543 lines
50 KiB
XML
543 lines
50 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5681917" ID-GBIF-Dataset="54b2150d-e662-450d-b3c2-73b51ac32041" ID-GBIF-Taxon="157069996" ID-Zenodo-Dep="5681917" checkinTime="1557324176605" checkinUser="plazi" docAuthor="Agosti, D." docDate="1994" docId="03A1879CFFD3FFB4CD75987266B6FD85" docLanguage="en" docName="jNatHist.28_ocr.pdf" docOrigin="Journal of Natural History 28" docStyle="DocumentStyle{}" docTitle="Cataglyphis hannae Agosti 1994, n. sp." docType="treatment" docVersion="10" lastPageId="5" lastPageNumber="918" masterDocId="FF98FFE4FFD2FFB1CE7990186062FFED" masterDocTitle="A new inquiline ant (Hymenoptera: Formicidae) in Cataglyphis and its phylogenetic relationship" masterLastPageNumber="919" masterPageNumber="913" pageId="1" pageNumber="914" updateTime="1636728940491" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new inquiline ant (Hymenoptera: Formicidae) in Cataglyphis and its phylogenetic relationship</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Agosti, D.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>1994</mods:date>
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<mods:detail type="pubDate">
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<mods:number>1993-10-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>28</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>913</mods:start>
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<mods:end>919</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.2949937</mods:identifier>
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<mods:identifier type="GBIF-Dataset">54b2150d-e662-450d-b3c2-73b51ac32041</mods:identifier>
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<mods:identifier type="Zenodo-Dep">2949937</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5681917" ID-GBIF-Taxon="157069996" ID-Zenodo-Dep="5681917" LSID="urn:lsid:plazi:treatment:03A1879CFFD3FFB4CD75987266B6FD85" httpUri="http://treatment.plazi.org/id/03A1879CFFD3FFB4CD75987266B6FD85" lastPageId="5" lastPageNumber="918" pageId="1" pageNumber="914">
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<subSubSection box="[780,1253,2154,2192]" pageId="1" pageNumber="914" type="nomenclature">
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<paragraph blockId="1.[261,1770,2154,2342]" box="[780,1253,2154,2192]" pageId="1" pageNumber="914">
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<heading box="[780,1253,2154,2192]" centered="true" fontSize="12" level="2" pageId="1" pageNumber="914" reason="4">
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<taxonomicName authority="Agosti, 1994" authorityName="Agosti" authorityYear="1994" box="[780,1142,2154,2192]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="917" phylum="Arthropoda" rank="species" species="hannae" status="n. sp.">
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<emphasis box="[780,1142,2154,2192]" italics="true" pageId="1" pageNumber="914">
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<emphasis bold="true" box="[780,992,2154,2192]" italics="true" pageId="1" pageNumber="914">Cataglyphis</emphasis>
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hannae
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</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1159,1253,2154,2192]" pageId="1" pageNumber="914" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="916" pageId="1" pageNumber="914" type="description">
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<paragraph blockId="1.[261,1770,2154,2342]" box="[262,599,2205,2243]" pageId="1" pageNumber="914">
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<emphasis box="[262,599,2205,2243]" italics="true" pageId="1" pageNumber="914">Material examined</emphasis>
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</paragraph>
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<paragraph blockId="1.[261,1770,2154,2342]" pageId="1" pageNumber="914">Holotype: <?, HL 1-40, HW 1-20, EL 0-50, SL 218, AL 2-63, CI 86, SI 181, El 41-7, FI 35-4, LI 100-8, TAI 114-3.</paragraph>
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<paragraph blockId="1.[261,1770,2406,2544]" box="[263,587,2407,2445]" pageId="1" pageNumber="914">
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<emphasis box="[263,587,2407,2445]" italics="true" pageId="1" pageNumber="914">Diagnosis of male</emphasis>
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</paragraph>
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<paragraph blockId="1.[261,1770,2406,2544]" pageId="1" pageNumber="914">
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Ant of the
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[531,883,2456,2494]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="914" phylum="Arthropoda" rank="species" species="bicolor">
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<emphasis box="[531,883,2456,2494]" italics="true" pageId="1" pageNumber="914">Cataglyphis bicolor</emphasis>
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</taxonomicName>
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species group (
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<bibRefCitation author="Agosti, D" box="[1168,1405,2456,2494]" firstAuthor="Agosti" journalOrPublisher="Journal of Natural History" pageId="1" pageNumber="914" pagination="1457 - 1505" part="24" refId="ref2824" refString="Agosti, D., 1990, Review and reclassification of Cataglyphis (Hymenoptera, Formicidae), Journal of Natural History, 24, 1457 - 1505." title="Review and reclassification of Cataglyphis (Hymenoptera, Formicidae)" type="journal article" year="1990">Agosti, 1990</bibRefCitation>
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), with the following diagnostic features:
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</paragraph>
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<paragraph blockId="1.[329,849,2592,2680]" box="[331,838,2593,2631]" pageId="1" pageNumber="914">(1) Black head and alitrunk.</paragraph>
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<paragraph blockId="1.[329,849,2592,2680]" box="[331,849,2642,2680]" pageId="1" pageNumber="914">(2) Alitrunk length 2-63 mm.</paragraph>
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<paragraph blockId="2.[281,1723,275,663]" pageId="2" pageNumber="915">(3) Subgenital plate distally trilobed with the median process triangular and not digitiform.</paragraph>
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<paragraph blockId="2.[281,1723,275,663]" box="[282,1520,375,413]" pageId="2" pageNumber="915">(4) In ventral view, median process with hair-carrying pits to the tip.</paragraph>
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<paragraph blockId="2.[281,1723,275,663]" pageId="2" pageNumber="915">(5) In lateral view, apicolateral appendix of sagitta not overreaching outline of apical, serrated plate.</paragraph>
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<paragraph blockId="2.[281,1723,275,663]" pageId="2" pageNumber="915">(6) Apicolateral appendix of sagitta short and terminally rounded; in ventral view only slightly raised above the plane of the apical, serrated plate.</paragraph>
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<paragraph blockId="2.[281,1723,275,663]" box="[282,885,625,663]" pageId="2" pageNumber="915">(7) Long hind tibiae (TAI 114-3).</paragraph>
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<paragraph blockId="2.[214,1721,698,1037]" pageId="2" pageNumber="915">
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<emphasis box="[306,459,704,734]" italics="true" pageId="2" pageNumber="915">Paratype</emphasis>
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: ♂ ♂ HL 1-30-1-43, HW 1-05-1-19, EL 0-44-0-50, SL 1-98-2-23, AL 2-48-2-73, CI 80-8-86-6, SI 175-5-195-6, El 41-1-43-0, FI 91-4-103-2, LI 101-7-105-5, TAI 108-3-116-0 (
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<emphasis box="[548,571,799,837]" italics="true" pageId="2" pageNumber="915">n</emphasis>
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= 8).
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</paragraph>
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<paragraph blockId="2.[214,1721,698,1037]" pageId="2" pageNumber="915">
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There is a slight variation in size, but the extremes of the range of absolute measurements are always far below those of the other males of the
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1452,1579,900,938]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="915" phylum="Arthropoda" rank="species" species="bicolor">
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<emphasis box="[1452,1579,900,938]" italics="true" pageId="2" pageNumber="915">bicolor</emphasis>
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</taxonomicName>
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species group. Compared with all the other males of the
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1062,1189,949,987]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="915" phylum="Arthropoda" rank="species" species="bicolor">
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<emphasis box="[1062,1189,949,987]" italics="true" pageId="2" pageNumber="915">bicolor</emphasis>
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</taxonomicName>
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group,
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<taxonomicName authorityName="Agosti" authorityYear="1994" box="[1323,1450,949,987]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="915" phylum="Arthropoda" rank="species" species="hannae">
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<emphasis box="[1323,1450,949,987]" italics="true" pageId="2" pageNumber="915">hannae</emphasis>
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</taxonomicName>
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has the longest hind tibiae (TAI> 108). Head and alitrunk always black in colour.
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</paragraph>
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<paragraph blockId="2.[214,1720,1099,1236]" box="[214,569,1099,1137]" pageId="2" pageNumber="915">
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<emphasis box="[214,569,1099,1137]" italics="true" pageId="2" pageNumber="915">Diagnosis offemale</emphasis>
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</paragraph>
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<paragraph blockId="2.[214,1720,1099,1236]" pageId="2" pageNumber="915">
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Ant of the
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[463,810,1148,1186]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="915" phylum="Arthropoda" rank="species" species="bicolor">
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<emphasis box="[463,810,1148,1186]" italics="true" pageId="2" pageNumber="915">Cataglyphis bicolor</emphasis>
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</taxonomicName>
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species group with the following diagnostic features (
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<figureCitation box="[230,360,1198,1236]" captionStart="Fig" captionStartId="2.[214,236,2623,2659]" captionTargetBox="[262,1676,1402,2588]" captionTargetPageId="2" captionText="Fig. 1. Lateral view ofthe females of C. bicolor (A) and its inquiline C. hannae n.sp. (B). Hairs are only drawn on the occiput." httpUri="https://zenodo.org/record/2949939/files/figure.png" pageId="2" pageNumber="915">Fig. 1B</figureCitation>
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):
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</paragraph>
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<paragraph blockId="2.[282,1211,1273,1360]" box="[282,901,1273,1311]" pageId="2" pageNumber="915">(1) Small size, alitrunk <3-2 mm.</paragraph>
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<paragraph blockId="2.[282,1211,1273,1360]" box="[282,1211,1322,1360]" pageId="2" pageNumber="915">(2) Scape much longer than head length (SI> 120).</paragraph>
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<caption httpUri="https://zenodo.org/record/2949939/files/figure.png" pageId="2" pageNumber="915" startId="2.[214,236,2623,2659]" targetBox="[262,1676,1402,2588]" targetPageId="2">
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<paragraph blockId="2.[214,1721,2623,2701]" pageId="2" pageNumber="915">
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Fig. 1. Lateral view of the females of
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[827,986,2623,2659]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="915" phylum="Arthropoda" rank="species" species="bicolor">
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<emphasis box="[827,854,2623,2659]" italics="true" pageId="2" pageNumber="915">C</emphasis>
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.
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<emphasis box="[871,986,2623,2659]" italics="true" pageId="2" pageNumber="915">bicolor</emphasis>
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</taxonomicName>
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(A) and its inquiline
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<taxonomicName authority="Agosti, 1994" authorityName="Agosti" authorityYear="1994" box="[1317,1476,2623,2659]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="915" phylum="Arthropoda" rank="species" species="hannae" status="n.sp.">
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<emphasis box="[1317,1476,2623,2659]" italics="true" pageId="2" pageNumber="915">C. hannae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1484,1557,2623,2659]" pageId="2" pageNumber="915" rank="species">n.sp.</taxonomicNameLabel>
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(B). Hairs are only drawn on the occiput.
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</paragraph>
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</caption>
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<paragraph blockId="3.[323,1493,271,457]" box="[324,910,271,308]" pageId="3" pageNumber="916">(3) Long hind tibiae (TAI> 90).</paragraph>
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<paragraph blockId="3.[323,1493,271,457]" box="[323,910,320,357]" pageId="3" pageNumber="916">(4) Head and alitrunk bright red.</paragraph>
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<paragraph blockId="3.[323,1493,271,457]" box="[324,1493,369,406]" pageId="3" pageNumber="916">(5) First funicular segment of intermediate length (28 <FI <35).</paragraph>
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<paragraph blockId="3.[323,1493,271,457]" box="[324,1046,420,457]" pageId="3" pageNumber="916">(6) Low rounded petiole in lateral view.</paragraph>
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<paragraph blockId="3.[257,1763,493,632]" pageId="3" pageNumber="916">
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Paratypes: ♀ ♀ HL 1-75-1-88, HW 1-65-1 -73, EL 0-48-0-50, SL 2-00-2-18, AL 2.95-3.13, CI 90-4—94-3, SI 121-2-129-9, FI 29-0-31-1, El 28-4-29-0, LI 101-8-107-1, TAI 93-5-100-8 (
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<emphasis box="[569,591,595,632]" italics="true" pageId="3" pageNumber="916">n</emphasis>
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= 5).
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</paragraph>
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<paragraph blockId="3.[258,1763,695,1082]" box="[258,746,695,732]" pageId="3" pageNumber="916">
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<emphasis box="[258,746,695,732]" italics="true" pageId="3" pageNumber="916">Deposition of type material</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="916" type="materials_examined">
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<paragraph blockId="3.[258,1763,695,1082]" pageId="3" pageNumber="916">
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<materialsCitation ID-GBIF-Occurrence="2239092173" collectedFrom="Oasis, at the edge of irrigated fields under Eucalyptus trees, in nest with Cataglyphis bicolor" collectingDate="0 8.06. 1992" collectionCode="MHNG" collectorName="D. Agosti" country="Tunisia" latitude="34.33" location="El Guettar" longitude="8.92" municipality="El Guettar" pageId="3" pageNumber="916" specimenCode="F92039" specimenCount="301" specimenCount-male="301" typeStatus="holotype">
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<typeStatus box="[324,508,744,781]" pageId="3" pageNumber="916">Holotype</typeStatus>
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:
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<specimenCount box="[536,546,744,781]" pageId="3" pageNumber="916" type="male">♂</specimenCount>
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,
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<emphasis bold="true" box="[590,739,744,781]" pageId="3" pageNumber="916">
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<collectingCountry box="[590,731,744,781]" name="Tunisia" pageId="3" pageNumber="916">Tunisia</collectingCountry>
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,
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</emphasis>
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<collectingMunicipality box="[753,939,744,781]" pageId="3" pageNumber="916">El Guettar</collectingMunicipality>
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,
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<geoCoordinate box="[960,1110,744,781]" pageId="3" pageNumber="916">34.33°N</geoCoordinate>
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,
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<geoCoordinate box="[1133,1254,744,781]" pageId="3" pageNumber="916">8.92°E</geoCoordinate>
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,
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<specimenCount box="[1276,1386,744,781]" pageId="3" pageNumber="916" type="male">
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<quantity box="[1276,1386,744,781]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="916" unit="m" value="300.0">300 m</quantity>
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</specimenCount>
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; in
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<collectedFrom pageId="3" pageNumber="916">
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Oasis, at the edge of irrigated fields under
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<taxonomicName box="[665,859,794,831]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="916" phylum="Tracheophyta" rank="genus">
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<emphasis box="[665,859,794,831]" italics="true" pageId="3" pageNumber="916">Eucalyptus</emphasis>
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</taxonomicName>
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trees
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</collectedFrom>
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,
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<collectorName box="[969,1138,794,831]" pageId="3" pageNumber="916">D. Agosti</collectorName>
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,
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<collectingDate box="[1154,1353,794,831]" pageId="3" pageNumber="916">0 8.06.1992</collectingDate>
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, sample
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<specimenCode box="[1501,1638,794,831]" pageId="3" pageNumber="916">F92039</specimenCode>
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,
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<collectedFrom pageId="3" pageNumber="916">
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in nest with
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[352,703,845,882]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="bicolor">
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<emphasis box="[352,703,845,882]" italics="true" pageId="3" pageNumber="916">Cataglyphis bicolor</emphasis>
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</taxonomicName>
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</collectedFrom>
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(
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<collectionCode box="[729,869,845,882]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="3" pageNumber="916">MHNG</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph blockId="3.[258,1763,695,1082]" pageId="3" pageNumber="916">
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<materialsCitation ID-GBIF-Occurrence="2239092172" box="[325,1419,893,930]" collectionCode="BMNH, CDA, CRW, MCZ, MHNG" collectorName="D. Agosti" country="Tunisia" location="Oasis" municipality="El Guettar" pageId="3" pageNumber="916" specimenCount="328" specimenCount-female="6" specimenCount-male="322" typeStatus="paratype">
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<typeStatus box="[325,521,893,930]" pageId="3" pageNumber="916">Paratypes</typeStatus>
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:
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<specimenCount box="[553,592,893,930]" pageId="3" pageNumber="916" type="female">5 ♀</specimenCount>
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<specimenCount box="[615,625,893,930]" pageId="3" pageNumber="916" type="female">♀</specimenCount>
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, (alates) and
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<specimenCount box="[921,996,893,930]" pageId="3" pageNumber="916" type="male">14 ♂</specimenCount>
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<specimenCount box="[1021,1031,893,930]" pageId="3" pageNumber="916" type="male">
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<emphasis box="[1021,1031,893,930]" italics="true" pageId="3" pageNumber="916">♂</emphasis>
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</specimenCount>
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, same as holotype.
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</materialsCitation>
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<materialsCitation ID-GBIF-Occurrence="2239092171" collectedFrom="Oasis, at the edge of an irrigated lucerne field, nest with Cataglyphis bicolor" collectingDate="0 8.06. 1992" collectionCode="BMNH, CDA, CRW, MCZ, MHNG" collectorName="D. Agosti" country="Tunisia" latitude="34.33" location="El Guettar" longitude="8.92" municipality="El Guettar" pageId="3" pageNumber="916" specimenCode="F92309" specimenCount="307" specimenCount-female="6" specimenCount-male="307" typeStatus="paratype">
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<specimenCount box="[1444,1592,893,930]" pageId="3" pageNumber="916" type="male">7 males</specimenCount>
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,
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<collectingCountry box="[1622,1756,893,930]" name="Tunisia" pageId="3" pageNumber="916">Tunisia</collectingCountry>
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,
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<collectingMunicipality box="[259,443,944,981]" pageId="3" pageNumber="916">El Guettar</collectingMunicipality>
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,
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<geoCoordinate box="[464,613,944,981]" pageId="3" pageNumber="916">34.33°N</geoCoordinate>
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,
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<geoCoordinate box="[635,756,944,981]" pageId="3" pageNumber="916">8.92°E</geoCoordinate>
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,
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<specimenCount box="[778,887,944,981]" pageId="3" pageNumber="916" type="male">
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<quantity box="[778,887,944,981]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="916" unit="m" value="300.0">300 m</quantity>
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</specimenCount>
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; in
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<collectedFrom box="[953,1756,944,981]" pageId="3" pageNumber="916">Oasis, at the edge of an irrigated lucerne field</collectedFrom>
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,
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<collectorName box="[259,427,994,1031]" pageId="3" pageNumber="916">D. Agosti</collectorName>
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,
|
|
<collectingDate box="[442,642,994,1031]" pageId="3" pageNumber="916">0 8.06.1992</collectingDate>
|
|
, sample
|
|
<specimenCode box="[789,925,994,1031]" pageId="3" pageNumber="916">F92309</specimenCode>
|
|
, in
|
|
<collectedFrom box="[982,1491,994,1031]" pageId="3" pageNumber="916">
|
|
nest with
|
|
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1146,1491,994,1031]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="bicolor">
|
|
<emphasis box="[1146,1491,994,1031]" italics="true" pageId="3" pageNumber="916">Cataglyphis bicolor</emphasis>
|
|
</taxonomicName>
|
|
</collectedFrom>
|
|
(
|
|
<collectionCode box="[1510,1647,994,1031]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="3" pageNumber="916">BMNH</collectionCode>
|
|
,
|
|
<collectionCode box="[1662,1756,994,1031]" pageId="3" pageNumber="916">CDA</collectionCode>
|
|
,
|
|
<collectionCode box="[259,361,1045,1082]" collectionName="CRW" pageId="3" pageNumber="916">CRW</collectionCode>
|
|
,
|
|
<collectionCode box="[384,481,1045,1082]" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" pageId="3" pageNumber="916">MCZ</collectionCode>
|
|
,
|
|
<collectionCode box="[503,642,1045,1082]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="3" pageNumber="916">MHNG</collectionCode>
|
|
).
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="3" pageNumber="916" type="diagnosis">
|
|
<paragraph blockId="3.[259,1769,1118,2693]" pageId="3" pageNumber="916">
|
|
<taxonomicName authorityName="Agosti" authorityYear="1994" box="[327,504,1118,1155]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="hannae">
|
|
<emphasis box="[327,504,1118,1155]" italics="true" pageId="3" pageNumber="916">C. hannae</emphasis>
|
|
</taxonomicName>
|
|
can be separated from all the other known species in the
|
|
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1525,1652,1118,1155]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="bicolor">
|
|
<emphasis box="[1525,1652,1118,1155]" italics="true" pageId="3" pageNumber="916">bicolor</emphasis>
|
|
</taxonomicName>
|
|
group by the above mentioned diagnostic characters, especially by the body size, the long antennal scape and hind tibiae, the low rounded petiole, and the male genitalia.
|
|
<emphasis box="[261,533,1268,1305]" italics="true" pageId="3" pageNumber="916">
|
|
<taxonomicName box="[261,526,1268,1305]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="abyssiniens">C. abyssiniens</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
the only species with small females, has a much shorter scape (SI <113), and shorter hind tibiae (TAI <85); no males of this species are known. The long black hairs on the occiput of the head as seen in
|
|
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1314,1441,1367,1404]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="bicolor">
|
|
<emphasis box="[1314,1441,1367,1404]" italics="true" pageId="3" pageNumber="916">bicolor</emphasis>
|
|
</taxonomicName>
|
|
are not present in
|
|
<emphasis box="[260,397,1416,1453]" italics="true" pageId="3" pageNumber="916">
|
|
<taxonomicName authorityName="Agosti" authorityYear="1994" box="[260,391,1416,1453]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="hannae">hannae</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
and thus exclude the possibility of
|
|
<taxonomicName authorityName="Agosti" authorityYear="1994" box="[1044,1171,1416,1453]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="hannae">
|
|
<emphasis box="[1044,1171,1416,1453]" italics="true" pageId="3" pageNumber="916">hannae</emphasis>
|
|
</taxonomicName>
|
|
being a microgyne of
|
|
<emphasis box="[1584,1716,1416,1453]" italics="true" pageId="3" pageNumber="916">
|
|
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1584,1711,1416,1453]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="bicolor">bicolor</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
</paragraph>
|
|
<paragraph blockId="3.[259,1769,1118,2693]" pageId="3" pageNumber="916">
|
|
<author box="[326,490,1466,1503]" firstAuthorName="Although" fns="*" inm="true" npo="*" pageId="3" pageNumber="916">Although</author>
|
|
>
|
|
<year box="[560,642,1466,1503]" pageId="3" pageNumber="916">1500</year>
|
|
different samples of species of the
|
|
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1287,1414,1466,1503]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="bicolor">
|
|
<emphasis box="[1287,1414,1466,1503]" italics="true" pageId="3" pageNumber="916">bicolor</emphasis>
|
|
</taxonomicName>
|
|
complex are in our collections, and most of them from Tunisia, no further
|
|
<taxonomicName authorityName="Agosti" authorityYear="1994" box="[1248,1375,1516,1553]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="hannae">
|
|
<emphasis box="[1248,1375,1516,1553]" italics="true" pageId="3" pageNumber="916">hannae</emphasis>
|
|
</taxonomicName>
|
|
specimens have been collected before. The host of
|
|
<taxonomicName authorityName="Agosti" authorityYear="1994" box="[782,908,1565,1602]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="hannae">
|
|
<emphasis box="[782,908,1565,1602]" italics="true" pageId="3" pageNumber="916">hannae</emphasis>
|
|
</taxonomicName>
|
|
is distributed along the southern foothills of the Atlas mountains in the transitional zone from the Mediterranean to the Desert region, not reaching the Atlantic in the West and not extending into Libya (Agosti
|
|
<emphasis box="[1662,1763,1665,1702]" italics="true" pageId="3" pageNumber="916">et al.,</emphasis>
|
|
in preparation). Thus,
|
|
<taxonomicName authorityName="Agosti" authorityYear="1994" box="[654,781,1714,1751]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="916" phylum="Arthropoda" rank="species" species="hannae">
|
|
<emphasis box="[654,781,1714,1751]" italics="true" pageId="3" pageNumber="916">hannae</emphasis>
|
|
</taxonomicName>
|
|
might be more common, certainly if one considers that the southern extension of the Mediterranean vegetation during the Pleistocene reached as far south as the Saharan mountains Tassili and Hoggar (
|
|
<bibRefCitation author="Quezel, P." bookContentInfo="333 pp." box="[1312,1555,1814,1851]" firstAuthor="Quezel" journalOrPublisher="du Tchad a la Mauritanie (Stuttgart: Gustav Fischer)" pageId="3" pageNumber="916" refId="ref3268" refString="Quezel, P., 1965, La vegetation du Sahara, du Tchad a la Mauritanie (Stuttgart: Gustav Fischer), 333 pp." title="La vegetation du Sahara" type="book" year="1965">Quezel, 1965</bibRefCitation>
|
|
). But social parasitic ants tend to have a clumped distribution (
|
|
<bibRefCitation author="Buschinger, A" box="[1134,1445,1863,1900]" firstAuthor="Buschinger" journalOrPublisher="Insectes Sociaux" pageId="3" pageNumber="916" pagination="321 - 324" part="32" refId="ref2931" refString="Buschinger, A., 1985, New records of rare parasitic ants (Hym., Form.) in the French Alps, Insectes Sociaux, 32, 321 - 324." title="New records of rare parasitic ants (Hym., Form.) in the French Alps" type="journal article" year="1985">Buschinger, 1985</bibRefCitation>
|
|
) and the restricted collecting area might truly reflect its distribution.
|
|
<journalOrPublisher jopScore="0.14357431" pageId="3" pageNumber="916" type="_" typeScore="1">Finally, these males and females were collected as alates, during a survey where</journalOrPublisher>
|
|
>
|
|
<pagination box="[1062,1127,1962,1999]" pageId="3" pageNumber="916" type="pagination">200</pagination>
|
|
nests were at least partially dug up.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="3" pageNumber="916" type="biology_ecology">
|
|
<paragraph blockId="3.[259,1769,1118,2693]" pageId="3" pageNumber="916">
|
|
The habitat was at the edge of an irrigated
|
|
<emphasis box="[1098,1391,2011,2048]" italics="true" pageId="3" pageNumber="916">Medicago sativa</emphasis>
|
|
field and on a sandy place in the shade of some tall
|
|
<emphasis box="[802,996,2061,2098]" italics="true" pageId="3" pageNumber="916">Eucalyptus</emphasis>
|
|
trees on the side of a track within the oasis. In this desert region, the occurrence of species of the
|
|
<emphasis box="[1166,1294,2111,2148]" italics="true" pageId="3" pageNumber="916">bicolor</emphasis>
|
|
group is restricted to oases.
|
|
</paragraph>
|
|
<paragraph blockId="3.[259,1769,1118,2693]" lastBlockId="5.[246,1752,280,619]" lastPageId="5" lastPageNumber="918" pageId="3" pageNumber="916">
|
|
The position of
|
|
<emphasis box="[603,730,2160,2197]" italics="true" pageId="3" pageNumber="916">hannae</emphasis>
|
|
within the
|
|
<emphasis box="[930,1057,2160,2197]" italics="true" pageId="3" pageNumber="916">bicolor</emphasis>
|
|
species group is established by the male genitalia and the funicular index of the female. More problematic, and challenging, is the sister group relationship within the
|
|
<emphasis box="[1049,1176,2259,2296]" italics="true" pageId="3" pageNumber="916">bicolor</emphasis>
|
|
group. A phylogenetic analysis using morphological characters of
|
|
<emphasis box="[891,994,2309,2346]" italics="true" pageId="3" pageNumber="916">nodus</emphasis>
|
|
(outgroup),
|
|
<emphasis italics="true" pageId="3" pageNumber="916">abyssinicus, bicolor, savignyi, viaticus</emphasis>
|
|
(
|
|
<tableCitation box="[436,585,2359,2396]" captionStart="Table 1" captionStartId="4.[205,294,416,452]" captionTargetBox="[303,1588,958,1314]" captionTargetPageId="4" captionText="Table 1. Data matrix. One ri: SI> 170 (character state 0), SI <170 (1); 2 Queen: CI> 95 (0), C K 95 (1); 3 Q: SI<105 (0); SI>105 (1); 4 Q, TL<3-5mm (0), TL<3-5mm (1); 5 Q, TAI <90 (0), TAI> 90 (1); 6, Q, white hairs on occiput of head (0), black hairs (1); 7, Q, head and alitrunk bright red (0), dark red to black (1); 8 M, lateral apical appendage of subgenital plate digitiforme, parallel sided (0), triangular (1); 9 M, median apical appendage of subgenital plate in crossection (at midlength) rounded (0), angular (1); 10 M, Sagitta with antero-lateral appendage convergent at base (0), parallel sided at base (1); 11 M, in lateral view, antero-lateral appendage of sagitta crossing outline of apical blade (0), not extended beyond (1); 12 M, median appendage of subgenital plate shorter as lateral appendages (0), of same length (1); 13 M, antero-lateral appendage of sagitta apically pointed (0), rounded (1). (A full discussion of the characters, and coding, based on> 1000 specimens, will be provided elsewhere (Agosti, in preparation)." httpUri="http://table.plazi.org/id/DF776602FFD6FFB5CEB491B8658DFC7D" pageId="3" pageNumber="916" tableUuid="DF776602FFD6FFB5CEB491B8658DFC7D">
|
|
<journalOrPublisher box="[436,537,2359,2396]" jopScore="0.0062278016" pageId="3" pageNumber="916">Table</journalOrPublisher>
|
|
1
|
|
</tableCitation>
|
|
) and the explicit enumeration procedure from the computer program Hennig86 (
|
|
<bibRefCitation author="Farris, S." box="[621,842,2408,2445]" firstAuthor="Farris" journalOrPublisher="Software and MSDOS program" pageId="3" pageNumber="916" refId="ref3094" refString="Farris, S., 1988, Hennig 86 version 1.5 manual, Software and MSDOS program." title="Hennig 86 version 1.5 manual, Software and MSDOS program" type="book" year="1988">Farris, 1988</bibRefCitation>
|
|
), resulted in 3 most parsimonious trees. Subsequent successive weighting established 2 most parsimonious trees (
|
|
<figureCitation box="[1410,1527,2458,2495]" captionStart="Fig" captionStartId="4.[205,227,2299,2335]" captionTargetBox="[643,1314,1482,2237]" captionTargetPageId="4" captionText="Fig. 2. Phylogenetic relationships of Cataglyphis hannae n. sp. Two most parsimonious trees shown with a consistency index (ci) of 98 and a retention index (ri) of 96. The synapomorphies for the sistergroup hannae + viaticus are small body size (characters 3 (1) in Table 1), and long antennal scape (4 (1)); for the sistergroup viaticus + (hannae + abyssinicus) the synapomorphies are bright red colour of head and alitrunk (7 (1)) and shape of median appendage of the male subgenital plate (9 (0))." httpUri="https://zenodo.org/record/2949941/files/figure.png" pageId="3" pageNumber="916">Fig. 2</figureCitation>
|
|
). Both these trees share the sister group
|
|
<emphasis box="[825,1217,2508,2545]" italics="true" pageId="3" pageNumber="916">hannae + abyssinicus,</emphasis>
|
|
and support a sister group
|
|
<emphasis box="[262,865,2558,2595]" italics="true" pageId="3" pageNumber="916">viaticus + (hannae + abyssinicus).</emphasis>
|
|
The sister group relationship of
|
|
<emphasis italics="true" pageId="3" pageNumber="916">hannae + abyssinicus</emphasis>
|
|
is so far only supported by female characters, as the males of
|
|
<emphasis box="[1566,1765,2607,2644]" italics="true" pageId="3" pageNumber="916">abyssinicus</emphasis>
|
|
are unknown. However, the long antennal scape and the small size of the females of
|
|
<emphasis box="[246,458,280,317]" italics="true" pageId="5" pageNumber="918">
|
|
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1904" box="[246,451,280,317]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="918" phylum="Arthropoda" rank="species" species="abyssinicus">abyssinicus</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
which is the same as in
|
|
<taxonomicName authorityName="Agosti" authorityYear="1994" box="[868,995,280,317]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="918" phylum="Arthropoda" rank="species" species="hannae">
|
|
<emphasis box="[868,995,280,317]" italics="true" pageId="5" pageNumber="918">hannae</emphasis>
|
|
</taxonomicName>
|
|
and unique in species of the
|
|
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1490,1617,280,317]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="918" phylum="Arthropoda" rank="species" species="bicolor">
|
|
<emphasis box="[1490,1617,280,317]" italics="true" pageId="5" pageNumber="918">bicolor</emphasis>
|
|
</taxonomicName>
|
|
species group, indicate an inquiline life style for this species too. A more detailed discussion will be provided by Agosti (in preparation).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption ID-Table-UUID="DF776602FFD6FFB5CEB491B8658DFC7D" httpUri="http://table.plazi.org/id/DF776602FFD6FFB5CEB491B8658DFC7D" pageId="4" pageNumber="917" targetBox="[303,1588,958,1314]" targetIsTable="true" targetPageId="4">
|
|
<paragraph blockId="4.[205,1711,416,912]" pageId="4" pageNumber="917">Table 1. Data matrix. One ri: SI> 170 (character state 0), SI <170 (1); 2 Queen: CI> 95 (0), C K 9 5 (1); 3 Q: S I<105 (0); SI>105 (1); 4 Q, TL<3-5mm (0), TL<3-5mm (1); 5 Q, TAI <90 (0), TAI> 90 (1); 6, Q, white hairs on occiput of head (0), black hairs (1); 7, Q, head and alitrunk bright red (0), dark red to black (1); 8 M, lateral apical appendage of subgenital plate digitiforme, parallel sided (0), triangular (1); 9 M, median apical appendage of subgenital plate in crossection (at midlength) rounded (0), angular (1); 10 M, Sagitta with antero-lateral appendage convergent at base (0), parallel sided at base (1); 11 M, in lateral view, antero-lateral appendage of sagitta crossing outline of apical blade (0), not extended beyond (1); 12 M, median appendage of subgenital plate shorter as lateral appendages (0), of same length (1); 13 M, antero-lateral appendage of sagitta apically pointed (0), rounded (1). (A full discussion of the characters, and coding, based on> 1000 specimens, will be provided elsewhere (Agosti, in preparation).</paragraph>
|
|
</caption>
|
|
<paragraph pageId="4" pageNumber="917">
|
|
<table box="[303,1588,958,1314]" gridcols="14" gridrows="8" pageId="4" pageNumber="917">
|
|
<tr box="[303,1588,958,995]" gridrow="0" pageId="4" pageNumber="917">
|
|
<th box="[303,491,958,995]" gridcol="0" gridrow="0" pageId="4" pageNumber="917">Species</th>
|
|
<th box="[563,1588,958,995]" colspan="13" colspanRight="12" gridcol="1" gridrow="0" pageId="4" pageNumber="917">Characters</th>
|
|
</tr>
|
|
<tr box="[303,1588,1028,1065]" gridrow="1" pageId="4" pageNumber="917" rowspan-0="1">
|
|
<td box="[563,588,1028,1065]" gridcol="1" gridrow="1" pageId="4" pageNumber="917">1</td>
|
|
<td box="[647,672,1028,1065]" gridcol="2" gridrow="1" pageId="4" pageNumber="917">2</td>
|
|
<td box="[730,755,1028,1065]" gridcol="3" gridrow="1" pageId="4" pageNumber="917">3</td>
|
|
<td box="[814,839,1028,1065]" gridcol="4" gridrow="1" pageId="4" pageNumber="917">4</td>
|
|
<td box="[898,923,1028,1065]" gridcol="5" gridrow="1" pageId="4" pageNumber="917">5</td>
|
|
<td box="[982,1006,1028,1065]" gridcol="6" gridrow="1" pageId="4" pageNumber="917">6</td>
|
|
<td box="[1066,1090,1028,1065]" gridcol="7" gridrow="1" pageId="4" pageNumber="917">7</td>
|
|
<td box="[1149,1174,1028,1065]" gridcol="8" gridrow="1" pageId="4" pageNumber="917">8</td>
|
|
<td box="[1233,1258,1028,1065]" gridcol="9" gridrow="1" pageId="4" pageNumber="917">9</td>
|
|
<td box="[1301,1341,1028,1065]" gridcol="10" gridrow="1" pageId="4" pageNumber="917">10</td>
|
|
<td box="[1385,1425,1028,1065]" gridcol="11" gridrow="1" pageId="4" pageNumber="917">11</td>
|
|
<td box="[1468,1509,1028,1065]" gridcol="12" gridrow="1" pageId="4" pageNumber="917">12</td>
|
|
<td box="[1548,1588,1028,1065]" gridcol="13" gridrow="1" pageId="4" pageNumber="917">13</td>
|
|
</tr>
|
|
<tr box="[303,1588,1070,1107]" gridrow="2" pageId="4" pageNumber="917">
|
|
<th box="[303,491,1070,1107]" gridcol="0" gridrow="2" pageId="4" pageNumber="917">
|
|
<taxonomicName baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[305,406,1071,1107]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="916" phylum="Arthropoda" rank="species" species="nodus">
|
|
<emphasis box="[305,406,1071,1107]" italics="true" pageId="4" pageNumber="917">Nodus</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[563,588,1070,1107]" gridcol="1" gridrow="2" pageId="4" pageNumber="917">1</td>
|
|
<td box="[647,672,1070,1107]" gridcol="2" gridrow="2" pageId="4" pageNumber="917">1</td>
|
|
<td box="[730,755,1070,1107]" gridcol="3" gridrow="2" pageId="4" pageNumber="917">0</td>
|
|
<td box="[814,839,1070,1107]" gridcol="4" gridrow="2" pageId="4" pageNumber="917">0</td>
|
|
<td box="[898,923,1070,1107]" gridcol="5" gridrow="2" pageId="4" pageNumber="917">0</td>
|
|
<td box="[982,1006,1070,1107]" gridcol="6" gridrow="2" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1066,1090,1070,1107]" gridcol="7" gridrow="2" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1149,1174,1070,1107]" gridcol="8" gridrow="2" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1233,1258,1070,1107]" gridcol="9" gridrow="2" pageId="4" pageNumber="917">1</td>
|
|
<td box="[1301,1341,1070,1107]" gridcol="10" gridrow="2" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1385,1425,1070,1107]" gridcol="11" gridrow="2" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1468,1509,1070,1107]" gridcol="12" gridrow="2" pageId="4" pageNumber="917">1</td>
|
|
<td box="[1548,1588,1070,1107]" gridcol="13" gridrow="2" pageId="4" pageNumber="917">1</td>
|
|
</tr>
|
|
<tr box="[303,1588,1111,1148]" gridrow="3" pageId="4" pageNumber="917">
|
|
<th box="[303,491,1111,1148]" gridcol="0" gridrow="3" pageId="4" pageNumber="917">
|
|
<emphasis box="[305,430,1112,1148]" italics="true" pageId="4" pageNumber="917">Hannae</emphasis>
|
|
</th>
|
|
<td box="[563,588,1111,1148]" gridcol="1" gridrow="3" pageId="4" pageNumber="917">0</td>
|
|
<td box="[647,672,1111,1148]" gridcol="2" gridrow="3" pageId="4" pageNumber="917">1</td>
|
|
<td box="[730,755,1111,1148]" gridcol="3" gridrow="3" pageId="4" pageNumber="917">1</td>
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<td box="[814,839,1111,1148]" gridcol="4" gridrow="3" pageId="4" pageNumber="917">1</td>
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|
<td box="[898,923,1111,1148]" gridcol="5" gridrow="3" pageId="4" pageNumber="917">1</td>
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|
<td box="[982,1006,1111,1148]" gridcol="6" gridrow="3" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1066,1090,1111,1148]" gridcol="7" gridrow="3" pageId="4" pageNumber="917">1</td>
|
|
<td box="[1149,1174,1111,1148]" gridcol="8" gridrow="3" pageId="4" pageNumber="917">1</td>
|
|
<td box="[1233,1258,1111,1148]" gridcol="9" gridrow="3" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1301,1341,1111,1148]" gridcol="10" gridrow="3" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1385,1425,1111,1148]" gridcol="11" gridrow="3" pageId="4" pageNumber="917">1</td>
|
|
<td box="[1468,1509,1111,1148]" gridcol="12" gridrow="3" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1548,1588,1111,1148]" gridcol="13" gridrow="3" pageId="4" pageNumber="917">0</td>
|
|
</tr>
|
|
<tr box="[303,1588,1153,1190]" gridrow="4" pageId="4" pageNumber="917">
|
|
<th box="[303,491,1153,1190]" gridcol="0" gridrow="4" pageId="4" pageNumber="917">
|
|
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1904" box="[303,491,1153,1189]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="918" phylum="Arthropoda" rank="species" species="abyssinicus">
|
|
<emphasis box="[303,491,1153,1189]" italics="true" pageId="4" pageNumber="917">Abyssinicus</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[563,588,1153,1190]" gridcol="1" gridrow="4" pageId="4" pageNumber="917">-</td>
|
|
<td box="[647,672,1153,1190]" gridcol="2" gridrow="4" pageId="4" pageNumber="917">1</td>
|
|
<td box="[730,755,1153,1190]" gridcol="3" gridrow="4" pageId="4" pageNumber="917">1</td>
|
|
<td box="[814,839,1153,1190]" gridcol="4" gridrow="4" pageId="4" pageNumber="917">1</td>
|
|
<td box="[898,923,1153,1190]" gridcol="5" gridrow="4" pageId="4" pageNumber="917">0</td>
|
|
<td box="[982,1006,1153,1190]" gridcol="6" gridrow="4" pageId="4" pageNumber="917">0</td>
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|
<td box="[1066,1090,1153,1190]" gridcol="7" gridrow="4" pageId="4" pageNumber="917">1</td>
|
|
<td box="[1149,1174,1153,1190]" gridcol="8" gridrow="4" pageId="4" pageNumber="917">-</td>
|
|
<td box="[1233,1258,1153,1190]" gridcol="9" gridrow="4" pageId="4" pageNumber="917">-</td>
|
|
<td box="[1301,1341,1153,1190]" gridcol="10" gridrow="4" pageId="4" pageNumber="917">-</td>
|
|
<td box="[1385,1425,1153,1190]" gridcol="11" gridrow="4" pageId="4" pageNumber="917">-</td>
|
|
<td box="[1468,1509,1153,1190]" gridcol="12" gridrow="4" pageId="4" pageNumber="917">-</td>
|
|
<td box="[1548,1588,1153,1190]" gridcol="13" gridrow="4" pageId="4" pageNumber="917">-</td>
|
|
</tr>
|
|
<tr box="[303,1588,1194,1231]" gridrow="5" pageId="4" pageNumber="917">
|
|
<th box="[303,491,1194,1231]" gridcol="0" gridrow="5" pageId="4" pageNumber="917">
|
|
<emphasis box="[305,424,1194,1230]" italics="true" pageId="4" pageNumber="917">Bicolor</emphasis>
|
|
</th>
|
|
<td box="[563,588,1194,1231]" gridcol="1" gridrow="5" pageId="4" pageNumber="917">1</td>
|
|
<td box="[647,672,1194,1231]" gridcol="2" gridrow="5" pageId="4" pageNumber="917">0</td>
|
|
<td box="[730,755,1194,1231]" gridcol="3" gridrow="5" pageId="4" pageNumber="917">0</td>
|
|
<td box="[814,839,1194,1231]" gridcol="4" gridrow="5" pageId="4" pageNumber="917">0</td>
|
|
<td box="[898,923,1194,1231]" gridcol="5" gridrow="5" pageId="4" pageNumber="917">0</td>
|
|
<td box="[982,1006,1194,1231]" gridcol="6" gridrow="5" pageId="4" pageNumber="917">1</td>
|
|
<td box="[1066,1090,1194,1231]" gridcol="7" gridrow="5" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1149,1174,1194,1231]" gridcol="8" gridrow="5" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1233,1258,1194,1231]" gridcol="9" gridrow="5" pageId="4" pageNumber="917">1</td>
|
|
<td box="[1301,1341,1194,1231]" gridcol="10" gridrow="5" pageId="4" pageNumber="917">1</td>
|
|
<td box="[1385,1425,1194,1231]" gridcol="11" gridrow="5" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1468,1509,1194,1231]" gridcol="12" gridrow="5" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1548,1588,1194,1231]" gridcol="13" gridrow="5" pageId="4" pageNumber="917">0</td>
|
|
</tr>
|
|
<tr box="[303,1588,1236,1273]" gridrow="6" pageId="4" pageNumber="917">
|
|
<th box="[303,491,1236,1273]" gridcol="0" gridrow="6" pageId="4" pageNumber="917">
|
|
<taxonomicName baseAuthorityName="Dufour" baseAuthorityYear="1862" box="[306,441,1237,1273]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="918" phylum="Arthropoda" rank="species" species="savignyi">
|
|
<emphasis box="[306,441,1237,1273]" italics="true" pageId="4" pageNumber="917">Savignyi</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[563,588,1236,1273]" gridcol="1" gridrow="6" pageId="4" pageNumber="917">0</td>
|
|
<td box="[647,672,1236,1273]" gridcol="2" gridrow="6" pageId="4" pageNumber="917">0</td>
|
|
<td box="[730,755,1236,1273]" gridcol="3" gridrow="6" pageId="4" pageNumber="917">0</td>
|
|
<td box="[814,839,1236,1273]" gridcol="4" gridrow="6" pageId="4" pageNumber="917">0</td>
|
|
<td box="[898,923,1236,1273]" gridcol="5" gridrow="6" pageId="4" pageNumber="917">0</td>
|
|
<td box="[982,1006,1236,1273]" gridcol="6" gridrow="6" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1066,1090,1236,1273]" gridcol="7" gridrow="6" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1149,1174,1236,1273]" gridcol="8" gridrow="6" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1233,1258,1236,1273]" gridcol="9" gridrow="6" pageId="4" pageNumber="917">1</td>
|
|
<td box="[1301,1341,1236,1273]" gridcol="10" gridrow="6" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1385,1425,1236,1273]" gridcol="11" gridrow="6" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1468,1509,1236,1273]" gridcol="12" gridrow="6" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1548,1588,1236,1273]" gridcol="13" gridrow="6" pageId="4" pageNumber="917">0</td>
|
|
</tr>
|
|
<tr box="[303,1588,1277,1314]" gridrow="7" pageId="4" pageNumber="917">
|
|
<th box="[303,491,1277,1314]" gridcol="0" gridrow="7" pageId="4" pageNumber="917">
|
|
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[308,434,1278,1314]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="917" phylum="Arthropoda" rank="species" species="viaticus">
|
|
<emphasis box="[308,434,1278,1314]" italics="true" pageId="4" pageNumber="917">Viaticus</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<td box="[563,588,1277,1314]" gridcol="1" gridrow="7" pageId="4" pageNumber="917">1</td>
|
|
<td box="[647,672,1277,1314]" gridcol="2" gridrow="7" pageId="4" pageNumber="917">0</td>
|
|
<td box="[730,755,1277,1314]" gridcol="3" gridrow="7" pageId="4" pageNumber="917">0</td>
|
|
<td box="[814,839,1277,1314]" gridcol="4" gridrow="7" pageId="4" pageNumber="917">0</td>
|
|
<td box="[898,923,1277,1314]" gridcol="5" gridrow="7" pageId="4" pageNumber="917">0</td>
|
|
<td box="[982,1006,1277,1314]" gridcol="6" gridrow="7" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1066,1090,1277,1314]" gridcol="7" gridrow="7" pageId="4" pageNumber="917">1</td>
|
|
<td box="[1149,1174,1277,1314]" gridcol="8" gridrow="7" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1233,1258,1277,1314]" gridcol="9" gridrow="7" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1301,1341,1277,1314]" gridcol="10" gridrow="7" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1385,1425,1277,1314]" gridcol="11" gridrow="7" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1468,1509,1277,1314]" gridcol="12" gridrow="7" pageId="4" pageNumber="917">0</td>
|
|
<td box="[1548,1588,1277,1314]" gridcol="13" gridrow="7" pageId="4" pageNumber="917">0</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<caption httpUri="https://zenodo.org/record/2949941/files/figure.png" pageId="4" pageNumber="917" startId="4.[205,227,2299,2335]" targetBox="[643,1314,1482,2237]" targetPageId="4">
|
|
<paragraph blockId="4.[205,1712,2299,2541]" pageId="4" pageNumber="917">
|
|
Fig. 2. Phylogenetic relationships of
|
|
<taxonomicName authority="Agosti, 1994" authorityName="Agosti" authorityYear="1994" box="[854,1180,2299,2335]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="917" phylum="Arthropoda" rank="species" species="hannae" status="n. sp.">
|
|
<emphasis box="[854,1180,2299,2335]" italics="true" pageId="4" pageNumber="917">Cataglyphis hannae</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[1200,1291,2299,2335]" pageId="4" pageNumber="917" rank="species">n. sp.</taxonomicNameLabel>
|
|
Two most parsimonious trees shown with a consistency index
|
|
<emphasis box="[937,990,2340,2376]" italics="true" pageId="4" pageNumber="917">(ci)</emphasis>
|
|
of 98 and a retention index
|
|
<emphasis box="[1473,1524,2340,2376]" italics="true" pageId="4" pageNumber="917">(ri)</emphasis>
|
|
of 96. The synapomorphies for the sistergroup
|
|
<emphasis box="[900,1186,2382,2418]" italics="true" pageId="4" pageNumber="917">
|
|
<taxonomicName authorityName="Agosti" authorityYear="1994" box="[900,1017,2382,2418]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="917" phylum="Arthropoda" rank="species" species="hannae">hannae</taxonomicName>
|
|
+
|
|
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[1064,1186,2382,2418]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="917" phylum="Arthropoda" rank="species" species="viaticus">viaticus</taxonomicName>
|
|
</emphasis>
|
|
are small body size (characters 3 (1) in Table 1), and long antennal scape (4 (1)); for the sistergroup
|
|
<emphasis italics="true" pageId="4" pageNumber="917">
|
|
<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[1546,1668,2423,2459]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="917" phylum="Arthropoda" rank="species" species="viaticus">viaticus</taxonomicName>
|
|
+ (
|
|
<taxonomicName authorityName="Agosti" authorityYear="1994" box="[317,435,2464,2500]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="917" phylum="Arthropoda" rank="species" species="hannae">hannae</taxonomicName>
|
|
+
|
|
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1904" box="[481,664,2464,2500]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="917" phylum="Arthropoda" rank="species" species="abyssinicus">abyssinicus</taxonomicName>
|
|
)
|
|
</emphasis>
|
|
the synapomorphies are bright red colour of head and alitrunk (7 (1)) and shape of median appendage of the male subgenital plate (9 (0)).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph blockId="5.[246,1752,280,619]" pageId="5" pageNumber="918">
|
|
As all the results are based on morphological characters, it would be interesting to test them using other characters such as DNA sequences, electrophoretic data or the chemical composition of glandular compounds, all of which have already showed their specificity at the species level (
|
|
<bibRefCitation author="Keegans, S. J. & Morgan, E. D. & Agosti, D. & Wehner, R." box="[818,1198,579,616]" etAl="et al." firstAuthor="Keegans" journalOrPublisher="Biochemical Systematics and Ecology" pageId="5" pageNumber="918" pagination="559 - 572" part="20" refId="ref3173" refString="Keegans, S. J., Morgan, E. D., Agosti, D. and Wehner, R., 1992, What do glands tell us about species? A chemical case study of Cataglyphis ants, Biochemical Systematics and Ecology, 20, 559 - 572." title="What do glands tell us about species? A chemical case study of Cataglyphis ants" type="journal article" year="1992">
|
|
Keegans
|
|
<emphasis box="[988,1020,579,616]" italics="true" pageId="5" pageNumber="918">et</emphasis>
|
|
al., 1992
|
|
</bibRefCitation>
|
|
; Agosti
|
|
<emphasis box="[1357,1458,579,616]" italics="true" pageId="5" pageNumber="918">et al.,</emphasis>
|
|
in preparation).
|
|
</paragraph>
|
|
</treatment>
|
|
</document> |