141 lines
17 KiB
XML
141 lines
17 KiB
XML
<document id="B7A5783EF1A767E1B763CF0EC43F9D62" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFD9FFD96194FACA00B6F5AE" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus velatus Perkins 2021, new species" docType="treatment" docVersion="4" lastPageNumber="39" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="38" updateTime="1698978858223" updateUser="plazi">
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<mods:titleInfo id="1696767472106CDB36F6FF80A9AD46AE">
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<mods:title id="FFDEABB92FD9492694576CB6AB86F351">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="81D765FEB3CB27D3A908CC18B537AE8D" type="personal">
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<mods:role id="B4C7E34481656D3B4FD788BDA7779D2D">
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<mods:roleTerm id="225F688A0D0BE17CE7915D38D326791F">Author</mods:roleTerm>
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<mods:namePart id="D02C148443774311CBD9363F04AE9529">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="1077180AF2612FDC56D6CAB8CABFD6C5">text</mods:typeOfResource>
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<mods:title id="7DE31319473593D27C1451A843EBA470">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="93EB9794842FD9492C28B16F70385CD4">
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<mods:date id="140511D2E46D1D43002592CDB6F1C400">2021</mods:date>
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<mods:detail id="03E9F8BA0CFC92F47FDE57745E01366F" type="pubDate">
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<mods:number id="938810FEE6A867880B558342F75ABB6F">2021-06-28</mods:number>
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</mods:detail>
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<mods:detail id="19F207E80300A01A39C82FEFF7B8827D" type="volume">
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<mods:number id="177D12C32E0BA641D412B005E0CFCF0F">4994</mods:number>
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</mods:detail>
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<mods:detail id="AC093ADB816C7FF1BBB44752875A0AC0" type="issue">
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<mods:number id="3F6A3C484F24B7DD1345C124D2BB0211">1</mods:number>
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<mods:start id="046828EBECFF8D07F09D4C91A11B7A70">1</mods:start>
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<mods:end id="432E0DB11995450B6CF0A055BBD0307F">93</mods:end>
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<mods:classification id="44E9C0230B178D59DCD331C9A66EF25D">journal article</mods:classification>
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<mods:identifier id="59AEC61DE082C7E0866B495266ED99B5" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
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<mods:identifier id="1C3C7EFEF415D257BEDF7F6C63B1A030" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="136032A4CEFC14B6FFE91B58FFC210B8" type="Zenodo-Dep">5043261</mods:identifier>
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<mods:identifier id="5AC2664C926351AE191F23B35B6E3DE1" type="ZooBank">107FCA64-345F-40A4-99D3-5C1441EEAD93</mods:identifier>
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<treatment id="03A187DEFFD9FFD96194FACA00B6F5AE" ID-DOI="http://doi.org/10.5281/zenodo.5043987" ID-Zenodo-Dep="5043987" LSID="urn:lsid:plazi:treatment:03A187DEFFD9FFD96194FACA00B6F5AE" httpUri="http://treatment.plazi.org/id/03A187DEFFD9FFD96194FACA00B6F5AE" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
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<subSubSection id="C3126543FFD9FFDA6194FACA0648F253" box="[151,535,1338,1365]" pageId="37" pageNumber="38" type="nomenclature">
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<paragraph id="8BB736C8FFD9FFDA6194FACA0648F253" blockId="37.[151,535,1338,1401]" box="[151,535,1338,1365]" pageId="37" pageNumber="38">
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<heading id="D0FF81A4FFD9FFDA6194FACA0648F253" bold="true" box="[151,535,1338,1365]" fontSize="11" level="2" pageId="37" pageNumber="38" reason="4">
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<emphasis id="B97CEADAFFD9FFDA6194FACA0648F253" bold="true" box="[151,535,1338,1365]" pageId="37" pageNumber="38">
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<taxonomicName id="4C084D4BFFD9FFDA6194FACA0526F252" ID-CoL="8Y4RV" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,377,1338,1365]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="velatus" status="sp. nov.">
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<emphasis id="B97CEADAFFD9FFDA6194FACA0526F252" bold="true" box="[151,377,1338,1365]" italics="true" pageId="37" pageNumber="38">Hydrochus velatus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24F57A1FFD9FFDA608BFACB0648F253" box="[392,535,1338,1365]" pageId="37" pageNumber="38" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFD9FFDA6194FAAF0568F27E" box="[151,311,1374,1401]" pageId="37" pageNumber="38" type="description">
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<paragraph id="8BB736C8FFD9FFDA6194FAAF0568F27E" blockId="37.[151,535,1338,1401]" box="[151,311,1374,1401]" pageId="37" pageNumber="38">
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<figureCitation id="13332A4DFFD9FFDA6194FAAF04BAF27E" box="[151,229,1375,1401]" captionStart="FIGURE 2" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[164,1405,189,1904]" captionTargetId="figure-15@7.[153,1434,181,1919]" captionTargetPageId="7" captionText="FIGURE 2. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043267" httpUri="https://zenodo.org/record/5043267/files/figure.png" pageId="37" pageNumber="38">Figs. 2</figureCitation>
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,
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<figureCitation id="13332A4DFFD9FFDA61F2FAAF0550F27E" box="[241,271,1375,1401]" captionStart="FIGURES 18–19" captionStartId="61.[151,264,1944,1968]" captionTargetBox="[170,1425,205,1900]" captionTargetId="figure-15@61.[151,1435,181,1920]" captionTargetPageId="61" captionText="FIGURES 18–19. Habitus and male genitalia of holotypes. 18. Hydrochus pronotulus n. sp. 19. Hydrochus velatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043296" httpUri="https://zenodo.org/record/5043296/files/figure.png" pageId="37" pageNumber="38">19</figureCitation>
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,
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<figureCitation id="13332A4DFFD9FFDA6018FAAE0568F27E" box="[283,311,1374,1401]" captionStart="FIGURES 59–60" captionStartId="82.[151,264,1677,1701]" captionTargetBox="[157,1431,181,1654]" captionTargetId="figure-28@82.[157,1431,181,1654]" captionTargetPageId="82" captionText="FIGURES 59–60. Geographical distributions. 59. Endemics and near endemics. 60. H. formosus." figureDoi="http://doi.org/10.5281/zenodo.5043360" httpUri="https://zenodo.org/record/5043360/files/figure.png" pageId="37" pageNumber="38">59</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFD9FFDA6194FA5705B2F2E2" pageId="37" pageNumber="38" type="materials_examined">
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<paragraph id="8BB736C8FFD9FFDA6194FA5705B2F2E2" blockId="37.[151,1436,1446,2013]" pageId="37" pageNumber="38">
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<emphasis id="B97CEADAFFD9FFDA6194FA570518F2C7" bold="true" box="[151,327,1446,1473]" pageId="37" pageNumber="38">Type Material.</emphasis>
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<materialsCitation id="3B603C95FFD9FFDA604FFA570090F2C6" box="[332,1231,1446,1473]" collectionCode="NHMUK" collectorName="F. Plaumann" country="Brazil" location="BRAZIL" pageId="37" pageNumber="38" specimenCount="1" typeStatus="holotype">
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<emphasis id="B97CEADAFFD9FFDA604FFA57064BF2C7" bold="true" box="[332,532,1446,1473]" pageId="37" pageNumber="38">
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<typeStatus id="54B3886AFFD9FFDA604FFA5705E6F2C6" box="[332,441,1447,1473]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
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(male):
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</emphasis>
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“
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<collectingCountry id="F31F7658FFD9FFDA6327FA5706CFF2C6" box="[548,656,1447,1473]" name="Brazil" pageId="37" pageNumber="38">BRAZIL</collectingCountry>
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: Matto Grosso, Corumba,
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<collectorName id="26FD531EFFD9FFDA62B1FA570060F2C6" box="[946,1087,1447,1473]" pageId="37" pageNumber="38">F. Plaumann</collectorName>
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” (
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<collectionCode id="ED19AE0DFFD9FFDA6555FA570098F2C6" box="[1110,1223,1447,1473]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="37" pageNumber="38" type="Museum">NHMUK</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B603C95FFD9FFDA65D4FA5705B5F2E2" collectionCode="NHMUK" collectorName="F. Plaumann" country="Brazil" location="BRAZIL" pageId="37" pageNumber="38" specimenCount="1" typeStatus="paratype">
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<emphasis id="B97CEADAFFD9FFDA65D4FA570107F2C6" bold="true" box="[1239,1368,1447,1473]" pageId="37" pageNumber="38">
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<typeStatus id="54B3886AFFD9FFDA65D4FA57010CF2C6" box="[1239,1363,1447,1473]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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Same data as holotype (2
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<collectionCode id="ED19AE0DFFD9FFDA6071FA3B05BEF2E2" box="[370,481,1483,1509]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="37" pageNumber="38" type="Museum">NHMUK</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFD9FFDA61C4FA1E006EF156" pageId="37" pageNumber="38" type="diagnosis">
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<paragraph id="8BB736C8FFD9FFDA61C4FA1E006EF156" blockId="37.[151,1436,1446,2013]" pageId="37" pageNumber="38">
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<emphasis id="B97CEADAFFD9FFDA61C4FA1E058BF10F" bold="true" box="[199,468,1518,1544]" pageId="37" pageNumber="38">Differential Diagnosis.</emphasis>
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Among Brazilian species, recognized by the combination of moderate size (ca.
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<quantity id="4CF09B2DFFD9FFDA6468FA1F0498F12A" metricMagnitude="-3" metricUnit="m" metricValue="3.34" pageId="37" pageNumber="38" unit="mm" value="3.34">3.34 mm</quantity>
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), the shallow pronotal depressions, and the male genitalia (
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<figureCitation id="13332A4DFFD9FFDA6259F9E307F2F12A" box="[858,941,1555,1581]" captionStart="FIGURES 18–19" captionStartId="61.[151,264,1944,1968]" captionTargetBox="[170,1425,205,1900]" captionTargetId="figure-15@61.[151,1435,181,1920]" captionTargetPageId="61" captionText="FIGURES 18–19. Habitus and male genitalia of holotypes. 18. Hydrochus pronotulus n. sp. 19. Hydrochus velatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043296" httpUri="https://zenodo.org/record/5043296/files/figure.png" pageId="37" pageNumber="38">Fig. 19</figureCitation>
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; described below). This species has a generalized
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<taxonomicName id="4C084D4BFFD9FFDA61E2F9C70504F156" authorityName="Leach" authorityYear="1817" box="[225,347,1591,1617]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CEADAFFD9FFDA61E2F9C70504F156" box="[225,347,1591,1617]" italics="true" pageId="37" pageNumber="38">Hydrochus</emphasis>
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</taxonomicName>
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habitus; reliable determinations will require dissection of males.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFD9FFD961C4F9AA04B9F582" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="description">
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<paragraph id="8BB736C8FFD9FFDA61C4F9AA05C8F02E" blockId="37.[151,1436,1446,2013]" pageId="37" pageNumber="38">
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<emphasis id="B97CEADAFFD9FFDA61C4F9AA050EF173" bold="true" box="[199,337,1626,1652]" pageId="37" pageNumber="38">Description</emphasis>
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. Size:
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<typeStatus id="54B3886AFFD9FFDA60A7F9AB0659F172" box="[420,518,1627,1653]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 3.34/1.18; head width 0.81; pronotum l/w 0.73/0.79; PA 0.73; PB 0.62; elytra 2.06/1.18. Habitus as illustrated (
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<figureCitation id="13332A4DFFD9FFDA654CF98F00FBF19E" box="[1103,1188,1663,1689]" captionStart="FIGURES 18–19" captionStartId="61.[151,264,1944,1968]" captionTargetBox="[170,1425,205,1900]" captionTargetId="figure-15@61.[151,1435,181,1920]" captionTargetPageId="61" captionText="FIGURES 18–19. Habitus and male genitalia of holotypes. 18. Hydrochus pronotulus n. sp. 19. Hydrochus velatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043296" httpUri="https://zenodo.org/record/5043296/files/figure.png" pageId="37" pageNumber="38">Fig. 19</figureCitation>
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). Body size moderate (ca.
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<quantity id="4CF09B2DFFD9FFDA61C4F9530570F1BA" box="[199,303,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="3.34" pageId="37" pageNumber="38" unit="mm" value="3.34">3.34 mm</quantity>
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). Dorsum dark brown to dark grey, with slight or no iridescence, elytra without black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 2x that of pronotum. Elytra interstriae ca. 0.5–1x strial puncture diameter. Interstria 9
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<superScript id="7C7D9B80FFD9FFDA6329F91A0668F1FF" attach="left" box="[554,567,1770,1784]" fontSize="6" pageId="37" pageNumber="38">th</superScript>
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more convex than others, overhanging 10
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<superScript id="7C7D9B80FFD9FFDA6510F91A007FF1FF" attach="left" box="[1043,1056,1770,1784]" fontSize="6" pageId="37" pageNumber="38">th</superScript>
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interstria. Usual area of callus on 5
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<superScript id="7C7D9B80FFD9FFDA61A6F8FE04EDF01B" attach="left" box="[165,178,1806,1820]" fontSize="6" pageId="37" pageNumber="38">th</superScript>
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very slightly raised.
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</paragraph>
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<paragraph id="8BB736C8FFD9FFDA61C4F8C306BAF092" blockId="37.[151,1436,1446,2013]" pageId="37" pageNumber="38">Pronotum length and width ca. equal, widest at anterior margin, narrowed at base, sides very weakly sinuate, smooth; anterior margin slightly bisinuate; depressions shallow, much more densely punctate than low reliefs between depressions, latter almost entirely impunctate.</paragraph>
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<paragraph id="8BB736C8FFD9FFDA61C4F86F0724F0DA" blockId="37.[151,1436,1446,2013]" pageId="37" pageNumber="38">Elytra with spaces between strial punctures ca. 0.5–1x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with slight angulation.</paragraph>
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<paragraph id="8BB736C8FFDAFFD961C4FF670156F7B6" blockId="38.[151,1437,151,681]" box="[199,1289,151,177]" pageId="38" pageNumber="39">Ventral characters: Mentum punctate, with central depression; submentum with two large foveae.</paragraph>
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<paragraph id="8BB736C8FFDAFFD961C4FF4B00CEF6D6" blockId="38.[151,1437,151,681]" pageId="38" pageNumber="39">
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Male genitalia (
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<figureCitation id="13332A4DFFDAFFD9607EFF4B058EF7D2" box="[381,465,187,213]" captionStart="FIGURES 18–19" captionStartId="61.[151,264,1944,1968]" captionTargetBox="[170,1425,205,1900]" captionTargetId="figure-15@61.[151,1435,181,1920]" captionTargetPageId="61" captionText="FIGURES 18–19. Habitus and male genitalia of holotypes. 18. Hydrochus pronotulus n. sp. 19. Hydrochus velatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043296" httpUri="https://zenodo.org/record/5043296/files/figure.png" pageId="38" pageNumber="39">Fig. 19</figureCitation>
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) general characters: genitalia elongate, length ca. 5x greatest width, basal piece and parameres ca. same length; paramere tips and aedeagus tip at same level, paramere tips arcing toward midline, tips widely separated, plsa absent, lateral margins of tips weakly arcuate; in lateral view parameres wide basally, narrow from ca. midlength to rounded apex; pseudoparameres large, distal end slightly past midlength of parameres; aedeagus wide in both dorsal/ventral and lateral views, in dorsal/ventral views almost parallel-sided for most of length, narrowing near base, alw just distal to adbl, agw at ca. proximal 1/3; basal piece in dorsal/ventral views tapering from distal end to base, in lateral view also tapering toward base, very slightly arcuate, greatest width in dorsal/ventral views slightly greater than lateral width, orifice large, slightly asymmetrical.
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</paragraph>
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<paragraph id="8BB736C8FFDAFFD961C4FE2B0489F53A" blockId="38.[151,1437,151,681]" pageId="38" pageNumber="39">Dorsal surface: adtl not apparent; medial part of parameres very thin, velum-like, pdmm slightly sinuate, separated from alm over ca. distal 2/3, proximal 1/3 turning toward midline and going beneath alm; adbl moderately large.</paragraph>
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<paragraph id="8BB736C8FFDAFFD961C4FDB704B9F582" blockId="38.[151,1437,151,681]" pageId="38" pageNumber="39">Ventral surface: avtl not apparent; pvmm slightly wider than pdmm in distal ½, otherwise approximating pdmm.</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFDAFFD961C4FD7F00B6F5AE" box="[199,1257,655,681]" pageId="38" pageNumber="39" type="etymology">
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<paragraph id="8BB736C8FFDAFFD961C4FD7F00B6F5AE" blockId="38.[151,1437,151,681]" box="[199,1257,655,681]" pageId="38" pageNumber="39">
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<emphasis id="B97CEADAFFDAFFD961C4FD7F0517F5AE" bold="true" box="[199,328,655,681]" pageId="38" pageNumber="39">Etymology</emphasis>
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. Named in reference to the very thin, velum-like medial margins of the parameres.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |