530 lines
70 KiB
XML
530 lines
70 KiB
XML
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<mods:title id="D11955994CBAF0DAD83DEFCE8AF0BB1F">Morphology and multilocus phylogeny of the Spiny-footed Lizard (Acanthodactylus erythrurus) complex reveal two new mountain species from the Moroccan Atlas</mods:title>
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<mods:namePart id="AFDB5ED70C88B94F0CFE5B1BAA5FC09F">Miralles, Aurélien</mods:namePart>
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<mods:namePart id="7D71ED8493ED8CBE71B68A89352E7B98">Geniez, Philippe</mods:namePart>
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<mods:namePart id="FA7749268564E7D816789FF2FAC74C59">Beddek, Menad</mods:namePart>
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<mods:namePart id="4EB140E3DFB8A42E90C62CA15A9B6572">Aranda, Daniel Mendez</mods:namePart>
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<mods:namePart id="5AF44824BAFD0AA84219361D96C35348">Brito, José Carlos</mods:namePart>
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<mods:namePart id="480F8C340A9006F3AB6FC3E57B95B4A7">Leblois, Raphaël</mods:namePart>
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<mods:namePart id="D2BA82E5E408FBF44DD37633DC3EA453">Crochet, Pierre-André</mods:namePart>
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<mods:title id="D47B722FCB39B93A97E5A5C2645D539A">Zootaxa</mods:title>
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<mods:date id="566373730EF01CF33A0D38B528DB8312">2020</mods:date>
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<mods:identifier id="021B43DED61BC42F100F09D9A515C48F" type="DOI">10.11646/zootaxa.4747.2.4</mods:identifier>
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<treatment id="039DEF5B5F590C0362AB6040F1AEFD90" ID-DOI="http://doi.org/10.5281/zenodo.3704095" ID-GBIF-Taxon="162381194" ID-Zenodo-Dep="3704095" LSID="urn:lsid:plazi:treatment:039DEF5B5F590C0362AB6040F1AEFD90" httpUri="http://treatment.plazi.org/id/039DEF5B5F590C0362AB6040F1AEFD90" lastPageId="20" lastPageNumber="322" pageId="14" pageNumber="316">
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<subSubSection id="C32E0DC65F590C1962AB6040F295FE00" pageId="14" pageNumber="316" type="nomenclature">
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<paragraph id="8B8B5E4D5F590C1962AB6040F12EFEEC" blockId="14.[151,582,333,395]" box="[151,582,333,360]" pageId="14" pageNumber="316">
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<emphasis id="B940825F5F590C1962AB6040F12EFEEC" bold="true" box="[151,582,333,360]" pageId="14" pageNumber="316">
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<taxonomicName id="4C3425CE5F590C1962AB6040F28AFEEC" authority="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet, 2020" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[151,482,333,360]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="316" phylum="Chordata" rank="species" species="montanus" status="sp. nov.">
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<emphasis id="B940825F5F590C1962AB6040F28AFEEC" bold="true" box="[151,482,333,360]" italics="true" pageId="14" pageNumber="316">Acanthodactylus montanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2733F245F590C1963D56040F12EFEEC" box="[489,582,333,359]" pageId="14" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8B5E4D5F590C1962AB607CF295FE00" blockId="14.[151,582,333,395]" box="[151,509,369,395]" pageId="14" pageNumber="316">
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(Appendixes I, III;
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<figureCitation id="130F42C85F590C196353607CF2D6FE00" box="[367,446,369,395]" captionStart="FIGURE 6" captionStartId="15.[151,250,1657,1682]" captionTargetBox="[159,1433,202,1601]" captionTargetId="figure@15.[151,1436,196,1605]" captionTargetPageId="15" captionText="FIGURE 6. Phenotypic variation in Acanthodactylus montanus sp. nov. and A. lacrymae sp. nov. (compare with figure 6 showing the phenotypic variation in the rest of the Acanthodactylus erythrurus group): (A) BEV.11795*, (B) BEV.11793*, (C) MNHN-RA-2018.0026 (holotype of Acanthodactylus montanus), (D) BEV.11797, (E) BEV.11988, (F) BEV.11989, (G) MNHN- RA- 2018.0027 (holotype of A. lacrymae), (H) BEV.11992, (I) BEV.11998, (J) BEV.12000, (K) BEV.11997, (L) BEV.12006. Asterisks indicate symmetrically reversed pictures. Photo credits: A to D (PAC), E to L (PG)." figureDoi="http://doi.org/10.5281/zenodo.3694836" httpUri="https://zenodo.org/record/3694836/files/figure.png" pageId="14" pageNumber="316">Figs. 6</figureCitation>
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,
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<figureCitation id="130F42C85F590C1963F6607CF2B1FE00" box="[458,473,369,395]" captionStart="FIGURE 8" captionStartId="17.[151,250,1935,1960]" captionTargetBox="[183,1406,522,1913]" captionTargetId="figure@17.[179,1409,517,1916]" captionTargetPageId="17" captionText="FIGURE 8. Holotypes of Acanthodactylus montanus sp. nov. (MNHN-RA-2018.0026, Tizi n’Tichka) and (B) Acanthodactylus lacrymae sp. nov. (MNHN-RA-2018.0027, Tislit) with a picture of the habitat of this species (Isli lake). Photo credits:A (PAC), B (PG)." figureDoi="http://doi.org/10.5281/zenodo.3694844" httpUri="https://zenodo.org/record/3694844/files/figure.png" pageId="14" pageNumber="316">8</figureCitation>
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,
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<figureCitation id="130F42C85F590C1963DA607CF29DFE00" box="[486,501,369,395]" captionStart="FIGURE 9" captionStartId="18.[151,250,1859,1884]" captionTargetBox="[198,1394,184,1829]" captionTargetId="figure@18.[185,1402,174,1840]" captionTargetPageId="18" captionText="FIGURE 9. Phenotypic variation in juveniles. Pileus of juveniles in A lacrymae sp. nov. (A, BEV.11981 & B, BEV. 11990, both from Tislit) and A. montanus sp. nov. (C, JCB.A546, Jbel Siroua & D, BEV.11792, Tizi n’Tichka). Caudal and cloacal colour patterns in juveniles of A. lacrymae sp. nov. (E, BEV.11986 & F, BEV.11990, both from Tislit), A. montanus sp. nov. (G, BEV.11798, Tizi n’Tichka & H, JCB.A546, Jbel Siroua) and A. erythrurus (I, BEV.11926, Dayet Ifrah & J, BEV.11595, Marrakech). Photo credits: A, B, E, F, I (PG), D, G, J (PAC), C, H (JCB)." figureDoi="http://doi.org/10.5281/zenodo.3694846" httpUri="https://zenodo.org/record/3694846/files/figure.png" pageId="14" pageNumber="316">9</figureCitation>
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)
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</paragraph>
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<subSubSection id="C32E0DC65F590C1962AB60B4F039FB28" pageId="14" pageNumber="316" type="materials_examined">
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<paragraph id="8B8B5E4D5F590C1962AB60B4F185FDB4" blockId="14.[151,1437,441,1979]" pageId="14" pageNumber="316">
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<materialsCitation id="3B5C54105F590C1962AB60B4F185FDB4" ID-GBIF-Occurrence="2573218768" collectingDate="2012-05-12" country="Morocco" county="southwestern High Atlas" elevation="2176" latitude="31.2879" location="Museum" longLatPrecision="6" longitude="-7.3824" municipality="Marrakech" pageId="14" pageNumber="316" specimenCount="1" typeStatus="holotype">
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<typeStatus id="548FE0EF5F590C1962AB60B4F26EFE58" box="[151,262,441,467]" pageId="14" pageNumber="316" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<location id="8EEB08965F590C19632E60B4F21CFE58" LSID="urn:lsid:plazi:treatment:039DEF5B5F590C0362AB6040F1AEFD90:8EEB08965F590C19632E60B4F21CFE58" box="[274,372,441,467]" country="Morocco" county="southwestern High Atlas" latitude="31.2879" longLatPrecision="6" longitude="-7.3824" municipality="Marrakech" name="Museum" pageId="14" pageNumber="316">Muséum</location>
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national
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<location id="8EEB08965F590C1963E160B4F127FE58" LSID="urn:lsid:plazi:treatment:039DEF5B5F590C0362AB6040F1AEFD90:8EEB08965F590C1963E160B4F127FE58" box="[477,591,441,467]" country="Morocco" county="southwestern High Atlas" latitude="31.2879" longLatPrecision="6" longitude="-7.3824" municipality="Marrakech" name="d'Histoire" pageId="14" pageNumber="316">d’Histoire</location>
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naturelle (Paris) MNHN-RA-2018.0026 (formerly BEV.11794, tissue sample in the BEV tissue collection, code T5771), adult male, collected on
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May 12
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<quantity id="4CCCF3A85F590C19662860D0F727FE7C" box="[1044,1103,477,503]" metricMagnitude="6" metricUnit="m" metricValue="2.0128" pageId="14" pageNumber="316" unit="m" value="2012800.0">2012</quantity>
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</collectingDate>
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</date>
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, 800 m after the
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<location id="8EEB08965F590C19673A60D0F6F4FE7C" LSID="urn:lsid:plazi:treatment:039DEF5B5F590C0362AB6040F1AEFD90:8EEB08965F590C19673A60D0F6F4FE7C" box="[1286,1436,477,503]" country="Morocco" county="southwestern High Atlas" latitude="31.2879" longLatPrecision="6" longitude="-7.3824" municipality="Marrakech" name="Tizi n'Tichka" pageId="14" pageNumber="316">Tizi n’Tichka</location>
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pass towards
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,
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<collectingCounty id="62EA26C15F590C196380630CF1BEFD90" box="[444,726,513,539]" pageId="14" pageNumber="316">southwestern High Atlas</collectingCounty>
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,
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<collectingCountry id="F3231EDD5F590C1960D8630CF024FD90" box="[740,844,513,539]" name="Morocco" pageId="14" pageNumber="316">Morocco</collectingCountry>
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,
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,
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(WGS84),
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a.s.l.
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</elevation>
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by Pierre-André Crochet and Raphaël Leblois (
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<figureCitation id="130F42C85F590C1960B96328F1B7FDB4" box="[645,735,549,575]" captionStart="FIGURE 8" captionStartId="17.[151,250,1935,1960]" captionTargetBox="[183,1406,522,1913]" captionTargetId="figure@17.[179,1409,517,1916]" captionTargetPageId="17" captionText="FIGURE 8. Holotypes of Acanthodactylus montanus sp. nov. (MNHN-RA-2018.0026, Tizi n’Tichka) and (B) Acanthodactylus lacrymae sp. nov. (MNHN-RA-2018.0027, Tislit) with a picture of the habitat of this species (Isli lake). Photo credits:A (PAC), B (PG)." figureDoi="http://doi.org/10.5281/zenodo.3694844" httpUri="https://zenodo.org/record/3694844/files/figure.png" pageId="14" pageNumber="316">Fig. 8A</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8B8B5E4D5F590C1962FB6344F08DFD0C" blockId="14.[151,1437,441,1979]" pageId="14" pageNumber="316">
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<typeStatus id="548FE0EF5F590C1962FB6344F229FDE8" box="[199,321,585,611]" pageId="14" pageNumber="316" type="paratype">Paratypes</typeStatus>
|
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.
|
||
</emphasis>
|
||
Nine individuals, BEV.11791 to 11793, 11795 to 11800 (tissue sample codes T5768–5770, 5772– 5777), collected at the type locality at the same date by the same collectors.
|
||
</paragraph>
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<paragraph id="8B8B5E4D5F590C1962FB639DF1E7FC9C" blockId="14.[151,1437,441,1979]" pageId="14" pageNumber="316">
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<emphasis id="B940825F5F590C1962FB639DF2D2FD20" bold="true" box="[199,442,656,683]" pageId="14" pageNumber="316">Additional material.</emphasis>
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||
Eight specimens from El Azib n-Iriri, Jbel Siroua,
|
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<geoCoordinate id="EE00388A5F590C1961CB639CF71CFD20" box="[1015,1140,657,683]" degrees="30.7470" direction="north" orientation="latitude" pageId="14" pageNumber="316" precision="5" value="30.747">30.7470°N</geoCoordinate>
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,
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,
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<quantity id="4CCCF3A85F590C1966C2639CF632FD20" box="[1278,1370,657,683]" metricMagnitude="3" metricUnit="m" metricValue="2.319" pageId="14" pageNumber="316" unit="m" value="2319.0">2,319 m</quantity>
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a.s.l., collected on
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<date id="FF8A788D5F590C19631963B8F2B8FD44" box="[293,464,692,719]" pageId="14" pageNumber="316" value="2005-04-23">
|
||
April 23
|
||
<superScript id="7C41F3055F590C1963BF63B9F2F9FD49" attach="left" box="[387,401,692,706]" fontSize="6" pageId="14" pageNumber="316">rd</superScript>
|
||
2005
|
||
</date>
|
||
by José C. Brito: BEV.10459–10460 (field number JCB.A544 & A547, tissue samples T11829 & T11834) and tissue samples + photos T11826 (JCB.A541),
|
||
<materialsCitation id="3B5C54105F590C1961F863D4F710FD78" ID-GBIF-Occurrence="2573218761" box="[964,1144,728,755]" pageId="14" pageNumber="316" specimenCount="1">T11827 (A542)</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B5C54105F590C1966B963D4F650FD78" ID-GBIF-Occurrence="2573218762" box="[1157,1336,728,755]" pageId="14" pageNumber="316" specimenCount="1">T11828 (A543)</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B5C54105F590C19677A63D4F382FC9C" ID-GBIF-Occurrence="2573218764" pageId="14" pageNumber="316" specimenCount="1">T11830 (A545)</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B5C54105F590C1962CA63F0F2CCFC9C" ID-GBIF-Occurrence="2573218766" box="[246,420,764,791]" pageId="14" pageNumber="316" specimenCount="1">T11831 (A546)</materialsCitation>
|
||
<materialsCitation id="3B5C54105F590C19639663F0F158FC9C" ID-GBIF-Occurrence="2573218767" box="[426,560,765,791]" pageId="14" pageNumber="316" specimenCount="1">and T11833</materialsCitation>
|
||
(A458).
|
||
</paragraph>
|
||
<paragraph id="8B8B5E4D5F590C1962FB622CF2F6FC40" blockId="14.[151,1437,441,1979]" pageId="14" pageNumber="316">
|
||
We identified the following specimens as
|
||
<taxonomicName id="4C3425CE5F590C196099622CF05BFCB0" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[677,819,801,827]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="316" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F590C196099622CF05BFCB0" box="[677,819,801,827]" italics="true" pageId="14" pageNumber="316">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
on the basis of morphology, distribution and ecology: BEV.4973 from
|
||
<quantity id="4CCCF3A85F590C1963716249F2EDFCD4" box="[333,389,836,863]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="316" unit="km" value="5.0">5 km</quantity>
|
||
south of Askaoun, Jbel Siroua,
|
||
<geoCoordinate id="EE00388A5F590C1960DF6248F02CFCD4" box="[739,836,837,863]" degrees="30.69" direction="north" orientation="latitude" pageId="14" pageNumber="316" precision="555" value="30.69">30.69°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE00388A5F590C1961736248F0CEFCD4" box="[847,934,837,863]" degrees="7.77" direction="west" orientation="longitude" pageId="14" pageNumber="316" precision="555" value="-7.77">7.77°W</geoCoordinate>
|
||
, collected on
|
||
<date id="FF8A788D5F590C1966016248F79EFCD4" box="[1085,1270,836,863]" pageId="14" pageNumber="316" value="1974-10-09">
|
||
October 9
|
||
<superScript id="7C41F3055F590C1966906249F7D1FCD9" attach="left" box="[1196,1209,836,850]" fontSize="6" pageId="14" pageNumber="316">th</superScript>
|
||
1974
|
||
</date>
|
||
by Gérard Délye; BEV.4974 from
|
||
<quantity id="4CCCF3A85F590C1963436264F2DFFC08" box="[383,439,873,899]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="316" unit="km" value="3.0">3 km</quantity>
|
||
west of Askaoun, Jbel Siroua,
|
||
<geoCoordinate id="EE00388A5F590C1961336264F018FC08" box="[783,880,873,899]" degrees="30.74" direction="north" orientation="latitude" pageId="14" pageNumber="316" precision="555" value="30.74">30.74°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE00388A5F590C1961476264F0BAFC08" box="[891,978,873,899]" degrees="7.81" direction="west" orientation="longitude" pageId="14" pageNumber="316" precision="555" value="-7.81">7.81°W</geoCoordinate>
|
||
, collected on
|
||
<date id="FF8A788D5F590C1966576264F64DFC08" box="[1131,1317,872,899]" pageId="14" pageNumber="316" value="1974-10-08">
|
||
October 8
|
||
<superScript id="7C41F3055F590C1966E66265F78FFCFD" attach="left" box="[1242,1255,872,886]" fontSize="6" pageId="14" pageNumber="316">th</superScript>
|
||
1974
|
||
</date>
|
||
by Gérard Délye; BEV.4975 from Tisfaldat mountain, Jbel Siroua,
|
||
<geoCoordinate id="EE00388A5F590C19612A6280F010FC2C" box="[790,888,909,935]" degrees="30.68" direction="north" orientation="latitude" pageId="14" pageNumber="316" precision="555" value="30.68">30.68°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE00388A5F590C1961B86281F0B3FC2C" box="[900,987,908,935]" degrees="7.58" direction="west" orientation="longitude" pageId="14" pageNumber="316" precision="555" value="-7.58">7.58°W</geoCoordinate>
|
||
,
|
||
<quantity id="4CCCF3A85F590C1961DB6280F72DFC2C" box="[999,1093,909,935]" metricMagnitude="3" metricUnit="m" metricValue="2.46" pageId="14" pageNumber="316" unit="m" value="2460.0">2,460 m</quantity>
|
||
a.s.l., collected on
|
||
<date id="FF8A788D5F590C19671E6280F3A7FC40" pageId="14" pageNumber="316" value="1994-03-13">
|
||
March 13
|
||
<superScript id="7C41F3055F590C1967AC6281F6F5FC11" attach="none" box="[1424,1437,908,922]" fontSize="6" pageId="14" pageNumber="316">th</superScript>
|
||
1994
|
||
</date>
|
||
by Tahar Slimani.
|
||
</paragraph>
|
||
<paragraph id="8B8B5E4D5F590C1962FB62D8F039FB28" blockId="14.[151,1437,441,1979]" pageId="14" pageNumber="316">
|
||
Lastly, a series of specimens exhibit morphological features of
|
||
<taxonomicName id="4C3425CE5F590C1961B262D8F772FC64" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[910,1050,981,1007]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="316" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F590C1961B262D8F772FC64" box="[910,1050,981,1007]" italics="true" pageId="14" pageNumber="316">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
but were apparently collected at a lower elevation than other specimens (around
|
||
<quantity id="4CCCF3A85F590C1960A962F4F198FB98" box="[661,752,1017,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="316" unit="m" value="1800.0">1,800 m</quantity>
|
||
a.s.l.) and slightly south of the main massif of the Jbel Siroua. They are tentatively identified as
|
||
<taxonomicName id="4C3425CE5F590C1960336510F1F3FBBC" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[527,667,1053,1079]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="316" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F590C1960336510F1F3FBBC" box="[527,667,1053,1079]" italics="true" pageId="14" pageNumber="316">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
here (white square in
|
||
<figureCitation id="130F42C85F590C1961A86510F08CFBBC" box="[916,996,1053,1079]" captionStart="FIGURE 10" captionStartId="19.[151,250,1622,1647]" captionTargetBox="[206,1380,802,1609]" captionTargetId="figure@19.[200,1387,798,1614]" captionTargetPageId="19" captionText="FIGURE 10. Distribution of Acanthodactylus montanus (squares) and Acanthodactylus lacrymae (diamonds) based on specimens examined in this study. Coloured dots represent specimens of the three main phenotypes (none is monophyletic in the mtDNA or nDNA trees) in Morocco based on Bons and Geniez 1995. Specimens with the “bellii” phenotype in the High Atlas are almost certainly montanus and lacrymae. White symbols represent specimens tentatively identified as A. montanus (square) or A. lacrymae (diamond, see text). The continuous white lines represent the currently known distribution of the new species while dotted white lines represent possible extensions of the distribution if tentative identifications are correct. The dotted dark blue line represents the southernmost extension of confirmed IM clade specimens based on our field sampling." figureDoi="http://doi.org/10.5281/zenodo.3694848" httpUri="https://zenodo.org/record/3694848/files/figure.png" pageId="14" pageNumber="316">Fig. 10</figureCitation>
|
||
) but this identification will require further data and possibly genetic confirmation before it can be firmly established: BEV.4932–4941 and 14898 from
|
||
<quantity id="4CCCF3A85F590C1967BC654CF3D3FBF4" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="316" unit="km" value="40.0">40 km</quantity>
|
||
past Tazenakht along road to Taroudant, approx.
|
||
<geoCoordinate id="EE00388A5F590C1960D66569F031FBF4" box="[746,857,1124,1151]" degrees="30.455" direction="north" orientation="latitude" pageId="14" pageNumber="316" precision="55" value="30.455">30.455°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE00388A5F590C1961596569F0A2FBF4" box="[869,970,1124,1151]" degrees="7.558" direction="west" orientation="longitude" pageId="14" pageNumber="316" precision="55" value="-7.558">7.558°W</geoCoordinate>
|
||
, around
|
||
<quantity id="4CCCF3A85F590C1966176568F7E0FBF4" box="[1067,1160,1125,1151]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="316" unit="m" value="1800.0">1,800 m</quantity>
|
||
a.s.l., collected on
|
||
<date id="FF8A788D5F590C19675D6568F399FB28" pageId="14" pageNumber="316" value="1960-04-05">
|
||
April 5
|
||
<superScript id="7C41F3055F590C1962996585F3DAFB1D" attach="left" box="[165,178,1160,1174]" fontSize="6" pageId="14" pageNumber="316">th</superScript>
|
||
1960
|
||
</date>
|
||
by Jacques Bons, Georges Pasteur and Bernard Girot.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E0DC65F590C1962FB65A0F260FA84" pageId="14" pageNumber="316" type="etymology">
|
||
<paragraph id="8B8B5E4D5F590C1962FB65A0F260FA84" blockId="14.[151,1437,441,1979]" pageId="14" pageNumber="316">
|
||
<emphasis id="B940825F5F590C1962FB65A0F226FB4C" bold="true" box="[199,334,1197,1223]" pageId="14" pageNumber="316">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="4C3425CE5F590C19607E65A3F1C7FB4C" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[578,687,1198,1223]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="316" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F590C19607E65A3F1C7FB4C" box="[578,687,1198,1223]" italics="true" pageId="14" pageNumber="316">montanus</emphasis>
|
||
</taxonomicName>
|
||
is a Latin adjective meaning “of mountains, belonging to mountains”, the high mountains of the Western High Atlas being the unique habitat where this species has been found until now.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E0DC65F590C0662FB6415F278FE7C" lastPageId="17" lastPageNumber="319" pageId="14" pageNumber="316" type="diagnosis">
|
||
<paragraph id="8B8B5E4D5F590C1962FB6415F18FF830" blockId="14.[151,1437,441,1979]" pageId="14" pageNumber="316">
|
||
<emphasis id="B940825F5F590C1962FB6415F22AFAB8" bold="true" box="[199,322,1304,1331]" pageId="14" pageNumber="316">Diagnosis.</emphasis>
|
||
A new species of the
|
||
<taxonomicName id="4C3425CE5F590C1960116414F00BFAB8" baseAuthorityName="Schinz" baseAuthorityYear="1833" box="[557,867,1305,1331]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="316" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="B940825F5F590C1960116414F00BFAB8" box="[557,867,1305,1331]" italics="true" pageId="14" pageNumber="316">Acanthodactylus erythrurus</emphasis>
|
||
</taxonomicName>
|
||
species-group (small flat or carinated dorsal scales; three series of scales around the fingers; three entire supraoculars; 8–10, sometimes 12 straight longitudinal row of ventrals; slightly pectinate toes; undertail and underside of the hind limbs red or reddish in juveniles, subadults and young adults) from the south-western part of the High Atlas and from the Jbel Siroua above
|
||
<quantity id="4CCCF3A85F590C1966986488F668FA14" box="[1188,1280,1413,1439]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="316" unit="m" value="2000.0">2,000 m</quantity>
|
||
a.s.l., characterized by the combination of the following characters: (1) head scalation usually of the
|
||
<emphasis id="B940825F5F590C19664E64A4F7A3FA48" box="[1138,1227,1449,1475]" italics="true" pageId="14" pageNumber="316">
|
||
“
|
||
<taxonomicName id="4C3425CE5F590C19664264A4F7A8FA48" authorityName="Gray" authorityYear="1845" box="[1150,1216,1449,1475]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="316" phylum="Chordata" rank="species" species="bellii">bellii</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
<typeStatus id="548FE0EF5F590C1966EE64A4F66AFA48" box="[1234,1282,1449,1475]" pageId="14" pageNumber="316">type</typeStatus>
|
||
(subocular in large contact with the upper lip, wedged between the 4
|
||
<superScript id="7C41F3055F590C1960C864C1F069FA51" attach="left" box="[756,769,1484,1498]" fontSize="6" pageId="14" pageNumber="316">th</superScript>
|
||
and 5
|
||
<superScript id="7C41F3055F590C19617864C1F039FA51" attach="left" box="[836,849,1484,1498]" fontSize="6" pageId="14" pageNumber="316">th</superScript>
|
||
supralabials), one internasal plate, no scales inserted between the prefrontals, but BEV.T11828 has no contact between the left supralabial and the lip; (2) a low number of dorsal scales (47–59 longitudinal rows around the body, mean 54.0), femoral pores (17–23, mean 19.9) on each side and subdigital lamellae (19–22, mean 19.9) on the 4
|
||
<superScript id="7C41F3055F590C1961306735F071F9CD" attach="left" box="[780,793,1592,1606]" fontSize="6" pageId="14" pageNumber="316">th</superScript>
|
||
toe; (3) one or one and a half row of supraciliary granules on each side with a very reduced number (15–46, mean 25.7) of scales and granules around the 2
|
||
<superScript id="7C41F3055F590C1966FB6751F7BFF9E1" attach="left" box="[1223,1239,1628,1642]" fontSize="6" pageId="14" pageNumber="316">nd</superScript>
|
||
and 3
|
||
<superScript id="7C41F3055F590C1967266751F640F9E1" attach="left" box="[1306,1320,1628,1642]" fontSize="6" pageId="14" pageNumber="316">rd</superScript>
|
||
supraoculars on each side; (4) temporals smooth or slightly keeled; (5) dorsal scales smooth or slightly keeled on the neck and progressively turning to distinctly tectiform or sometimes keeled on the back; (6) stocky proportions, especially in adults, with a relatively short tail (57.9–65.9% of the total length, mean 60.8%, almost always longer for all the other
|
||
<taxonomicName id="4C3425CE5F590C1962E467E0F2FBF88C" box="[216,403,1773,1799]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="316" phylum="Chordata" rank="genus">
|
||
<emphasis id="B940825F5F590C1962E467E0F2FBF88C" box="[216,403,1773,1799]" italics="true" pageId="14" pageNumber="316">Acanthodactylus</emphasis>
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="4C3425CE5F590C1963D867E0F132F88C" baseAuthorityName="Schinz" baseAuthorityYear="1833" box="[484,602,1773,1799]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="316" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="B940825F5F590C1963D867E0F132F88C" box="[484,602,1773,1799]" italics="true" pageId="14" pageNumber="316">erythrurus</emphasis>
|
||
</taxonomicName>
|
||
group except
|
||
<taxonomicName id="4C3425CE5F590C1960C067E0F00CF88C" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[764,868,1773,1799]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="316" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F590C1960C067E0F00CF88C" box="[764,868,1773,1799]" italics="true" pageId="14" pageNumber="316">lacrymae</emphasis>
|
||
</taxonomicName>
|
||
); (7) base of the tail of adult males very thick; (8) in juveniles, underparts of the tail orange red to intense coral red, slightly paler under the base of the tail, even paler under the rear of the thighs, this colour usually not reaching the rows of femoral pores; (9) some marginal ventral plates yellow of yellowish in adult and subadult males and females, sometimes in juveniles; (10) a series of small round ocelli on the flanks (each one containing 4–9 brightly coloured scales), usually yellow greenish in adults, which tend to disappear completely in adult females.
|
||
</paragraph>
|
||
<caption id="DF4B0EC55F580C1862AB6774F72FF8A9" ID-DOI="http://doi.org/10.5281/zenodo.3694836" ID-Zenodo-Dep="3694836" httpUri="https://zenodo.org/record/3694836/files/figure.png" pageId="15" pageNumber="317" startId="15.[151,250,1657,1682]" targetBox="[159,1433,202,1601]" targetPageId="15">
|
||
<paragraph id="8B8B5E4D5F580C1862AB6774F72FF8A9" blockId="15.[151,1437,1657,1826]" pageId="15" pageNumber="317">
|
||
<emphasis id="B940825F5F580C1862AB6774F27EF919" bold="true" box="[151,278,1657,1682]" pageId="15" pageNumber="317">FIGURE 6.</emphasis>
|
||
Phenotypic variation in
|
||
<taxonomicName id="4C3425CE5F580C1860206777F059F919" authority="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet, 2020" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[540,817,1658,1682]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="317" phylum="Chordata" rank="species" species="montanus" status="sp. nov.">
|
||
<emphasis id="B940825F5F580C1860206777F059F919" box="[540,817,1658,1682]" italics="true" pageId="15" pageNumber="317">Acanthodactylus montanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B940825F5F580C1861066777F0E6F919" bold="true" box="[826,910,1658,1682]" pageId="15" pageNumber="317">
|
||
<taxonomicNameLabel id="A2733F245F580C1861066777F0E6F919" box="[826,910,1658,1682]" pageId="15" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C3425CE5F580C1861FA6777F72BF919" authority="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet, 2020" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[966,1091,1658,1682]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="317" phylum="Chordata" rank="species" species="lacrymae" status="sp. nov.">
|
||
<emphasis id="B940825F5F580C1861FA6777F72BF919" box="[966,1091,1658,1682]" italics="true" pageId="15" pageNumber="317">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B940825F5F580C1866706777F7C8F919" bold="true" box="[1100,1184,1658,1682]" pageId="15" pageNumber="317">
|
||
<taxonomicNameLabel id="A2733F245F580C1866706777F7C8F919" box="[1100,1184,1658,1682]" pageId="15" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(compare with figure 6 showing the phenotypic variation in the rest of the
|
||
<taxonomicName id="4C3425CE5F580C1860A06793F0DEF93D" baseAuthorityName="Schinz" baseAuthorityYear="1833" box="[668,950,1694,1718]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="317" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="B940825F5F580C1860A06793F0DEF93D" box="[668,950,1694,1718]" italics="true" pageId="15" pageNumber="317">Acanthodactylus erythrurus</emphasis>
|
||
</taxonomicName>
|
||
group): (A) BEV.11795*, (B) BEV.11793*, (C) MNHN-RA-2018.0026 (holotype of
|
||
<taxonomicName id="4C3425CE5F580C18603667CFF073F951" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[522,795,1730,1754]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="317" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F580C18603667CFF073F951" box="[522,795,1730,1754]" italics="true" pageId="15" pageNumber="317">Acanthodactylus montanus</emphasis>
|
||
</taxonomicName>
|
||
), (D) BEV.11797, (E) BEV.11988, (F) BEV.11989, (G) MNHN- RA- 2018.0027 (holotype of
|
||
<taxonomicName id="4C3425CE5F580C1863F867EBF157F975" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[452,575,1766,1790]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="317" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F580C1863F867EBF157F975" box="[452,575,1766,1790]" italics="true" pageId="15" pageNumber="317">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
), (H) BEV.11992, (I) BEV.11998, (J) BEV.12000, (K) BEV.11997, (L) BEV.12006. Asterisks indicate symmetrically reversed pictures. Photo credits: A to D (PAC), E to L (PG).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8B5E4D5F580C0762FB665DF67FFEEC" blockId="15.[151,1436,1872,2043]" lastBlockId="16.[151,1437,153,359]" lastPageId="16" lastPageNumber="318" pageId="15" pageNumber="317">
|
||
Body coloration similar to the other members of the
|
||
<taxonomicName id="4C3425CE5F580C186133665DF72EF8E1" baseAuthorityName="Schinz" baseAuthorityYear="1833" box="[783,1094,1872,1898]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="317" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="B940825F5F580C186133665DF72EF8E1" box="[783,1094,1872,1898]" italics="true" pageId="15" pageNumber="317">Acanthodactylus erythrurus</emphasis>
|
||
</taxonomicName>
|
||
group. Juveniles are very dark (black or blackish) with three pale continuous lines on each side of the body (one on lower flanks, one at the junction between the dorsum and the flanks and one on the dorsum joining with the opposite one on the tail base) and a dark vertebral area dissected by a paler narrow vertebral area, frequently split on the nape by dark spots or a dark line and disappearing before the base of the tail; one or two series of pale ocelli are found in the dark interspaces between these stripes (juveniles of the other lineages are usually paler and lack the bifurcation of the pale central vertebral area on the nape). Adults are medium brown with faint pale lines on the dorsum and upper flanks (corresponding to the juvenile pattern) and a better marked line on the lower flanks; irregular dark marks are present on the back between the pale lines, sometimes invading the area on the centre of the dorsum. The flanks typically appear darker due to the large dark areas surrounding each pale ocellus. The pileus presents a complex pattern of pale and dark reticulations in juvenile; in adults it is of the same colour as the back with isolated irregular dark spots.
|
||
</paragraph>
|
||
<caption id="DF4B0EC55F470C0762AB6624F188F859" ID-DOI="http://doi.org/10.5281/zenodo.3694840" ID-Zenodo-Dep="3694840" httpUri="https://zenodo.org/record/3694840/files/figure.png" pageId="16" pageNumber="318" startId="16.[151,250,1833,1858]" targetBox="[157,1435,382,1805]" targetPageId="16">
|
||
<paragraph id="8B8B5E4D5F470C0762AB6624F188F859" blockId="16.[151,1437,1833,2002]" pageId="16" pageNumber="318">
|
||
<emphasis id="B940825F5F470C0762AB6624F27BF8C9" bold="true" box="[151,275,1833,1858]" pageId="16" pageNumber="318">FIGURE 7.</emphasis>
|
||
Phenotypic variation in the rest of the
|
||
<taxonomicName id="4C3425CE5F470C0760A66627F0DCF8C9" baseAuthorityName="Schinz" baseAuthorityYear="1833" box="[666,948,1834,1858]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="318" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="B940825F5F470C0760A66627F0DCF8C9" box="[666,948,1834,1858]" italics="true" pageId="16" pageNumber="318">Acanthodactylus erythrurus</emphasis>
|
||
</taxonomicName>
|
||
group (IM, CA and AT clades). (A) BEV.11830, (B) BEV.11833, (C) BEV.11619*, (D) BEV.11615, (E) BEV.11968, (F) BEV.11972, (G) BEV.11845*, (H) BEV.11849*, (I) BEV.11826*, (J) BEV.11825*, (K) PGe.1539, (L) PGe.1538, (M) BEV.T4232* from Senalba, and (N) uncatalogued specimen* from Theniet.Asterisks indicate symmetrically reversed pictures. Photo credits: A to D (PAC), E to F (PG), G to J (AM), K to L (Jean-Michel Bompar), M (Vincent Rivière) and N (MB).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8B5E4D5F460C0662FB6194F278FE7C" blockId="17.[151,1437,153,503]" pageId="17" pageNumber="319">
|
||
Very similar to the allopatric
|
||
<taxonomicName id="4C3425CE5F460C0660316194F05FFF38" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[525,823,153,179]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="319" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F460C0660316194F05FFF38" box="[525,823,153,179]" italics="true" pageId="17" pageNumber="319">Acanthodactylus lacrymae</emphasis>
|
||
</taxonomicName>
|
||
and single individuals are not always possible to separate but differs on average by the following characters: (1) in juveniles, parietals are less clearly striped but exhibit a more reticulated pattern; (2) orange red coloration on the undertail fading less toward tail base and more extensive around the cloacal slit; (3) supralabials usually with diffuse dark vertical stripes (uniformly pale creamy, even in juveniles, in
|
||
<taxonomicName id="4C3425CE5F460C0663766024F2BBFEC8" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[330,467,297,323]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="319" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F460C0663766024F2BBFEC8" box="[330,467,297,323]" italics="true" pageId="17" pageNumber="319">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
); (4) contrast between the lower border of the flanks and tail side and under surface of the belly and tail usually less marked in adults, especially along the sides of the tail; (5) temporal less flat and dorsal scales slightly killed in adults (smooth in
|
||
<taxonomicName id="4C3425CE5F460C066081607CF02DFE00" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[701,837,369,395]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="319" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F460C066081607CF02DFE00" box="[701,837,369,395]" italics="true" pageId="17" pageNumber="319">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
except sometimes on the rear of the back); (6) some specimens have one and a half row of supraciliary granules on each side (always one row in
|
||
<taxonomicName id="4C3425CE5F460C06669F6098F643FE24" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[1187,1323,405,431]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="319" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F460C06669F6098F643FE24" box="[1187,1323,405,431]" italics="true" pageId="17" pageNumber="319">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
) and with a larger number of scales and granules around the 2
|
||
<superScript id="7C41F3055F460C0660EA60B5F18EFE4D" attach="left" box="[726,742,440,454]" fontSize="6" pageId="17" pageNumber="319">nd</superScript>
|
||
and 3
|
||
<superScript id="7C41F3055F460C06611660B5F050FE4D" attach="left" box="[810,824,440,454]" fontSize="6" pageId="17" pageNumber="319">rd</superScript>
|
||
supraoculars (maximum 46 against maximum
|
||
<quantity id="4CCCF3A85F460C06677960B4F616FE58" box="[1349,1406,441,467]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" pageId="17" pageNumber="319" unit="in" value="29.0">29 in</quantity>
|
||
<taxonomicName id="4C3425CE5F460C0667B860B4F397FE7C" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="319" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F460C0667B860B4F397FE7C" italics="true" pageId="17" pageNumber="319">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4B0EC55F460C0662AB6682F38FF87B" ID-DOI="http://doi.org/10.5281/zenodo.3694844" ID-Zenodo-Dep="3694844" httpUri="https://zenodo.org/record/3694844/files/figure.png" pageId="17" pageNumber="319" startId="17.[151,250,1935,1960]" targetBox="[183,1406,522,1913]" targetPageId="17">
|
||
<paragraph id="8B8B5E4D5F460C0662AB6682F38FF87B" blockId="17.[151,1436,1935,2032]" pageId="17" pageNumber="319">
|
||
<emphasis id="B940825F5F460C0662AB6682F27AF823" bold="true" box="[151,274,1935,1960]" pageId="17" pageNumber="319">FIGURE 8.</emphasis>
|
||
Holotypes of
|
||
<taxonomicName id="4C3425CE5F460C06639C669DF1D9F82C" authority="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet, 2020" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[416,689,1936,1960]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="319" phylum="Chordata" rank="species" species="montanus" status="sp. nov.">
|
||
<emphasis id="B940825F5F460C06639C669DF1D9F82C" box="[416,689,1936,1960]" italics="true" pageId="17" pageNumber="319">Acanthodactylus montanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B940825F5F460C06608A669DF06EF823" bold="true" box="[694,774,1936,1960]" pageId="17" pageNumber="319">
|
||
<taxonomicNameLabel id="A2733F245F460C06608A669DF06EF823" box="[694,774,1936,1960]" pageId="17" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(MNHN-RA-2018.0026, Tizi n’Tichka) and (B)
|
||
<taxonomicName id="4C3425CE5F460C0666CF669DF39DF847" authority="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet, 2020" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="319" phylum="Chordata" rank="species" species="lacrymae" status="sp. nov.">
|
||
<emphasis id="B940825F5F460C0666CF669DF39DF847" italics="true" pageId="17" pageNumber="319">Acanthodactylus lacrymae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B940825F5F460C0662C766B9F223F847" bold="true" box="[251,331,1972,1996]" pageId="17" pageNumber="319">
|
||
<taxonomicNameLabel id="A2733F245F460C0662C766B9F223F847" box="[251,331,1972,1996]" pageId="17" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(MNHN-RA-2018.0027, Tislit) with a picture of the habitat of this species (Isli lake). Photo credits: A (PAC), B (PG).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4B0EC55F450C0562AB664EF03BF867" ID-DOI="http://doi.org/10.5281/zenodo.3694846" ID-Zenodo-Dep="3694846" httpUri="https://zenodo.org/record/3694846/files/figure.png" pageId="18" pageNumber="320" startId="18.[151,250,1859,1884]" targetBox="[198,1394,184,1829]" targetPageId="18">
|
||
<paragraph id="8B8B5E4D5F450C0562AB664EF03BF867" blockId="18.[151,1436,1859,2028]" pageId="18" pageNumber="320">
|
||
<emphasis id="B940825F5F450C0562AB664EF27DF8D7" bold="true" box="[151,277,1859,1884]" pageId="18" pageNumber="320">FIGURE 9.</emphasis>
|
||
Phenotypic variation in juveniles. Pileus of juveniles in
|
||
<taxonomicName id="4C3425CE5F450C05615D6648F0BEF8D7" authority="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet, 2020" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[865,982,1860,1884]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="320" phylum="Chordata" rank="species" species="lacrymae" status="sp. nov.">
|
||
<emphasis id="B940825F5F450C05615D6648F0BEF8D7" box="[865,982,1860,1884]" italics="true" pageId="18" pageNumber="320">A lacrymae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B940825F5F450C0561E36649F759F8D7" bold="true" box="[991,1073,1860,1884]" pageId="18" pageNumber="320">
|
||
<taxonomicNameLabel id="A2733F245F450C0561E36649F759F8D7" box="[991,1073,1860,1884]" pageId="18" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A, BEV.11981 & B, BEV. 11990, both from Tislit) and
|
||
<taxonomicName id="4C3425CE5F450C05634B6664F290F80B" authority="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet, 2020" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[375,504,1897,1920]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="320" phylum="Chordata" rank="species" species="montanus" status="sp. nov.">
|
||
<emphasis id="B940825F5F450C05634B6664F290F80B" box="[375,504,1897,1920]" italics="true" pageId="18" pageNumber="320">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B940825F5F450C05603C6665F13BF80B" bold="true" box="[512,595,1896,1920]" pageId="18" pageNumber="320">
|
||
<taxonomicNameLabel id="A2733F245F450C05603C6665F13BF80B" box="[512,595,1896,1920]" pageId="18" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(C, JCB.A546, Jbel Siroua & D, BEV.11792, Tizi n’Tichka). Caudal and cloacal colour patterns in juveniles of
|
||
<taxonomicName id="4C3425CE5F450C0563F56680F12BF82F" authority="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet, 2020" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[457,579,1932,1956]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="320" phylum="Chordata" rank="species" species="lacrymae" status="sp. nov.">
|
||
<emphasis id="B940825F5F450C0563F56680F12BF82F" box="[457,579,1932,1956]" italics="true" pageId="18" pageNumber="320">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B940825F5F450C0560756681F1F1F82F" bold="true" box="[585,665,1932,1956]" pageId="18" pageNumber="320">
|
||
<taxonomicNameLabel id="A2733F245F450C0560756681F1F1F82F" box="[585,665,1932,1956]" pageId="18" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(E, BEV.11986 & F, BEV.11990, both from Tislit),
|
||
<taxonomicName id="4C3425CE5F450C05669D6680F648F82F" authority="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet, 2020" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[1185,1312,1933,1956]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="320" phylum="Chordata" rank="species" species="montanus" status="sp. nov.">
|
||
<emphasis id="B940825F5F450C05669D6680F648F82F" box="[1185,1312,1933,1956]" italics="true" pageId="18" pageNumber="320">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B940825F5F450C0567196681F61DF82F" bold="true" box="[1317,1397,1932,1956]" pageId="18" pageNumber="320">
|
||
<taxonomicNameLabel id="A2733F245F450C0567196681F61DF82F" box="[1317,1397,1932,1956]" pageId="18" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(G, BEV.11798, Tizi n’Tichka & H, JCB.A546, Jbel Siroua) and
|
||
<taxonomicName id="4C3425CE5F450C05613666BCF0FAF843" baseAuthorityName="Schinz" baseAuthorityYear="1833" box="[778,914,1968,1992]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="320" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="B940825F5F450C05613666BCF0FAF843" box="[778,914,1968,1992]" italics="true" pageId="18" pageNumber="320">A. erythrurus</emphasis>
|
||
</taxonomicName>
|
||
(I, BEV.11926, Dayet Ifrah & J, BEV.11595, Marrakech). Photo credits: A, B, E, F, I (PG), D, G, J (PAC), C, H (JCB).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32E0DC65F440C0462FB6195F186FC9C" pageId="19" pageNumber="321" type="description">
|
||
<paragraph id="8B8B5E4D5F440C0462FB6195F614FE58" blockId="19.[151,1437,152,791]" pageId="19" pageNumber="321">
|
||
<emphasis id="B940825F5F440C0462FB6195F17DFF38" bold="true" box="[199,533,152,179]" pageId="19" pageNumber="321">
|
||
Description of the
|
||
<typeStatus id="548FE0EF5F440C0463946194F179FF38" box="[424,529,153,179]" pageId="19" pageNumber="321" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
An adult male (
|
||
<figureCitation id="130F42C85F440C0460F36194F040FF38" box="[719,808,153,179]" captionStart="FIGURE 8" captionStartId="17.[151,250,1935,1960]" captionTargetBox="[183,1406,522,1913]" captionTargetId="figure@17.[179,1409,517,1916]" captionTargetPageId="17" captionText="FIGURE 8. Holotypes of Acanthodactylus montanus sp. nov. (MNHN-RA-2018.0026, Tizi n’Tichka) and (B) Acanthodactylus lacrymae sp. nov. (MNHN-RA-2018.0027, Tislit) with a picture of the habitat of this species (Isli lake). Photo credits:A (PAC), B (PG)." figureDoi="http://doi.org/10.5281/zenodo.3694844" httpUri="https://zenodo.org/record/3694844/files/figure.png" pageId="19" pageNumber="321">Fig. 8A</figureCitation>
|
||
) measuring 60.0 mm of snout-vent length,
|
||
<quantity id="4CCCF3A85F440C04672B6194F615FF38" box="[1303,1405,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="19" pageNumber="321" unit="mm" value="14.3">14.3 mm</quantity>
|
||
of pileus length,
|
||
<quantity id="4CCCF3A85F440C04630B61B0F2E7FF5C" box="[311,399,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="19" pageNumber="321" unit="mm" value="7.7">7.7 mm</quantity>
|
||
of pileus width,
|
||
<quantity id="4CCCF3A85F440C04606C61B0F1CFFF5C" box="[592,679,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="19" pageNumber="321" unit="mm" value="8.4">8.4 mm</quantity>
|
||
of head height,
|
||
<quantity id="4CCCF3A85F440C04615D61B1F0AEFF5C" box="[865,966,188,215]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="19" pageNumber="321" unit="mm" value="19.5">19.5 mm</quantity>
|
||
of forelimb length, 31.0 mm of hindlimb length, with missing tail tip, and having the following scalation features: 57 longitudinal rows of dorsal scales at mid-body which are smooth on the neck but carinated on the back, 25 gular scales along a line from the contact between the fourth pair of maxillary scales to the collar, 26 transversal rows of ventral plates from the collar to the anal plate, 21 and 20 femoral pores on the left and right sides respectively, 20 subdigital lamellae beneath the fourth toe, one and a half row of supraciliary granules on each side, with 41 (left) and 46 (right) scales and granules around the 2
|
||
<superScript id="7C41F3055F440C0462996099F3DDFE29" attach="left" box="[165,181,404,418]" fontSize="6" pageId="19" pageNumber="321">nd</superScript>
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||
and 3
|
||
<superScript id="7C41F3055F440C0462C46099F26EFE29" attach="left" box="[248,262,404,418]" fontSize="6" pageId="19" pageNumber="321">rd</superScript>
|
||
supraoculars, one entire internasal, no scale inserted between the two prefrontals, two supralabials in contact with the subocular, the latter one in large contact with the upper lip on each side. Base of the tail very thick.
|
||
</paragraph>
|
||
<paragraph id="8B8B5E4D5F440C0462FB60D0F186FC9C" blockId="19.[151,1437,152,791]" pageId="19" pageNumber="321">Coloration in life: general ground colour light rufous brown with 3 longitudinal light grey stripes on each side; a series of irregular transverse black marks on the sides of the dorsum between the dorsolateral and dorsal pale lines and dark marks on the edge of the vertebral area between the two dorsal pale lines; a series of 7–8 small pale greenish-yellow ocelli, each surrounded a large black spot that are connected by black coloration, are present along the flanks; pileus roughly of the same colour as the dorsum but more intensely rufous, with very few dark marks; underparts pure white except for the tail and thighs (whitish with pale cream/salmon tinge) and some grey marks on the outer ventrals, 5 spots of yellow-green scales on the outer ventrals/lower dorsals; a poorly contrasted and fragmented dark longitudinal stripe runs along each side of the tail; tail base invaded by yellow-green spots of the same colour as the ones on the flanks and outer belly.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4B0EC55F440C0462AB675BF792F8B8" ID-DOI="http://doi.org/10.5281/zenodo.3694848" ID-Zenodo-Dep="3694848" httpUri="https://zenodo.org/record/3694848/files/figure.png" pageId="19" pageNumber="321" startId="19.[151,250,1622,1647]" targetBox="[206,1380,802,1609]" targetPageId="19">
|
||
<paragraph id="8B8B5E4D5F440C0462AB675BF792F8B8" blockId="19.[151,1436,1622,1843]" pageId="19" pageNumber="321">
|
||
<emphasis id="B940825F5F440C0462AB675BF249F9E4" bold="true" box="[151,289,1622,1647]" pageId="19" pageNumber="321">FIGURE 10.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C3425CE5F440C0463F9675AF1B0F9E4" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[453,728,1623,1647]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="321" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F440C0463F9675AF1B0F9E4" box="[453,728,1623,1647]" italics="true" pageId="19" pageNumber="321">Acanthodactylus montanus</emphasis>
|
||
</taxonomicName>
|
||
(squares) and
|
||
<taxonomicName id="4C3425CE5F440C046153675AF715F9E4" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[879,1149,1623,1647]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="321" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F440C046153675AF715F9E4" box="[879,1149,1623,1647]" italics="true" pageId="19" pageNumber="321">Acanthodactylus lacrymae</emphasis>
|
||
</taxonomicName>
|
||
(diamonds) based on specimens examined in this study. Coloured dots represent specimens of the three main phenotypes (none is monophyletic in the mtDNA or nDNA trees) in Morocco based on
|
||
<bibRefCitation id="EFA523BC5F440C0460516795F03FF924" author="Bons, J. & Geniez, P." box="[621,855,1686,1711]" pageId="19" pageNumber="321" pagination="271 - 280" refId="ref13957" refString="Bons, J. & Geniez, P. (1995) Contribution to the systematics of the lizard Acanthodactylus erythrurus (Sauria, Lacertidae) in Morocco. Herpetological Journal, 5, 271 - 280." type="journal article" year="1995">Bons and Geniez 1995</bibRefCitation>
|
||
. Specimens with the “bellii” phenotype in the High Atlas are almost certainly
|
||
<taxonomicName id="4C3425CE5F440C04635867B5F2AFF944" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[356,455,1720,1743]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="321" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F440C04635867B5F2AFF944" box="[356,455,1720,1743]" italics="true" pageId="19" pageNumber="321">montanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3425CE5F440C0463CB67BAF13DF944" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[503,597,1719,1743]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="321" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F440C0463CB67BAF13DF944" box="[503,597,1719,1743]" italics="true" pageId="19" pageNumber="321">lacrymae</emphasis>
|
||
</taxonomicName>
|
||
. White symbols represent specimens tentatively identified as
|
||
<taxonomicName id="4C3425CE5F440C0466F967BAF62CF944" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[1221,1348,1719,1743]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="321" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F440C0466F967BAF62CF944" box="[1221,1348,1719,1743]" italics="true" pageId="19" pageNumber="321">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
(square) or
|
||
<taxonomicName id="4C3425CE5F440C04628867DAF247F964" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[180,303,1751,1775]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="321" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F440C04628867DAF247F964" box="[180,303,1751,1775]" italics="true" pageId="19" pageNumber="321">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
(diamond, see text). The continuous white lines represent the currently known distribution of the new species while dotted white lines represent possible extensions of the distribution if tentative identifications are correct. The dotted dark blue line represents the southernmost extension of confirmed IM clade specimens based on our field sampling.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32E0DC65F440C0362FB666CF1AEFD90" lastPageId="20" lastPageNumber="322" pageId="19" pageNumber="321" type="distribution">
|
||
<paragraph id="8B8B5E4D5F440C0362FB666CF04BFEC8" blockId="19.[151,1436,1889,2024]" lastBlockId="20.[151,1437,153,539]" lastPageId="20" lastPageNumber="322" pageId="19" pageNumber="321">
|
||
<emphasis id="B940825F5F440C0462FB666CF209F8F7" bold="true" box="[199,353,1889,1916]" pageId="19" pageNumber="321">Distribution.</emphasis>
|
||
Known only from high altitudes (between
|
||
<quantity id="4CCCF3A85F440C046107666FF0FEF8F7" box="[827,918,1890,1916]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="321" unit="m" value="2000.0">2,000 m</quantity>
|
||
and
|
||
<quantity id="4CCCF3A85F440C0461F6666CF74DF8F7" box="[970,1061,1889,1916]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="321" unit="m" value="2500.0">2,500 m</quantity>
|
||
a.s.l. but precise elevation known for a very small number of localities only) in the southwestern High Atlas (Tizi n’Tichka) and in the Jbel Siroua massif,
|
||
<quantity id="4CCCF3A85F440C0462D166A4F236F848" box="[237,350,1961,1987]" metricMagnitude="4" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="19" pageNumber="321" unit="km" value="65.0" valueMax="70.0" valueMin="60.0">60–70 km</quantity>
|
||
southwest of the Tzin-n-Tichka area (
|
||
<figureCitation id="130F42C85F440C04613F66A4F03EF84F" box="[771,854,1961,1988]" captionStart="FIGURE 10" captionStartId="19.[151,250,1622,1647]" captionTargetBox="[206,1380,802,1609]" captionTargetId="figure@19.[200,1387,798,1614]" captionTargetPageId="19" captionText="FIGURE 10. Distribution of Acanthodactylus montanus (squares) and Acanthodactylus lacrymae (diamonds) based on specimens examined in this study. Coloured dots represent specimens of the three main phenotypes (none is monophyletic in the mtDNA or nDNA trees) in Morocco based on Bons and Geniez 1995. Specimens with the “bellii” phenotype in the High Atlas are almost certainly montanus and lacrymae. White symbols represent specimens tentatively identified as A. montanus (square) or A. lacrymae (diamond, see text). The continuous white lines represent the currently known distribution of the new species while dotted white lines represent possible extensions of the distribution if tentative identifications are correct. The dotted dark blue line represents the southernmost extension of confirmed IM clade specimens based on our field sampling." figureDoi="http://doi.org/10.5281/zenodo.3694848" httpUri="https://zenodo.org/record/3694848/files/figure.png" pageId="19" pageNumber="321">Fig. 10</figureCitation>
|
||
). Based on observations of animals with a
|
||
<taxonomicName id="4C3425CE5F440C04671366A4F601F848" authorityName="Gray" authorityYear="1845" box="[1327,1385,1961,1987]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="321" phylum="Chordata" rank="species" species="bellii">
|
||
<emphasis id="B940825F5F440C04671366A4F601F848" box="[1327,1385,1961,1987]" italics="true" pageId="19" pageNumber="321">bellii</emphasis>
|
||
</taxonomicName>
|
||
-like subocular scalation in the western High Atlas (
|
||
<bibRefCitation id="EFA523BC5F440C04609466C3F0C4F863" author="Bons, J. & Geniez, P." box="[680,940,1997,2024]" pageId="19" pageNumber="321" pagination="271 - 280" refId="ref13957" refString="Bons, J. & Geniez, P. (1995) Contribution to the systematics of the lizard Acanthodactylus erythrurus (Sauria, Lacertidae) in Morocco. Herpetological Journal, 5, 271 - 280." type="journal article" year="1995">Bons and Geniez 1995</bibRefCitation>
|
||
) and on habitat availability, the distribution probably extends rather continuously from just east of Tizi n’Tichka to Jbel Siroua but maybe not much further east. For example, no member of the
|
||
<taxonomicName id="4C3425CE5F430C0363C561B0F047FF5C" baseAuthorityName="Schinz" baseAuthorityYear="1833" box="[505,815,189,215]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="322" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="B940825F5F430C0363C561B0F047FF5C" box="[505,815,189,215]" italics="true" pageId="20" pageNumber="322">Acanthodactylus erythrurus</emphasis>
|
||
</taxonomicName>
|
||
complex has ever been found at Oukaimeden in spite of the availability of similar habitats and of numerous visits by many herpetologists. If specimens of doubtful identity from between Tazenakht and Taroundant prove to be
|
||
<taxonomicName id="4C3425CE5F430C0360CD6008F016FE94" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[753,894,261,287]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="322" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F430C0360CD6008F016FE94" box="[753,894,261,287]" italics="true" pageId="20" pageNumber="322">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
(white square in
|
||
<figureCitation id="130F42C85F430C03667F6008F7FCFE94" box="[1091,1172,261,287]" captionStart="FIGURE 10" captionStartId="19.[151,250,1622,1647]" captionTargetBox="[206,1380,802,1609]" captionTargetId="figure@19.[200,1387,798,1614]" captionTargetPageId="19" captionText="FIGURE 10. Distribution of Acanthodactylus montanus (squares) and Acanthodactylus lacrymae (diamonds) based on specimens examined in this study. Coloured dots represent specimens of the three main phenotypes (none is monophyletic in the mtDNA or nDNA trees) in Morocco based on Bons and Geniez 1995. Specimens with the “bellii” phenotype in the High Atlas are almost certainly montanus and lacrymae. White symbols represent specimens tentatively identified as A. montanus (square) or A. lacrymae (diamond, see text). The continuous white lines represent the currently known distribution of the new species while dotted white lines represent possible extensions of the distribution if tentative identifications are correct. The dotted dark blue line represents the southernmost extension of confirmed IM clade specimens based on our field sampling." figureDoi="http://doi.org/10.5281/zenodo.3694848" httpUri="https://zenodo.org/record/3694848/files/figure.png" pageId="20" pageNumber="322">Fig. 10</figureCitation>
|
||
) the distribution would extend to the northernmost parts of the eastern Anti-Atlas.
|
||
</paragraph>
|
||
<paragraph id="8B8B5E4D5F430C0362FB6040F1AEFD90" blockId="20.[151,1437,153,539]" pageId="20" pageNumber="322">
|
||
<emphasis id="B940825F5F430C0362FB6040F2EDFEEC" bold="true" box="[199,389,332,359]" pageId="20" pageNumber="322">Natural history.</emphasis>
|
||
At the Tizi n’Tichka locality, the habitat was very similar to the habitats occupied by
|
||
<taxonomicName id="4C3425CE5F430C0367076040F3AEFE00" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="322" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F430C0367076040F3AEFE00" italics="true" pageId="20" pageNumber="322">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
further east: earthy slopes with bare gravel ground or short grass interspaced with dense low bushes and some stones. Population density was rather high and the animals were relatively easy to catch (compared to many populations of the IM clade) due to their more confident behaviour and slower escape. The animals were active in sunny and calm weather and their behaviour was typical of the
|
||
<taxonomicName id="4C3425CE5F430C03612660D0F0D8FE7C" baseAuthorityName="Schinz" baseAuthorityYear="1833" box="[794,944,477,503]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="322" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="B940825F5F430C03612660D0F0D8FE7C" box="[794,944,477,503]" italics="true" pageId="20" pageNumber="322">A. erythrurus</emphasis>
|
||
</taxonomicName>
|
||
complex, actively foraging in the open and retreating to the shelter of bushes when disturbed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |