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<document id="73D242F88807BC404ECD723E340926E1" ID-CLB-Dataset="8644" ID-DOI="10.5852/ejt.2021.732.1213" ID-GBIF-Dataset="e98cc372-5374-45f3-b965-c57d9a916566" ID-ISSN="2118-9773" ID-Zenodo-Dep="4471717" ID-ZooBank="39F0C998-10AE-44BF-B1ED-7593AD4AE3AA" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611647697284" checkinUser="plazi" docAuthor="Gonçalves, Ana Rita, Grootaert, Patrick, Andrade, Rui, Paulo, Octávio S. &amp; Mengual, Ximo" docDate="2021" docId="03A187E12064FFA8FDA9FE65FC028D36" docLanguage="en" docName="ejt-732_1-56.pdf" docOrigin="European Journal of Taxonomy 732" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tachydromia pieltaini" docType="treatment" docVersion="11" lastPageNumber="35" masterDocId="FF98FF992044FF8AFFB7FFC9FF9A8C66" masterDocTitle="Revision of the morphology, phylogenetic relationships, behaviour and diversity of the Iberian and Italian ant-like Tachydromia Meigen, 1803 (Diptera: Hybotidae)" masterLastPageNumber="56" masterPageNumber="1" pageNumber="33" updateTime="1698925265419" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9D058AF7C459B5D0D748E693D80427FF">Revision of the morphology, phylogenetic relationships, behaviour and diversity of the Iberian and Italian ant-like Tachydromia Meigen, 1803 (Diptera: Hybotidae)</mods:title>
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<mods:namePart id="34B8821C40B4B0B17BE116383CB83D3E">Gonçalves, Ana Rita</mods:namePart>
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<mods:affiliation id="CF279F56D8E6A1DCE68F38D84769593D">Computational Biology and Population Genomics Group, cE 3 c, Faculdade de Ciências, Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal.</mods:affiliation>
<mods:nameIdentifier id="4E9BE5B4B86E5B70282DB123377F62E5" type="email">ana.goncalves@mail.mcgill.ca</mods:nameIdentifier>
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<mods:namePart id="7F77C2C5043940CC0788654FC504D7E2">Grootaert, Patrick</mods:namePart>
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<mods:affiliation id="EA81F45FA8343308137AAB7453B2939B">Royal Belgian Institute of Natural Sciences, Brussels, Belgium and Lee Kong Chian Natural History Museum, National University of Singapore, Singapore.</mods:affiliation>
<mods:nameIdentifier id="3047DBC0A33B452A19B0B5968D08BFF9" type="email">pgrootaert@yahoo.co.uk</mods:nameIdentifier>
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<mods:namePart id="512CD9786A9ABA9FC1637ED068AE472C">Andrade, Rui</mods:namePart>
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<mods:affiliation id="8834C826FF9B844ECAEB552E26A069BA">Rua Calouste Gulbenkian 237 4 H 3, 4050 - 145 Porto, Portugal.</mods:affiliation>
<mods:nameIdentifier id="81F915816D846CCD1E70229FF4A4FEC4" type="email">ruiamandrade@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="96FCF126233B52DE120396AC48F5DE25">Paulo, Octávio S.</mods:namePart>
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<mods:affiliation id="118658EDCB1606D6D3291461AE621CC2">Computational Biology and Population Genomics Group, cE 3 c, Faculdade de Ciências, Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal.</mods:affiliation>
<mods:nameIdentifier id="211E1A9CB5CE9FBA7172C41D2EBCD76F" type="email">ofpaulo@fc.ul.pt</mods:nameIdentifier>
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<mods:namePart id="A5716FCAEAEF7D93CD0A1A212EDCEB2B">Mengual, Ximo</mods:namePart>
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<mods:affiliation id="A7A35D8D20BC6CC292E6793EA492140F">Zoologisches Forschungsmuseum Alexander Koenig, Leibniz-Institut f ̹ r Biodiversität der Tiere, Adenauerallee 160, 53113, Bonn, Germany.</mods:affiliation>
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<mods:title id="A31C4F2D424976AA0D8AF97316F48A05">European Journal of Taxonomy</mods:title>
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<mods:date id="0D7F95E76350D2CA2B06C832A665248D">2021</mods:date>
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<mods:number id="D50B91650359ADFD9F5A4C62E3747960">2021-01-25</mods:number>
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<mods:identifier id="19B36CE9496FC13A010B2C07898A9D81" type="DOI">10.5852/ejt.2021.732.1213</mods:identifier>
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<treatment id="03A187E12064FFA8FDA9FE65FC028D36" ID-DOI="http://doi.org/10.5281/zenodo.4472575" ID-GBIF-Taxon="177132285" ID-Zenodo-Dep="4472575" LSID="urn:lsid:plazi:treatment:03A187E12064FFA8FDA9FE65FC028D36" httpUri="http://treatment.plazi.org/id/03A187E12064FFA8FDA9FE65FC028D36" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33">
<subSubSection id="C312657C2064FFAAFDA9FE65FB8F8DA1" box="[542,1045,428,455]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BB736F72064FFAAFDA9FE65FB8F8DA1" blockId="32.[542,1045,428,487]" box="[542,1045,428,455]" pageId="32" pageNumber="33">
<heading id="D0FF819B2064FFAAFDA9FE65FB8F8DA1" box="[542,1045,428,455]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="33" reason="2">
<taxonomicName id="4C084D742064FFAAFDA9FE65FB8F8DA1" authority="(Gil Collado, 1936)" baseAuthorityName="Gil Collado" baseAuthorityYear="1936" box="[542,1045,428,455]" class="Insecta" family="Hybotidae" genus="Tachydromia" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pieltaini">
<emphasis id="B97CEAE52064FFAAFDA9FE65FCBD8DA1" bold="true" box="[542,807,428,455]" italics="true" pageId="32" pageNumber="33">Tachydromia pieltaini</emphasis>
(Gil Collado, 1936)
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</paragraph>
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<subSubSection id="C312657C2064FFAAFD98FE04FB9F8D81" box="[559,1029,461,487]" pageId="32" pageNumber="33" type="description">
<paragraph id="8BB736F72064FFAAFD98FE04FB9F8D81" blockId="32.[542,1045,428,487]" box="[559,1029,461,487]" pageId="32" pageNumber="33">
<figureCitation id="13332A722064FFAAFD98FE04FDE28D81" box="[559,632,461,487]" captionStart="Fig" captionStartId="4.[189,232,1160,1186]" captionTargetBox="[189,1398,265,1120]" captionTargetId="figure-313@4.[189,1398,265,1120]" captionTargetPageId="4" captionText="Fig. 1. Currently known distribution of the Iberian ant-like Tachydromia Meigen, 1803. Each dot represents a presence point, with each colour corresponding to a different species. When two species co-occur in the same area, their presence is represented by a smaller dot on top of a dot of regular dimension, each of those with the colour corresponding to the co-occurring species. The dots surrounded by a black circle with a vertical line represent localities previously known." figureDoi="http://doi.org/10.5281/zenodo.4471719" httpUri="https://zenodo.org/record/4471719/files/figure.png" pageId="32" pageNumber="33">Figs 1</figureCitation>
,
<figureCitation id="13332A722064FFAAFD31FE04FD3C8D81" box="[646,678,461,487]" captionStart="Fig" captionStartId="33.[189,232,1927,1953]" captionTargetBox="[195,1388,1285,1888]" captionTargetId="figure-393@33.[189,1398,1279,1890]" captionTargetPageId="33" captionText="Fig. 12. Terminalia of Tachydromia pieltaini (Gil Collado, 1936) from Spain, Asturias, Covadonga (RBINS). A. Right surstylus and right epandrial lamella. B. Epandrium with cerci. C. Left epandrial lamella and left surstylus. D. Right surstylus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4471741" httpUri="https://zenodo.org/record/4471741/files/figure.png" pageId="32" pageNumber="33">12</figureCitation>
,
<figureCitation id="13332A722064FFAAFD05FE04FD578D81" box="[690,717,461,487]" captionStart="Fig" captionStartId="39.[189,232,1892,1918]" captionTargetBox="[340,1246,265,1864]" captionTargetId="figure-66@39.[340,1247,265,1864]" captionTargetPageId="39" captionText="Fig. 16. Habitus of live specimens of the Iberian ant-like Tachydromia Meigen, 1803 and the Italian flightless Tachydromia. Males are pictured in the left column, females in the right.A–B. T. iberica (Arias, 1919). C–D. T. semiaptera (Gil Collado, 1923). E–F. T. pieltaini (Gil Collado, 1936). G–H. T. pandellei (Séguy, 1941). I–J. T. apterygon Plant &amp; Deeming, 2006 (Italy)." figureDoi="http://doi.org/10.5281/zenodo.4471749" httpUri="https://zenodo.org/record/4471749/files/figure.png" pageId="32" pageNumber="33">16</figureCitation>
E–F,
<figureCitation id="13332A722064FFAAFCBAFE04FCB28D81" box="[781,808,461,487]" captionStart="Fig" captionStartId="41.[189,232,1890,1916]" captionTargetBox="[340,1247,265,1856]" captionTargetId="figure-63@41.[340,1247,265,1856]" captionTargetPageId="41" captionText="Fig. 18. Mounted specimens of the Iberian ant-like Tachydromia Meigen, 1803. Males are pictured in the left column, females in the right. A–B. T. cantabrica Gonçalves, Grootaert &amp; Andrade sp. nov. C–D. T. iberica (Arias, 1919). E–F. T. semiaptera (Gil Collado, 1923). G–H. T. pieltaini (Gil Collado, 1936). I–J. T. pandellei (Séguy, 1941). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4471755" httpUri="https://zenodo.org/record/4471755/files/figure.png" pageId="32" pageNumber="33">18</figureCitation>
G–H,
<figureCitation id="13332A722064FFAAFCC5FE04FC0A8D81" box="[882,912,461,487]" captionStart="Fig" captionStartId="42.[189,232,1823,1849]" captionTargetBox="[312,1276,265,1775]" captionTargetId="figure-104@42.[312,1276,265,1775]" captionTargetPageId="42" captionText="Fig. 19. Images obtained by scanning electron microscopy of the spur-like structures present on the apical portion of the male mid tibia. A. T. lusitanica (Grootaert, Shamshev &amp; Andrade, 2009). B. T. pandellei (Séguy, 1941). C. T. pieltaini (Gil Collado, 1936). D. T. ebejeri Gonçalves, Grootaert &amp; Andrade sp. nov. E–F. T. semiaptera (Gil Collado, 1923). G. T. stenoptera Gonçalves, Grootaert &amp; Andrade sp. nov. H. T. cantabrica Gonçalves, Grootaert &amp; Andrade sp. nov. I. T. nigrohirta Gonçalves, Grootaert &amp; Andrade sp. nov. J. T. apterygon Plant &amp; Deeming, 2006. Scale bars: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4471757" httpUri="https://zenodo.org/record/4471757/files/figure.png" pageId="32" pageNumber="33">19</figureCitation>
C,
<figureCitation id="13332A722064FFAAFC06FE04FC568D81" box="[945,972,461,487]" captionStart="Fig" captionStartId="44.[189,232,1857,1883]" captionTargetBox="[294,1292,267,1813]" captionTargetId="figure-76@44.[294,1294,262,1813]" captionTargetPageId="44" captionText="Fig. 21. Drawings of the tip of stenopterous wings and images obtained by scanning electron microscope (SEM) of the micropterous wings. Males are pictured in the left column, females in the right. A, C. T. pandellei (Séguy, 1941). B–C. T. pieltaini (Gil Collado, 1936). D–E. T. semiaptera (Gil Collado, 1923). F–G. T. stenoptera Gonçalves, Grootaert &amp; Andrade sp. nov. Scale bars: A–D, F = 50 µm; E, G = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4471761" httpUri="https://zenodo.org/record/4471761/files/figure.png" pageId="32" pageNumber="33">21</figureCitation>
B–C
</paragraph>
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<subSubSection id="C312657C2064FFAAFF0AFDC5FB968E00" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="8BB736F72064FFAAFF0AFDC5FD358E41" blockId="32.[189,687,524,551]" box="[189,687,524,551]" pageId="32" pageNumber="33">
<treatmentCitationGroup id="AB1811D92064FFAAFF0AFDC5FD318E41" box="[189,683,524,551]" pageId="32" pageNumber="33">
<treatmentCitation id="0AA910E62064FFAAFF0AFDC5FD318E41" box="[189,683,524,551]" pageId="32" pageNumber="33">
<taxonomicName id="4C084D742064FFAAFF0AFDC5FD318E41" ID-CoL="GLTZ" authority="Gil Collado, 1936: 191" authorityName="Gil Collado" authorityPageNumber="191" authorityYear="1936" box="[189,683,524,551]" class="Insecta" family="Hybotidae" genus="Ariasella" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pieltaini">
<emphasis id="B97CEAE52064FFAAFF0AFDC5FE088E40" box="[189,402,524,550]" italics="true" pageId="32" pageNumber="33">Ariasella pieltaini</emphasis>
<bibRefCitation id="EF994B062064FFAAFE2EFDC5FD318E41" author="Gil J." box="[409,683,524,551]" pageId="32" pageNumber="33" pagination="191 - 201" refId="ref28509" refString="Gil J. 1936. Una nueva especie del genero Ariasella Gil, con breves consideraciones sobre la reduccion del torax en los Taquidrominos apteros. Eos (Revista Espanola de Entomologia) 11 (3): 191 - 201." type="journal article" year="1936">Gil Collado, 1936: 191</bibRefCitation>
</taxonomicName>
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</treatmentCitationGroup>
.
</paragraph>
<paragraph id="8BB736F72064FFAAFF0AFD85FB968E00" blockId="32.[189,1036,588,615]" box="[189,1036,588,615]" pageId="32" pageNumber="33">
<treatmentCitationGroup id="AB1811D92064FFAAFF0AFD85FB968E00" box="[189,1036,588,615]" pageId="32" pageNumber="33">
<taxonomicName id="4C084D742064FFAAFF0AFD85FE258E00" baseAuthorityName="Gil Collado" baseAuthorityYear="1936" box="[189,447,588,614]" class="Insecta" family="Hybotidae" genus="Tachydromia" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pieltaini">
<emphasis id="B97CEAE52064FFAAFF0AFD85FE258E00" box="[189,447,588,614]" italics="true" pageId="32" pageNumber="33">Tachydromia pieltaini</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AA910E62064FFAAFE6BFD85FCF28E01" author="Shamshev I. V. &amp; Grootaert P." box="[476,872,588,615]" page="425" pageId="32" pageNumber="33" year="2018">
<bibRefCitation id="EF994B062064FFAAFE6BFD85FCF28E01" author="Shamshev &amp; Grootaert" box="[476,872,588,615]" firstAuthor="Shamshev" pageId="32" pageNumber="33" pagination="425 - 434" refId="ref29649" refString="Shamshev I. V. &amp; Grootaert P. 2018. Proposed changes in systematics and status of some genera of Tachydromiini (Diptera: Hybotidae: Tachydromiinae), with description of a new species of Tachypeza Meigen from Canada and USA. Russian Entomological Journal 27 (4): 425 - 434. https: // doi. org / 10.15298 / rusentj. 27.4.10" type="journal article" year="2018">Shamshev &amp; Grootaert 2018: 425</bibRefCitation>
</treatmentCitation>
(comb. nov.).
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</paragraph>
</subSubSection>
<subSubSection id="C312657C2064FFAAFF0AFD42FE808FFE" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8BB736F72064FFAAFF0AFD42FEAC8EC3" blockId="32.[189,310,651,677]" box="[189,310,651,677]" pageId="32" pageNumber="33">
<heading id="D0FF819B2064FFAAFF0AFD42FEAC8EC3" bold="true" box="[189,310,651,677]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B97CEAE52064FFAAFF0AFD42FEAC8EC3" bold="true" box="[189,310,651,677]" pageId="32" pageNumber="33">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736F72064FFAAFF0AFD71FE808FFE" blockId="32.[189,1399,696,920]" pageId="32" pageNumber="33">Shiny black with distinct yellow parts. Wing dimorphic: male stenopterous; wing with oval apex; stalklike process yellow to light brown; lobed apex with hyaline cells, veins and distal margin black; female stenopterous, stalk-like portion yellowish with minute lobed apex pale brown. Proboscis and antennae black; palpi yellow; postpedicel roundish, approximately as long as pedicel. Legs overall yellow only with the distal tarsomeres black. Abdomen black, tergites and sternites overall covered with setae; grey microtrichia present at last apical tergite and most sternites, resulting in shiny appearance of dorsal surface.</paragraph>
</subSubSection>
<subSubSection id="C312657C2064FFAAFF0AFC77FE3088B5" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BB736F72064FFAAFF0AFC77FCAA8862" blockId="32.[189,816,957,1029]" pageId="32" pageNumber="33">
<materialsCitation id="3B603CAA2064FFAAFF0AFC77FCAA8862" ID-GBIF-Occurrence="3026462440" collectorName="C. Bolivar" country="Spain" location="Porra de Enol" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Asturias" typeStatus="type">
<emphasis id="B97CEAE52064FFAAFF0AFC77FEF68FB1" bold="true" box="[189,364,957,984]" pageId="32" pageNumber="33">
<typeStatus id="54B388552064FFAAFF0AFC77FF608FBE" box="[189,250,958,984]" pageId="32" pageNumber="33">Type</typeStatus>
material
</emphasis>
(not examined, lost)
<collectingCountry id="F31F76672064FFAAFF0AFC23FE958862" box="[189,271,1002,1028]" name="Spain" pageId="32" pageNumber="33">SPAIN</collectingCountry>
•
<specimenCount id="9D0EFD7E2064FFAAFE9FFC23FECF8863" box="[296,341,1002,1029]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49CCF8152064FFAAFED5FC23FE5D8862" box="[354,455,1002,1028]" country="Spain" name="Asturias" pageId="32" pageNumber="33">Asturias</collectingRegion>
,
<location id="8ED7602C2064FFAAFE65FC22FDED8862" LSID="urn:lsid:plazi:treatment:03A187E12064FFA8FDA9FE65FC028D36:8ED7602C2064FFAAFE65FC22FDED8862" box="[466,631,1002,1029]" country="Spain" name="Porra de Enol" pageId="32" pageNumber="33" stateProvince="Asturias">Porra de Enol</location>
;
<collectorName id="26FD53212064FFAAFD34FC23FD678862" box="[643,765,1002,1028]" pageId="32" pageNumber="33">C. Bolívar</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BB736F72064FFAAFF0AFBE0FDAE8825" blockId="32.[189,1399,1065,1235]" box="[189,564,1065,1091]" pageId="32" pageNumber="33">
<heading id="D0FF819B2064FFAAFF0AFBE0FDAE8825" bold="true" box="[189,564,1065,1091]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B97CEAE52064FFAAFF0AFBE0FDAE8825" bold="true" box="[189,564,1065,1091]" pageId="32" pageNumber="33">Additional material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736F72064FFAAFF0AFB9FFE3088B5" blockId="32.[189,1399,1065,1235]" pageId="32" pageNumber="33">
<materialsCitation id="3B603CAA2064FFAAFF0AFB9FFE5A88F7" ID-GBIF-Occurrence="3026462311" collectingDate="2011-04-26" collectorName="R. Andrade" country="Spain" latitude="43.311775" location="Covadonga" longLatPrecision="1" longitude="-5.057778" municipality="Cangas de Onis" pageId="32" pageNumber="33" specimenCount="6" specimenCount-female="3" stateProvince="Asturias">
<collectingCountry id="F31F76672064FFAAFF0AFB9FFE958816" box="[189,271,1110,1136]" name="Spain" pageId="32" pageNumber="33">SPAIN</collectingCountry>
•
<specimenCount id="9D0EFD7E2064FFAAFE9BFB9FFEE88816" box="[300,370,1110,1136]" pageId="32" pageNumber="33">3 ♂♂</specimenCount>
,
<specimenCount id="9D0EFD7E2064FFAAFE37FB9FFE5F8816" box="[384,453,1110,1136]" pageId="32" pageNumber="33" type="female">3 ♀♀</specimenCount>
;
<collectingRegion id="49CCF8152064FFAAFE63FB9FFDA38816" box="[468,569,1110,1136]" country="Spain" name="Asturias" pageId="32" pageNumber="33">Asturias</collectingRegion>
,
<collectingMunicipality id="6BD3AC8D2064FFAAFDF1FB9FFC9E8816" box="[582,772,1110,1136]" pageId="32" pageNumber="33">Cangas de Onís</collectingMunicipality>
,
<location id="8ED7602C2064FFAAFCA5FB9FFC018816" LSID="urn:lsid:plazi:treatment:03A187E12064FFA8FDA9FE65FC028D36:8ED7602C2064FFAAFCA5FB9FFC018816" box="[786,923,1110,1136]" country="Spain" latitude="43.311775" longLatPrecision="1" longitude="-5.057778" municipality="Cangas de Onis" name="Covadonga" pageId="32" pageNumber="33" stateProvince="Asturias">Covadonga</location>
;
<geoCoordinate id="EE3C50302064FFAAFC1EFB9DFBCD8816" box="[937,1111,1108,1136]" degrees="43" direction="north" minutes="18" orientation="latitude" pageId="32" pageNumber="33" precision="1" seconds="42.4" value="43.311775">43°18′42.4″ N</geoCoordinate>
,
<geoCoordinate id="EE3C50302064FFAAFBD2FB9DFA938816" box="[1125,1289,1108,1136]" degrees="5" direction="west" minutes="03" orientation="longitude" pageId="32" pageNumber="33" precision="1" seconds="28.0" value="-5.057778">5°03′28.0″ W</geoCoordinate>
;
<date id="FFB610372064FFAAFAA0FB9FFF6088F7" pageId="32" pageNumber="33" value="2011-04-26">
<collectingDate id="EFF2E9DF2064FFAAFAA0FB9FFF6088F7" pageId="32" pageNumber="33" value="2011-04-26">26. Apr. 2011</collectingDate>
</date>
;
<collectorName id="26FD53212064FFAAFEB0FBB1FE1788F7" box="[263,397,1143,1170]" pageId="32" pageNumber="33">R. Andrade</collectorName>
leg.
</materialsCitation>
•
<materialsCitation id="3B603CAA2064FFAAFE6DFBBEFB0288F4" ID-GBIF-Occurrence="3026462387" box="[474,1176,1143,1170]" collectingDate="2011-04-26" collectionCode="ZFMK" collectorName="R. Andrade" country="Spain" latitude="43.311775" location="Covadonga" longLatPrecision="1" longitude="-5.057778" municipality="Cangas de Onis" pageId="32" pageNumber="33" specimenCount="4" specimenCount-female="2" stateProvince="Asturias">
<specimenCount id="9D0EFD7E2064FFAAFE6DFBBEFD8588F4" box="[474,543,1143,1170]" pageId="32" pageNumber="33">2 ♂♂</specimenCount>
,
<specimenCount id="9D0EFD7E2064FFAAFD9AFBBEFDEA88F4" box="[557,624,1143,1170]" pageId="32" pageNumber="33" type="female">2 ♀♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED19AE322064FFAAFBF0FBB1FB0288F4" box="[1095,1176,1144,1170]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="32" pageNumber="33" type="Museum">ZFMK</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B603CAA2064FFAAFB04FBBEFE3088B5" ID-GBIF-Occurrence="3026462412" collectingDate="2016-04-09" collectionCode="RBINS" collectorName="A. R. Goncalves" country="Spain" latitude="43.354053" location="Cangas de Onis" longLatPrecision="1" longitude="-5.1238613" pageId="32" pageNumber="33" specimenCount="6" specimenCount-female="4" stateProvince="Asturias">
<specimenCount id="9D0EFD7E2064FFAAFB04FBBEFB6D88F4" box="[1203,1271,1143,1170]" pageId="32" pageNumber="33">2 ♂♂</specimenCount>
,
<specimenCount id="9D0EFD7E2064FFAAFAB2FBBEFADD88F4" box="[1285,1351,1143,1170]" pageId="32" pageNumber="33" type="female">4 ♀♀</specimenCount>
(all male terminalia studied,
<collectionCode id="ED19AE322064FFAAFE68FB51FDAD88D4" box="[479,567,1176,1202]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="32" pageNumber="33" type="Museum">RBINS</collectionCode>
);
<collectingRegion id="49CCF8152064FFAAFDFAFB51FD2888D4" box="[589,690,1176,1202]" country="Spain" name="Asturias" pageId="32" pageNumber="33">Asturias</collectingRegion>
,
<location id="8ED7602C2064FFAAFD0AFB51FCE288D4" LSID="urn:lsid:plazi:treatment:03A187E12064FFA8FDA9FE65FC028D36:8ED7602C2064FFAAFD0AFB51FCE288D4" box="[701,888,1176,1202]" country="Spain" latitude="43.354053" longLatPrecision="1" longitude="-5.1238613" name="Cangas de Onis" pageId="32" pageNumber="33" stateProvince="Asturias">Cangas de Onís</location>
;
<geoCoordinate id="EE3C50302064FFAAFC33FB5EFBB788D5" box="[900,1069,1175,1203]" degrees="43" direction="north" minutes="21" orientation="latitude" pageId="32" pageNumber="33" precision="1" seconds="14.6" value="43.354053">43º21′14.6″ N</geoCoordinate>
,
<geoCoordinate id="EE3C50302064FFAAFB8EFB5EFB4388D5" box="[1081,1241,1175,1203]" degrees="5" direction="west" minutes="07" orientation="longitude" pageId="32" pageNumber="33" precision="1" seconds="25.9" value="-5.1238613">5º07′25.9″ W</geoCoordinate>
;
<date id="FFB610372064FFAAFB52FB51FAE888D4" box="[1253,1394,1176,1202]" pageId="32" pageNumber="33" value="2016-04-09">
<collectingDate id="EFF2E9DF2064FFAAFB52FB51FAE888D4" box="[1253,1394,1176,1202]" pageId="32" pageNumber="33" value="2016-04-09">9 Apr. 2016</collectingDate>
</date>
;
<collectorName id="26FD53212064FFAAFF0AFB70FEE288B5" box="[189,376,1209,1235]" pageId="32" pageNumber="33">A.R. Gonçalves</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C312657C2064FFABFF0AFB31FD5388CE" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="8BB736F72064FFAAFF0AFB31FEF18974" blockId="32.[189,363,1272,1298]" box="[189,363,1272,1298]" pageId="32" pageNumber="33">
<heading id="D0FF819B2064FFAAFF0AFB31FEF18974" bold="true" box="[189,363,1272,1298]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B97CEAE52064FFAAFF0AFB31FEF18974" bold="true" box="[189,363,1272,1298]" pageId="32" pageNumber="33">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736F72064FFAAFF0AFAECFC7E8959" blockId="32.[189,996,1317,1344]" box="[189,996,1317,1344]" pageId="32" pageNumber="33">
Similar to
<taxonomicName id="4C084D742064FFAAFE8EFAECFDC28959" authority="Gonçalves &amp; Grootaert &amp; Andrade &amp; Paulo &amp; Mengual, 2021" authorityName="Gonçalves &amp; Grootaert &amp; Andrade &amp; Paulo &amp; Mengual" authorityYear="2021" box="[313,600,1317,1343]" class="Insecta" family="Hybotidae" genus="Tachydromia" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="B97CEAE52064FFAAFE8EFAECFDC28959" box="[313,600,1317,1343]" italics="true" pageId="32" pageNumber="33">Tachydromia cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F579E2064FFAAFDE8FAEFFD208926" box="[607,698,1318,1344]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
except for the following:
</paragraph>
<paragraph id="8BB736F72064FFAAFF5DFAACFDCC8919" blockId="32.[189,1178,1380,1440]" box="[234,598,1380,1407]" pageId="32" pageNumber="33">
<emphasis id="B97CEAE52064FFAAFF5DFAACFEB08919" bold="true" box="[234,298,1381,1407]" pageId="32" pageNumber="33">Male</emphasis>
(previously undescribed)
</paragraph>
<paragraph id="8BB736F72064FFAAFF0AFA4FFB0089C6" blockId="32.[189,1178,1380,1440]" box="[189,1178,1413,1440]" pageId="32" pageNumber="33">
MEASUREMENTS. Body length:
<quantity id="4CF09B122064FFAAFDACFA4CFD3289C6" box="[539,680,1413,1440]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="32" pageNumber="33" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
(n = 5). Wing length:
<quantity id="4CF09B122064FFAAFC19FA4FFBA189C6" box="[942,1083,1414,1440]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="32" pageNumber="33" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
(n = 5).
</paragraph>
<paragraph id="8BB736F72064FFAAFF0AFA0CFD398A66" blockId="32.[189,1399,1477,1536]" pageId="32" pageNumber="33">HEAD. Ocellar tubercle glabrous. Lateroclinate ocellars 2 times as long as postpedicel. Proclinate verticals 1.8 times as long as postpedicel.</paragraph>
<paragraph id="8BB736F72064FFAAFF0AF9EFFC4B8A06" blockId="32.[189,1398,1573,1633]" pageId="32" pageNumber="33">ANTENNA. Yellow. Postpedicel roundish, as long as wide and, approximately as long as pedicel; stylus almost 4 times as long as scape, pedicel and postpedicel combined.</paragraph>
<paragraph id="8BB736F72064FFAAFF0AF94FFC2B8AC6" blockId="32.[189,945,1670,1696]" box="[189,945,1670,1696]" pageId="32" pageNumber="33">PALPUS. About 4 times as long as wide. Yellow in ground colour.</paragraph>
<paragraph id="8BB736F72064FFAAFF0AF90FFDC88B02" blockId="32.[189,1399,1733,1892]" pageId="32" pageNumber="33">THORAX. Postpronotal lobe, notopleuron, episternum, anepisternum, sternopleuron, katepimeron and meron weakly differentiated. Scutellum short, postscutellum longer and metanotum well developed. Prosternum and basal half of episternum densely covered by grey microtrichia. Postpronotal lobe bearing 7 setae laterally, 5 acrostichal setae, irregularly placed; notopleuron with 1 strong, prominent, very long seta, plus 3 smaller ones</paragraph>
<paragraph id="8BB736F72064FFAAFF0AF840FDDA8B8E" blockId="32.[189,1399,1929,2025]" pageId="32" pageNumber="33">
WING (
<figureCitation id="13332A722064FFAAFEA5F840FEE58BC5" box="[274,383,1929,1955]" captionStart="Fig" captionStartId="44.[189,232,1857,1883]" captionTargetBox="[294,1292,267,1813]" captionTargetId="figure-76@44.[294,1294,262,1813]" captionTargetPageId="44" captionText="Fig. 21. Drawings of the tip of stenopterous wings and images obtained by scanning electron microscope (SEM) of the micropterous wings. Males are pictured in the left column, females in the right. A, C. T. pandellei (Séguy, 1941). B–C. T. pieltaini (Gil Collado, 1936). D–E. T. semiaptera (Gil Collado, 1923). F–G. T. stenoptera Gonçalves, Grootaert &amp; Andrade sp. nov. Scale bars: A–D, F = 50 µm; E, G = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4471761" httpUri="https://zenodo.org/record/4471761/files/figure.png" pageId="32" pageNumber="33">Fig. 21B</figureCitation>
). Lobed apex oval. Stalk-like portion rigid, yellow to light brown; lobed apex with hyaline cells, veins and distal margin black. Venation very reduced (costal, longitudinal and cross veins distinguishable), forming 4 cells.
</paragraph>
<paragraph id="8BB736F72065FFABFF0AFEC0FD3E8E7F" blockId="33.[188,1400,265,538]" pageId="33" pageNumber="34">
LEGS. Overall yellow, with the following brown to black: dorsal strip on fore and mid femora, posterior coxae, tarsomere 5 of fore and mid-legs, apical portions of tarsomeres 2 and 3, tarsomeres 4 and 5 of hind leg. Mid trochanter with several black anteroventral marginal setae. Mid femur with small roundish posteroventral projection on basal surface with several black denticle-like setae and ca 3 black setae. Mid tibia anteroventral apical spur produced into distinct, long elongation (
<figureCitation id="13332A722065FFABFBF0FE5FFB288DD6" box="[1095,1202,406,432]" captionStart="Fig" captionStartId="42.[189,232,1823,1849]" captionTargetBox="[312,1276,265,1775]" captionTargetId="figure-104@42.[312,1276,265,1775]" captionTargetPageId="42" captionText="Fig. 19. Images obtained by scanning electron microscopy of the spur-like structures present on the apical portion of the male mid tibia. A. T. lusitanica (Grootaert, Shamshev &amp; Andrade, 2009). B. T. pandellei (Séguy, 1941). C. T. pieltaini (Gil Collado, 1936). D. T. ebejeri Gonçalves, Grootaert &amp; Andrade sp. nov. E–F. T. semiaptera (Gil Collado, 1923). G. T. stenoptera Gonçalves, Grootaert &amp; Andrade sp. nov. H. T. cantabrica Gonçalves, Grootaert &amp; Andrade sp. nov. I. T. nigrohirta Gonçalves, Grootaert &amp; Andrade sp. nov. J. T. apterygon Plant &amp; Deeming, 2006. Scale bars: 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4471757" httpUri="https://zenodo.org/record/4471757/files/figure.png" pageId="33" pageNumber="34">Fig. 19C</figureCitation>
); slightly longer than diameter of tibia at apex. Spur covered with microtrichia and&gt; 10 short, blunt denticle-like setae; additionally, with 6 longer, stout, curved setae along lateral margin. Elongation mostly glabrous with a few setae arising marginally and apically.
</paragraph>
<paragraph id="8BB736F72065FFABFF0AFD88FF658EA3" blockId="33.[189,1400,577,709]" pageId="33" pageNumber="34">ABDOMEN. Shiny black, with grey microtrichia covering only last tergite and almost all sternites, except for sternites 1 and 2. Tergites and sternites overall covered with setae, except for dorsal surface of all tergites and most of sternite 1 (distal margin with a few setae); apical sternite and tergite with longer setae.</paragraph>
<paragraph id="8BB736F72065FFABFF0AFD25FD4A8FD0" blockId="33.[189,1399,748,950]" pageId="33" pageNumber="34">
TERMINALIA (
<figureCitation id="13332A722065FFABFEE0FD25FE288F60" box="[343,434,748,774]" captionStart="Fig" captionStartId="33.[189,232,1927,1953]" captionTargetBox="[195,1388,1285,1888]" captionTargetId="figure-393@33.[189,1398,1279,1890]" captionTargetPageId="33" captionText="Fig. 12. Terminalia of Tachydromia pieltaini (Gil Collado, 1936) from Spain, Asturias, Covadonga (RBINS). A. Right surstylus and right epandrial lamella. B. Epandrium with cerci. C. Left epandrial lamella and left surstylus. D. Right surstylus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4471741" httpUri="https://zenodo.org/record/4471741/files/figure.png" pageId="33" pageNumber="34">Fig. 12</figureCitation>
). Right surstylus short with irregular margins, narrowing distally, bearing long setae, with several (ca 8) distinct denticle-like setae aggregated on distal margin. Right cercus longer than left. Right cercus with setae on apical ½; left cercus with setae of unequal lengths on apical half and 2 long apical setae. Left epandrial lamella ½ shorter than left cercus, with setae of similar length on apical half of ventral surface and on apex of dorsal margin. Left surstylus longer than right cercus, with very long setae on lateral and apical margins.
</paragraph>
<paragraph id="8BB736F72065FFABFF5DFC14FEDF8F91" blockId="33.[188,1399,989,1192]" box="[234,325,989,1015]" pageId="33" pageNumber="34">
<emphasis id="B97CEAE52065FFABFF5DFC14FEDF8F91" bold="true" box="[234,325,989,1015]" pageId="33" pageNumber="34">Female</emphasis>
</paragraph>
<paragraph id="8BB736F72065FFABFF0AFBC9FD5388CE" blockId="33.[188,1399,989,1192]" pageId="33" pageNumber="34">
Similar to male, except for following features: stenopterous (length of wing (n = 2):
<quantity id="4CF09B122065FFABFB2AFBC8FAB4887D" box="[1181,1326,1025,1051]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="33" pageNumber="34" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
), with very minute lobed apex, stalk-like portion yellowish with lobe light brown, covered by grey microtrichia (
<figureCitation id="13332A722065FFABFF73FB8EFEAF8807" box="[196,309,1095,1121]" captionStart="Fig" captionStartId="44.[189,232,1857,1883]" captionTargetBox="[294,1292,267,1813]" captionTargetId="figure-76@44.[294,1294,262,1813]" captionTargetPageId="44" captionText="Fig. 21. Drawings of the tip of stenopterous wings and images obtained by scanning electron microscope (SEM) of the micropterous wings. Males are pictured in the left column, females in the right. A, C. T. pandellei (Séguy, 1941). B–C. T. pieltaini (Gil Collado, 1936). D–E. T. semiaptera (Gil Collado, 1923). F–G. T. stenoptera Gonçalves, Grootaert &amp; Andrade sp. nov. Scale bars: A–D, F = 50 µm; E, G = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.4471761" httpUri="https://zenodo.org/record/4471761/files/figure.png" pageId="33" pageNumber="34">Fig. 21C</figureCitation>
); legs covered with regular rows of setae, with distinct circle of setae on fore femur and anteroapical setae on coxae; without other distinctive hairs or setae; spurs on mid tibia absent; cercus pale brown with grey microtrichia and setae.
</paragraph>
</subSubSection>
<caption id="DF77667F2065FFABFF0AF84EFC288B8E" ID-DOI="http://doi.org/10.5281/zenodo.4471741" ID-Zenodo-Dep="4471741" httpUri="https://zenodo.org/record/4471741/files/figure.png" pageId="33" pageNumber="34" startId="33.[189,232,1927,1953]" targetBox="[195,1388,1285,1888]" targetPageId="33">
<paragraph id="8BB736F72065FFABFF0AF84EFC288B8E" blockId="33.[189,1399,1927,2024]" pageId="33" pageNumber="34">
<emphasis id="B97CEAE52065FFABFF0AF84EFE848BC7" bold="true" box="[189,286,1927,1953]" pageId="33" pageNumber="34">Fig. 12.</emphasis>
Terminalia of
<taxonomicName id="4C084D742065FFABFE60F84EFC428BC4" authority="(Gil Collado, 1936)" baseAuthorityName="Gil Collado" baseAuthorityYear="1936" box="[471,984,1927,1954]" class="Insecta" family="Hybotidae" genus="Tachydromia" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pieltaini">
<emphasis id="B97CEAE52065FFABFE60F84EFD478BC7" box="[471,733,1927,1953]" italics="true" pageId="33" pageNumber="34">Tachydromia pieltaini</emphasis>
(Gil Collado, 1936)
</taxonomicName>
from Spain, Asturias, Covadonga (RBINS).
<emphasis id="B97CEAE52065FFABFE8FF863FED78BA2" bold="true" box="[312,333,1962,1988]" pageId="33" pageNumber="34">A</emphasis>
. Right surstylus and right epandrial lamella.
<emphasis id="B97CEAE52065FFABFCD8F863FC198BA2" bold="true" box="[879,899,1962,1988]" pageId="33" pageNumber="34">B</emphasis>
. Epandrium with cerci.
<emphasis id="B97CEAE52065FFABFB1FF863FB278BA2" bold="true" box="[1192,1213,1962,1988]" pageId="33" pageNumber="34">C</emphasis>
. Left epandrial lamella and left surstylus.
<emphasis id="B97CEAE52065FFABFE45F807FD9D8B8E" bold="true" box="[498,519,1998,2024]" pageId="33" pageNumber="34">D</emphasis>
. Right surstylus. Scale bar: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C312657C2066FFA8FF0AFEC1FC028D36" box="[189,920,264,336]" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BB736F72066FFA8FF0AFEC1FECC8D44" blockId="34.[189,342,264,290]" box="[189,342,264,290]" pageId="34" pageNumber="35">
<heading id="D0FF819B2066FFA8FF0AFEC1FECC8D44" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B97CEAE52066FFA8FF0AFEC1FECC8D44" bold="true" box="[189,342,264,290]" pageId="34" pageNumber="35">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736F72066FFA8FF0AFEFCFC028D36" blockId="34.[189,920,309,336]" box="[189,920,309,336]" pageId="34" pageNumber="35">
<collectingCountry id="F31F76672066FFA8FF0AFEFCFE9E8D29" box="[189,260,309,335]" name="Spain" pageId="34" pageNumber="35">Spain</collectingCountry>
. Only known from the Picos de Europa mountain range.
</paragraph>
</subSubSection>
</treatment>
</document>
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