649 lines
79 KiB
XML
649 lines
79 KiB
XML
<document id="A22419710DC7086854AAC3D7B2C97277" ID="10.11646/zootaxa.4204.1.1" ID-DOI="10.11646/zootaxa.4204.1.1" ID-GBIF-Dataset="3f8866d2-529e-43ad-b971-29fc52a13858" ID-ISSN="1175-5326" ID-Zenodo-Dep="192302" ID-ZooBank="C050058A-774D-49C0-93F9-7A055B51C2A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1480667017343" checkinUser="plazi" docAuthor="Williams, Paul H., Huang, Jiaxing, Rasmont, Pierre & An, Jiandong" docDate="2016" docId="03AE67547C7A3317B090A017A2EEFBB6" docLanguage="en" docName="zootaxa.4204.1.1.pdf" docOrigin="Zootaxa 4204 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Bombus superbus Tkalcu" docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="FF971F2C7C6D330CB007A424A336FFA4" masterDocTitle="Early-diverging bumblebees from across the roof of the world: the high-mountain subgenus Mendacibombus revised from species’ gene coalescents and morphology (Hymenoptera, Apidae)" masterLastPageNumber="72" masterPageNumber="1" pageNumber="24" updateTime="1698395689147" updateUser="plazi">
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<mods:title id="E72326EFC11CF8B50DCBA979FF5B600D">Early-diverging bumblebees from across the roof of the world: the high-mountain subgenus Mendacibombus revised from species’ gene coalescents and morphology (Hymenoptera, Apidae)</mods:title>
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<mods:namePart id="CEA79B8B4212FAEDCA8DFC2263C7CAEF">Williams, Paul H.</mods:namePart>
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<mods:roleTerm id="EF87D13734034F586FB4ABB724DA6334">Author</mods:roleTerm>
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<mods:namePart id="4A5390F3F4DEF18041704AB612E64410">Huang, Jiaxing</mods:namePart>
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<mods:namePart id="0F8B345FAD280A621F7F67FA6498FBF7">Rasmont, Pierre</mods:namePart>
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<mods:namePart id="494929EDE2EFBBD678FD5EBDBCBB377A">An, Jiandong</mods:namePart>
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<mods:typeOfResource id="E62A5223D7041E2D0647AC652794D52C">text</mods:typeOfResource>
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<mods:title id="FA5C7B5340579F6ED53E59B2EB9498E0">Zootaxa</mods:title>
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<mods:date id="A6DC25E0FCE39962960AEE9034DBC117">2016</mods:date>
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<mods:detail id="9F85FE08F5590DF2F23AEBEF9ED192CD" type="volume">
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<mods:number id="0499C793DC6FF60F2F86DC3D0B38A207">4204</mods:number>
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<treatment id="03AE67547C7A3317B090A017A2EEFBB6" ID-DOI="http://doi.org/10.5281/zenodo.5625285" ID-GBIF-Taxon="124940391" ID-Zenodo-Dep="5625285" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6" httpUri="http://treatment.plazi.org/id/03AE67547C7A3317B090A017A2EEFBB6" lastPageId="27" lastPageNumber="28" pageId="23" pageNumber="24">
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<subSubSection id="C31D85C97C7A331BB090A017A2C9FBE9" box="[151,511,1074,1101]" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="8BB8D6427C7A331BB090A017A2C9FBE9" blockId="23.[151,511,1074,1134]" box="[151,511,1074,1101]" pageId="23" pageNumber="24">
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<heading id="D0F0612E7C7A331BB090A017A2C9FBE9" bold="true" box="[151,511,1074,1101]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<emphasis id="B9730A507C7A331BB090A017A2C9FBE9" bold="true" box="[151,511,1074,1101]" pageId="23" pageNumber="24">
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1.
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<taxonomicName id="4C07ADC17C7A331BB0B3A016A2C9FBE9" ID-CoL="MFW5" authority="Tkalcu" authorityName="Tkalcu" box="[180,511,1074,1101]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="superbus">
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<emphasis id="B9730A507C7A331BB0B3A016A2BAFBE8" bold="true" box="[180,396,1074,1100]" italics="true" pageId="23" pageNumber="24">Bombus superbus</emphasis>
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(Tkalců)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D85C97C7A331BB090A072A24DFBCA" box="[151,379,1109,1134]" pageId="23" pageNumber="24" type="description">
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<paragraph id="8BB8D6427C7A331BB090A072A24DFBCA" blockId="23.[151,511,1074,1134]" box="[151,379,1109,1134]" pageId="23" pageNumber="24">
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(
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<figureCitation id="133CCAC77C7A331BB0A7A071A3C3FBCA" box="[160,245,1109,1134]" captionStart="FIGURES 24 ‒ 35" captionStartId="26.[151,264,1867,1889]" captionTargetBox="[233,1387,193,1789]" captionTargetId="figure@26.[233,1390,193,1894]" captionTargetPageId="26" captionText="FIGURES 24 ‒ 35. Morphology of the male genitalia for species of the subgenus Mendacibombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 24, B. superbus, China MD # 339; 25, B. waltoni, China MD # 216; 26, B. convexus, China MD # 215; 27, B. makarjini, Kazakhstan MD # 1240; 28, B. marussinus, Pakistan MD # 408; 29, B. avinoviellus, India MD # 404; 30, B. himalayanus, India MD # 426; 31, B. turkestanicus, China MD # 351; 32, B. defector, Kyrgyzstan MD # 367; 33, B. margreiteri, China MD # 294; 34, B. handlirschianus, Russia MD # 835; 35, B. mendax, Spain MD # 689." httpUri="https://zenodo.org/record/192312/files/figure.png" pageId="23" pageNumber="24">Figs 24</figureCitation>
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,
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<figureCitation id="133CCAC77C7A331BB106A072A216FBCA" box="[257,288,1110,1134]" captionStart="FIGURES 36 ‒ 55" captionStartId="29.[151,264,1875,1897]" captionTargetBox="[151,1415,299,1829]" captionTargetId="figure@29.[151,1436,193,1853]" captionTargetPageId="29" captionText="FIGURES 36 ‒ 55. Morphology of parts of the male genitalia from the dorsal aspect of the left side, anterior to the left of the image, posterior to the right, white bars indicating morphological characters referred to in the key to males: 36, B. waltoni; 37, B. mendax; 38, B. superbus; 39, B. convexus; 40, B. waltoni; 41, B. convexus; 42, B. defector; 43, B. mendax; 44, B. mendax; 45, B. turkestanicus; 46, B. avinoviellus; 47, B. mendax; 48, B. mendax; 49, B. makarjini; 50, B. handlirschianus; 51, B. himalayanus; 52, B. turkestanicus; 53, B. himalayanus; 54, B. himalayanus; 55, B. marussinus." httpUri="https://zenodo.org/record/192314/files/figure.png" pageId="23" pageNumber="24">38</figureCitation>
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,
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<figureCitation id="133CCAC77C7A331BB12CA071A27CFBCA" box="[299,330,1109,1134]" captionStart="FIGURES 56 ‒ 67" captionStartId="35.[151,264,1891,1913]" captionTargetBox="[320,1245,193,1835]" captionTargetId="figure@35.[320,1245,193,1843]" captionTargetPageId="35" captionText="FIGURES 56 ‒ 67. Maps of the global distributions of species of the subgenus Mendacibombus from the material examined, with grey spots for specimen records and black spots for the samples with COI sequences (each species is unknown from regions outside the map boundaries shown); grey crosses for records of all species of Mendacibombus combined. Relief map with hill shading, Cartesian orthonormal projection, grey lines show borders to national administration according to UN maps, north at the top of the map. Image created in ArcGIS using World _ Shaded _ Relief basemap © 2014 Esri." httpUri="https://zenodo.org/record/192315/files/figure.png" pageId="23" pageNumber="24">56</figureCitation>
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,
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<figureCitation id="133CCAC77C7A331BB151A071A245FBCA" box="[342,371,1109,1134]" captionStart="FIGURES 69 ‒ 70" captionStartId="59.[151,264,1618,1640]" captionTargetBox="[151,1436,259,1554]" captionTargetId="figure@59.[151,1436,193,1653]" captionTargetPageId="59" captionText="FIGURES 69 ‒ 70. Photos of (69) a worker of B. superbus (MD # 4123) from (left) the dorsal and (right) the left lateral aspect (scale bar 10 mm), from the sample site (70) in the Tanggula mountains, Xizang, China (16. viii. 2014, 32.86142 ° N 91.91739 ° E, GPS elevation ca 5220 m). Photos by (69) NHM digital imaging unit and (70) Z. - Y. Miao." httpUri="https://zenodo.org/record/192317/files/figure.png" pageId="23" pageNumber="24">69</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D85C97C7A331BB090A0B9A148F937" pageId="23" pageNumber="24" type="reference_group">
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<paragraph id="8BB8D6427C7A331BB090A0B9A071FB54" blockId="23.[151,1436,1181,1326]" pageId="23" pageNumber="24">
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<treatmentCitationGroup id="AB17F16C7C7A331BB090A0B9A071FB54" infered="true" pageId="23" pageNumber="24">
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<taxonomicName id="4C07ADC17C7A331BB090A0B9A17BFB17" ID-CoL="3ZPKW" authority="Tkalcu 1968:22" authorityName="Tkalcu" authorityPageNumber="22" authorityYear="1968" box="[151,589,1181,1203]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="superbus">
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<emphasis id="B9730A507C7A331BB090A0B9A297FB17" box="[151,417,1181,1203]" italics="true" pageId="23" pageNumber="24">Mendacibombus superbus</emphasis>
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<treatmentCitation id="0AA6F0537C7A331BB1AEA0B9A17BFB17" author="Tkalcu" box="[425,589,1181,1203]" page="22" pageId="23" pageNumber="24" year="1968">
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<bibRefCitation id="EF96ABB37C7A331BB1AEA0B9A11BFB17" author="Tkalcu" box="[425,557,1181,1203]" pageId="23" pageNumber="24" refString="Tkalcu, B. (1968) Neue Arten der Unterfamilie Bombinae der palaarktischen Region (Hymenoptera, Apoidea). Sbornik Entomologickeho oddeleni Narodniho musea v Praze, 65, 21 - 51." type="journal article" year="1968">Tkalců 1968</bibRefCitation>
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:22
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</treatmentCitation>
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</taxonomicName>
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, type-locality citation ‘Mongolei Monda’.
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<materialsCitation id="3B6FDC1F7C7A331BB402A0B9A071FB54" ID-GBIF-Occurrence="1413110440" collectionCode="MNHU" country="China" county="Note" location="Mongolei" municipality="Monda'" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Qinghai" typeStatus="holotype">
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<typeStatus id="54BC68E07C7A331BB402A0B9A755FB17" box="[1029,1123,1181,1203]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
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queen by original designation
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<collectionCode id="ED164E877C7A331BB0C0A099A222FB76" box="[199,276,1213,1234]" pageId="23" pageNumber="24">MNHU</collectionCode>
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examined, ‘
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||
<location id="8ED880997C7A331BB18AA098A2C0FB76" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB18AA098A2C0FB76" box="[397,502,1212,1234]" country="China" county="Note" municipality="Monda'" name="Mongolei" pageId="23" pageNumber="24" stateProvince="Qinghai">Mongolei</location>
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,
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<collectingMunicipality id="6BDC4C387C7A331BB206A098A167FB76" box="[513,593,1212,1234]" pageId="23" pageNumber="24">Monda’</collectingMunicipality>
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believed incorrect (probably
|
||
<collectingRegion id="49C318A07C7A331BB37BA098A0F8FB76" box="[892,974,1212,1234]" country="China" name="Qinghai" pageId="23" pageNumber="24">Qinghai</collectingRegion>
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or
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<collectingRegion id="49C318A07C7A331BB3F7A098A70AFB76" box="[1008,1084,1212,1234]" country="China" name="Xizang" pageId="23" pageNumber="24">Xizang</collectingRegion>
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||
,
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<collectingCountry id="F31096D27C7A331BB441A098A7B1FB76" box="[1094,1159,1212,1234]" name="China" pageId="23" pageNumber="24">China</collectingCountry>
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).
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<collectingCounty id="62D9AECE7C7A331BB490A099A7FEFB76" box="[1175,1224,1213,1234]" pageId="23" pageNumber="24">Note</collectingCounty>
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1. [
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<taxonomicName id="4C07ADC17C7A331BB098A0FEA2CFFB54" authority="Tkalcu, 1968:24" authorityName="Tkalcu" authorityPageNumber="24" authorityYear="1968" box="[159,505,1242,1264]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="wagneri">
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<emphasis id="B9730A507C7A331BB098A0FEA27CFB54" box="[159,330,1242,1264]" italics="true" pageId="23" pageNumber="24">Bombus wagneri</emphasis>
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<treatmentCitation id="0AA6F0537C7A331BB157A0FEA2CFFB54" author="Tkalcu" box="[336,505,1242,1264]" page="24" pageId="23" pageNumber="24" year="1968">
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<bibRefCitation id="EF96ABB37C7A331BB157A0FEA2EFFB54" author="Tkalcu" box="[336,473,1242,1264]" pageId="23" pageNumber="24" refString="Tkalcu, B. (1968) Neue Arten der Unterfamilie Bombinae der palaarktischen Region (Hymenoptera, Apoidea). Sbornik Entomologickeho oddeleni Narodniho musea v Praze, 65, 21 - 51." type="journal article" year="1968">Tkalců, 1968</bibRefCitation>
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:24
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</treatmentCitation>
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</taxonomicName>
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, published as a junior synonym.]
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</materialsCitation>
|
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</treatmentCitationGroup>
|
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</paragraph>
|
||
<paragraph id="8BB8D6427C7A331BB090A0DDA2EFFA8A" blockId="23.[151,1436,1181,1326]" pageId="23" pageNumber="24">
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||
<treatmentCitationGroup id="AB17F16C7C7A331BB090A0DDA2EFFA8A" pageId="23" pageNumber="24">
|
||
<taxonomicName id="4C07ADC17C7A331BB090A0DDA159FAAB" ID-CoL="MFW5" authority="Tkalcu" authorityName="Tkalcu" box="[151,623,1273,1295]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="superbus" subGenus="Mendacibombus">
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<emphasis id="B9730A507C7A331BB090A0DDA13BFAAB" box="[151,525,1273,1295]" italics="true" pageId="23" pageNumber="24">Bombus (Mendacibombus) superbus</emphasis>
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(Tkalců)
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</taxonomicName>
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;
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<treatmentCitation id="0AA6F0537C7A331BB27BA0DEA05FFAAB" author="Williams" box="[636,873,1273,1295]" page="42" pageId="23" pageNumber="24" year="1991">
|
||
<bibRefCitation id="EF96ABB37C7A331BB27BA0DEA07FFAAB" author="Williams" box="[636,841,1273,1295]" pageId="23" pageNumber="24" refString="Williams, P. H. (1991) The bumble bees of the Kashmir Himalaya (Hymenoptera: Apidae, Bombini). Bulletin of the British Museum (Natural History) (Entomology), 60, 1 - 204." type="journal article" year="1991">P.H. Williams 1991</bibRefCitation>
|
||
:42
|
||
</treatmentCitation>
|
||
; S.-
|
||
<treatmentCitation id="0AA6F0537C7A331BB388A0DDA7A9FAAB" author="Wang" box="[911,1183,1273,1295]" page="303" pageId="23" pageNumber="24" year="1996">
|
||
<bibRefCitation id="EF96ABB37C7A331BB388A0DDA745FAAB" author="Wang" box="[911,1139,1273,1295]" pageId="23" pageNumber="24" refString="Wang, S. - F. & Yao, J. (1996) Hymenoptera: Apodae [sic] - Bombini. In: F. - S. Huang (Ed), Insects of the Karakorum - Kunlun mountains. Science Press, Beijing, pp. 303 - 309." type="book chapter" year="1996">F. Wang & Yao 1996</bibRefCitation>
|
||
:303
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA6F0537C7A331BB4ACA0DEA6AEFAAB" author="Williams" box="[1195,1432,1273,1295]" page="99" pageId="23" pageNumber="24" year="1998">
|
||
<bibRefCitation id="EF96ABB37C7A331BB4ACA0DEA64FFAAB" author="Williams" box="[1195,1401,1273,1295]" pageId="23" pageNumber="24" refString="Williams, P. H. (1998) An annotated checklist of bumble bees with an analysis of patterns of description (Hymenoptera: Apidae, Bombini). Bulletin of The Natural History Museum (Entomology), 67, 79 - 152. [updated at www. nhm. ac. uk / bombus / accessed 2015]." type="journal article" year="1998">P.H. Williams 1998</bibRefCitation>
|
||
:99
|
||
</treatmentCitation>
|
||
;
|
||
<bibRefCitation id="EF96ABB37C7A331BB0C0A13DA2A7FA8A" author="Williams" box="[199,401,1304,1326]" pageId="23" pageNumber="24" refString="Williams, P. H. (2004) Genus Bombus Latreille. In: R. K. Gupta (Ed), An annotated catalogue of the bee species of the Indian Region. Jai Narain Vyas University, Jodhpur, p. Web page providing a list of bumblebees of India and adjacent countries with common synonyms and a summary of their distributions within the region." type="book" year="2004">P.H. Williams 2004</bibRefCitation>
|
||
:no. 29.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C7A331BB090A17FA148F937" blockId="23.[151,1437,1370,2007]" pageId="23" pageNumber="24">
|
||
<materialsCitation id="3B6FDC1F7C7A331BB090A17FA061F9A7" ID-GBIF-Occurrence="1413110438" collectionCode="MNHU" collectorName="H. Friese" country="Mongolia" county="Note" location="Weiske'" municipality="Tkalcu's" pageId="23" pageNumber="24" specimenCount="2" stateProvince="Mus" typeStatus="holotype">
|
||
<collectingCounty id="62D9AECE7C7A331BB090A17FA3FBFAD6" box="[151,205,1371,1394]" pageId="23" pageNumber="24">Note</collectingCounty>
|
||
1 (
|
||
<taxonomicName id="4C07ADC17C7A331BB0F5A17EA261FAD7" ID-CoL="MFW5" box="[242,343,1370,1395]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C7A331BB0F5A17EA261FAD7" box="[242,343,1370,1395]" italics="true" pageId="23" pageNumber="24">superbus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<collectingMunicipality id="6BDC4C387C7A331BB169A17EA2F9FAD7" box="[366,463,1370,1395]" pageId="23" pageNumber="24">Tkalců’s</collectingMunicipality>
|
||
original description of the taxon
|
||
<taxonomicName id="4C07ADC17C7A331BB345A17EA091FAD7" ID-CoL="MFW5" box="[834,935,1370,1395]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C7A331BB345A17EA091FAD7" box="[834,935,1370,1395]" italics="true" pageId="23" pageNumber="24">superbus</emphasis>
|
||
</taxonomicName>
|
||
cites ‘
|
||
<collectingCountry id="F31096D27C7A331BB3EAA17EA769FAD7" box="[1005,1119,1370,1395]" name="Mongolia" pageId="23" pageNumber="24">Mongolei</collectingCountry>
|
||
Monda 6. 0 8
|
||
<location id="8ED880997C7A331BB4FDA17EA663FAD7" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB4FDA17EA663FAD7" box="[1274,1365,1370,1395]" country="Mongolia" county="Note" municipality="Tkalcu's" name="Weiske'" pageId="23" pageNumber="24" stateProvince="Mus">Weiske’</location>
|
||
as the
|
||
<typeStatus id="54BC68E07C7A331BB090A15BA3F1FA33" box="[151,199,1407,1431]" pageId="23" pageNumber="24">type</typeStatus>
|
||
locality.
|
||
<location id="8ED880997C7A331BB135A15AA268FA33" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB135A15AA268FA33" box="[306,350,1406,1431]" country="Mongolia" county="Note" municipality="Tkalcu's" name="The" pageId="23" pageNumber="24" stateProvince="Mus">The</location>
|
||
<collectionCode id="ED164E877C7A331BB161A15BA28DFA32" box="[358,443,1407,1430]" pageId="23" pageNumber="24">MNHU</collectionCode>
|
||
collection studied by
|
||
<location id="8ED880997C7A331BB2BDA15AA03EFA33" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB2BDA15AA03EFA33" box="[698,776,1406,1431]" country="Mongolia" county="Note" municipality="Tkalcu's" name="Tkalcu" pageId="23" pageNumber="24" stateProvince="Mus">Tkalců</location>
|
||
contains a queen that agrees with the original description and carries the labels: (1) white, printed ‘
|
||
<collectingCountry id="F31096D27C7A331BB276A186A1D5FA1F" box="[625,739,1442,1467]" name="Mongolia" pageId="23" pageNumber="24">Mongolei</collectingCountry>
|
||
/
|
||
<location id="8ED880997C7A331BB2F9A186A07BFA1F" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB2F9A186A07BFA1F" box="[766,845,1442,1467]" country="Mongolia" county="Note" municipality="Tkalcu's" name="Monda" pageId="23" pageNumber="24" stateProvince="Mus">Monda</location>
|
||
/ 6. 0 8 /
|
||
<location id="8ED880997C7A331BB3B9A186A72BFA1F" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB3B9A186A72BFA1F" box="[958,1053,1442,1467]" country="Mongolia" county="Note" municipality="Tkalcu's" name="Weiske'" pageId="23" pageNumber="24" stateProvince="Mus">Weiske’</location>
|
||
; (2) white, handwritten by
|
||
<location id="8ED880997C7A331BB55FA186A6ABFA1F" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB55FA186A6ABFA1F" box="[1368,1437,1442,1467]" country="Mongolia" county="Note" municipality="Tkalcu's" name="Friese" pageId="23" pageNumber="24" stateProvince="Mus">Friese</location>
|
||
‘
|
||
<taxonomicName id="4C07ADC17C7A331BB0A7A1E2A3CDFA7B" ID-CoL="62H8K" box="[160,251,1478,1503]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C7A331BB0A7A1E2A3CDFA7B" box="[160,251,1478,1503]" italics="true" pageId="23" pageNumber="24">Bombus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C07ADC17C7A331BB108A1E3A25DFA7A" box="[271,363,1479,1502]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="wagneri">
|
||
<emphasis id="B9730A507C7A331BB108A1E3A25DFA7A" box="[271,363,1479,1502]" italics="true" pageId="23" pageNumber="24">wagneri</emphasis>
|
||
</taxonomicName>
|
||
/ [female] 1909 Friese
|
||
<emphasis id="B9730A507C7A331BB277A1E3A1BDFA7A" box="[624,651,1479,1502]" italics="true" pageId="23" pageNumber="24">Fr</emphasis>
|
||
. Det.’ (
|
||
<collectorName id="26F2B3947C7A331BB2E6A1E3A07CFA7B" box="[737,842,1478,1503]" pageId="23" pageNumber="24">H. Friese</collectorName>
|
||
, unpublished); (3) maroon, printed ‘
|
||
<location id="8ED880997C7A331BB4DCA1E3A612FA7B" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB4DCA1E3A612FA7B" box="[1243,1316,1479,1503]" country="Mongolia" county="Note" municipality="Tkalcu's" name="Type'" pageId="23" pageNumber="24" stateProvince="Mus">Type’</location>
|
||
; (4) white, printed ‘Zool.
|
||
<collectingRegion id="49C318A07C7A331BB139A1CFA245F9A6" box="[318,371,1515,1538]" country="Turkey" name="Mus" pageId="23" pageNumber="24">Mus</collectingRegion>
|
||
. /
|
||
<location id="8ED880997C7A331BB195A1CEA2D3F9A7" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB195A1CEA2D3F9A7" box="[402,485,1514,1539]" country="Mongolia" county="Note" municipality="Tkalcu's" name="Berlin'" pageId="23" pageNumber="24" stateProvince="Mus">Berlin’</location>
|
||
; (5) red, printed ‘Holo- /
|
||
<typeStatus id="54BC68E07C7A331BB317A1CFA061F9A7" box="[784,855,1515,1539]" pageId="23" pageNumber="24">typus</typeStatus>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B6FDC1F7C7A331BB35CA1CFA0AAF9CB" ID-GBIF-Occurrence="1413110449" country="Mongolia" determinerName="Tkalcu & PH Williams" location="Tkalcu" pageId="23" pageNumber="24" specimenCount="2" stateProvince="Mus" typeStatus="holotype">
|
||
’; (6) white, handwritten by
|
||
<location id="8ED880997C7A331BB49BA1CEA7DCF9A7" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB49BA1CEA7DCF9A7" box="[1180,1258,1514,1539]" country="Mongolia" name="Tkalcu" pageId="23" pageNumber="24" stateProvince="Mus">Tkalců</location>
|
||
‘
|
||
<emphasis id="B9730A507C7A331BB4FAA1CFA6BCF9A6" box="[1277,1418,1515,1538]" italics="true" pageId="23" pageNumber="24">
|
||
<typeStatus id="54BC68E07C7A331BB4FAA1CFA6BCF9A6" box="[1277,1418,1515,1538]" pageId="23" pageNumber="24" type="holotype">HOLOTYPE</typeStatus>
|
||
</emphasis>
|
||
/
|
||
<taxonomicName id="4C07ADC17C7A331BB090A22AA266F983" box="[151,336,1550,1575]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C7A331BB090A22AA266F983" box="[151,336,1550,1575]" italics="true" pageId="23" pageNumber="24">Mendacibombus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C07ADC17C7A331BB161A22AA2C0F983" ID-CoL="MFW5" authority="Tk." authorityName="Tk." box="[358,502,1550,1575]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C7A331BB161A22AA2C0F983" box="[358,502,1550,1575]" italics="true" pageId="23" pageNumber="24">superbus Tk.</emphasis>
|
||
</taxonomicName>
|
||
/ [female]
|
||
<emphasis id="B9730A507C7A331BB276A22AA1D1F983" box="[625,743,1550,1575]" italics="true" pageId="23" pageNumber="24">
|
||
<determinerName id="7E643A867C7A331BB276A22AA188F983" box="[625,702,1550,1575]" pageId="23" pageNumber="24">Tkalců</determinerName>
|
||
det
|
||
</emphasis>
|
||
.’; (7) green, printed ‘
|
||
<taxonomicName id="4C07ADC17C7A331BB3DDA22AA7A5F983" box="[986,1171,1550,1575]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C7A331BB3DDA22AA7A5F983" box="[986,1171,1550,1575]" italics="true" pageId="23" pageNumber="24">Mendacibombus</emphasis>
|
||
</taxonomicName>
|
||
/ MD# 338 det. PHW’; (8) red, printed ‘
|
||
<typeStatus id="54BC68E07C7A331BB149A217A2D1F9EF" box="[334,487,1587,1611]" pageId="23" pageNumber="24" type="holotype">HOLOTYPE</typeStatus>
|
||
[female] /
|
||
<taxonomicName id="4C07ADC17C7A331BB262A216A028F9EF" box="[613,798,1586,1611]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C7A331BB262A216A028F9EF" box="[613,798,1586,1611]" italics="true" pageId="23" pageNumber="24">Mendacibombus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C07ADC17C7A331BB331A216A0ADF9EF" ID-CoL="MFW5" box="[822,923,1586,1611]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C7A331BB331A216A0ADF9EF" box="[822,923,1586,1611]" italics="true" pageId="23" pageNumber="24">superbus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<bibRefCitation id="EF96ABB37C7A331BB3B4A216A77EF9EF" author="Tkalcu" box="[947,1096,1586,1611]" pageId="23" pageNumber="24" refString="Tkalcu, B. (1968) Neue Arten der Unterfamilie Bombinae der palaarktischen Region (Hymenoptera, Apoidea). Sbornik Entomologickeho oddeleni Narodniho musea v Praze, 65, 21 - 51." type="journal article" year="1968">Tkalců, 1968</bibRefCitation>
|
||
/ det.
|
||
<determinerName id="7E643A867C7A331BB496A217A615F9EF" box="[1169,1315,1586,1611]" pageId="23" pageNumber="24">PH Williams</determinerName>
|
||
2012’; (9) white, printed ‘[female]
|
||
<taxonomicName id="4C07ADC17C7A331BB1AEA272A132F9CB" ID-CoL="62H8K" box="[425,516,1622,1647]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C7A331BB1AEA272A132F9CB" box="[425,516,1622,1647]" italics="true" pageId="23" pageNumber="24">Bombus</emphasis>
|
||
</taxonomicName>
|
||
/ (
|
||
<taxonomicName id="4C07ADC17C7A331BB225A272A1EDF9CB" box="[546,731,1622,1647]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C7A331BB225A272A1EDF9CB" box="[546,731,1622,1647]" italics="true" pageId="23" pageNumber="24">Mendacibombus</emphasis>
|
||
</taxonomicName>
|
||
) /
|
||
<taxonomicName id="4C07ADC17C7A331BB2FCA272A056F9CB" ID-CoL="MFW5" box="[763,864,1622,1647]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C7A331BB2FCA272A056F9CB" box="[763,864,1622,1647]" italics="true" pageId="23" pageNumber="24">superbus</emphasis>
|
||
</taxonomicName>
|
||
/ det
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B6FDC1F7C7A331BB3A1A273A14CF937" ID-GBIF-Occurrence="1413110448" country="Philippines" location="Tkalcu's" municipality="This" pageId="23" pageNumber="24" specimenCount="1" typeStatus="holotype">
|
||
<collectingCountry id="F31096D27C7A331BB3A1A273A0FCF9CA" box="[934,970,1623,1646]" name="Philippines" pageId="23" pageNumber="24">PH</collectingCountry>
|
||
Williams 2012’.
|
||
<collectingMunicipality id="6BDC4C387C7A331BB48AA272A789F9CB" box="[1165,1215,1622,1647]" pageId="23" pageNumber="24">This</collectingMunicipality>
|
||
specimen, which is complete, is regarded as
|
||
<location id="8ED880997C7A331BB1AAA25EA13BF937" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB1AAA25EA13BF937" box="[429,525,1658,1683]" country="Philippines" municipality="This" name="Tkalcu's" pageId="23" pageNumber="24">Tkalců’s</location>
|
||
<typeStatus id="54BC68E07C7A331BB213A25EA14CF937" box="[532,634,1658,1683]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C7A3314B0C0A2BAA7C5FDB3" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
|
||
<paragraph id="8BB8D6427C7A3314B0C0A2BAA7C5FDB3" blockId="23.[151,1437,1370,2007]" lastBlockId="24.[151,1437,151,2013]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
|
||
<materialsCitation id="3B6FDC1F7C7A331BB0C0A2BAA20AF82B" ID-GBIF-Occurrence="1413110434" collectorName="E. Weiske & A. Ebmer & G. Potanin's & Komarov & However & Mondy & Tkalcu & Monda" country="Russia" county="western Nepal" elevation="1300" latitude="51.67281" location="The" longLatPrecision="1" longitude="100.98458" municipality="Mongolian" pageId="23" pageNumber="24" specimenCount="2" stateProvince="Buryatia" typeStatus="holotype">
|
||
<location id="8ED880997C7A331BB0C0A2BAA3C5F913" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB0C0A2BAA3C5F913" box="[199,243,1694,1719]" country="Russia" county="western Nepal" latitude="51.67281" longLatPrecision="1" longitude="100.98458" municipality="Mongolian" name="The" pageId="23" pageNumber="24" stateProvince="Buryatia">The</location>
|
||
<typeStatus id="54BC68E07C7A331BB0FDA2BBA21CF913" box="[250,298,1695,1719]" pageId="23" pageNumber="24">type</typeStatus>
|
||
locality ‘
|
||
<location id="8ED880997C7A331BB195A2BAA132F913" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB195A2BAA132F913" box="[402,516,1694,1719]" country="Russia" county="western Nepal" latitude="51.67281" longLatPrecision="1" longitude="100.98458" municipality="Mongolian" name="Mongolei" pageId="23" pageNumber="24" stateProvince="Buryatia">Mongolei</location>
|
||
/
|
||
<location id="8ED880997C7A331BB21CA2BAA142F913" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB21CA2BAA142F913" box="[539,628,1694,1719]" country="Russia" county="western Nepal" latitude="51.67281" longLatPrecision="1" longitude="100.98458" municipality="Mongolian" name="Monda'" pageId="23" pageNumber="24" stateProvince="Buryatia">Monda’</location>
|
||
and the style of the printed label appear to refer to collections made by the
|
||
<collectorName id="26F2B3947C7A331BB0C6A2E7A204F97F" box="[193,306,1730,1755]" pageId="23" pageNumber="24">E. Weiske</collectorName>
|
||
1908 expedition (
|
||
<bibRefCitation id="EF96ABB37C7A331BB1F9A2E6A19FF97F" author="Kerzhner" box="[510,681,1730,1755]" pageId="23" pageNumber="24" refString="Kerzhner, I. M. (1972) Historical survey of studies of the insect fauna of the Mongolian People's Republic. Nasekomye Mongolii, Leningrad, 1, 57 - 112." type="journal article" year="1972">
|
||
<location id="8ED880997C7A331BB1F9A2E6A15EF97F" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB1F9A2E6A15EF97F" box="[510,616,1730,1755]" country="Russia" county="western Nepal" latitude="51.67281" longLatPrecision="1" longitude="100.98458" municipality="Mongolian" name="Kerzhner" pageId="23" pageNumber="24" stateProvince="Buryatia">Kerzhner</location>
|
||
1972
|
||
</bibRefCitation>
|
||
) near the relatively well known town of
|
||
<location id="8ED880997C7A331BB471A2E6A7FEF97F" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C7A331BB471A2E6A7FEF97F" box="[1142,1224,1730,1755]" country="Russia" county="western Nepal" latitude="51.67281" longLatPrecision="1" longitude="100.98458" municipality="Mongolian" name="Mondy" pageId="23" pageNumber="24" stateProvince="Buryatia">Mondy</location>
|
||
,
|
||
<collectingRegion id="49C318A07C7A331BB4D4A2E6A600F97F" box="[1235,1334,1730,1755]" country="Russia" name="Buryatiya" pageId="23" pageNumber="24">Buryatia</collectingRegion>
|
||
, near the
|
||
<collectingMunicipality id="6BDC4C387C7A331BB090A2C2A224F95B" box="[151,274,1766,1791]" pageId="23" pageNumber="24">Mongolian</collectingMunicipality>
|
||
border (
|
||
<collectingCountry id="F31096D27C7A331BB177A2C2A289F95B" box="[368,447,1766,1791]" name="Russia" pageId="23" pageNumber="24">Russia</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE33B0857C7A331BB1CEA2C2A164F95B" box="[457,594,1766,1791]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="51.67281">51.67281°N</geoCoordinate>
|
||
<geoCoordinate id="EE33B0857C7A331BB25FA2C2A1D8F95B" box="[600,750,1766,1791]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="100.98458">100.98458°E</geoCoordinate>
|
||
,
|
||
<quantity id="4CFF7BA77C7A331BB2FEA2C3A067F95B" box="[761,849,1767,1791]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="24" unit="m" value="1300.0">
|
||
<elevation id="002A31717C7A331BB2FEA2C3A067F95B" box="[761,849,1767,1791]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="24" unit="m" value="1300.0">1300 m</elevation>
|
||
</quantity>
|
||
).
|
||
<collectorName id="26F2B3947C7A331BB363A2C3A0E7F95B" box="[868,977,1766,1791]" pageId="23" pageNumber="24">A. Ebmer</collectorName>
|
||
(in litt.) also points to the information on
|
||
<collectorName id="26F2B3947C7A331BB090A32FA228F887" box="[151,286,1802,1827]" pageId="23" pageNumber="24">G. Potanin’s</collectorName>
|
||
itinerary in 1880 given by
|
||
<bibRefCitation id="EF96ABB37C7A331BB257A32FA03AF887" author="Komarov" box="[592,780,1803,1827]" pageId="23" pageNumber="24" refString="Komarov, V. L. (1928) Les itineraires botaniques des principales expeditions russes en Asie Centrale. Itineraires de G. N. Potanine (1876 - 1899) Trudy Imperatorskago S. - Peterburgskago botanicheskago sada, 34, 201 - 404." type="journal article" year="1928">
|
||
<collectorName id="26F2B3947C7A331BB257A32FA18CF886" box="[592,698,1803,1826]" pageId="23" pageNumber="24">Komarov</collectorName>
|
||
(1928)
|
||
</bibRefCitation>
|
||
as evidence that these names both refer to the same place.
|
||
<collectorName id="26F2B3947C7A331BB090A30BA3C9F8E2" box="[151,255,1839,1862]" pageId="23" pageNumber="24">However</collectorName>
|
||
, the interpretation that the
|
||
<typeStatus id="54BC68E07C7A331BB257A30BA1B6F8E3" box="[592,640,1839,1863]" pageId="23" pageNumber="24">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C07ADC17C7A331BB312A30BA097F8E3" box="[789,929,1838,1863]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C7A331BB312A30BA097F8E3" box="[789,929,1838,1863]" italics="true" pageId="23" pageNumber="24">B. superbus</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<collectorName id="26F2B3947C7A331BB3D7A30AA717F8E3" box="[976,1057,1838,1863]" pageId="23" pageNumber="24">Mondy</collectorName>
|
||
was challenged in the original publication by
|
||
<bibRefCitation id="EF96ABB37C7A331BB143A376A2DEF8CF" author="Tkalcu" box="[324,488,1874,1899]" pageId="23" pageNumber="24" refString="Tkalcu, B. (1968) Neue Arten der Unterfamilie Bombinae der palaarktischen Region (Hymenoptera, Apoidea). Sbornik Entomologickeho oddeleni Narodniho musea v Praze, 65, 21 - 51." type="journal article" year="1968">
|
||
<collectorName id="26F2B3947C7A331BB143A376A2A4F8CF" box="[324,402,1874,1899]" pageId="23" pageNumber="24">Tkalců</collectorName>
|
||
(1968)
|
||
</bibRefCitation>
|
||
, because the spelling
|
||
<collectorName id="26F2B3947C7A331BB2E1A376A003F8CF" box="[742,821,1874,1899]" pageId="23" pageNumber="24">Monda</collectorName>
|
||
was known to him for a locality in the mountains of
|
||
<collectingCounty id="62D9AECE7C7A331BB090A353A20AF82B" box="[151,316,1910,1935]" pageId="23" pageNumber="24">western Nepal</collectingCounty>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B6FDC1F7C7A3314B142A353A7C5FDB3" ID-GBIF-Occurrence="1413110436" collectionCode="IAR, IZB, MNHU" collectorName="The & H. Friese & Ebmer & Monda' & Mondy & Friese & V. Roborovski & P. Kozlov & This & Roborovski & Kozlov & Bretschneider & Therefore & Qinghai-Tibetan & Even" country="Russia" elevation="4600" lastPageId="24" lastPageNumber="25" location="eastern Qinghai-Tibetan" pageId="23" pageNumber="24" specimenCount="6" stateProvince="Qinghai and Xizang" typeStatus="holotype">
|
||
A likely Nepalese site with this name could not be identified by us. The long hair of the
|
||
<typeStatus id="54BC68E07C7A331BB51BA352A648F82B" box="[1308,1406,1910,1935]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C07ADC17C7A331BB090A3B8A229F817" box="[151,287,1946,1971]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C7A331BB090A3B8A229F817" box="[151,287,1946,1971]" italics="true" pageId="23" pageNumber="24">B. superbus</emphasis>
|
||
</taxonomicName>
|
||
resembles the hair of other species from very high elevations (much higher than Mondy). This is consistent with the extreme habitat of the other known specimens of this species (
|
||
<collectionCode id="ED164E877C7A331BB45DA39BA7B8F873" box="[1114,1166,1983,2007]" pageId="23" pageNumber="24">IAR</collectionCode>
|
||
, MD#4110‒4131;
|
||
<collectionCode id="ED164E877C7A331BB56FA39BA6AEF872" box="[1384,1432,1983,2006]" pageId="23" pageNumber="24">IZB</collectionCode>
|
||
, MD#339‒342), which are from above
|
||
<quantity id="4CFF7BA77C753314B248A4B3A193FF0B" box="[591,677,151,176]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="24" pageNumber="25" unit="m" value="4600.0">
|
||
<elevation id="002A31717C753314B248A4B3A193FF0B" box="[591,677,151,176]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="24" pageNumber="25" unit="m" value="4600.0">4600 m</elevation>
|
||
</quantity>
|
||
in
|
||
<collectingRegion id="49C318A07C753314B2CCA4B3A082FF14" box="[715,948,151,176]" isEnumeration="true" pageId="24" pageNumber="25">Qinghai and Xizang</collectingRegion>
|
||
.
|
||
<collectorName id="26F2B3947C753314B3B8A4B3A0DDFF14" box="[959,1003,151,176]" pageId="24" pageNumber="25">The</collectorName>
|
||
<typeStatus id="54BC68E07C753314B3F4A4B3A763FF14" box="[1011,1109,151,176]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
specimen is in the
|
||
<collectorName id="26F2B3947C753314B533A4BCA6ABFF14" box="[1332,1437,151,176]" pageId="24" pageNumber="25">H. Friese</collectorName>
|
||
collection (
|
||
<collectionCode id="ED164E877C753314B112A498A259FF70" box="[277,367,188,212]" pageId="24" pageNumber="25">MNHU</collectionCode>
|
||
, MD#338), a collection that is known to have a number of bumblebee specimens with dubious locality data (see the notes on the nominal taxa
|
||
<emphasis id="B9730A507C753314B2DAA4C4A07CFF5C" box="[733,842,224,248]" italics="true" pageId="24" pageNumber="25">rufitarsus</emphasis>
|
||
and
|
||
<taxonomicName id="4C07ADC17C753314B38BA4FBA0ECFF5C" box="[908,986,223,248]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="asellus">
|
||
<emphasis id="B9730A507C753314B38BA4FBA0ECFF5C" box="[908,986,223,248]" italics="true" pageId="24" pageNumber="25">asellus</emphasis>
|
||
</taxonomicName>
|
||
under
|
||
<taxonomicName id="4C07ADC17C753314B434A4C5A79DFF5C" box="[1075,1195,223,248]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C753314B434A4C5A79DFF5C" box="[1075,1195,223,248]" italics="true" pageId="24" pageNumber="25">B. waltoni</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<bibRefCitation id="EF96ABB37C753314B4CFA4FBA65CFF5C" author="Ebmer" box="[1224,1386,223,248]" pageId="24" pageNumber="25" refString="Ebmer, A. W. (2008) Die westpalaarktischen Arten der Gattung Dufourea LEPELETIER 1841 (Insecta: Hymenoptera: Apoidea: Halictidae: Rophitinae) Funfter Nachtrag. Linzer biologische Beitrage, 40, 581 - 625." type="journal article" year="2008">
|
||
<collectorName id="26F2B3947C753314B4CFA4FBA625FF5C" box="[1224,1299,223,248]" pageId="24" pageNumber="25">Ebmer</collectorName>
|
||
(2008)
|
||
</bibRefCitation>
|
||
has described how for other bees with ‘
|
||
<collectorName id="26F2B3947C753314B21AA520A14CFEB9" box="[541,634,260,285]" pageId="24" pageNumber="25">Monda’</collectorName>
|
||
labels, while in some cases they appear to refer genuinely to
|
||
<collectorName id="26F2B3947C753314B52EA520A64DFEB9" box="[1321,1403,260,285]" pageId="24" pageNumber="25">Mondy</collectorName>
|
||
, in other cases these labels appear to be spurious later additions and he discussed how this might have arisen.
|
||
<collectorName id="26F2B3947C753314B555A503A6ABFEE4" box="[1362,1437,295,320]" pageId="24" pageNumber="25">Ebmer</collectorName>
|
||
suggests that
|
||
<collectorName id="26F2B3947C753314B12AA568A244FEC1" box="[301,370,332,357]" pageId="24" pageNumber="25">Friese</collectorName>
|
||
described some of the material collected by the
|
||
<collectorName id="26F2B3947C753314B38CA569A71AFEC1" box="[907,1068,332,357]" pageId="24" pageNumber="25">V. Roborovski</collectorName>
|
||
and
|
||
<collectorName id="26F2B3947C753314B465A569A7F8FEC1" box="[1122,1230,332,357]" pageId="24" pageNumber="25">P. Kozlov</collectorName>
|
||
expeditions of the 1890s and that some specimens from these expeditions ended up erroneously with ‘
|
||
<collectorName id="26F2B3947C753314B466A54BA788FE2C" box="[1121,1214,367,392]" pageId="24" pageNumber="25">Monda’</collectorName>
|
||
labels.
|
||
<collectorName id="26F2B3947C753314B51BA54BA678FE2C" box="[1308,1358,367,392]" pageId="24" pageNumber="25">This</collectorName>
|
||
seems likely in this case.
|
||
<collectorName id="26F2B3947C753314B16FA5B0A2DDFE09" box="[360,491,404,429]" pageId="24" pageNumber="25">Roborovski</collectorName>
|
||
and
|
||
<collectorName id="26F2B3947C753314B226A5B0A142FE09" box="[545,628,404,429]" pageId="24" pageNumber="25">Kozlov</collectorName>
|
||
took part in expeditions that visited the
|
||
<location id="8ED880997C753314B431A5B1A672FE09" LSID="urn:lsid:plazi:treatment:03AE67547C7A3317B090A017A2EEFBB6:8ED880997C753314B431A5B1A672FE09" box="[1078,1348,404,429]" country="Russia" name="eastern Qinghai-Tibetan" pageId="24" pageNumber="25" stateProvince="Qinghai and Xizang">eastern Qinghai-Tibetan</location>
|
||
plateau in the summers of 1879, 1880, 1884, 1894, and 1895 (
|
||
<bibRefCitation id="EF96ABB37C753314B2F4A593A0E2FE74" author="Bretschneider" box="[755,980,439,464]" pageId="24" pageNumber="25" refString="Bretschneider, E. (1981) History of European botanical discoveries in China (Vol. 2). Leipzig, Zentral - Antiquariat der DDR." type="journal volume" year="1981">
|
||
<collectorName id="26F2B3947C753314B2F4A593A0A6FE74" box="[755,912,439,464]" pageId="24" pageNumber="25">Bretschneider</collectorName>
|
||
1981
|
||
</bibRefCitation>
|
||
).
|
||
<collectorName id="26F2B3947C753314B3E0A593A763FE74" box="[999,1109,439,464]" pageId="24" pageNumber="25">Therefore</collectorName>
|
||
we interpret the origin of the
|
||
<typeStatus id="54BC68E07C753314B090A5F8A3CFFE51" box="[151,249,476,501]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
and the
|
||
<typeStatus id="54BC68E07C753314B150A5F9A2B1FE51" box="[343,391,477,501]" pageId="24" pageNumber="25">type</typeStatus>
|
||
locality as most likely to have been on the
|
||
<collectorName id="26F2B3947C753314B36EA5F8A717FE51" box="[873,1057,476,501]" pageId="24" pageNumber="25">Qinghai-Tibetan</collectorName>
|
||
plateau.
|
||
<collectorName id="26F2B3947C753314B481A5F9A7F6FE50" box="[1158,1216,477,500]" pageId="24" pageNumber="25">Even</collectorName>
|
||
so, the precise
|
||
<typeStatus id="54BC68E07C753314B56BA5F9A6AAFE51" box="[1388,1436,477,501]" pageId="24" pageNumber="25">type</typeStatus>
|
||
locality remains very uncertain and consequently no location for the
|
||
<typeStatus id="54BC68E07C753314B39EA5DBA0CDFDBC" box="[921,1019,511,536]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
is shown on our map.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C753314B0C0A600A087FCF9" pageId="24" pageNumber="25" type="etymology">
|
||
<paragraph id="8BB8D6427C753314B0C0A600A083FDC4" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9730A507C753314B0C0A600A278FD99" bold="true" box="[199,334,548,573]" pageId="24" pageNumber="25">Etymology.</emphasis>
|
||
The species is named from the Latin
|
||
<taxonomicName id="4C07ADC17C753314B310A600A04AFD99" box="[791,892,548,573]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C753314B310A600A04AFD99" box="[791,892,548,573]" italics="true" pageId="24" pageNumber="25">superbus</emphasis>
|
||
</taxonomicName>
|
||
for its resplendent (‘prangend’) or sumptuous (‘prächtig’) appearance according to the original (German) description.
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C753314B0C0A648A087FCF9" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9730A507C753314B0C0A648A2C0FD21" bold="true" box="[199,502,620,645]" pageId="24" pageNumber="25">Taxonomy and variation.</emphasis>
|
||
This species shows a single colour pattern of the hair with little variation. This is the pattern in the original description, which is unique within the subgenus. All known specimens of the species have a yellow-banded and none has a white-banded colour pattern. The males from the Tanggula mountains (MD#339, 4131) have the yellow anterior collar of the thoracic dorsum narrower, barely reaching posteriorly to the tegula, and with many black hairs intermixed, and the metasomal terga 4‒7 have many of the hairs white-tipped. The queens and workers are unusually large for species of this subgenus, although the males are similar in size to those of the other species. The form of the male genitalia is diagnostic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C753314B0C0A743A76EFA39" pageId="24" pageNumber="25" type="description">
|
||
<paragraph id="8BB8D6427C753314B0C0A743A76EFA39" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9730A507C753314B0C0A743A2E9FC24" bold="true" box="[199,479,871,896]" pageId="24" pageNumber="25">Diagnostic description.</emphasis>
|
||
Wings nearly clear. Hair long, uneven, and slightly sparse.
|
||
<emphasis id="B9730A507C753314B4B1A743A3C7FC00" italics="true" pageId="24" pageNumber="25">Female hair colour pattern:</emphasis>
|
||
(
|
||
<figureCitation id="133CCAC77C753314B103A7A8A26FFC01" box="[260,345,908,933]" captionStart="FIGURES 69 ‒ 70" captionStartId="59.[151,264,1618,1640]" captionTargetBox="[151,1436,259,1554]" captionTargetId="figure@59.[151,1436,193,1653]" captionTargetPageId="59" captionText="FIGURES 69 ‒ 70. Photos of (69) a worker of B. superbus (MD # 4123) from (left) the dorsal and (right) the left lateral aspect (scale bar 10 mm), from the sample site (70) in the Tanggula mountains, Xizang, China (16. viii. 2014, 32.86142 ° N 91.91739 ° E, GPS elevation ca 5220 m). Photos by (69) NHM digital imaging unit and (70) Z. - Y. Miao." httpUri="https://zenodo.org/record/192317/files/figure.png" pageId="24" pageNumber="25">Fig. 69</figureCitation>
|
||
) generally black, the head sometimes with some yellow hairs dorsally posterior to the ocelli, bright yellow hair in a transverse band in almost the anterior half on the thoracic dorsum and extending laterally and ventrally to just below the wing bases, and on metasomal T1‒2, although T2 with a few black hairs intermixed along the posterior margin, T3‒6 predominantly black with some white tips or sometimes a few hairs entirely white (
|
||
<emphasis id="B9730A507C753314B0A7A038A382FB90" box="[160,180,1052,1076]" italics="true" pageId="24" pageNumber="25">cf</emphasis>
|
||
. all other
|
||
<taxonomicName id="4C07ADC17C753314B12CA038A2D2FB91" box="[299,484,1052,1077]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C753314B12CA038A2D2FB91" box="[299,484,1052,1077]" italics="true" pageId="24" pageNumber="25">Mendacibombus</emphasis>
|
||
</taxonomicName>
|
||
species). Midleg basitarsus with the hair predominantly orange, with long black or orange hairs arising from the outer surface in the proximal third (more apparent in workers than queens); hindleg tibia with the corbicular fringes black, the posterior fringe with some hairs with orange tips or distally some hairs entirely orange (more for queens than for workers); hindleg basitarsus with the hair predominantly orange, with the long posterior fringe orange.
|
||
<emphasis id="B9730A507C753314B1EEA088A1E1FB61" box="[489,727,1196,1221]" italics="true" pageId="24" pageNumber="25">Female morphology:</emphasis>
|
||
labrum with the basal depression very narrow, the transverse ridge very broad and low, medially subsiding but not interrupted, with large punctures in the median third, lateral tubercles with few punctures. T2 without a posteriorly-directed convexity of its median posterior edge (even for the queens,
|
||
<emphasis id="B9730A507C753314B0F3A13CA2A8FA94" box="[244,414,1304,1328]" italics="true" pageId="24" pageNumber="25">
|
||
cf.
|
||
<taxonomicName id="4C07ADC17C753314B110A13DA2A8FA94" box="[279,414,1305,1328]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="convexus">B. convexus</taxonomicName>
|
||
</emphasis>
|
||
).
|
||
<emphasis id="B9730A507C753314B1B1A133A149FA94" box="[438,639,1303,1328]" italics="true" pageId="24" pageNumber="25">Male morphology</emphasis>
|
||
: beard of the mandible long, dense and black; genitalia (
|
||
<figureCitation id="133CCAC77C753314B500A133A66CFA94" box="[1287,1370,1303,1328]" captionStart="FIGURES 24 ‒ 35" captionStartId="26.[151,264,1867,1889]" captionTargetBox="[233,1387,193,1789]" captionTargetId="figure@26.[233,1390,193,1894]" captionTargetPageId="26" captionText="FIGURES 24 ‒ 35. Morphology of the male genitalia for species of the subgenus Mendacibombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 24, B. superbus, China MD # 339; 25, B. waltoni, China MD # 216; 26, B. convexus, China MD # 215; 27, B. makarjini, Kazakhstan MD # 1240; 28, B. marussinus, Pakistan MD # 408; 29, B. avinoviellus, India MD # 404; 30, B. himalayanus, India MD # 426; 31, B. turkestanicus, China MD # 351; 32, B. defector, Kyrgyzstan MD # 367; 33, B. margreiteri, China MD # 294; 34, B. handlirschianus, Russia MD # 835; 35, B. mendax, Spain MD # 689." httpUri="https://zenodo.org/record/192312/files/figure.png" pageId="24" pageNumber="25">Fig. 24</figureCitation>
|
||
) with the volsella at its broadest near the midpoint of its length, the dorsal surface just distal to this point with a raised curved ridge, just inside the inner margin, running for 0.5× the remaining distal length of the volsella; volsella distally a rounded right angle and barely curled back dorsally and not at all anteriorly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D85C97C753314B0C0A183A2FEF879" pageId="24" pageNumber="25" type="materials_examined">
|
||
<paragraph id="8BB8D6427C753314B0C0A183A02BFA40" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9730A507C753314B0C0A183A286FA64" bold="true" box="[199,432,1447,1472]" pageId="24" pageNumber="25">Material examined.</emphasis>
|
||
3 queens
|
||
<specimenCount id="9D011DCB7C753314B227A18CA1ABFA64" box="[544,669,1447,1472]" pageId="24" pageNumber="25" type="worker">22 workers</specimenCount>
|
||
<specimenCount id="9D011DCB7C753314B2A4A18CA1CAFA64" box="[675,764,1447,1472]" pageId="24" pageNumber="25" type="male">2 males</specimenCount>
|
||
, from
|
||
<collectingCountry id="F31096D27C753314B343A183A0B1FA64" box="[836,903,1447,1472]" name="China" pageId="24" pageNumber="25">China</collectingCountry>
|
||
(
|
||
<figureCitation id="133CCAC77C753314B391A183A0DCFA64" box="[918,1002,1447,1472]" captionStart="FIGURES 56 ‒ 67" captionStartId="35.[151,264,1891,1913]" captionTargetBox="[320,1245,193,1835]" captionTargetId="figure@35.[320,1245,193,1843]" captionTargetPageId="35" captionText="FIGURES 56 ‒ 67. Maps of the global distributions of species of the subgenus Mendacibombus from the material examined, with grey spots for specimen records and black spots for the samples with COI sequences (each species is unknown from regions outside the map boundaries shown); grey crosses for records of all species of Mendacibombus combined. Relief map with hill shading, Cartesian orthonormal projection, grey lines show borders to national administration according to UN maps, north at the top of the map. Image created in ArcGIS using World _ Shaded _ Relief basemap © 2014 Esri." httpUri="https://zenodo.org/record/192315/files/figure.png" pageId="24" pageNumber="25">Fig. 56</figureCitation>
|
||
:
|
||
<collectionCode id="ED164E877C753314B3F2A18CA71EFA64" box="[1013,1064,1448,1472]" pageId="24" pageNumber="25">IAR</collectionCode>
|
||
,
|
||
<collectionCode id="ED164E877C753314B434A18CA755FA1B" box="[1075,1123,1448,1471]" pageId="24" pageNumber="25">IZB</collectionCode>
|
||
,
|
||
<collectionCode id="ED164E877C753314B469A18CA7FEFA64" box="[1134,1224,1448,1472]" pageId="24" pageNumber="25">MNHU</collectionCode>
|
||
), with
|
||
<specimenCount id="9D011DCB7C753314B514A18CA6ABFA64" box="[1299,1437,1447,1472]" pageId="24" pageNumber="25" type="generic">7 specimens</specimenCount>
|
||
sequenced (interpretable sequences listed in
|
||
<figureCitation id="133CCAC77C753314B28EA1E8A026FA40" box="[649,784,1484,1509]" captionStart-0="FIGURES 11 ‒ 12" captionStart-1="FIGURE 13" captionStartId-0="15.[151,264,1630,1652]" captionStartId-1="16.[151,250,1733,1755]" captionTargetBox-0="[300,1121,202,1610]" captionTargetBox-1="[314,1260,194,1721]" captionTargetId-0="figure@15.[292,1266,193,1625]" captionTargetId-1="figure@16.[314,1261,193,1722]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 11 ‒ 12. Recognising species from an estimate of phylogeny for 30 unique COI haplotypes of the subgenus Mendacibombus by BEAST, using GMYC models and the PTP procedure. From the GMYC models (11), the single threshold T is shown by the vertical grey bar so that the intersecting lineages are interpreted as indicating 12 separate species (black spots at coalescent nodes; outgroups not shown). Values below the nodes of the maximum clade-credibility tree are Bayesian posterior probabilities showing branch support (groups with values <0.8 are considered unreliable) from the BEAST analysis with two billion MCMC generations and a 1 % burn-in. The x - scale axis is calibrated in substitutions per nucleotide site. From the PTP procedure (12), the Bayesian PTP solution with highest support also indicates 12 species. Values above the nodes are PTP Bayesian support values that all daughter haplotypes are parts of a single species; values below the nodes are Bayesian posterior probabilities showing branch support. The scale bar is calibrated in substitutions per nucleotide site. Each unique haplotype is represented by one of the longest available sample sequences, labelled with: the sequence length in number of base pairs, a taxon name or name for an unpublished colour pattern (light / dark / yellow / white / pale), and then a code that consists of a specimen identifier (MD #) from the project database and (after the hyphen) a sample identifier from BOLD or from GenBank, followed with its geographic origin." captionText-1="FIGURE 13. Assigning names to species from an estimate of phylogeny for 130 COI sequences of the subgenus Mendacibombus. Based on a BEAST analysis of the 30 unique haplotypes (Fig. 11), with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is added after the first matching sequence, with the addition of the two short-sequence yellow unbanded specimens from Pakistan, MD # 4058, 4059). The sequence labels follow the format used in Figs 11, 12. Lineages accepted as species from the GMYC models (Fig. 11) are shown as thick branches, the most recent common ancestors of each species are shown with a black spot, and branches within species are shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names (Table 1). The proxy sample for the type specimen for the oldest available name (the valid name) for each species (from Fig. 11) is marked in bold. The interpretations of the species with their valid names are shown in grey to the right." httpUri-0="https://zenodo.org/record/192308/files/figure.png" httpUri-1="https://zenodo.org/record/192309/files/figure.png" pageId="24" pageNumber="25">Figs. 11–13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C753314B0C0A1CBA2FEF879" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="25">
|
||
<emphasis id="B9730A507C753314B0C0A1CBA2C4F9AC" bold="true" box="[199,498,1519,1544]" pageId="24" pageNumber="25">Habitat and distribution.</emphasis>
|
||
Flower-rich alpine grassland, at elevations 4800‒(5167)‒
|
||
<quantity id="4CFF7BA77C753314B472A1CBA7FDF9A3" box="[1141,1227,1519,1544]" metricMagnitude="3" metricUnit="m" metricValue="5.22" pageId="24" pageNumber="25" unit="m" value="5220.0">5220 m</quantity>
|
||
a.s.l.. A species of the high Qinghai-Tibetan plateau, known so far definitely only from the Hohxilshan (= Kekexili; collected 1990, X.-Z. Zhang) within the eastern Kunlun mountain system and from the Tanggulashan (collected 2014, Z.-Y. Miao,
|
||
<figureCitation id="133CCAC77C753314B090A278A22EF9D0" box="[151,280,1628,1653]" captionStart="FIGURES 69 ‒ 70" captionStartId="59.[151,264,1618,1640]" captionTargetBox="[151,1436,259,1554]" captionTargetId="figure@59.[151,1436,193,1653]" captionTargetPageId="59" captionText="FIGURES 69 ‒ 70. Photos of (69) a worker of B. superbus (MD # 4123) from (left) the dorsal and (right) the left lateral aspect (scale bar 10 mm), from the sample site (70) in the Tanggula mountains, Xizang, China (16. viii. 2014, 32.86142 ° N 91.91739 ° E, GPS elevation ca 5220 m). Photos by (69) NHM digital imaging unit and (70) Z. - Y. Miao." httpUri="https://zenodo.org/record/192317/files/figure.png" pageId="24" pageNumber="25">Figs 69, 70</figureCitation>
|
||
) on the Xizang-Qinghai border. This apparent rarity could be explained by the very limited sampling for bumblebees at the highest elevations in the central and northern Qinghai-Tibetan plateau, because access in this region is difficult, with few roads. In this case, the mapped distribution in
|
||
<figureCitation id="133CCAC77C753314B3FAA280A779F919" box="[1021,1103,1700,1725]" captionStart="FIGURES 56 ‒ 67" captionStartId="35.[151,264,1891,1913]" captionTargetBox="[320,1245,193,1835]" captionTargetId="figure@35.[320,1245,193,1843]" captionTargetPageId="35" captionText="FIGURES 56 ‒ 67. Maps of the global distributions of species of the subgenus Mendacibombus from the material examined, with grey spots for specimen records and black spots for the samples with COI sequences (each species is unknown from regions outside the map boundaries shown); grey crosses for records of all species of Mendacibombus combined. Relief map with hill shading, Cartesian orthonormal projection, grey lines show borders to national administration according to UN maps, north at the top of the map. Image created in ArcGIS using World _ Shaded _ Relief basemap © 2014 Esri." httpUri="https://zenodo.org/record/192315/files/figure.png" pageId="24" pageNumber="25">Fig. 56</figureCitation>
|
||
could be a small part of the species’ true distribution, which could be broader, although patchy, at very high elevations. Alternatively, the map may be a more precise reflection of the species’ true distribution, limited by aridity (in the Qiangtang semi-arid area: C.-Z.
|
||
<bibRefCitation id="EF96ABB37C753314B11EA334A2EBF88C" author="Wang" box="[281,477,1807,1832]" pageId="24" pageNumber="25" refString="Wang, C. - Z., Guo, H. - D., Zhang, L., Qiu, Y. - B., Sun, Z. - C., Liao, J. - J., et al. (2013) Improved alpine grassland mapping in the Tibetan Plateau with MODIS time series: a phenology perspective. International Journal of Digital Earth." type="book" year="2013">
|
||
Wang
|
||
<emphasis id="B9730A507C753314B165A335A2ADF88C" box="[354,411,1807,1832]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
) to a few small patches in the central and northern mountain groups where there are sufficiently large permanent streams to provide water to sustain flower-rich grassland reliably throughout every summer (P.H.
|
||
<bibRefCitation id="EF96ABB37C753314B13CA373A1FEF8D4" author="Williams" box="[315,712,1879,1904]" pageId="24" pageNumber="25" refString="Williams, P. H., Bystriakova, N., Huang, J. - X., Miao, Z. - Y. & An, J. - D. (2015) Bumblebees, climate and glaciers across the Tibetan plateau (Apidae: Bombus Latreille). Systematics and Biodiversity, 13, 164 - 181." type="journal article" year="2015">
|
||
Williams, Bystriakova,
|
||
<emphasis id="B9730A507C753314B24FA37DA1B4F8D4" box="[584,642,1879,1904]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
). For example, suitable patches may occur around the large ice caps further west in the Tanggulashan.
|
||
<taxonomicName id="4C07ADC17C753314B256A358A02EF831" box="[593,792,1916,1941]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C753314B256A358A02EF831" box="[593,792,1916,1941]" italics="true" pageId="24" pageNumber="25">Bombus superbus</emphasis>
|
||
</taxonomicName>
|
||
may be one of the few truly endemic bumblebee species of the extreme habitats of the high Qinghai-Tibetan plateau. This species overlaps with
|
||
<taxonomicName id="4C07ADC17C753314B465A385A7E2F81D" box="[1122,1236,1952,1977]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C753314B465A385A7E2F81D" box="[1122,1236,1952,1977]" italics="true" pageId="24" pageNumber="25">B. waltoni</emphasis>
|
||
</taxonomicName>
|
||
in the Hohxil and in the Tanggula mountains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7886CA7C743315B090A318A3CFF84F" httpUri="https://zenodo.org/record/192311/files/figure.png" pageId="25" pageNumber="26" targetBox="[186,1380,193,1766]" targetPageId="25">
|
||
<paragraph id="8BB8D6427C743315B090A318A3CFF84F" blockId="25.[151,1436,1852,2027]" pageId="25" pageNumber="26">
|
||
<emphasis id="B9730A507C743315B090A318A262F8F6" bold="true" box="[151,340,1852,1874]" pageId="25" pageNumber="26">FIGURES 15‒23.</emphasis>
|
||
Morphology of the female (queen) head from the ventral aspect (labrum and mandibles): 15,
|
||
<taxonomicName id="4C07ADC17C743315B50BA319A6AAF8F6" box="[1292,1436,1853,1874]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="marussinus">
|
||
<emphasis id="B9730A507C743315B50BA319A6AAF8F6" box="[1292,1436,1853,1874]" italics="true" pageId="25" pageNumber="26">B. marussinus</emphasis>
|
||
</taxonomicName>
|
||
queen, Pakistan MD#482; 16,
|
||
<taxonomicName id="4C07ADC17C743315B1D4A378A15DF8D5" box="[467,619,1883,1905]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="avinoviellus">
|
||
<emphasis id="B9730A507C743315B1D4A378A15DF8D5" box="[467,619,1883,1905]" italics="true" pageId="25" pageNumber="26">B. avinoviellus</emphasis>
|
||
</taxonomicName>
|
||
queen, India MD#598; 17,
|
||
<taxonomicName id="4C07ADC17C743315B389A378A702F8D5" box="[910,1076,1883,1905]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="turkestanicus">
|
||
<emphasis id="B9730A507C743315B389A378A702F8D5" box="[910,1076,1883,1905]" italics="true" pageId="25" pageNumber="26">B. turkestanicus</emphasis>
|
||
</taxonomicName>
|
||
queen, Tajikistan MD#320; 18,
|
||
<taxonomicName id="4C07ADC17C743315B581A378A3DCF82B" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="defector">
|
||
<emphasis id="B9730A507C743315B581A378A3DCF82B" italics="true" pageId="25" pageNumber="26">B. defector</emphasis>
|
||
</taxonomicName>
|
||
queen, Kyrgyzstan MD#1080; 19,
|
||
<taxonomicName id="4C07ADC17C743315B269A35EA035F82B" box="[622,771,1914,1935]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="margreiteri">
|
||
<emphasis id="B9730A507C743315B269A35EA035F82B" box="[622,771,1914,1935]" italics="true" pageId="25" pageNumber="26">B. margreiteri</emphasis>
|
||
</taxonomicName>
|
||
queen, Mongolia MD#319; 20,
|
||
<taxonomicName id="4C07ADC17C743315B46FA35EA63AF82B" box="[1128,1292,1913,1935]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="himalayanus">
|
||
<emphasis id="B9730A507C743315B46FA35EA63AF82B" box="[1128,1292,1913,1935]" italics="true" pageId="25" pageNumber="26">B. himalayanus</emphasis>
|
||
</taxonomicName>
|
||
queen, India MD#453. Morphology of the female (queen) left hindleg tibia from the left lateral aspect, dorsal at the top of the image, anterior to the left: 21,
|
||
<taxonomicName id="4C07ADC17C743315B183A39CA2C4F868" box="[388,498,1974,1997]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="defector">
|
||
<emphasis id="B9730A507C743315B183A39CA2C4F868" box="[388,498,1974,1997]" italics="true" pageId="25" pageNumber="26">B. defector</emphasis>
|
||
</taxonomicName>
|
||
, China MD#1251; 22,
|
||
<taxonomicName id="4C07ADC17C743315B2E7A39CA046F869" box="[736,880,1976,1997]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="margreiteri">
|
||
<emphasis id="B9730A507C743315B2E7A39CA046F869" box="[736,880,1976,1997]" italics="true" pageId="25" pageNumber="26">B. margreiteri</emphasis>
|
||
</taxonomicName>
|
||
, Mongolia MD#319; 23,
|
||
<taxonomicName id="4C07ADC17C743315B471A39CA62CF868" box="[1142,1306,1974,1997]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="turkestanicus">
|
||
<emphasis id="B9730A507C743315B471A39CA62CF868" box="[1142,1306,1974,1997]" italics="true" pageId="25" pageNumber="26">B. turkestanicus</emphasis>
|
||
</taxonomicName>
|
||
, Kyrgyzstan MD#324.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7886CA7C773316B090A36FA667F87F" httpUri="https://zenodo.org/record/192312/files/figure.png" pageId="26" pageNumber="27" targetBox="[233,1387,193,1789]" targetPageId="26">
|
||
<paragraph id="8BB8D6427C773316B090A36FA667F87F" blockId="26.[151,1436,1867,2012]" pageId="26" pageNumber="27">
|
||
<emphasis id="B9730A507C773316B090A36FA261F8C5" bold="true" box="[151,343,1867,1889]" pageId="26" pageNumber="27">FIGURES 24‒35.</emphasis>
|
||
Morphology of the male genitalia for species of the subgenus
|
||
<taxonomicName id="4C07ADC17C773316B3FEA36FA796F8C5" box="[1017,1184,1867,1889]" class="Insecta" family="Apidae" genus="Mendacibombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9730A507C773316B3FEA36FA796F8C5" box="[1017,1184,1867,1889]" italics="true" pageId="26" pageNumber="27">Mendacibombus</emphasis>
|
||
</taxonomicName>
|
||
from the dorsal aspect, anterior to the left of the image, posterior to the right: 24,
|
||
<taxonomicName id="4C07ADC17C773316B2E1A34EA068F8DB" box="[742,862,1897,1919]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="superbus">
|
||
<emphasis id="B9730A507C773316B2E1A34EA068F8DB" box="[742,862,1897,1919]" italics="true" pageId="26" pageNumber="27">B. superbus</emphasis>
|
||
</taxonomicName>
|
||
, China MD#339; 25,
|
||
<taxonomicName id="4C07ADC17C773316B439A34EA790F8DB" box="[1086,1190,1897,1919]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="waltoni">
|
||
<emphasis id="B9730A507C773316B439A34EA790F8DB" box="[1086,1190,1897,1919]" italics="true" pageId="26" pageNumber="27">B. waltoni</emphasis>
|
||
</taxonomicName>
|
||
, China MD#216; 26,
|
||
<taxonomicName id="4C07ADC17C773316B581A34EA3C2F83A" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="convexus">
|
||
<emphasis id="B9730A507C773316B581A34EA3C2F83A" italics="true" pageId="26" pageNumber="27">B. convexus</emphasis>
|
||
</taxonomicName>
|
||
, China MD#215; 27,
|
||
<taxonomicName id="4C07ADC17C773316B1D4A3ADA167F83A" box="[467,593,1928,1950]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="makarjini">
|
||
<emphasis id="B9730A507C773316B1D4A3ADA167F83A" box="[467,593,1928,1950]" italics="true" pageId="26" pageNumber="27">B. makarjini</emphasis>
|
||
</taxonomicName>
|
||
, Kazakhstan MD#1240; 28,
|
||
<taxonomicName id="4C07ADC17C773316B370A3ADA730F83A" box="[887,1030,1929,1950]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="marussinus">
|
||
<emphasis id="B9730A507C773316B370A3ADA730F83A" box="[887,1030,1929,1950]" italics="true" pageId="26" pageNumber="27">B. marussinus</emphasis>
|
||
</taxonomicName>
|
||
, Pakistan MD#408; 29,
|
||
<taxonomicName id="4C07ADC17C773316B4F8A3ADA6A3F83A" box="[1279,1429,1928,1950]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="avinoviellus">
|
||
<emphasis id="B9730A507C773316B4F8A3ADA6A3F83A" box="[1279,1429,1928,1950]" italics="true" pageId="26" pageNumber="27">B. avinoviellus</emphasis>
|
||
</taxonomicName>
|
||
, India MD#404; 30,
|
||
<taxonomicName id="4C07ADC17C773316B175A38CA122F819" box="[370,532,1959,1981]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="himalayanus">
|
||
<emphasis id="B9730A507C773316B175A38CA122F819" box="[370,532,1959,1981]" italics="true" pageId="26" pageNumber="27">B. himalayanus</emphasis>
|
||
</taxonomicName>
|
||
, India MD#426; 31,
|
||
<taxonomicName id="4C07ADC17C773316B305A38CA09DF819" box="[770,939,1959,1981]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="turkestanicus">
|
||
<emphasis id="B9730A507C773316B305A38CA09DF819" box="[770,939,1959,1981]" italics="true" pageId="26" pageNumber="27">B. turkestanicus</emphasis>
|
||
</taxonomicName>
|
||
, China MD#351; 32,
|
||
<taxonomicName id="4C07ADC17C773316B4A6A38CA623F819" box="[1185,1301,1959,1981]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="defector">
|
||
<emphasis id="B9730A507C773316B4A6A38CA623F819" box="[1185,1301,1959,1981]" italics="true" pageId="26" pageNumber="27">B. defector</emphasis>
|
||
</taxonomicName>
|
||
, Kyrgyzstan MD#367; 33,
|
||
<taxonomicName id="4C07ADC17C773316B122A3E3A283F878" box="[293,437,1991,2012]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="margreiteri">
|
||
<emphasis id="B9730A507C773316B122A3E3A283F878" box="[293,437,1991,2012]" italics="true" pageId="26" pageNumber="27">B. margreiteri</emphasis>
|
||
</taxonomicName>
|
||
, China MD#294; 34,
|
||
<taxonomicName id="4C07ADC17C773316B294A3E3A064F87F" box="[659,850,1989,2012]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="handlirschianus">
|
||
<emphasis id="B9730A507C773316B294A3E3A064F87F" box="[659,850,1989,2012]" italics="true" pageId="26" pageNumber="27">B. handlirschianus</emphasis>
|
||
</taxonomicName>
|
||
, Russia MD#835; 35,
|
||
<taxonomicName id="4C07ADC17C773316B430A3E3A794F87F" box="[1079,1186,1989,2012]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="mendax">
|
||
<emphasis id="B9730A507C773316B430A3E3A794F87F" box="[1079,1186,1989,2012]" italics="true" pageId="26" pageNumber="27">B. mendax</emphasis>
|
||
</taxonomicName>
|
||
, Spain MD#689.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31D85C97C763317B090A713A2EEFBB6" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="8BB8D6427C763317B090A713A2E4FC55" blockId="27.[151,468,823,845]" lastBlockId="27.[151,1436,917,1042]" pageId="27" pageNumber="28">
|
||
<figureCitation id="133CCAC77C763317B090A713A27BFCE9" box="[151,333,823,845]" captionStart="FIGURES 24 ‒ 35" captionStartId="26.[151,264,1867,1889]" captionTargetBox="[233,1387,193,1789]" captionTargetId="figure@26.[233,1390,193,1894]" captionTargetPageId="26" captionText="FIGURES 24 ‒ 35. Morphology of the male genitalia for species of the subgenus Mendacibombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 24, B. superbus, China MD # 339; 25, B. waltoni, China MD # 216; 26, B. convexus, China MD # 215; 27, B. makarjini, Kazakhstan MD # 1240; 28, B. marussinus, Pakistan MD # 408; 29, B. avinoviellus, India MD # 404; 30, B. himalayanus, India MD # 426; 31, B. turkestanicus, China MD # 351; 32, B. defector, Kyrgyzstan MD # 367; 33, B. margreiteri, China MD # 294; 34, B. handlirschianus, Russia MD # 835; 35, B. mendax, Spain MD # 689." httpUri="https://zenodo.org/record/192312/files/figure.png" pageId="27" pageNumber="28">
|
||
<emphasis id="B9730A507C763317B090A713A27BFCE9" bold="true" box="[151,333,823,845]" pageId="27" pageNumber="28">FIGURES 24‒35</emphasis>
|
||
</figureCitation>
|
||
. (Continued)
|
||
<emphasis id="B9730A507C763317B0C0A7B1A25DFC0A" bold="true" box="[199,363,917,942]" pageId="27" pageNumber="28">Food plants.</emphasis>
|
||
(
|
||
<taxonomicName id="4C07ADC17C763317B180A7B2A13FFC0A" box="[391,521,918,942]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||
)
|
||
<taxonomicName id="4C07ADC17C763317B222A7B2A09EFC0A" authority="C. Winkler" authorityName="C. Winkler" box="[549,936,917,942]" class="Magnoliopsida" family="Asteraceae" genus="Saussurea" kingdom="Animalia" order="Asterales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="tibetica">
|
||
<emphasis id="B9730A507C763317B222A7B2A032FC0A" box="[549,772,917,942]" italics="true" pageId="27" pageNumber="28">Saussurea tibetica</emphasis>
|
||
C. Winkler
|
||
</taxonomicName>
|
||
; (
|
||
<taxonomicName id="4C07ADC17C763317B3CEA7B1A751FC0A" box="[969,1127,917,942]" class="Magnoliopsida" family="Gentianaceae" kingdom="Plantae" order="Gentianales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="family">Gentianaceae</taxonomicName>
|
||
)
|
||
<taxonomicName id="4C07ADC17C763317B484A7B2A6A1FC0A" authority="Pall." authorityName="Pall." box="[1155,1431,917,942]" class="Magnoliopsida" family="Gentianaceae" genus="Gentiana" kingdom="Plantae" order="Gentianales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="algida">
|
||
<emphasis id="B9730A507C763317B484A7B2A67AFC0A" box="[1155,1356,917,942]" italics="true" pageId="27" pageNumber="28">Gentiana algida</emphasis>
|
||
Pall.
|
||
</taxonomicName>
|
||
; (Leguminosae)
|
||
<taxonomicName id="4C07ADC17C763317B14BA792A1C7FC6B" authority="Ulbrich" authorityName="Ulbrich" box="[332,753,950,975]" class="Magnoliopsida" family="Fabaceae" genus="Hedysarum" kingdom="Plantae" order="Fabales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="pseudastragalus">
|
||
<emphasis id="B9730A507C763317B14BA792A1A5FC6B" box="[332,659,950,975]" italics="true" pageId="27" pageNumber="28">Hedysarum pseudastragalus</emphasis>
|
||
Ulbrich
|
||
</taxonomicName>
|
||
(Z.-Y. Miao:
|
||
<date id="FFB9F0827C763317B397A792A728FC6B" box="[912,1054,950,975]" pageId="27" pageNumber="28" value="2014-08-16">16.viii.2014</date>
|
||
, in the Tanggula mountains
|
||
<quantity id="4CFF7BA77C763317B564A792A386FC54" metricMagnitude="3" metricUnit="m" metricValue="5.22" pageId="27" pageNumber="28" unit="m" value="5220.0">5220 m</quantity>
|
||
,
|
||
<collectingRegion id="49C318A07C763317B0BBA7FCA227FC55" box="[188,273,984,1009]" country="China" name="Xizang" pageId="27" pageNumber="28">Xizang</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31096D27C763317B11AA7FCA252FC55" box="[285,356,984,1009]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
|
||
,
|
||
<figureCitation id="133CCAC77C763317B168A7FCA2F3FC55" box="[367,453,984,1009]" captionStart="FIGURES 69 ‒ 70" captionStartId="59.[151,264,1618,1640]" captionTargetBox="[151,1436,259,1554]" captionTargetId="figure@59.[151,1436,193,1653]" captionTargetPageId="59" captionText="FIGURES 69 ‒ 70. Photos of (69) a worker of B. superbus (MD # 4123) from (left) the dorsal and (right) the left lateral aspect (scale bar 10 mm), from the sample site (70) in the Tanggula mountains, Xizang, China (16. viii. 2014, 32.86142 ° N 91.91739 ° E, GPS elevation ca 5220 m). Photos by (69) NHM digital imaging unit and (70) Z. - Y. Miao." httpUri="https://zenodo.org/record/192317/files/figure.png" pageId="27" pageNumber="28">Fig. 70</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB8D6427C763317B0C0A7DDA2EEFBB6" blockId="27.[151,1436,917,1042]" box="[199,472,1017,1042]" pageId="27" pageNumber="28">
|
||
<emphasis id="B9730A507C763317B0C0A7DDA27DFBB6" bold="true" box="[199,331,1017,1042]" pageId="27" pageNumber="28">Behaviour.</emphasis>
|
||
No records.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |