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<document id="4D3A4DA20D41507CC756164BFF45CAA7" ID-DOI="10.5281/zenodo.3746744" ID-GBIF-Dataset="3ab3ca17-b957-454a-bdcf-d664740802e6" ID-ISSN="0253-116X" ID-Zenodo-Dep="3746744" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1586462215853" checkinUser="valdenar" docAuthor="Opitz, Weston" docDate="2019" docId="03A1879CD842FFC23CB4FBB33142FAC9" docLanguage="en" docName="Linzer.biol.Beitr.51.2.0959-1076.pdf.imf" docOrigin="Linzer biologische Beiträge 51 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Madoniella delta, nov.sp." docType="treatment" docVersion="7" lastPageNumber="983" masterDocId="FF98FFE4D855FFDA3C23FFBA323FFB66" masterDocTitle="Descriptions of new genera and new species of Western Hemisphere checkered beetles (Coleoptera, Cleroidea, Cleridae)" masterLastPageNumber="1076" masterPageNumber="959" pageNumber="982" updateTime="1698736766143" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7E9C2278AC3DA731492CD83C2A48F2BD">Descriptions of new genera and new species of Western Hemisphere checkered beetles (Coleoptera, Cleroidea, Cleridae)</mods:title>
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<mods:namePart id="884DFC569AFCCA0CDE56FDBD0C8C586E">Opitz, Weston</mods:namePart>
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<mods:title id="06C8C1917AD7FC36233C9D52D0307012">Linzer biologische Beiträge</mods:title>
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<mods:date id="363D43D7624FEEF026762549DCA93B50">2019</mods:date>
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<mods:number id="C826826E4424F4BED612DA9DEE32B81E">2019-12-20</mods:number>
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<mods:identifier id="6B6758FBDDAB90063D37C7EA0B378853" type="DOI">10.5281/zenodo.3746744</mods:identifier>
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<treatment id="03A1879CD842FFC23CB4FBB33142FAC9" ID-DOI="http://doi.org/10.5281/zenodo.3809160" ID-GBIF-Taxon="163544771" ID-Zenodo-Dep="3809160" LSID="urn:lsid:plazi:treatment:03A1879CD842FFC23CB4FBB33142FAC9" httpUri="http://treatment.plazi.org/id/03A1879CD842FFC23CB4FBB33142FAC9" lastPageId="24" lastPageNumber="983" pageId="23" pageNumber="982">
<subSubSection id="C3126501D842FFCD3CB4FBB330E0FF79" box="[151,735,1033,1055]" pageId="23" pageNumber="982" type="nomenclature">
<paragraph id="8BB7368AD842FFCD3CB4FBB330E0FF79" blockId="23.[151,735,1033,1055]" box="[151,735,1033,1055]" pageId="23" pageNumber="982">
<heading id="D0FF81E6D842FFCD3CB4FBB330E0FF79" box="[151,735,1033,1055]" fontSize="8" level="2" pageId="23" pageNumber="982" reason="2">
<emphasis id="B97CEA98D842FFCD3CB4FBB330E0FF79" bold="true" box="[151,735,1033,1055]" pageId="23" pageNumber="982">
<taxonomicName id="4C084D09D842FFCD3CB4FBB33377FF79" authority="Opitz, 2019" box="[151,328,1033,1055]" class="Insecta" family="Cleridae" genus="Madoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="982" phylum="Arthropoda" rank="species" species="delta">
<emphasis id="B97CEA98D842FFCD3CB4FBB33377FF79" bold="true" box="[151,328,1033,1055]" italics="true" pageId="23" pageNumber="982">Madoniella delta</emphasis>
</taxonomicName>
OPITZ
<taxonomicNameLabel id="A24F57E3D842FFCD3DB4FBB333DCFF78" box="[407,483,1033,1054]" pageId="23" pageNumber="982" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="13332A0FD842FFCD3DD3FBB3307FFF78" box="[496,576,1033,1055]" captionStart="Figs 61-91" captionStartId="73.[151,191,1343,1365]" captionTargetBox="[151,1033,198,1310]" captionTargetId="figure@73.[151,1059,198,1328]" captionTargetPageId="73" captionText="Figs 61-91: Antennae; (61) Madoniella muralla; (62) M. nota; (63) M. acicula; (64) M. aspera; (65) M. baja; (66) M. delta; (67) Amboakis bermejo; (68) Pujoliclerus variolus; (69) Megatrachys irrasis; (70) Cregya arima; (71) Nelsopelonium aspikelum; (72) Cregya vicosa; (73) Pujoliclerus abditus; (74) P. dexus; (75) Cregya triguttula; (76) Pyticeroides brevitubulus; (77) Pujoliclerus lasiosus; (78) Cregya biguttula; (79) Sirpa acara; (80) Parapelonides naranjo; (81) Cregya livida; (82) Amboakis collucoatis; (83) Ampleris jatai; (84) Megaphloeus yasuni; (85) Pelonium implacide, male; (86) Epiphloeus prolixicornus; (87) Pelonium implacide, female; (88) Epiphloeus mantillerii; (89) Amboakis mitaraka; (90) Ellipotoma makrosa; (91) Cregya maura." figureDoi="http://doi.org/10.5281/zenodo.3746752" httpUri="https://zenodo.org/record/3746752/files/figure.png" pageId="23" pageNumber="982">Figs 66</figureCitation>
,
<figureCitation id="13332A0FD842FFCD3E68FBB3304DFF78" box="[587,626,1033,1054]" captionStart="Figs 179-193" captionStartId="81.[151,191,1359,1381]" captionTargetBox="[167,1032,201,1342]" captionTargetId="figure@81.[152,1058,201,1346]" captionTargetPageId="81" captionText="Figs 179-193: Pronota; 179) Phonius versus; (180) Perilypus arsus; (181) A. nodus; (182) Pyticara frontegalba; (183) Pelonium implacide; (184) Neorthoplera quintana; (185) Perilypus virgulatus; (186) P. alpinus; (187) Amboakis expandosa; (188) Ampleris jatai; (189) Megaphloeus jasuni; (190) Amboakis bermejo (191) Madoniella delta; (192) Megaphloeus imparilus; (193) Pyticeroides brevitubulus." figureDoi="http://doi.org/10.5281/zenodo.3746768" httpUri="https://zenodo.org/record/3746768/files/figure.png" pageId="23" pageNumber="982">191</figureCitation>
,
<figureCitation id="13332A0FD842FFCD3E5DFBB3309AFF78" box="[638,677,1033,1054]" captionStart="Figs 219-224" captionStartId="86.[151,191,1278,1300]" captionTargetBox="[153,1061,198,1263]" captionTargetId="figure@86.[153,1061,198,1263]" captionTargetPageId="86" captionText="Figs 219-224: Aedeagi; (219) Amboakis mitaraka; (220) Epiphloeus mantillerii; (221) E. prolixicornus; (222) Ichnea cerinis; (223) Madoniella delta; (224) M. muralla." figureDoi="http://doi.org/10.5281/zenodo.3746778" httpUri="https://zenodo.org/record/3746778/files/figure.png" pageId="23" pageNumber="982">223</figureCitation>
,
<figureCitation id="13332A0FD842FFCD3E93FBB330E8FF79" box="[688,727,1033,1055]" captionStart="Figs 294-297" captionStartId="101.[151,191,1525,1547]" captionTargetBox="[153,1062,200,1513]" captionTargetId="figure@101.[153,1062,200,1514]" captionTargetPageId="101" captionText="Figs 294-297: Habitus; (294) Madoniella aspera; (295) M. baja; (296) M. delta; (297) M. muralla." figureDoi="http://doi.org/10.5281/zenodo.3746808" httpUri="https://zenodo.org/record/3746808/files/figure.png" pageId="23" pageNumber="982">296</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3126501D842FFC23CB4FB943142FAC9" lastPageId="24" lastPageNumber="983" pageId="23" pageNumber="982" type="materials_examined">
<paragraph id="8BB7368AD842FFCD3CB4FB94330BFF3D" blockId="23.[151,1058,1070,1115]" pageId="23" pageNumber="982">
<materialsCitation id="3B603CD7D842FFCD3CB4FB94330BFF3D" ID-GBIF-Occurrence="2597529827" collectingDate="2015-06-07" collectingDateMax="2015-06-13" collectingDateMin="2015-06-07" collectionCode="FSCA" collectorName="Morris &amp; Wappes" elevation="1300" location="Cerro Jesus" pageId="23" pageNumber="982" specimenCount="1" stateProvince="Nueva Segovia" typeStatus="holotype">
<typeStatus id="54B38828D842FFCD3CB4FB9432D0FF22" box="[151,239,1070,1092]" pageId="23" pageNumber="982" type="holotype">Holotype</typeStatus>
: ♂. Type locality: NIC:
<collectingRegion id="49CCF868D842FFCD3DFBFB943058FF22" box="[472,615,1070,1092]" country="Nicaragua" name="Nueva Segovia" pageId="23" pageNumber="982">Nueva Segovia</collectingRegion>
,
<location id="8ED76051D842FFCD3E51FB9430DFFF25" LSID="urn:lsid:plazi:treatment:03A1879CD842FFC23CB4FBB33142FAC9:8ED76051D842FFCD3E51FB9430DFFF25" box="[626,736,1070,1091]" name="Cerro Jesus" pageId="23" pageNumber="982" stateProvince="Nueva Segovia">Cerro Jesus</location>
,
<quantity id="4CF09B6FD842FFCD3ECFFB943109FF25" box="[748,822,1070,1091]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="982" unit="m" value="1300.0">
<elevation id="0025D1B9D842FFCD3ECFFB943109FF25" box="[748,822,1070,1091]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="982" unit="m" value="1300.0">1300 m</elevation>
</quantity>
,
<date id="FFB6104AD842FFCD3F62FB943180FF22" box="[833,959,1070,1092]" pageId="23" pageNumber="982" value="2015-06-07" valueMax="2015-06-13" valueMin="2015-06-07">
<collectingDate id="EFF2E9A2D842FFCD3F62FB943180FF22" box="[833,959,1070,1092]" pageId="23" pageNumber="982" value="2015-06-07" valueMax="2015-06-13" valueMin="2015-06-07">VI/7-13/2015</collectingDate>
</date>
,
<collectorName id="26FD535CD842FFCD3FE8FB943636FF22" box="[971,1033,1070,1092]" pageId="23" pageNumber="982">Morris</collectorName>
&amp;
<collectorName id="26FD535CD842FFCD3CB4FBFF32DFFF3C" box="[151,224,1093,1114]" pageId="23" pageNumber="982">Wappes</collectorName>
(
<collectionCode id="ED19AE4FD842FFCD3CCFFBFF3315FF3D" box="[236,298,1093,1115]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="23" pageNumber="982" type="Museum">FSCA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB7368AD842FFCD3CB4FBD3313FFF92" blockId="23.[151,1058,1129,1268]" pageId="23" pageNumber="982">
D i a g n o s i s: The genus
<taxonomicName id="4C084D09D842FFCD3DF3FBD3307AFF18" box="[464,581,1129,1150]" class="Insecta" family="Cleridae" genus="Madoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="982" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEA98D842FFCD3DF3FBD3307AFF18" box="[464,581,1129,1150]" italics="true" pageId="23" pageNumber="982">Madoniella</emphasis>
</taxonomicName>
PIC was revised in 2011 (
<bibRefCitation id="EF994B7BD842FFCD3F7AFBD331DEFF19" author="OPITZ W" box="[857,993,1129,1151]" pageId="23" pageNumber="982" pagination="133 - 248" refId="ref53228" refString="OPITZ W. (2011 b): Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part X. The genus Madoniella PIC. - Entomologica Basiliensia et Collectionis Frey 33: 133 - 248." type="journal article" year="2011">OPITZ 2011b</bibRefCitation>
). This work includes a key to species.
<taxonomicName id="4C084D09D842FFCD3DD1FB3C309BFFFD" box="[498,676,1158,1179]" class="Insecta" family="Cleridae" genus="Madoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="982" phylum="Arthropoda" rank="species" species="delta">
<emphasis id="B97CEA98D842FFCD3DD1FB3C309BFFFD" box="[498,676,1158,1179]" italics="true" pageId="23" pageNumber="982">Madoniella delta</emphasis>
</taxonomicName>
specimens key out to
<emphasis id="B97CEA98D842FFCD3FB8FB3D361DFFFD" box="[923,1058,1158,1180]" italics="true" pageId="23" pageNumber="982">M. kuhlorum</emphasis>
OPITZ, from which the
<taxonomicName id="4C084D09D842FFCD3DA7FB1E33E6FFDE" box="[388,473,1187,1209]" class="Insecta" family="Cleridae" genus="Madoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="982" phylum="Arthropoda" rank="species" species="delta">
<emphasis id="B97CEA98D842FFCD3DA7FB1E33E6FFDE" box="[388,473,1187,1209]" italics="true" pageId="23" pageNumber="982">M. delta</emphasis>
</taxonomicName>
specimen differs by lacking a posterolateral extension of the anterior block. Also, the phallobasic apodeme is very short in the available specimen of
<taxonomicName id="4C084D09D842FFCD3C90FB653337FF95" box="[179,264,1246,1268]" class="Insecta" family="Cleridae" genus="Madoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="982" phylum="Arthropoda" rank="species" species="delta">
<emphasis id="B97CEA98D842FFCD3C90FB653337FF95" box="[179,264,1246,1268]" italics="true" pageId="23" pageNumber="982">M. delta</emphasis>
</taxonomicName>
, which is not the case in
<emphasis id="B97CEA98D842FFCD3E24FB6530B5FF95" box="[519,650,1246,1268]" italics="true" pageId="23" pageNumber="982">M. kuhlorum</emphasis>
specimens.
</paragraph>
<paragraph id="8BB7368AD842FFC23CB4FAB9311FFA65" blockId="23.[151,1059,1282,1584]" lastBlockId="24.[151,1058,210,260]" lastPageId="24" lastPageNumber="983" pageId="23" pageNumber="982">
D e s c r i p t i o n: Size: Length 5.0 mm; width
<quantity id="4CF09B6FD842FFCD3E82FAB830CEFE7E" box="[673,753,1282,1304]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="982" unit="mm" value="1.5">1.5 mm</quantity>
. Form: As in
<figureCitation id="13332A0FD842FFCD3F5CFAB831EBFE7E" box="[895,980,1282,1304]" captionStart="Figs 294-297" captionStartId="101.[151,191,1525,1547]" captionTargetBox="[153,1062,200,1513]" captionTargetId="figure@101.[153,1062,200,1514]" captionTargetPageId="101" captionText="Figs 294-297: Habitus; (294) Madoniella aspera; (295) M. baja; (296) M. delta; (297) M. muralla." figureDoi="http://doi.org/10.5281/zenodo.3746808" httpUri="https://zenodo.org/record/3746808/files/figure.png" pageId="23" pageNumber="982">Fig. 296</figureCitation>
. Color: Cranium red-castaneous; antenna brown, except scape testaceous; pronotum bicolored, mostly black, with a triangular red mark that extends laterally along pronotal arch; pterothorax black; elytra mostly dark brown, insignia well developed, anterolateral and anterocentral extensions of anterior block poorly developed, posterolateral extension of anterior block absent; legs and abdomen brown. Head: Cranium coarsely punctate; antenna (
<figureCitation id="13332A0FD842FFCD3CDDFA10336CFEA6" box="[254,339,1450,1472]" captionStart="Figs 61-91" captionStartId="73.[151,191,1343,1365]" captionTargetBox="[151,1033,198,1310]" captionTargetId="figure@73.[151,1059,198,1328]" captionTargetPageId="73" captionText="Figs 61-91: Antennae; (61) Madoniella muralla; (62) M. nota; (63) M. acicula; (64) M. aspera; (65) M. baja; (66) M. delta; (67) Amboakis bermejo; (68) Pujoliclerus variolus; (69) Megatrachys irrasis; (70) Cregya arima; (71) Nelsopelonium aspikelum; (72) Cregya vicosa; (73) Pujoliclerus abditus; (74) P. dexus; (75) Cregya triguttula; (76) Pyticeroides brevitubulus; (77) Pujoliclerus lasiosus; (78) Cregya biguttula; (79) Sirpa acara; (80) Parapelonides naranjo; (81) Cregya livida; (82) Amboakis collucoatis; (83) Ampleris jatai; (84) Megaphloeus yasuni; (85) Pelonium implacide, male; (86) Epiphloeus prolixicornus; (87) Pelonium implacide, female; (88) Epiphloeus mantillerii; (89) Amboakis mitaraka; (90) Ellipotoma makrosa; (91) Cregya maura." figureDoi="http://doi.org/10.5281/zenodo.3746752" httpUri="https://zenodo.org/record/3746752/files/figure.png" pageId="23" pageNumber="982">Fig. 66</figureCitation>
) capitate, funicular antennomeres filiform, progressively shorter towards capitulum, capitular antennomeres 8 and 9 triangular, antennomere 10 ovoid; eyes large, ommatidia small, eye notch large, eye narrower than frons (EW/FW 20/30). Thorax: Pronotum (
<figureCitation id="13332A0FD842FFCD3D47FA443382FD72" box="[356,445,1534,1556]" captionStart="Figs 179-193" captionStartId="81.[151,191,1359,1381]" captionTargetBox="[167,1032,201,1342]" captionTargetId="figure@81.[152,1058,201,1346]" captionTargetPageId="81" captionText="Figs 179-193: Pronota; 179) Phonius versus; (180) Perilypus arsus; (181) A. nodus; (182) Pyticara frontegalba; (183) Pelonium implacide; (184) Neorthoplera quintana; (185) Perilypus virgulatus; (186) P. alpinus; (187) Amboakis expandosa; (188) Ampleris jatai; (189) Megaphloeus jasuni; (190) Amboakis bermejo (191) Madoniella delta; (192) Megaphloeus imparilus; (193) Pyticeroides brevitubulus." figureDoi="http://doi.org/10.5281/zenodo.3746768" httpUri="https://zenodo.org/record/3746768/files/figure.png" pageId="23" pageNumber="982">Fig. 191</figureCitation>
) quadrate, coarsely punctured (PW/PL 62/62), disc deeply concave paralaterally near pronotal collar; discal and lateral trichobothria prominent; elytral asetiferous punctures striate (EL/EW 155/40); anterior margin of protibia with 4 spines. Abdomen: Pygidium scutiform; aedeagus as in
<figureCitation id="13332A0FD84DFFC23EE7FF54311FFA65" box="[708,800,238,260]" captionStart="Figs 219-224" captionStartId="86.[151,191,1278,1300]" captionTargetBox="[153,1061,198,1263]" captionTargetId="figure@86.[153,1061,198,1263]" captionTargetPageId="86" captionText="Figs 219-224: Aedeagi; (219) Amboakis mitaraka; (220) Epiphloeus mantillerii; (221) E. prolixicornus; (222) Ichnea cerinis; (223) Madoniella delta; (224) M. muralla." figureDoi="http://doi.org/10.5281/zenodo.3746778" httpUri="https://zenodo.org/record/3746778/files/figure.png" pageId="24" pageNumber="983">Fig. 223.</figureCitation>
</paragraph>
<paragraph id="8BB7368AD84DFFC23CB4FEAE32D4FA20" blockId="24.[151,1058,275,326]" pageId="24" pageNumber="983">
N a t u r a l H i s t o r y: The available specimen was collected during June, at
<quantity id="4CF09B6FD84DFFC23CB4FE8B32D9FA20" box="[151,230,305,326]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="24" pageNumber="983" unit="m" value="1300.0">1300 m</quantity>
.
</paragraph>
<paragraph id="8BB7368AD84DFFC23CB4FEED312CFA0A" blockId="24.[151,787,342,364]" box="[151,787,342,364]" pageId="24" pageNumber="983">
D i s t r i b u t i o n: This species is known from
<collectingCountry id="F31F761AD84DFFC23E87FEEC3130FA0A" box="[676,783,342,364]" name="Nicaragua" pageId="24" pageNumber="983">Nicaragua</collectingCountry>
.
</paragraph>
<paragraph id="8BB7368AD84DFFC23CB4FEC63142FAC9" blockId="24.[151,1058,379,431]" pageId="24" pageNumber="983">
E t y m o l o g y: The trivial name,
<emphasis id="B97CEA98D84DFFC23E03FEC1306CFAF6" box="[544,595,379,400]" italics="true" pageId="24" pageNumber="983">delta</emphasis>
, is the Greek capital letter Δ; with reference to the shape of the red making at the anterior-middle of the pronotal disc.
</paragraph>
</subSubSection>
</treatment>
</document>
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