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<document id="6A86F1FF1CFE8C17C4F848A48E8E23C0" ID-DOI="10.11646/zootaxa.3914.5.2" ID-GBIF-Dataset="c4987457-cba4-494d-b137-20035790c30d" ID-ISSN="1175-5326" ID-Zenodo-Dep="240826" ID-ZooBank="623E7F1A-2866-44BB-8B67-69971D8054C6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460964277127" checkinUser="plazi" docAuthor="Marshall, S. A." docDate="2015" docId="039D9771FF961A2F43E3DD86FC18FC02" docLanguage="en" docName="zt03914p540.pdf" docOrigin="Zootaxa 3914 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mesoconius zadbi Marshall, 2015, new species" docType="treatment" docVersion="9" lastPageNumber="540" masterDocId="FFA4EF09FF981A204374DD11FFE3FFCA" masterDocTitle="Mesoconius Enderlein (Diptera, Micropezidae, Taeniapterinae) of Central America" masterLastPageNumber="540" masterPageNumber="525" pageNumber="539" updateTime="1698557623514" updateUser="plazi">
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<mods:title id="F8B3838F6E7452BE15C009F6CBE20A99">Mesoconius Enderlein (Diptera, Micropezidae, Taeniapterinae) of Central America</mods:title>
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<mods:namePart id="739D8CBFF20369A5CB9E5C8A837A9454">Marshall, S. A.</mods:namePart>
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<mods:date id="9FD8BE33001D04C375F93AF7537CCC79">2015</mods:date>
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<mods:number id="A6477FEB58E4B9F817A089F041AC765F">3914</mods:number>
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<mods:classification id="072C63FC9AFE87486D4D52924C358149">journal article</mods:classification>
<mods:identifier id="592EF281C722606684A8BFE5ECFCA1AF" type="DOI">10.11646/zootaxa.3914.5.2</mods:identifier>
<mods:identifier id="6609306941600D19E4890C3CE00EF659" type="GBIF-Dataset">c4987457-cba4-494d-b137-20035790c30d</mods:identifier>
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<treatment id="039D9771FF961A2F43E3DD86FC18FC02" ID-DOI="http://doi.org/10.5281/zenodo.3503146" ID-GBIF-Taxon="119593351" ID-Zenodo-Dep="3503146" LSID="urn:lsid:plazi:treatment:039D9771FF961A2F43E3DD86FC18FC02" httpUri="http://treatment.plazi.org/id/039D9771FF961A2F43E3DD86FC18FC02" lastPageId="15" lastPageNumber="540" pageId="14" pageNumber="539">
<subSubSection id="C32E75ECFF961A2E43E3DD86FEC4FF19" pageId="14" pageNumber="539" type="nomenclature">
<paragraph id="8B8B2667FF961A2E43E3DD86FDE5FF78" blockId="14.[151,518,151,211]" box="[151,518,151,178]" pageId="14" pageNumber="539">
<heading id="D0C3910BFF961A2E43E3DD86FDE5FF78" bold="true" box="[151,518,151,178]" fontSize="11" level="1" pageId="14" pageNumber="539" reason="1">
<emphasis id="B940FA75FF961A2E43E3DD86FDE5FF78" bold="true" box="[151,518,151,178]" pageId="14" pageNumber="539">
<taxonomicName id="4C345DE4FF961A2E43E3DD86FE8DFF7B" ID-CoL="9HWNQ" box="[151,366,151,177]" class="Insecta" family="Micropezidae" genus="Mesoconius" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="539" phylum="Arthropoda" rank="species" species="zadbi" status="sp. nov.">
<emphasis id="B940FA75FF961A2E43E3DD86FE8DFF7B" bold="true" box="[151,366,151,177]" italics="true" pageId="14" pageNumber="539">Mesoconius zadbi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273470EFF961A2E4202DD89FDE5FF78" box="[374,518,152,178]" pageId="14" pageNumber="539" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B2667FF961A2E43E3DDABFEC4FF19" blockId="14.[151,518,151,211]" box="[151,295,186,211]" pageId="14" pageNumber="539">
<figureCitation id="130F3AE2FF961A2E43E3DDABFEC4FF19" box="[151,295,186,211]" captionStart="FIGURE 6 A – D" captionStartId="14.[151,250,1588,1610]" captionTargetBox="[150,1435,515,1566]" captionTargetId="figure@14.[150,1436,515,1567]" captionTargetPageId="14" captionText="FIGURE 6 A – D. Mesoconius zadbi. A, head; B, female; C, thorax – abdomen junction to show pointed katatergite; D, spermatheca and associated structures." httpUri="https://zenodo.org/record/240831/files/figure.png" pageId="14" pageNumber="539">Figs. 6A–6D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32E75ECFF961A2F43E3DC13FEA5FE3F" lastPageId="15" lastPageNumber="540" pageId="14" pageNumber="539" type="description">
<paragraph id="8B8B2667FF961A2E43E3DC13FC56FE05" blockId="14.[151,1437,258,463]" pageId="14" pageNumber="539">
<emphasis id="B940FA75FF961A2E43E3DC13FEC2FED1" bold="true" box="[151,289,258,283]" pageId="14" pageNumber="539">Description</emphasis>
(females only): Length (antenna to wing tip)
<quantity id="4CCC8B82FF961A2E4059DC13FC63FED0" box="[813,896,258,283]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="539" unit="mm" value="15.0">15 mm</quantity>
. Color: Black to dark brown except as follows: gena, extreme anteroventral corner of face and parafacial silvery pruinose, palpus orange–yellow, margin of frontal vitta silvery pruinose. Face and antenna reddish brown. Fore tarsus white with white setae, hind tarsus white with white setae on tarsomeres 1–2, mixed white and brown setae on tarsomere 3, brown setae on tarsomeres 4–5; fore femur narrowly orange basally, black distally; mid and hind femora black with orange apex. Fore tibia black, mid tibia and hind tibia orange to yellow. Distal abdominal scerites reddish.
</paragraph>
<caption id="DF4B76EFFF961A2E43E3DB25FDFCF9A3" httpUri="https://zenodo.org/record/240831/files/figure.png" pageId="14" pageNumber="539" targetBox="[150,1435,515,1566]" targetPageId="14">
<paragraph id="8B8B2667FF961A2E43E3DB25FDFCF9A3" blockId="14.[151,1435,1588,1641]" pageId="14" pageNumber="539">
<emphasis id="B940FA75FF961A2E43E3DB25FDFEF980" bold="true" box="[151,541,1588,1610]" pageId="14" pageNumber="539">
FIGURE
<emphasis id="B940FA75FF961A2E4272DB25FDFEF980" bold="true" box="[262,541,1588,1610]" italics="true" pageId="14" pageNumber="539">
6A–D.
<taxonomicName id="4C345DE4FF961A2E4221DB25FDFAF980" box="[341,537,1588,1610]" class="Insecta" family="Micropezidae" genus="Mesoconius" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="539" phylum="Arthropoda" rank="species" species="zadbi">Mesoconius zadbi</taxonomicName>
.
</emphasis>
</emphasis>
A, head; B, female; C, thorax–abdomen junction to show pointed katatergite; D, spermatheca and associated structures.
</paragraph>
</caption>
<paragraph id="8B8B2667FF961A2E43B3DB84FEE8F84C" blockId="14.[151,1437,1685,2034]" pageId="14" pageNumber="539">Head: Epicephalon and paracephalon with a microscopic pattern of longitudinal striations, shining, smooth, sharply contrasting with orbits and lower frons; frontal vitta velvety black, sharply constricted behind ocelli, then expanded to a densely microtrichose area between postocellar bristles; widely separate from anterior margin of frons. Outer vertical bristles absent, inner vertical bristles strong, postocellar bristles strong. Fronto-orbital bristles (2) strong and widely separated. Upper face entirely microsetulose, strongly and broadly carinate; parafacials, gena, and outer corner of face strongly silvery pruinose, subantennal areas shining reddish brown. Palpus yellowish.</paragraph>
<paragraph id="8B8B2667FF961A2F43B3DA80FCF6FF1F" blockId="14.[151,1437,1685,2034]" lastBlockId="15.[151,1437,151,968]" lastPageId="15" lastPageNumber="540" pageId="14" pageNumber="539">Thorax: Cervical sclerite with a vertical groove separating a microtrichose, subquadrate posterior portion from a small bare anterior portion. Notum microtrichose, without acrostichal setulae but with a distinct row of small dorsocentral setulae; postpronotal lobe microtrichose with some scattered small setulae, anterior margin forming a vertical, shining face. Katatergite very prominent, with a tapered microtrichose apex. Vertical row of katepisternal bristles yellow. Coxae with anteroventral golden bristles.</paragraph>
<paragraph id="8B8B2667FF971A2F43B3DDCEFDBAFED6" blockId="15.[151,1437,151,968]" pageId="15" pageNumber="540">Abdomen: Syntergite 1+2 very narrow basally, constriction between syntergites about half as wide as base of T3; length of T1+2 double length of T3.</paragraph>
<paragraph id="8B8B2667FF971A2F43B3DC36FEA5FE3F" blockId="15.[151,1437,151,968]" pageId="15" pageNumber="540">Female abdomen: Pleuron dark on dorsal half of segments 2–5, otherwise pale. Oviscape reddish brown, paler apically. Bursa copulatrix large, ventral receptacle small. Single spermatheca small, slightly broader than duct; paired spermathecal duct similar in diameter to paired duct but longer and divided into a broad striate distal half and a narrower smooth distal half, distally dividing into long spermathecal bases each with a subbasal elongate oval swelling and a distal sinuate section with 2–3 deep S–bends. Paired spermathecae elongate, distinctly striate, tapered basally.</paragraph>
</subSubSection>
<subSubSection id="C32E75ECFF971A2F43B3DCEEFB81FD4E" pageId="15" pageNumber="540" type="materials_examined">
<paragraph id="8B8B2667FF971A2F43B3DCEEFB81FD4E" blockId="15.[151,1437,151,968]" pageId="15" pageNumber="540">
<emphasis id="B940FA75FF971A2F43B3DCEEFE8EFDD2" bold="true" box="[199,365,511,536]" pageId="15" pageNumber="540">
<typeStatus id="548F98C5FF971A2F43B3DCEEFEE1FDD2" box="[199,258,511,536]" pageId="15" pageNumber="540">Type</typeStatus>
material
</emphasis>
(all
<collectionCode id="ED25BEA2FF971A2F42EBDF11FE0BFDD2" box="[415,488,512,536]" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="15" pageNumber="540">INBC</collectionCode>
):
<typeStatus id="548F98C5FF971A2F428DDCEEFD82FDD2" box="[505,609,511,536]" pageId="15" pageNumber="540" type="holotype">Holotype</typeStatus>
(♀):
<collectingCountry id="F32366F7FF971A2F41EADF11FCA6FDD2" box="[670,837,512,536]" name="Costa Rica" pageId="15" pageNumber="540">COSTA RICA</collectingCountry>
. Cartago, Rio Macho forest reserve, La Esperanza del Guarco, end of Esperanza road,
<quantity id="4CCC8B82FF971A2F4170DF35FDBEFDF6" box="[516,605,548,572]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="15" pageNumber="540" unit="m" value="2400.0">2400 m</quantity>
.
<date id="FF8A00A7FF971A2F411DDF35FD12FDF7" box="[617,753,548,573]" pageId="15" pageNumber="540" value="1998-06-01" valueMax="1998-06-02" valueMin="1998-06-01">1–2.vi.1998</date>
, S.A.
<collectingCountry id="F32366F7FF971A2F404CDF35FC7CFDF7" box="[824,927,548,573]" name="Marshall Islands" pageId="15" pageNumber="540">Marshall</collectingCountry>
.
<typeStatus id="548F98C5FF971A2F40DEDF34FBF8FDF7" box="[938,1051,549,573]" pageId="15" pageNumber="540" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F32366F7FF971A2F4753DF35FB2CFDF6" box="[1063,1231,548,572]" name="Costa Rica" pageId="15" pageNumber="540">COSTA RICA</collectingCountry>
. Cartago, Paraíso, PN Tapantí, Macizo de la Muerte,
<quantity id="4CCC8B82FF971A2F415ADF56FD6AFD95" box="[558,649,583,608]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="540" unit="m" value="1500.0">1500 m</quantity>
.
<date id="FF8A00A7FF971A2F41E3DF59FCEBFDAA" box="[663,776,584,608]" pageId="15" pageNumber="540" value="2000-05-19">19.v.2000</date>
, malaise, C. Hanson, D. Rubi (1 ♀). San José, Moravia, Zurquí de Moravia, tower path,
<quantity id="4CCC8B82FF971A2F4289DF7DFDB6FD4E" box="[509,597,620,645]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="540" unit="m" value="1600.0">1600 m</quantity>
.
<date id="FF8A00A7FF971A2F4114DF7DFD19FD4E" box="[608,762,620,644]" pageId="15" pageNumber="540" value="2013-05-24" valueMax="2013-05-31" valueMin="2013-05-24">24–31.v.2013</date>
, Malaise, project
<taxonomicName id="4C345DE4FF971A2F40B6DF7DFBF4FD4E" box="[962,1047,620,644]" class="Insecta" family="Micropezidae" genus="Mesoconius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="540" phylum="Arthropoda" rank="species" species="zadbi">
<collectionCode id="ED25BEA2FF971A2F40B6DF7DFBF4FD4E" box="[962,1047,620,644]" pageId="15" pageNumber="540">ZADBI</collectionCode>
</taxonomicName>
(1 ♀).
</paragraph>
</subSubSection>
<subSubSection id="C32E75ECFF971A2F43B3DF9EFDC1FD3A" pageId="15" pageNumber="540" type="etymology">
<paragraph id="8B8B2667FF971A2F43B3DF9EFDC1FD3A" blockId="15.[151,1437,151,968]" pageId="15" pageNumber="540">
<emphasis id="B940FA75FF971A2F43B3DF9EFEADFD62" bold="true" box="[199,334,655,680]" pageId="15" pageNumber="540">Etymology.</emphasis>
The specific name refers to Project
<taxonomicName id="4C345DE4FF971A2F407ADF81FC85FD62" box="[782,870,656,680]" class="Insecta" family="Micropezidae" genus="Mesoconius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="540" phylum="Arthropoda" rank="species" species="zadbi">ZADBI</taxonomicName>
, the Zurquí All
<taxonomicName id="4C345DE4FF971A2F4745DF9EFA7EFD62" authority="Biodiversity Inventory." authorityName="Biodiversity Inventory." box="[1073,1437,655,680]" kingdom="Plantae" order="Diptera" pageId="15" pageNumber="540" rank="order">Diptera Biodiversity Inventory.</taxonomicName>
<taxonomicName id="4C345DE4FF971A2F43E3DFA4FE83FD06" box="[151,352,692,716]" class="Insecta" family="Micropezidae" genus="Mesoconius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="540" phylum="Arthropoda" rank="species" species="zadbi">
<emphasis id="B940FA75FF971A2F43E3DFA4FE83FD06" box="[151,352,692,716]" italics="true" pageId="15" pageNumber="540">Mesoconius zadbi</emphasis>
</taxonomicName>
and most of the other
<taxonomicName id="4C345DE4FF971A2F4128DFA4FD01FD06" box="[604,738,693,716]" class="Insecta" family="Micropezidae" genus="Mesoconius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="B940FA75FF971A2F4128DFA4FD01FD06" box="[604,738,693,716]" italics="true" pageId="15" pageNumber="540">Mesoconius</emphasis>
</taxonomicName>
species newly described here are described partly on the basis of specimens from Project
<taxonomicName id="4C345DE4FF971A2F42B1DFC9FDFDFD3A" box="[453,542,728,752]" class="Insecta" family="Micropezidae" genus="Mesoconius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="540" phylum="Arthropoda" rank="species" species="zadbi">ZADBI</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E75ECFF971A2F43B3DFEDFC18FC02" pageId="15" pageNumber="540" type="discussion">
<paragraph id="8B8B2667FF971A2F43B3DFEDFEEDFC97" blockId="15.[151,1437,151,968]" pageId="15" pageNumber="540">
<emphasis id="B940FA75FF971A2F43B3DFEDFEACFCDF" bold="true" box="[199,335,764,789]" pageId="15" pageNumber="540">Comments.</emphasis>
Although this species remains known only from males,
<taxonomicName id="4C345DE4FF971A2F40A2DFECFBDAFCDE" box="[982,1081,764,788]" class="Insecta" family="Micropezidae" genus="Mesoconius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="540" phylum="Arthropoda" rank="species" species="zadbi">
<emphasis id="B940FA75FF971A2F40A2DFECFBDAFCDE" box="[982,1081,764,788]" italics="true" pageId="15" pageNumber="540">M. zadbi</emphasis>
</taxonomicName>
clearly belongs in
<taxonomicName id="4C345DE4FF971A2F4662DFECFA7FFCDE" box="[1302,1436,765,788]" class="Insecta" family="Micropezidae" genus="Mesoconius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="B940FA75FF971A2F4662DFECFA7FFCDE" box="[1302,1436,765,788]" italics="true" pageId="15" pageNumber="540">Mesoconius</emphasis>
</taxonomicName>
and can be placed in a clade characterized by a paired spermathecal duct with a striate basal half and a smooth distal half.
</paragraph>
<paragraph id="8B8B2667FF971A2F43B3DE76FC18FC02" blockId="15.[151,1437,151,968]" pageId="15" pageNumber="540">
This species is similar to the South American
<taxonomicName id="4C345DE4FF971A2F4195DE79FC24FC4A" box="[737,967,872,896]" class="Insecta" family="Micropezidae" genus="Mesoconius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="540" phylum="Arthropoda" rank="species" species="infestus">
<emphasis id="B940FA75FF971A2F4195DE79FC24FC4A" box="[737,967,872,896]" italics="true" pageId="15" pageNumber="540">Mesoconius infestus</emphasis>
</taxonomicName>
group, but differs most obviously by its black mid and hind femora with contrasting orange apices.
<taxonomicName id="4C345DE4FF971A2F4044DE9CFC71FC6E" box="[816,914,908,932]" class="Insecta" family="Micropezidae" genus="Mesoconius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="540" phylum="Arthropoda" rank="species" species="zadbi">
<emphasis id="B940FA75FF971A2F4044DE9CFC71FC6E" box="[816,914,908,932]" italics="true" pageId="15" pageNumber="540">M. zadbi</emphasis>
</taxonomicName>
is probably most closely related to
<emphasis id="B940FA75FF971A2F4656DE9CFA7FFC6E" box="[1314,1436,908,932]" italics="true" pageId="15" pageNumber="540">
<taxonomicName id="4C345DE4FF971A2F4656DE9CFA7BFC6E" box="[1314,1432,908,932]" class="Insecta" family="Micropezidae" genus="Mesoconius" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="540" phylum="Arthropoda" rank="species" species="bicolor">M. bicolor</taxonomicName>
,
</emphasis>
but characters of the as yet unknown males are needed to test that hypothesis.
</paragraph>
</subSubSection>
</treatment>
</document>
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