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<document ID-DOI="http://doi.org/10.5281/zenodo.3792106" ID-GBIF-Dataset="09827415-df1c-4129-9bc1-39321f933d5d" ID-GBIF-Taxon="163794397" ID-ISSN="13132970" ID-Zenodo-Dep="3792106" ID-ZooBank="531CF885-FED5-4B1B-806B-02C9E5051929" LSID="urn:lsid:zoobank.org:act:CBC97796-8CBD-4AFF-B9CC-95F4FFD3232C" checkinTime="1588009417727" checkinUser="plazi" docAuthor="Giupponi, Alessandro, de Vasconcelos, Eduardo &amp; Lourenco, Wilson" docDate="2009" docId="03DC0060FF823C74DC8CFCF05B88FAE0" docLanguage="en" docName="ZK_article_1998.pdf" docOrigin="ZooKeys 13 (13)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@18c8ecf5" docStyleName="zookeys.2008.journal_article" docTitle="Ananteris chagasi Giupponi &amp; de Vasconcelos &amp; Lourenco 2009, sp. n." docType="treatment" docUuid="CBC97796-8CBD-4AFF-B9CC-95F4FFD3232C" docUuidSource="ZooBank" docVersion="9" lastPageId="4" lastPageNumber="33" masterDocId="FFE57818FF803C70DC00FFB35804FFC7" masterDocTitle="The genus Ananteris Thorell, 1891 (Scorpiones, Buthidae) in southeast Brazil, with the description of three new species" masterLastPageNumber="41" masterPageNumber="29" pageId="2" pageNumber="31" updateTime="1636545320677" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The genus Ananteris Thorell, 1891 (Scorpiones, Buthidae) in southeast Brazil, with the description of three new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Giupponi, Alessandro</mods:namePart>
<mods:affiliation>,, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>de Vasconcelos, Eduardo</mods:namePart>
<mods:affiliation>Museu Nacional - UFRJ, Rio de Janeiro ,, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lourenco, Wilson</mods:namePart>
<mods:affiliation>Museum national d Histoire naturelle, Paris, France</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-07-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>13</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>13</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>29</mods:start>
<mods:end>41</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.13.125</mods:identifier>
<mods:identifier type="GBIF-Dataset">09827415-df1c-4129-9bc1-39321f933d5d</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576464</mods:identifier>
<mods:identifier type="ZooBank">531CF885-FED5-4B1B-806B-02C9E5051929</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792106" ID-GBIF-Taxon="163794397" ID-Zenodo-Dep="3792106" LSID="urn:lsid:zoobank.org:act:CBC97796-8CBD-4AFF-B9CC-95F4FFD3232C" httpUri="http://treatment.plazi.org/id/03DC0060FF823C74DC8CFCF05B88FAE0" lastPageId="4" lastPageNumber="33" pageId="2" pageNumber="31">
<subSubSection box="[140,421,835,862]" pageId="2" pageNumber="31" type="nomenclature">
<paragraph blockId="2.[140,997,835,932]" box="[140,421,835,862]" pageId="2" pageNumber="31">
<heading box="[140,421,835,862]" fontSize="11" level="2" pageId="2" pageNumber="31" reason="3">
<emphasis bold="true" box="[140,421,835,862]" pageId="2" pageNumber="31">
<taxonomicName authority="Giupponi &amp; de Vasconcelos &amp; Lourenco, 2009" authorityName="Giupponi &amp; de Vasconcelos &amp; Lourenco" authorityYear="2009" box="[140,348,835,862]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="chagasi" status="sp. n.">
<emphasis bold="true" box="[140,348,835,862]" italics="true" pageId="2" pageNumber="31">
<emphasis box="[140,255,835,861]" inLineHeading="true" pageId="2" pageNumber="31" reason="1">Ananteris</emphasis>
chagasi
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[356,421,835,861]" pageId="2" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="31" type="description">
<paragraph blockId="2.[140,997,835,932]" box="[140,997,870,897]" pageId="2" pageNumber="31">
<uri box="[140,997,870,897]" pageId="2" pageNumber="31">
urn:lsid:zoobank.org:act:
<uuid box="[417,997,870,897]" pageId="2" pageNumber="31">CBC97796-8CBD-4AFF-B9CC-95F4FFD3232C</uuid>
</uri>
</paragraph>
<paragraph blockId="2.[140,997,835,932]" box="[140,274,906,932]" pageId="2" pageNumber="31">
<figureCitation box="[140,232,906,932]" captionStart="Figures 1-9" captionStartId="3.[140,222,1538,1560]" captionTargetBox="[159,1102,165,1507]" captionTargetId="figure@3.[140,1108,165,1519]" captionTargetPageId="3" captionText="Figures 1-9. Ananteris chagasi sp. n., male holotype. 1 Carapace dorsal view. 2 Pectines. 3 Chelicera, dorsal view. 4 Metasoma, segments IV-V and telson, lateral view. 5 Chela, dorsal view. 6 Manus, ventral view. 7 Pedipalpal femur, dorsal view. 8-9 Pedipalpal patella, dorsal and external view. Scale bars: 1-4 and 6-9 = 1mm; 5 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3814813" httpUri="https://zenodo.org/record/3814813/files/figure.png" pageId="2" pageNumber="31">Figs 1-9</figureCitation>
,
<figureCitation box="[245,274,906,932]" captionStart="Figure 28" captionStartId="7.[139,211,1576,1598]" captionTargetId="figure@7.[139,1108,165,1556]" captionTargetPageId="7" captionText="Figure 28. Map of distribution of the species of the genus Ananteris in Brazil." figureDoi="http://doi.org/10.5281/zenodo.3814817" httpUri="https://zenodo.org/record/3814817/files/figure.png" pageId="2" pageNumber="31">28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="31" type="materials_examined">
<paragraph blockId="2.[140,1108,976,1671]" pageId="2" pageNumber="31">
<emphasis bold="true" box="[140,316,976,1002]" pageId="2" pageNumber="31">
<emphasis box="[140,198,976,1002]" inLineHeading="true" pageId="2" pageNumber="31" reason="1">Type</emphasis>
material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608997649" collectingDate="2006-05-26" collectorName="Male. &amp; State &amp; Itacarambi &amp; Olhos &amp; Giupponi &amp; Chagas-Junior &amp; Baptista &amp; Silvia" country="Brazil" location="Museu Nacional" municipality="Deposited" pageId="2" pageNumber="31" specimenCount="1" stateProvince="Minas Gerais" typeStatus="holotype">
<emphasis bold="true" box="[328,527,976,1002]" pageId="2" pageNumber="31">
<typeStatus box="[328,444,976,1002]" pageId="2" pageNumber="31">Holotype</typeStatus>
.
<collectorName box="[460,527,976,1002]" pageId="2" pageNumber="31">Male.</collectorName>
</emphasis>
(sub-adult).
<collectingCountry box="[684,752,976,1002]" name="Brazil" pageId="2" pageNumber="31">Brazil</collectingCountry>
,
<collectorName box="[768,823,976,1002]" pageId="2" pageNumber="31">State</collectorName>
of
<collectingRegion box="[870,1028,976,1002]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="31">Minas Gerais</collectingRegion>
,
<collectorName pageId="2" pageNumber="31">Itacarambi</collectorName>
, around
<collectorName box="[320,389,1011,1037]" pageId="2" pageNumber="31">Olhos</collectorName>
dágua cave (
<collectorName box="[555,670,1011,1037]" pageId="2" pageNumber="31">Giupponi</collectorName>
,
<collectorName box="[687,851,1011,1037]" pageId="2" pageNumber="31">Chagas-Junior</collectorName>
,
<collectorName box="[867,961,1011,1037]" pageId="2" pageNumber="31">Baptista</collectorName>
,
<collectorName box="[978,1039,1011,1037]" pageId="2" pageNumber="31">Silvia</collectorName>
leg.),
<date box="[140,271,1046,1072]" pageId="2" pageNumber="31" value="2006-05-26">
<collectingDate box="[140,271,1046,1072]" pageId="2" pageNumber="31" value="2006-05-26">26/V/2006</collectingDate>
</date>
.
<collectingMunicipality box="[284,400,1046,1072]" pageId="2" pageNumber="31">Deposited</collectingMunicipality>
in the arachnological collection of the
<location LSID="urn:lsid:plazi:treatment:03DC0060FF823C74DC8CFCF05B88FAE0:8EAAE7ADFF823C72DF5DFBA45C18FBF7" box="[861,1052,1046,1073]" country="Brazil" municipality="Deposited" name="Museu Nacional" pageId="2" pageNumber="31" stateProvince="Minas Gerais">Museu Nacional</location>
,
<collectingRegion country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="31">Rio de Janeiro</collectingRegion>
,
<collectingCountry box="[268,337,1082,1108]" name="Brazil" pageId="2" pageNumber="31">Brazil</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="31" type="diagnosis">
<paragraph blockId="2.[140,1108,976,1671]" pageId="2" pageNumber="31">
<emphasis bold="true" box="[185,304,1117,1143]" inLineHeading="true" pageId="2" pageNumber="31" reason="1">Diagnosis</emphasis>
. Species of small to moderate size when compared with the average size of the other species of the genus (
<quantity box="[511,616,1152,1179]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="2" pageNumber="31" unit="mm" value="16.3">16.3 mm</quantity>
in total length; see
<tableCitation box="[832,912,1152,1179]" captionStart="Table I" captionStartId="8.[140,200,166,188]" captionTargetBox="[147,1074,211,1131]" captionTargetPageId="8" captionText="Table I. Morphometric values (in mm) of the new Ananteris species described in this paper" httpUri="http://table.plazi.org/id/DF0AE1FEFF883C78DC8CFF155BD8FF7C" pageId="2" pageNumber="31" tableUuid="DF0AE1FEFF883C78DC8CFF155BD8FF7C">Table I</tableCitation>
). General coloration yellowish with intense variegated dark pigmentation. Pedipalps short and slender; fingers with 6 rows of granules; male pectines with 15-15 teeth. Th e new species can be distinguished from other known species of the genus, and in particular from
<taxonomicName authorityName="Thorell" authorityYear="1891" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="balzanii">
<emphasis italics="true" pageId="2" pageNumber="31">A. balzanii</emphasis>
</taxonomicName>
, which is also distributed in the State of
<collectingRegion box="[637,788,1293,1319]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="31">Minas Gerais</collectingRegion>
by: (i) a different pattern of pigmentation; fingers of pedipalp chela in
<taxonomicName authorityName="Thorell" authorityYear="1891" box="[613,734,1328,1354]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="balzanii">
<emphasis box="[613,734,1328,1354]" italics="true" pageId="2" pageNumber="31">A. balzanii</emphasis>
</taxonomicName>
are yellow, whereas these are very dark in the new species; chelicerae in
<taxonomicName authorityName="Thorell" authorityYear="1891" box="[565,687,1363,1390]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="balzanii">
<emphasis box="[565,687,1363,1390]" italics="true" pageId="2" pageNumber="31">A. balzanii</emphasis>
</taxonomicName>
are totally covered by reticular spots, whereas in the new species these cover only the anterior and lateral edges, (ii) pedipalp fingers with have 7 rows of granules in
<taxonomicName authorityName="Thorell" authorityYear="1891" box="[588,711,1434,1460]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="balzanii">
<emphasis box="[588,711,1434,1460]" italics="true" pageId="2" pageNumber="31">A. balzanii</emphasis>
</taxonomicName>
, against only
<quantity box="[871,917,1434,1460]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="31" unit="in" value="6.0">6 in</quantity>
the new species, (iii) male pectines with 15 teeth, in contrast to
<quantity box="[670,768,1469,1495]" metricMagnitude="-1" metricUnit="m" metricValue="4.445" metricValueMax="4.826" metricValueMin="4.064" pageId="2" pageNumber="31" unit="in" value="17.5" valueMax="19.0" valueMin="16.0">16-19 in</quantity>
<taxonomicName authorityName="Thorell" authorityYear="1891" box="[775,897,1469,1495]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="balzanii">
<emphasis box="[775,897,1469,1495]" italics="true" pageId="2" pageNumber="31">A. balzanii</emphasis>
</taxonomicName>
, (iv) some distinct morphometric values (see
<tableCitation box="[435,516,1504,1531]" captionStart="Table I" captionStartId="8.[140,200,166,188]" captionTargetBox="[147,1074,211,1131]" captionTargetPageId="8" captionText="Table I. Morphometric values (in mm) of the new Ananteris species described in this paper" httpUri="http://table.plazi.org/id/DF0AE1FEFF883C78DC8CFF155BD8FF7C" pageId="2" pageNumber="31" tableUuid="DF0AE1FEFF883C78DC8CFF155BD8FF7C">Table I</tableCitation>
). Th e new species is possibly an endemic element to the state of
<collectingRegion box="[271,425,1539,1565]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="31">Minas Gerais</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="31" type="etymology">
<paragraph blockId="2.[140,1108,976,1671]" pageId="2" pageNumber="31">
<emphasis bold="true" box="[185,320,1575,1601]" pageId="2" pageNumber="31">
<emphasis box="[185,315,1575,1601]" inLineHeading="true" pageId="2" pageNumber="31" reason="1">Etymology</emphasis>
.
</emphasis>
Patronym is in honor of Dr. Amazonas Chagas Júnior. of the Museu Nacional,
<collectingRegion box="[256,421,1610,1636]" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="31">Rio de Janeiro</collectingRegion>
,
<collectingCountry box="[433,502,1610,1636]" name="Brazil" pageId="2" pageNumber="31">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="33" pageId="2" pageNumber="31" type="description">
<paragraph blockId="2.[140,1108,976,1671]" box="[185,921,1645,1671]" pageId="2" pageNumber="31">
<emphasis bold="true" box="[185,335,1645,1671]" pageId="2" pageNumber="31">
<emphasis box="[185,330,1645,1671]" inLineHeading="true" pageId="2" pageNumber="31" reason="1">Description</emphasis>
.
</emphasis>
Based on male
<typeStatus box="[513,612,1645,1671]" pageId="2" pageNumber="31">holotype</typeStatus>
(measurements in
<tableCitation box="[826,907,1645,1671]" captionStart="Table I" captionStartId="8.[140,200,166,188]" captionTargetBox="[147,1074,211,1131]" captionTargetPageId="8" captionText="Table I. Morphometric values (in mm) of the new Ananteris species described in this paper" httpUri="http://table.plazi.org/id/DF0AE1FEFF883C78DC8CFF155BD8FF7C" pageId="2" pageNumber="31" tableUuid="DF0AE1FEFF883C78DC8CFF155BD8FF7C">Table I</tableCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3814813" ID-Zenodo-Dep="3814813" httpUri="https://zenodo.org/record/3814813/files/figure.png" pageId="3" pageNumber="32" startId="3.[140,222,1538,1560]" targetBox="[159,1102,165,1507]" targetPageId="3">
<paragraph blockId="3.[140,1108,1538,1656]" pageId="3" pageNumber="32">
<emphasis bold="true" box="[140,271,1538,1560]" pageId="3" pageNumber="32">Figures 1-9.</emphasis>
<taxonomicName authority="Giupponi &amp; de Vasconcelos &amp; Lourenco, 2009" authorityName="Giupponi &amp; de Vasconcelos &amp; Lourenco" authorityYear="2009" box="[279,434,1538,1560]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="chagasi" status="sp. n.">
<emphasis box="[279,434,1538,1560]" italics="true" pageId="3" pageNumber="32">Ananteris chagasi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[442,495,1539,1560]" pageId="3" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
, male
<typeStatus box="[559,643,1538,1560]" pageId="3" pageNumber="32">holotype</typeStatus>
.
<emphasis bold="true" box="[654,667,1538,1560]" pageId="3" pageNumber="32">1</emphasis>
Carapace dorsal view.
<emphasis bold="true" box="[884,897,1538,1560]" pageId="3" pageNumber="32">2</emphasis>
Pectines.
<emphasis bold="true" box="[993,1006,1538,1560]" pageId="3" pageNumber="32">3</emphasis>
Chelicera, dorsal view.
<emphasis bold="true" box="[254,267,1570,1592]" pageId="3" pageNumber="32">4</emphasis>
Metasoma, segments IV-V and telson, lateral view.
<emphasis bold="true" box="[748,761,1570,1592]" pageId="3" pageNumber="32">5</emphasis>
Chela, dorsal view.
<emphasis bold="true" box="[948,961,1570,1592]" pageId="3" pageNumber="32">6</emphasis>
Manus, ventral view.
<emphasis bold="true" box="[192,205,1602,1624]" pageId="3" pageNumber="32">7</emphasis>
Pedipalpal femur, dorsal view.
<emphasis bold="true" box="[490,524,1602,1624]" pageId="3" pageNumber="32">8-9</emphasis>
Pedipalpal patella, dorsal and external view. Scale bars: 1-4 and 6-9 =
<quantity box="[196,248,1634,1655]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="32" unit="mm" value="1.0">1mm</quantity>
; 5 =
<quantity box="[293,370,1634,1656]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="32" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
</caption>
<paragraph blockId="4.[140,1108,166,1319]" pageId="4" pageNumber="33">
<emphasis bold="true" box="[185,318,166,192]" pageId="4" pageNumber="33">
<emphasis box="[185,313,166,192]" inLineHeading="true" pageId="4" pageNumber="33" reason="1">Coloration</emphasis>
.
</emphasis>
Generally yellowish with dark brown to blackish pigmented zones on the body and its appendages. Prosoma: carapace yellowish with dark brown spots on the lateral and posterior edges; eyes surrounded by black pigment. Mesosoma: yellowish with confluent blackish zones on the posterior and lateral edges of tergites. Metasoma: segments I to V yellowish; all segments intensely marked with dark brown spots. Vesicle yellowish without spots; the base of the aculeus yellowish and the tip reddish. Venter yellowish; sternite VII with very diffused brownish spots. Chelicerae yellowish with variegated blackish spots over the anterior and lateral surfaces; fingers with blackish spots; teeth reddish. Pedipalps: yellowish; femur and patella with densely marked blackish-brown spots; chela hand almost entirely blackish-brown; fingers very dark. Legs yellowish, with several blackish-brown spots.
</paragraph>
<paragraph blockId="4.[140,1108,166,1319]" pageId="4" pageNumber="33">
<emphasis bold="true" box="[185,336,519,545]" pageId="4" pageNumber="33">
<emphasis box="[185,331,519,545]" inLineHeading="true" pageId="4" pageNumber="33" reason="1">Morphology</emphasis>
.
</emphasis>
Carapace with thin but intense granulation; anterior margin not emarginated, almost straight. Anterior median superciliary and posterior median carinae weak or absent. All furrows moderate to weak. Median ocular tubercle distinctly anterior to the centre of the carapace; median eyes separated by approximately one ocular diameter. Three pairs of lateral eyes. Sternum subpentagonal to pentagonal. Mesosoma: tergites with thin and sparse granulation. Median carina moderate to weak in all tergites. Tergite VII pentacarinate. Venter: genital operculum divided longitudinally, each plate more or less subtriangular in shape. Pectines: pectinal tooth count 15-15; basal middle lamellae of the pectines not dilated; fulcra absent. Sternites smooth; stigmata weakly elongate; setation moderate; sternite VII with very weakly marked carinae. Metasoma: segments I and II with 10 carinae, moderately crenulate. Segments III and IV with 8 carinae, moderately crenulate. Intercarinal spaces slightly granular, almost smooth. Segment V slightly rounded with 5 carinae. Telson moderately elongated and without granulations; with one ventral carina weakly marked; aculeus short and weakly curved; subaculear tooth strong and spinoid. Cheliceral dentition characteristic of the family
<taxonomicName authority="(Vachon 1963)" baseAuthorityName="Vachon" baseAuthorityYear="1963" box="[650,921,1011,1038]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="33" phylum="Arthropoda" rank="family">
Buthidae (
<bibRefCitation author="Vachon M" box="[762,913,1011,1038]" pageId="4" pageNumber="33" pagination="161 - 166" refId="ref5096" refString="Vachon M (1963) De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris 2 e ser., 35 (2): 161 - 166." type="journal article" year="1963">Vachon 1963</bibRefCitation>
)
</taxonomicName>
; fixed finger with two moderate basal teeth; movable finger with two weak basal teeth; ventral aspect of both finger and manus with dense, long setae. Pedipalps: femur pentacarinate; patella and chela with weak to vestigial carinae; internal face of patella with 5 minute spinoid granules; all faces slightly granular, almost smooth. Fixed and movable fingers with 6 almost linear rows of granules; two small external and one internal accessory granule present at the base of each row; three granules in the extremity of the fingers; Trichobothriotaxy; orthobothriotaxy A-ss-beta (
<bibRefCitation author="Vachon M" box="[330,480,1258,1284]" pageId="4" pageNumber="33" pagination="857 - 958" refId="ref5139" refString="Vachon M (1974) Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris, 3 e ser., n ° 140, Zool. 104: 857 - 958." type="journal article" year="1974">Vachon 1974</bibRefCitation>
,
<bibRefCitation author="Vachon M" box="[490,551,1257,1284]" pageId="4" pageNumber="33" pagination="1597 - 1599" refId="ref5204" refString="Vachon M (1975) Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des Scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus des seances de l'Academie des Sciences, Paris, ser. D, 281: 1597 - 1599." type="journal article" year="1975">1975</bibRefCitation>
). Legs: tarsus with very numerous fine median setae ventrally. Tibial spurs moderately developed on leg IV; weak on leg III.
</paragraph>
</subSubSection>
</treatment>
</document>