetPageId="6">
<paragraph id="8BFE3695FF85FFDAFF5330F6FB55D5DB" blockId="6.[152,1229,152,177]" box="[152,1229,152,177]" pageId="6" pageNumber="499">
<emphasis id="B935EA87FF85FFDAFF5330F6FE97D5DA" bold="true" box="[152,271,152,176]" pageId="6" pageNumber="499">TABLE 1.</emphasis>
Comparative morphology of species close to 
<taxonomicName id="4C414D16FF85FFDAFD3330F7FCC6D5DA" authorityName="Vilela &amp; Rodrigues &amp; Lencioni" authorityYear="2022" box="[760,862,153,176]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="roquei">
<emphasis id="B935EA87FF85FFDAFD3330F7FCC6D5DA" box="[760,862,153,176]" italics="true" pageId="6" pageNumber="499">H. roquei</emphasis>
</taxonomicName>
. All characters shown in Figure 4.
</paragraph>
</caption>
<paragraph id="8BFE3695FF85FFDAFF6D30AEFA90D43B" pageId="6" pageNumber="499">
<table id="F941C435FF850023FF6D30AEFAB6D43B" box="[166,1326,192,337]" gridcols="4" gridrows="4" pageId="6" pageNumber="499">
<tr id="357134D7FF850023FF6D30AEFAB6D5B2" box="[166,1326,192,216]" gridrow="0" pageId="6" pageNumber="499">
<th id="76A05DABFF850023FF6D30AEFEAFD5B2" box="[166,311,192,216]" gridcol="0" gridrow="0" pageId="6" pageNumber="499">
<emphasis id="B935EA87FF85FFDAFF6D30AEFF6DD5B2" bold="true" box="[166,245,192,216]" pageId="6" pageNumber="499">Species</emphasis>
</th>
<th id="76A05DABFF850023FE4330AEFE4DD5B2" box="[392,469,192,216]" gridcol="1" gridrow="0" pageId="6" pageNumber="499">
<emphasis id="B935EA87FF85FFDAFE4330AEFE21D5B2" bold="true" box="[392,441,192,216]" pageId="6" pageNumber="499">DbE</emphasis>
</th>
<th id="76A05DABFF850023FD9A30AEFC1CD5B2" box="[593,900,192,216]" gridcol="2" gridrow="0" pageId="6" pageNumber="499">
<emphasis id="B935EA87FF85FFDAFD9A30AEFD24D5B2" bold="true" box="[593,700,192,216]" pageId="6" pageNumber="499">VbE/VbA</emphasis>
</th>
<th id="76A05DABFF850023FC1030AEFAB6D5B2" box="[987,1326,192,216]" gridcol="3" gridrow="0" pageId="6" pageNumber="499">
<emphasis id="B935EA87FF85FFDAFC1030AEFB21D5B2" bold="true" box="[987,1209,192,216]" pageId="6" pageNumber="499">MP sclerotized ridge</emphasis>
</th>
</tr>
<tr id="357134D7FF850023FF6D3086FAB6D46A" box="[166,1326,232,256]" gridrow="1" pageId="6" pageNumber="499">
<th id="76A05DABFF850023FF6D3086FEAFD46A" box="[166,311,232,256]" gridcol="0" gridrow="1" pageId="6" pageNumber="499">
<emphasis id="B935EA87FF85FFDAFF6D3087FEAFD46A" box="[166,311,232,256]" italics="true" pageId="6" pageNumber="499">H. triangulare</emphasis>
</th>
<td id="76A05DABFF850023FE433086FE4DD46A" box="[392,469,232,256]" gridcol="1" gridrow="1" pageId="6" pageNumber="499">Broad</td>
<td id="76A05DABFF850023FD9A3086FC1CD46A" box="[593,900,232,256]" gridcol="2" gridrow="1" pageId="6" pageNumber="499">VbA present</td>
<td id="76A05DABFF850023FC103086FAB6D46A" box="[987,1326,232,256]" gridcol="3" gridrow="1" pageId="6" pageNumber="499">Single ridge, not reaching margin</td>
</tr>
<tr id="357134D7FF850023FF6D317FFAB6D443" box="[166,1326,273,297]" gridrow="2" pageId="6" pageNumber="499">
<th id="76A05DABFF850023FF6D317FFEAFD443" box="[166,311,273,297]" gridcol="0" gridrow="2" pageId="6" pageNumber="499">
<emphasis id="B935EA87FF85FFDAFF6D317CFE95D443" box="[166,269,273,297]" italics="true" pageId="6" pageNumber="499">H. gracile</emphasis>
</th>
<td id="76A05DABFF850023FE43317FFE4DD443" box="[392,469,273,297]" gridcol="1" gridrow="2" pageId="6" pageNumber="499">Narrow</td>
<td id="76A05DABFF850023FD9A317FFC1CD443" box="[593,900,273,297]" gridcol="2" gridrow="2" pageId="6" pageNumber="499">VbE present, broad</td>
<td id="76A05DABFF850023FC10317FFAB6D443" box="[987,1326,273,297]" gridcol="3" gridrow="2" pageId="6" pageNumber="499">Two ridges, not reaching margin</td>
</tr>
<tr id="357134D7FF850023FF6D3157FAB6D43B" box="[166,1326,313,337]" gridrow="3" pageId="6" pageNumber="499">
<th id="76A05DABFF850023FF6D3157FEAFD43B" box="[166,311,313,337]" gridcol="0" gridrow="3" pageId="6" pageNumber="499">
<emphasis id="B935EA87FF85FFDAFF6D3154FE9ED43B" box="[166,262,314,337]" italics="true" pageId="6" pageNumber="499">H. roquei</emphasis>
</th>
<td id="76A05DABFF850023FE433157FE4DD43B" box="[392,469,313,337]" gridcol="1" gridrow="3" pageId="6" pageNumber="499">Broad</td>
<td id="76A05DABFF850023FD9A3157FC1CD43B" box="[593,900,313,337]" gridcol="2" gridrow="3" pageId="6" pageNumber="499">VbE present, subequal to DbE</td>
<td id="76A05DABFF850023FC103157FAB6D43B" box="[987,1326,313,337]" gridcol="3" gridrow="3" pageId="6" pageNumber="499">Single ridge, reaching margin</td>
</tr>
</table>
</paragraph>
<paragraph id="8BFE3695FF85FFDAFF0C31EEFF68D741" blockId="6.[151,1436,384,555]" pageId="6" pageNumber="499">
<emphasis id="B935EA87FF85FFDAFF0C31EEFE20D4F1" bold="true" box="[199,440,384,411]" pageId="6" pageNumber="499">Habitat and ecology.</emphasis>
At the 
<typeStatus id="54FA8837FF85FFDAFDCE31EFFDADD4F1" box="[517,565,385,411]" pageId="6" pageNumber="499">type</typeStatus>
locality (
<figureCitation id="137A2A10FF85FFDAFD5031EFFCBCD4F1" box="[667,804,384,411]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="5.[151,250,1978,2002]" captionStartId-1="6.[151,250,1774,1798]" captionTargetBox-0="[227,1361,453,1954]" captionTargetBox-1="[227,1361,579,1751]" captionTargetId-0="figure-165@5.[227,1361,453,1954]" captionTargetId-1="figure-164@6.[227,1361,579,1751]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 5. (a−c) Habitat of Heteragrion roquei sp. nov. in Fazenda Araruna, Una municipality, Bahia state, Brazil; (d) Heteragrion roquei sp. nov. male holotype perched on leaves from its natural habitat." captionText-1="FIGURE 6. Map showing Bahia state in Brazil (bottom left in red), then the Una municipality in Bahia state (highlighted in red). Adapted from: https://cutt.ly/6JgZ47t." figureDoi-0="http://doi.org/10.5281/zenodo.7034062" figureDoi-1="http://doi.org/10.5281/zenodo.7034064" httpUri-0="https://zenodo.org/record/7034062/files/figure.png" httpUri-1="https://zenodo.org/record/7034064/files/figure.png" pageId="6" pageNumber="499">Figures 5−6</figureCitation>
), 
<taxonomicName id="4C414D16FF85FFDAFCFC31EFFB96D4F0" authority="Vilela &amp; Rodrigues &amp; Lencioni, 2022" authorityName="Vilela &amp; Rodrigues &amp; Lencioni" authorityYear="2022" box="[823,1038,385,410]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="roquei" status="sp. nov.">
<emphasis id="B935EA87FF85FFDAFCFC31EFFB96D4F0" box="[823,1038,385,410]" italics="true" pageId="6" pageNumber="499">Heteragrion roquei</emphasis>
</taxonomicName>
<emphasis id="B935EA87FF85FFDAFBD831EFFBF3D4F1" bold="true" box="[1043,1131,385,411]" pageId="6" pageNumber="499">
<taxonomicNameLabel id="A20657FCFF85FFDAFBD831EFFBF3D4F1" box="[1043,1131,385,411]" pageId="6" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
inhabits lotic environments placed within gallery forests with plenty of shaded areas, where they can be spotted on leaves exposed to the sunlight. Such habitats are common places to find 
<taxonomicName id="4C414D16FF85FFDAFD1431A7FC5CD489" box="[735,964,457,483]" pageId="6" pageNumber="499">
<emphasis id="B935EA87FF85FFDAFD1431A7FCF1D488" box="[735,873,457,482]" italics="true" pageId="6" pageNumber="499">Heteragrion</emphasis>
species
</taxonomicName>
(Ávila-Jr. 
<emphasis id="B935EA87FF85FFDAFB8A31A7FBEDD489" box="[1089,1141,457,483]" italics="true" pageId="6" pageNumber="499">et al</emphasis>
. 2017, 2020; 
<bibRefCitation id="EFD04B64FF85FFDAFADD31A7FF47D76D" author="Vilela, D. S. &amp; Farias, A. B. S. &amp; Santos, J. C." pageId="6" pageNumber="499" pagination="476 - 482" refId="ref3295" refString="Vilela, D. S., Farias, A. B. S. &amp; Santos, J. C. (2021 a) Heteragrion lencionii (Odonata: Heteragrionidae) sp. nov. from Serra de Itabaiana National Park, Northeastern Brazil. Zootaxa, 4966 (4), 476 - 482. https: // doi. org / 10.11646 / zootaxa. 4966.4.6" type="journal article" year="2021">
Vilela 
<emphasis id="B935EA87FF85FFDAFAA931A7FA0DD489" box="[1378,1429,457,483]" italics="true" pageId="6" pageNumber="499">et al</emphasis>
. 2021a
</bibRefCitation>
; 
<bibRefCitation id="EFD04B64FF85FFDAFF203183FDA5D76D" author="Mendoza-Penagos, C. C. &amp; Juen, L. &amp; Vilela, D. S." box="[235,573,493,519]" pageId="6" pageNumber="499" pagination="223 - 229" refId="ref2928" refString="Mendoza-Penagos, C. C., Juen, L. &amp; Vilela, D. S. (2022) Heteragrion calafatiensis (Odonata: Heteragrionidae) sp. nov. from Northern Brazil. Zootaxa, 5124 (2), 223 - 229. https: // doi. org / 10.11646 / zootaxa. 5124.2.7" type="journal article" year="2022">
Mendoza-Penagos 
<emphasis id="B935EA87FF85FFDAFE0A3183FE62D76D" box="[449,506,493,519]" italics="true" pageId="6" pageNumber="499">et al.</emphasis>
2022
</bibRefCitation>
), and in that same locality 
<taxonomicName id="4C414D16FF85FFDAFCA63183FBAFD76D" authority="Hagen" authorityName="Hagen" box="[877,1079,493,519]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="consors">
<emphasis id="B935EA87FF85FFDAFCA63183FC7ED76C" box="[877,998,493,518]" italics="true" pageId="6" pageNumber="499">H. consors</emphasis>
Hagen
</taxonomicName>
<emphasis id="B935EA87FF85FFDAFBF43183FBCDD76C" box="[1087,1109,493,518]" italics="true" pageId="6" pageNumber="499">in</emphasis>
Selys, 1862 was a sympatric species.
</paragraph>
</subSubSection>
</treatment>
</document>