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<document id="2566566417A8886FAECE98BAD685DB6D" ID-CLB-Dataset="55280" ID-DOI="10.5733/afin.055.0108" ID-GBIF-Dataset="00de24a2-1da8-4926-9425-9eb5c7b8dc35" ID-ISSN="2305-2562" ID-Zenodo-Dep="7661736" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1676998126033" checkinUser="felipe" docAuthor="Marshall, Stephen A." docDate="2014" docId="03C8892BFFA0787DFE651209FC0888C5" docLanguage="en" docName="AfrInvertebr.55.1.143-155.pdf" docOrigin="African Invertebrates 55 (1)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.055.0108" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.4:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="4" docTitle="Aristobatina metamelasma Marshall 2014, sp. n." docType="treatment" docUuid="BB07D97C-D5AF-41B7-AB6A-55EDAF512264" docUuidSource="ZooBank" docVersion="8" lastPageNumber="147" masterDocId="FFF1F153FFA37879FFB6135EFFA68D10" masterDocTitle="A review of the Afrotropical genus Aristobatina Verbeke (Diptera: Micropezidae: Taeniapterinae), with descriptions of four new species from the Eastern Arc Mountains of Tanzania" masterLastPageNumber="143" masterPageNumber="143" pageNumber="146" updateTime="1699475542377" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="22DA6949F059199952DB678C038C6BDF">A review of the Afrotropical genus Aristobatina Verbeke (Diptera: Micropezidae: Taeniapterinae), with descriptions of four new species from the Eastern Arc Mountains of Tanzania</mods:title>
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<mods:namePart id="D80CE2914F24368C9A71F11A79862564">Marshall, Stephen A.</mods:namePart>
<mods:affiliation id="1ACD36453CBD40F42BC7A49A75CDFA3C">School of Environmental Sciences, University of Guelph, Guelph, Ontario, Canada N 1 G 2 W 1</mods:affiliation>
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<mods:title id="7671F327012D225E9C94CFEC9A5584B5">African Invertebrates</mods:title>
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<mods:date id="FEAAC6875D59A42D48BC78F9D92266A7">2014</mods:date>
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<treatment id="03C8892BFFA0787DFE651209FC0888C5" ID-DOI="http://doi.org/10.5281/zenodo.7661979" ID-GBIF-Taxon="209342710" ID-Zenodo-Dep="7661979" LSID="urn:lsid:zoobank.org:act:BB07D97C-D5AF-41B7-AB6A-55EDAF512264" httpUri="http://treatment.plazi.org/id/03C8892BFFA0787DFE651209FC0888C5" lastPageId="4" lastPageNumber="147" pageId="3" pageNumber="146">
<subSubSection id="C37B6BB6FFA0787AFE651209FCFE8C62" box="[467,856,343,370]" pageId="3" pageNumber="146" type="nomenclature">
<paragraph id="8BDE383DFFA0787AFE651209FCFE8C62" blockId="3.[181,1142,343,1669]" box="[467,856,343,370]" pageId="3" pageNumber="146">
<heading id="D0968F51FFA0787AFE651209FCFE8C62" box="[467,856,343,370]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="146" reason="2">
<taxonomicName id="4C6143BEFFA0787AFE651209FCB08C62" ID-CoL="9HN23" authority="Marshall, 2014" authorityName="Marshall" authorityYear="2014" box="[467,790,343,370]" class="Insecta" family="Micropezidae" genus="Aristobatina" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="metamelasma" status="sp. nov.">
<emphasis id="B915E42FFFA0787AFE651209FCB08C62" bold="true" box="[467,790,343,370]" pageId="3" pageNumber="146">Aristobatina metamelasma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2265954FFA0787AFCAA1206FCFE8C62" box="[796,856,344,370]" pageId="3" pageNumber="146" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C37B6BB6FFA0787DFDD512DEFC0888C5" lastPageId="4" lastPageNumber="147" pageId="3" pageNumber="146" type="description">
<paragraph id="8BDE383DFFA0787AFDD512DEFD6E8C8A" blockId="3.[181,1142,343,1669]" box="[611,712,384,410]" pageId="3" pageNumber="146">
<figureCitation id="135A24B8FFA0787AFDD512DEFD6E8C8A" box="[611,712,384,410]" captionStart="Figs 16" captionStartId="4.[144,183,1178,1200]" captionTargetBox="[196,1036,204,1157]" captionTargetId="figure-111@4.[196,1036,204,1162]" captionTargetPageId="4" captionText="Figs 16. Aristobatina metamelasma sp.n.: (1) spermathecae and bursa copulatrix; (2) female, dorsal view; male: (3) lateral view; (4) sternite 5; (5) terminalia, dorsolateral view; (6) terminalia, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7661738" httpUri="https://zenodo.org/record/7661738/files/figure.png" pageId="3" pageNumber="146">Figs 16</figureCitation>
</paragraph>
<paragraph id="8BDE383DFFA0787AFF0312F6FBC68CF3" blockId="3.[181,1142,343,1669]" pageId="3" pageNumber="146">
Etymology: From the Greek words
<emphasis id="B915E42FFFA0787AFDF212F7FDDC8CD2" box="[580,634,425,450]" italics="true" pageId="3" pageNumber="146">meta</emphasis>
(near) and
<taxonomicName id="4C6143BEFFA0787AFD4C12F6FCF88CD2" authorityName="Marshall" authorityYear="2014" box="[762,862,424,450]" class="Insecta" family="Micropezidae" genus="Aristobatina" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="melasma">
<emphasis id="B915E42FFFA0787AFD4C12F6FCF88CD2" box="[762,862,424,450]" italics="true" pageId="3" pageNumber="146">melasma</emphasis>
</taxonomicName>
(black spot), referring to the distinctive discal macula on the wing of this and the closely related
<emphasis id="B915E42FFFA0787AFC601297FBC68CF3" box="[982,1120,457,483]" italics="true" pageId="3" pageNumber="146">
<taxonomicName id="4C6143BEFFA0787AFC601297FBFA8CF3" authorityName="Marshall" authorityYear="2014" box="[982,1116,457,483]" class="Insecta" family="Micropezidae" genus="Aristobatina" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="melasma">A. melasma</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BDE383DFFA0787AFF0312AFFEE78F1B" blockId="3.[181,1142,343,1669]" box="[181,321,497,523]" pageId="3" pageNumber="146">Description:</paragraph>
<paragraph id="8BDE383DFFA0787AFF031147FDC48F24" blockId="3.[181,1142,343,1669]" box="[181,610,537,564]" pageId="3" pageNumber="146">
<emphasis id="B915E42FFFA0787AFF031147FEA58F23" box="[181,259,537,563]" italics="true" pageId="3" pageNumber="146">Length</emphasis>
(head to wing tip):
<quantity id="4C9995D8FFA0787AFE571147FDF88F24" box="[481,606,537,564]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="3" pageNumber="146" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">1315 mm</quantity>
.
</paragraph>
<paragraph id="8BDE383DFFA0787AFF03111FFC128E96" blockId="3.[181,1142,343,1669]" pageId="3" pageNumber="146">
<emphasis id="B915E42FFFA0787AFF03111FFEA28F4B" box="[181,260,577,603]" italics="true" pageId="3" pageNumber="146">Colour</emphasis>
: Head orange anteriorly, reddish brown posteriorly, swollen part of frontal vitta dark reddish brown; thorax dark reddish brown; abdomen shiny blue­black; fore femur orange in basal ⅔, brown distally; mid and hind femur uniformly orange except for a small dark area at apex; mid tarsus dark brown or black; basal ½ of tarsomere 1 of hind leg pale, tarsus otherwise dark brown to black; abdominal pleurae grey on pinned
<typeStatus id="54DA869FFFA0787AFB85119DFBD48FCD" box="[1075,1138,707,733]" pageId="3" pageNumber="146">types</typeStatus>
.
<emphasis id="B915E42FFFA0787AFF0311B5FF548E15" box="[181,242,747,773]" italics="true" pageId="3" pageNumber="146">Head</emphasis>
: Scape bare except for marginal ring of short setae; pedicel short and setose, with some longer ventral apical setae; frontal vitta with strongly convex elongate oval area surrounding ocelli and extending half the distance from anterior ocellus to anterior margin of frons; 2 pairs of fronto-orbital setae, 1 large and above level of ocelli, 1 smaller below; postocellar, outer and inner vertical setae well­developed.
</paragraph>
<paragraph id="8BDE383DFFA0787AFF0310CAFE108920" blockId="3.[181,1142,343,1669]" pageId="3" pageNumber="146">
<emphasis id="B915E42FFFA0787AFF0310CAFEA08EBE" box="[181,262,916,942]" italics="true" pageId="3" pageNumber="146">Thorax</emphasis>
: Cervical sclerite flat, dull, microtrichose; proepisternum with a few long marginal ventral setae on posterior ½; katepisternum with double row of thin black setae, anterior row with only 5 setae, barely overlapping with lower end of posterior row; scutellum with 1 pair of marginal setae only, 1 small to minute seta between postalar and dorsocentral setae.
</paragraph>
<paragraph id="8BDE383DFFA0787AFF661768FE1989C1" blockId="3.[181,1142,343,1669]" pageId="3" pageNumber="146">
<emphasis id="B915E42FFFA0787AFF661768FEAC895F" box="[208,266,1078,1103]" italics="true" pageId="3" pageNumber="146">Wing</emphasis>
:
<emphasis id="B915E42FFFA0787AFEAE1768FEF48946" box="[280,338,1078,1110]" italics="true" pageId="3" pageNumber="146">
CuA
<subScript id="17E53A78FFA0787AFEFC1716FEF48946" attach="left" box="[330,338,1096,1110]" fontSize="6" pageId="3" pageNumber="146">2</subScript>
</emphasis>
virtually at right angle to and in line with
<emphasis id="B915E42FFFA0787AFC951768FCCF8940" box="[803,873,1078,1104]" italics="true" pageId="3" pageNumber="146">bm­cu</emphasis>
, wing membrane with a distinctly infuscated tip and a small, circular discal macula extending from just anterior to
<emphasis id="B915E42FFFA0787AFF671729FF5B8987" box="[209,253,1143,1175]" italics="true" pageId="3" pageNumber="146">
R
<subScript id="17E53A78FFA0787AFF5417D7FF5B8987" attach="left" box="[226,253,1161,1175]" fontSize="6" pageId="3" pageNumber="146">4+5</subScript>
</emphasis>
to
<emphasis id="B915E42FFFA0787AFEA81729FE938980" box="[286,309,1143,1168]" italics="true" pageId="3" pageNumber="146">M</emphasis>
; tegula, basicosta and stem vein brown, similar to adjacent wing base, stem vein dorsally microsetulose at base; wing base with 3 long costagial setae, inner one very long and inclinate.
</paragraph>
<paragraph id="8BDE383DFFA0787AFF031781FC8C886A" blockId="3.[181,1142,343,1669]" pageId="3" pageNumber="146">
<emphasis id="B915E42FFFA0787AFF031781FED289E9" box="[181,372,1247,1273]" italics="true" pageId="3" pageNumber="146">Female abdomen</emphasis>
: Paired spermathecae elongate egg­shaped, 1.9× as long as wide, with a constriction in basal ⅓, surface otherwise smooth, spermathecal duct densely covered with knob­like processes beyond the division of the common duct into 2 straight branches (
<figureCitation id="135A24B8FFA0787AFF08161EFF58884A" box="[190,254,1344,1370]" captionStart="Figs 16" captionStartId="4.[144,183,1178,1200]" captionTargetBox="[196,1036,204,1157]" captionTargetId="figure-111@4.[196,1036,204,1162]" captionTargetPageId="4" captionText="Figs 16. Aristobatina metamelasma sp.n.: (1) spermathecae and bursa copulatrix; (2) female, dorsal view; male: (3) lateral view; (4) sternite 5; (5) terminalia, dorsolateral view; (6) terminalia, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7661738" httpUri="https://zenodo.org/record/7661738/files/figure.png" pageId="3" pageNumber="146">Fig. 1</figureCitation>
). Single spermatheca on a much shorter and smaller duct, body densely covered with processes much like the paired spermathecal ducts.
</paragraph>
<paragraph id="8BDE383DFFA0787DFF0316D6FE5E8854" blockId="3.[181,1142,343,1669]" lastBlockId="4.[144,1103,1290,1348]" lastPageId="4" lastPageNumber="147" pageId="3" pageNumber="146">
<emphasis id="B915E42FFFA0787AFF0316D6FEFB88B2" box="[181,349,1416,1442]" italics="true" pageId="3" pageNumber="146">Male abdomen</emphasis>
: Pleuron with large dome­like differentiated area on segment 4 (occupying approximately ventral ¾ of pleuron 4); hypandrium with a very short anterior plate and strong posterodorsal arms connecting to phallic plate, dorsolateral margin of posterodorsal arm with a prominent lobe (
<figureCitation id="135A24B8FFA0787AFD3616B7FD648B14" box="[640,706,1513,1540]" captionStart="Figs 16" captionStartId="4.[144,183,1178,1200]" captionTargetBox="[196,1036,204,1157]" captionTargetId="figure-111@4.[196,1036,204,1162]" captionTargetPageId="4" captionText="Figs 16. Aristobatina metamelasma sp.n.: (1) spermathecae and bursa copulatrix; (2) female, dorsal view; male: (3) lateral view; (4) sternite 5; (5) terminalia, dorsolateral view; (6) terminalia, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7661738" httpUri="https://zenodo.org/record/7661738/files/figure.png" pageId="3" pageNumber="146">Fig. 5</figureCitation>
). Basiphallus elongate and tapered posteriorly, extending well beyond base of distiphallus; postgonites equal, small and finely spinulose ventrally; distiphallus with tubular basal part bifurcating before transition into strongly dextrally recurved membranous distal part, apex with a finely spinulose glans (
<figureCitation id="135A24B8FFA0787AFF081534FEA58B95" box="[190,259,1642,1669]" captionStart="Figs 16" captionStartId="4.[144,183,1178,1200]" captionTargetBox="[196,1036,204,1157]" captionTargetId="figure-111@4.[196,1036,204,1162]" captionTargetPageId="4" captionText="Figs 16. Aristobatina metamelasma sp.n.: (1) spermathecae and bursa copulatrix; (2) female, dorsal view; male: (3) lateral view; (4) sternite 5; (5) terminalia, dorsolateral view; (6) terminalia, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7661738" httpUri="https://zenodo.org/record/7661738/files/figure.png" pageId="3" pageNumber="146">Fig. 6</figureCitation>
); ejaculatory apodeme slightly larger than epandrium; genital fork (sternite 5) with gap Y­shaped at base (
<figureCitation id="135A24B8FFA7787DFE731654FDAC8834" box="[453,522,1290,1316]" captionStart="Figs 16" captionStartId="4.[144,183,1178,1200]" captionTargetBox="[196,1036,204,1157]" captionTargetId="figure-111@4.[196,1036,204,1162]" captionTargetPageId="4" captionText="Figs 16. Aristobatina metamelasma sp.n.: (1) spermathecae and bursa copulatrix; (2) female, dorsal view; male: (3) lateral view; (4) sternite 5; (5) terminalia, dorsolateral view; (6) terminalia, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.7661738" httpUri="https://zenodo.org/record/7661738/files/figure.png" pageId="4" pageNumber="147">Fig. 4</figureCitation>
), inner surfaces of arms densely spinose and with a right-angled bend in the middle.
</paragraph>
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<paragraph id="8BDE383DFFA7787DFF2617C4FEE589F0" blockId="4.[144,1105,1178,1248]" pageId="4" pageNumber="147">
Figs 16.
<taxonomicName id="4C6143BEFFA7787DFF5E17C4FE7D89BF" authority="Marshall, 2014" authorityName="Marshall" authorityYear="2014" box="[232,475,1178,1199]" class="Insecta" family="Micropezidae" genus="Aristobatina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="metamelasma" status="sp. nov.">
<emphasis id="B915E42FFFA7787DFF5E17C4FE7D89BF" box="[232,475,1178,1199]" italics="true" pageId="4" pageNumber="147">Aristobatina metamelasma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2265954FFA7787DFE6917C4FDB789BF" box="[479,529,1178,1199]" pageId="4" pageNumber="147" rank="species">sp. n.</taxonomicNameLabel>
: (1) spermathecae and bursa copulatrix; (2) female, dorsal view; male: (3) lateral view; (4) sternite 5; (5) terminalia, dorsolateral view; (6) terminalia, ventral view. Not to scale.
</paragraph>
</caption>
<paragraph id="8BDE383DFFA7787DFF261608FF7A889C" blockId="4.[144,1104,1366,1493]" pageId="4" pageNumber="147">
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<typeStatus id="54DA869FFFA7787DFF261608FF42887C" box="[144,228,1366,1388]" pageId="4" pageNumber="147" type="holotype">Holotype</typeStatus>
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and
</materialsCitation>
<materialsCitation id="3B093260FFA7787DFE951608FF7A889C" collectionCode="CNCI" country="Tanzania" elevation="1650" location="Uluguru Mountains" pageId="4" pageNumber="147" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D67F3B4FFA7787DFE951608FE99887B" box="[291,319,1366,1387]" pageId="4" pageNumber="147" type="female">3♀</specimenCount>
<typeStatus id="54DA869FFFA7787DFEF51608FE3B887B" box="[323,413,1366,1387]" pageId="4" pageNumber="147" type="paratype">paratypes</typeStatus>
:
<collectingCountry id="F37678ADFFA7787DFE121608FDBD887B" box="[420,539,1366,1387]" name="Tanzania" pageId="4" pageNumber="147">TANZANIA</collectingCountry>
:
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: “Tanganyika, Ulguru Mts,
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<elevation id="004CDF0EFFA7787DFC7B1608FBE1887C" box="[973,1095,1366,1388]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="4" pageNumber="147" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">15001800m</elevation>
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” (
<collectionCode id="ED70A0F8FFA7787DFF201628FF77889C" box="[150,209,1398,1420]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="4" pageNumber="147">CNCI</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BDE383DFFA7787DFF2616C4FC0888C5" blockId="4.[144,1104,1366,1493]" pageId="4" pageNumber="147">
Comments: This species is closely related to
<taxonomicName id="4C6143BEFFA7787DFDC616C5FD4988A4" authorityName="Marshall" authorityYear="2014" box="[624,751,1434,1461]" class="Insecta" family="Micropezidae" genus="Aristobatina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="melasma">
<emphasis id="B915E42FFFA7787DFDC616C5FD4988A4" box="[624,751,1434,1461]" italics="true" pageId="4" pageNumber="147">A. melasma</emphasis>
</taxonomicName>
, from which it differs in spermathecal shape and sculpturing and in the pigmentation of the hind tarsus.
</paragraph>
</subSubSection>
</treatment>
</document>