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<document id="3C4BF1E7236B2CCCE73E5B49E06BB3E5" ID-DOI="10.1111/j.1096-3642.2005.00189.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5435260" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630434122956" checkinUser="diego" docAuthor="Daugeron, Christophe &amp; Grootaert, Patrick" docDate="2005" docId="039D7838FFCAFFDB3FCED97DFE51FE48" docLanguage="en" docName="j.1096-3642.2005.00189.x.pdf" docOrigin="Zoological Journal of the Linnean Society 145 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2005.00189.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Empis (Coptophlebia) aspina Daugeron &amp; Grootaert 2005" docType="treatment" docVersion="3" lastPageNumber="359" masterDocId="FFA40040FFD8FFCF3F28DF3EFFC4FFD9" masterDocTitle="Phylogenetic systematics of the Empis (Coptophlebia) hyalea-group (Insecta: Diptera: Empididae)" masterLastPageNumber="391" masterPageNumber="339" pageNumber="357" updateTime="1699269823662" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="C4088220CE089B6918836F8C058B8A27">Phylogenetic systematics of the Empis (Coptophlebia) hyalea-group (Insecta: Diptera: Empididae)</mods:title>
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<mods:namePart id="508BF52F4B508F1870109AF9B5CBA747">Daugeron, Christophe</mods:namePart>
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<mods:namePart id="0CAC5D14FD80D4DBE17432D649C74801">Grootaert, Patrick</mods:namePart>
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<mods:title id="B1BF66DC486EA586AF9BC1F9BA8CE1A5">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="8AE293E4D3F2D896A0F0F142231F7508">2005</mods:date>
<mods:detail id="7A33BCDDFF421A94236B19BD9373362C" type="pubDate">
<mods:number id="699349AC2E19348B0BFF70930B50800E">2005-11-30</mods:number>
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<mods:number id="9EFC0B059097EDDE287A484E5FBACBDD">145</mods:number>
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<mods:identifier id="8AB2A8B0C5D6F5EC5DB22FB2761B03A4" type="DOI">10.1111/j.1096-3642.2005.00189.x</mods:identifier>
<mods:identifier id="77E4D7465D9315DF7D43F7EADD081DCA" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="0F203CC73ACDD46AD116627BC0DD89E9" type="Zenodo-Dep">5435260</mods:identifier>
</mods:mods>
<treatment id="039D7838FFCAFFDB3FCED97DFE51FE48" LSID="urn:lsid:plazi:treatment:039D7838FFCAFFDB3FCED97DFE51FE48" httpUri="http://treatment.plazi.org/id/039D7838FFCAFFDB3FCED97DFE51FE48" lastPageId="20" lastPageNumber="359" pageId="18" pageNumber="357">
<subSubSection id="C32E9AA5FFCAFFDD3FCED97DFE18F93F" pageId="18" pageNumber="357" type="nomenclature">
<paragraph id="8B8BC92EFFCAFFDD3FCED97DFD5AF980" blockId="18.[230,670,1603,1627]" box="[230,670,1603,1627]" pageId="18" pageNumber="357">
<heading id="D0C37E42FFCAFFDD3FCED97DFD5AF980" box="[230,670,1603,1627]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="357" reason="2">
<taxonomicName id="4C34B2ADFFCAFFDD3FCED97DFD89F980" ID-CoL="8BZ3K" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[230,589,1603,1627]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="361" phylum="Arthropoda" rank="species" species="aspina" subGenus="Coptophlebia">
<emphasis id="B940153CFFCAFFDD3FCED97DFEEAF980" box="[230,302,1603,1626]" italics="true" pageId="18" pageNumber="357">EMPIS</emphasis>
(
<emphasis id="B940153CFFCAFFDD3E17D97DFE2FF980" box="[319,491,1603,1627]" italics="true" pageId="18" pageNumber="357">COPTOPHLEBIA</emphasis>
)
<emphasis id="B940153CFFCAFFDD3ED5D978FD89F980" box="[509,589,1606,1625]" italics="true" pageId="18" pageNumber="357">ASPINA</emphasis>
</taxonomicName>
CLADE
</heading>
</paragraph>
<paragraph id="8B8BC92EFFCAFFDD3FA6D94DFE18F93F" blockId="18.[142,758,1651,1766]" pageId="18" pageNumber="357">
<emphasis id="B940153CFFCAFFDD3FA6D94DFED6F951" box="[142,274,1651,1672]" italics="true" pageId="18" pageNumber="357">Monophyly:</emphasis>
The monophyly of this clade is supported by one character found in the proboscis: labium strongly sclerotized, bare or only with sparse minute bristly hairs [character 1(1)].
</paragraph>
</subSubSection>
<subSubSection id="C32E9AA5FFCAFFDB3FA6D9C3FEBAFEAA" lastPageId="20" lastPageNumber="359" pageId="18" pageNumber="357" type="distribution">
<paragraph id="8B8BC92EFFCAFFDD3FA6D9C3FE9FF8B6" blockId="18.[142,758,1789,1903]" pageId="18" pageNumber="357">
<emphasis id="B940153CFFCAFFDD3FA6D9C3FE4CF8CB" box="[142,392,1789,1811]" italics="true" pageId="18" pageNumber="357">
Distribution (
<figureCitation id="130FD5ABFFCAFFDD3E02D9C0FEBEF8CB" box="[298,378,1789,1811]" captionStart="Figure 15" captionStartId="22.[142,220,942,961]" captionTargetBox="[311,1250,198,907]" captionTargetId="figure-345@22.[298,1258,186,918]" captionTargetPageId="22" captionText="Figure 15. Distribution of the Empis (Coptophlebia) aspina clade: E. (C.) aspina sp. nov. = 1; E. (C.) miranda Daugeron &amp; Grootaert = 4; E. (C.) nganga Daugeron &amp; Grootaert = 6; E. (C.) projecta sp. nov. = 2; E. (C.) pulchra Daugeron &amp; Grootaert = 5; E. (C.) ratburiensis Daugeron &amp; Grootaert = 3; E. (C.) urumae sp. nov. = 7." figureDoi="http://doi.org/10.5281/zenodo.5435294" httpUri="https://zenodo.org/record/5435294/files/figure.png" pageId="18" pageNumber="357">Fig. 15</figureCitation>
):
</emphasis>
Species included in this clade are recorded from South-east Asia (
<collectingCountry id="F32389BEFFCAFFDD3D16D822FDB2F8E8" box="[574,630,1820,1841]" name="Laos" pageId="18" pageNumber="357">Laos</collectingCountry>
,
<collectingCountry id="F32389BEFFCAFFDD3DAED822FD2AF8EB" box="[646,750,1820,1842]" name="Thailand" pageId="18" pageNumber="357">Thailand</collectingCountry>
) with one representative [
<taxonomicName id="4C34B2ADFFCAFFDD3EE1D805FDB4F889" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[457,624,1851,1872]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="357" phylum="Arthropoda" rank="species" species="urumae" subGenus="Coptophlebia">
<emphasis id="B940153CFFCAFFDD3EE1D805FE26F889" box="[457,482,1851,1872]" italics="true" pageId="18" pageNumber="357">E.</emphasis>
(
<emphasis id="B940153CFFCAFFDD3ED9D805FDCDF889" box="[497,521,1851,1872]" italics="true" pageId="18" pageNumber="357">C.</emphasis>
)
<emphasis id="B940153CFFCAFFDD3D30D805FDB4F889" box="[536,624,1851,1872]" italics="true" pageId="18" pageNumber="357">urumae</emphasis>
</taxonomicName>
] in
<collectingCountry id="F32389BEFFCAFFDD3D98D805FD32F889" box="[688,758,1851,1872]" name="Japan" pageId="18" pageNumber="357">Japan</collectingCountry>
(Ryukyu Islands).
</paragraph>
<paragraph id="8B8BC92EFFCAFFDD3C4FDC97FA89FC18" blockId="18.[871,1357,937,993]" box="[871,1357,937,961]" pageId="18" pageNumber="357">
<taxonomicName id="4C34B2ADFFCAFFDD3C4FDC97FB24FC66" authority="Daugeron &amp; Grootaert, 2005" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[871,1248,937,961]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="361" phylum="Arthropoda" rank="species" species="aspina" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B940153CFFCAFFDD3C4FDC97FC76FC66" bold="true" box="[871,946,937,961]" italics="true" pageId="18" pageNumber="357">EMPIS</emphasis>
(
<emphasis id="B940153CFFCAFFDD3CEBDC97FBBEFC66" bold="true" box="[963,1146,937,961]" italics="true" pageId="18" pageNumber="357">COPTOPHLEBIA</emphasis>
) ASPINA
</taxonomicName>
<emphasis id="B940153CFFCAFFDD3BC1DC92FA89FC18" bold="true" box="[1257,1357,937,961]" pageId="18" pageNumber="357">
<taxonomicNameLabel id="A273A847FFCAFFDD3BC1DC92FA89FC18" box="[1257,1357,937,961]" pageId="18" pageNumber="357" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B8BC92EFFCAFFDD3B00DCF7FB48FC38" blockId="18.[871,1357,937,993]" box="[1064,1164,969,993]" pageId="18" pageNumber="357">
(
<figureCitation id="130FD5ABFFCAFFDD3B18DCF7FB47FC38" box="[1072,1155,969,993]" captionStart="Figure 12" captionStartId="19.[162,240,1489,1508]" captionTargetBox="[338,1266,798,1455]" captionTargetId="figure-232@19.[317,1277,793,1480]" captionTargetPageId="19" captionText="Figure 12. The Empis (Coptophlebia) aspina clade, male of E. (C.) aspina sp. nov. A, epandrium and cercus in lateral view; B, hypandrium and phallus in lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435288" httpUri="https://zenodo.org/record/5435288/files/figure.png" pageId="18" pageNumber="357">FIG. 12</figureCitation>
)
</paragraph>
<paragraph id="8B8BC92EFFCAFFDD3C0EDCC4FAB7FB94" blockId="18.[806,1422,1017,1101]" pageId="18" pageNumber="357">
<emphasis id="B940153CFFCAFFDD3C0EDCC4FC0FFBD7" box="[806,971,1017,1039]" italics="true" pageId="18" pageNumber="357">Type material:</emphasis>
<materialsCitation id="3B5CC373FFCAFFDD3CF0DCC7FB75FBF7" collectingDate="1965-12" collectionCode="BPBM" country="Laos" location="Sayaboury Province" municipality="Sayaboury" pageId="18" pageNumber="357" specimenCount="1" typeStatus="holotype">
<typeStatus id="548F778CFFCAFFDD3CF0DCC7FB85FBD6" box="[984,1089,1017,1039]" pageId="18" pageNumber="357" type="holotype">Holotype</typeStatus>
: male,
<collectingCountry id="F32389BEFFCAFFDD3BA7DCC4FB02FBD6" box="[1167,1222,1018,1039]" name="Laos" pageId="18" pageNumber="357">Laos</collectingCountry>
,
<location id="8EEB9FF5FFCAFFDD3BF9DCC7FC92FBF7" LSID="urn:lsid:plazi:treatment:039D7838FFCAFFDB3FCED97DFE51FE48:8EEB9FF5FFCAFFDD3BF9DCC7FC92FBF7" country="Laos" municipality="Sayaboury" name="Sayaboury Province" pageId="18" pageNumber="357">Sayaboury Province</location>
,
<collectingMunicipality id="6BEF5354FFCAFFDD3C4DDB26FC1AFBF7" box="[869,990,1048,1070]" pageId="18" pageNumber="357">Sayaboury</collectingMunicipality>
,
<date id="FF8AEFEEFFCAFFDD3CC4DB26FB82FBF7" box="[1004,1094,1048,1070]" pageId="18" pageNumber="357" value="1965-12">
<collectingDate id="EFCE1606FFCAFFDD3CC4DB26FB82FBF7" box="[1004,1094,1048,1070]" pageId="18" pageNumber="357" value="1965-12">xii.1965</collectingDate>
</date>
(
<collectionCode id="ED2551EBFFCAFFDD3B71DB26FB6DFBF7" box="[1113,1193,1048,1070]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="18" pageNumber="357" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B5CC373FFCAFFDD3B97DB26FAB4FB94" collectingDate="1965-09-18" collectionCode="BPBM" collectorName="J. A. Rondon" country="Laos" location="Nongtevada" pageId="18" pageNumber="357" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="548F778CFFCAFFDD3B97DB26FAEDFBF4" box="[1215,1321,1048,1069]" pageId="18" pageNumber="357" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D3202A7FFCAFFDD3A1FDB26FA4DFBF7" box="[1335,1417,1048,1070]" pageId="18" pageNumber="357" type="male">1 male</specimenCount>
,
<collectingCountry id="F32389BEFFCAFFDD3C0EDB09FC99FB95" box="[806,861,1079,1100]" name="Laos" pageId="18" pageNumber="357">Laos</collectingCountry>
,
<location id="8EEB9FF5FFCAFFDD3C42DB09FC32FB94" LSID="urn:lsid:plazi:treatment:039D7838FFCAFFDB3FCED97DFE51FE48:8EEB9FF5FFCAFFDD3C42DB09FC32FB94" box="[874,1014,1079,1101]" country="Laos" name="Nongtevada" pageId="18" pageNumber="357">Nongtevada</location>
,
<date id="FF8AEFEEFFCAFFDD3B2ADB09FBBDFB94" box="[1026,1145,1079,1101]" pageId="18" pageNumber="357" value="1965-09-18">
<collectingDate id="EFCE1606FFCAFFDD3B2ADB09FBBDFB94" box="[1026,1145,1079,1101]" pageId="18" pageNumber="357" value="1965-09-18">18.ix.1965</collectingDate>
</date>
,
<collectorName id="26C1ACF8FFCAFFDD3BADDB09FAC3FB94" box="[1157,1287,1079,1101]" pageId="18" pageNumber="357">J.A.Rondon</collectorName>
(
<collectionCode id="ED2551EBFFCAFFDD3A3FDB09FAA3FB94" box="[1303,1383,1079,1101]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="18" pageNumber="357" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BC92EFFCAFFDD3C0EDB5BFCB1FB0F" blockId="18.[806,1422,1125,1238]" pageId="18" pageNumber="357">
<emphasis id="B940153CFFCAFFDD3C0EDB5BFC3AFBA3" box="[806,1022,1125,1146]" italics="true" pageId="18" pageNumber="357">
Etymology:
<taxonomicName id="4C34B2ADFFCAFFDD3C99DB5BFC3AFBA3" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[945,1022,1125,1146]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="357" phylum="Arthropoda" rank="species" species="aspina">aspina</taxonomicName>
</emphasis>
is a combination of
<emphasis id="B940153CFFCAFFDD3A2BDB5BFAD5FBA3" box="[1283,1297,1125,1146]" italics="true" pageId="18" pageNumber="357">a</emphasis>
(privative sense) and the Latin word
<emphasis id="B940153CFFCAFFDD3B78DBBAFB4BFB40" box="[1104,1167,1156,1177]" italics="true" pageId="18" pageNumber="357">spina</emphasis>
meaning
<emphasis id="B940153CFFCAFFDD3BD6DBBAFA80FB40" box="[1278,1348,1156,1177]" italics="true" pageId="18" pageNumber="357">spines</emphasis>
in reference to the absence of minute spines at tip of male cercus.
</paragraph>
<paragraph id="8B8BC92EFFCAFFDB3C0EDBCBFEBAFEAA" blockId="18.[806,1423,1269,1904]" lastBlockId="20.[142,758,195,401]" lastPageId="20" lastPageNumber="359" pageId="18" pageNumber="357">
<emphasis id="B940153CFFCAFFDD3C0EDBCBFC6AFAD3" box="[806,942,1269,1290]" italics="true" pageId="18" pageNumber="357">Description:</emphasis>
Blackish-brown species of rather large size with dark brown wing. Scape and pedicel pale brown, first flagellomere lengthened, labium strongly sclerotized, bare, fore tibia with characteristic strong dorsal bristles apically, hind tibia ventrally and dorsally covered with numerous strong, long bristles, cercus without spines at tip, epandrium with pair of long dorsal projections. MALE: HEAD. Occiput black. Ocellar triangle black. Face shiny black to dusty grey in lower part. Palpus blackish. Scape and pedicel dark brown, flagellum black, first flagellomere somewhat conical, second and third flagellomeres aristiform. Proboscis brown to black. Labrum length twice head height. Labium strongly sclerotized with minute bristly hairs. THORAX blackish. All bristles black. Antepronotum with short lateral bristles. Proepisternum and prosternum with 2 distinct bristles. Acrostichals uni- to biserial, fine. Dorsocentrals uni- to biserial, fine, ending with 3 strong, long bristles, 2 of which in prescutellar depression. Anterior and posterior spiracles brown. LEGS dark brown-black, fore and mid femora yellowish-brown. Fore tibia with
<emphasis id="B940153CFFCBFFDC3DF3D979FD28F985" box="[731,748,1607,1628]" italics="true" pageId="19" pageNumber="358">c.</emphasis>
6 strong, long dorsal bristles apically, densely covered with short ventral bristly hairs; fore tarsomeres densely covered with short ventral bristly hairs; first fore tarsomere with long antero- and posterodorsal bristles; first three fore tarsomeres with apical circlet of 3 strong bristles, last three tarsomeres with 2 ventral rows of spine-like short bristles. Mid tibia with dorsal row of 4 strong, long bristles, 1 strong, long ventral at middle; mid tarsomeres with strong spine-like ventrals and apical circlet of strong bristles. Hind femur with 2 ventral rows of fine, rather long bristles; hind tibia enlarged apically, ventrally and dorsally covered with very strong, long bristles especially in apical two-thirds; hind tarsomeres thickened; first hind tarsomere with same pilosity pattern as hind tibia; other tarsomeres with 2 ventral rows of short spine-like bristles. WING (length = 5) deeply brown. A1 complete, feebly sclerotized apically. Halter black to yellowish at base. ABDOMEN dark brown-blackish. Sternite 8 with strong, long posterior bristles. HYPOPYGIUM (
<figureCitation id="130FD5ABFFCCFFDB3E1BDFDCFE40FF21" box="[307,388,226,248]" captionStart="Figure 12" captionStartId="19.[162,240,1489,1508]" captionTargetBox="[338,1266,798,1455]" captionTargetId="figure-232@19.[317,1277,793,1480]" captionTargetPageId="19" captionText="Figure 12. The Empis (Coptophlebia) aspina clade, male of E. (C.) aspina sp. nov. A, epandrium and cercus in lateral view; B, hypandrium and phallus in lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435288" httpUri="https://zenodo.org/record/5435288/files/figure.png" pageId="20" pageNumber="359">Fig. 12</figureCitation>
).
<taxonomicName id="4C34B2ADFFCCFFDB3EB1DFDCFE2CFF2E" box="[409,488,226,247]" class="Insecta" family="Kateretidae" genus="Cercus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="345" phylum="Arthropoda" rank="genus">Cercus</taxonomicName>
without minute spines at tip. Epandrium with pair of pointed anterodorsal projections bearing fine bristles; 4 strong, long bristles at tip. Hypandrium reduced ventrally. Phallus truncate at tip, recurved.
</paragraph>
</subSubSection>
<table id="F9343B8EFFCB00303F8ADFFDFA67FD0E" box="[162,1443,195,727]" pageId="19" pageNumber="358">
<paragraph id="8B8BC92EFFCBFFDC3E43DFEBFA50FEF7" blockId="19.[363,1243,213,237]" lastBlockId="19.[179,1428,254,713]" pageId="19" pageNumber="358">
KEY TO SPECIES OF THE
<taxonomicName id="4C34B2ADFFCBFFDC3DA0DFEBFC2BFF32" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[648,1007,213,237]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="361" phylum="Arthropoda" rank="species" species="aspina" subGenus="Coptophlebia">
<emphasis id="B940153CFFCBFFDC3DA0DFEBFD14FF32" box="[648,720,213,236]" italics="true" pageId="19" pageNumber="358">EMPIS</emphasis>
(
<emphasis id="B940153CFFCBFFDC3DC9DFEBFC4AFF32" box="[737,910,213,237]" italics="true" pageId="19" pageNumber="358">COPTOPHLEBIA</emphasis>
)
<emphasis id="B940153CFFCBFFDC3CB7DFE6FC2BFF32" box="[927,1007,216,235]" italics="true" pageId="19" pageNumber="358">ASPINA</emphasis>
</taxonomicName>
CLADE (MALE ONLY) 1. Legs with pennate bristles (
<bibRefCitation id="EFA5B4DFFFCBFFDC3ED2DFC0FCFCFEC8" author="Daugeron C &amp; Grootaert P" box="[506,824,254,273]" pageId="19" pageNumber="358" pagination="167 - 179" refId="ref32630" refString="Daugeron C, Grootaert P. 2003 a. The Empis (Coptophlebia) hyalea - group from Thailand, with a discussion on the world distribution of this species-group (Diptera: Empididae: Empidinae). European Journal of Entomology 100: 167 - 179." type="journal article" year="2003">Daugeron &amp; Grootaert, 2003a</bibRefCitation>
: fig. 4).................................................................................. 2 Legs without pennate bristles............................................................................................................................................ 3
</paragraph>
<paragraph id="8B8BC92EFFCBFFDC3F9BDE07FA56FE18" blockId="19.[179,1428,254,713]" pageId="19" pageNumber="358">
2. Legs partly yellow; first mid tarsomere with anterolateral pennate bristles; hind tibia with dorsal pennate bristles apically; first hind tarsomere, ventrally and dorsally covered with pennate bristles; mid tibia without pennate bristle ..............................................................................................................
<taxonomicName id="4C34B2ADFFCBFFDC3CA6DE4AFA56FE5F" authority="Daugeron &amp; Grootaert" authorityName="Daugeron &amp; Grootaert" authorityYear="2003" box="[910,1426,371,391]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="358" phylum="Arthropoda" rank="species" species="pulchra" subGenus="Coptophlebia">
<emphasis id="B940153CFFCBFFDC3CA6DE4AFC61FE5F" box="[910,933,372,390]" italics="true" pageId="19" pageNumber="358">E.</emphasis>
(
<emphasis id="B940153CFFCBFFDC3C9ADE4AFC0CFE5E" box="[946,968,372,391]" italics="true" pageId="19" pageNumber="358">C.</emphasis>
)
<emphasis id="B940153CFFCBFFDC3CFEDE4DFBE3FE5F" box="[982,1063,371,390]" italics="true" pageId="19" pageNumber="358">pulchra</emphasis>
Daugeron &amp; Grootaert (
<collectingCountry id="F32389BEFFCBFFDC3A03DE4DFA4FFE5F" box="[1323,1419,371,390]" name="Thailand" pageId="19" pageNumber="358">Thailand</collectingCountry>
)
</taxonomicName>
Legs brown (hind tarsus can be yellowish); only mid tibia with antero- and posteroventral pennate bristles at apical third (
<bibRefCitation id="EFA5B4DFFFCBFFDC3E3DDE90FD97FE18" author="Daugeron C &amp; Grootaert P" box="[277,595,430,449]" pageId="19" pageNumber="358" pagination="167 - 179" refId="ref32630" refString="Daugeron C, Grootaert P. 2003 a. The Empis (Coptophlebia) hyalea - group from Thailand, with a discussion on the world distribution of this species-group (Diptera: Empididae: Empidinae). European Journal of Entomology 100: 167 - 179." type="journal article" year="2003">Daugeron &amp; Grootaert, 2003a</bibRefCitation>
: fig. 4)
<emphasis id="B940153CFFCBFFDC3DB3DE91FCBAFE18" box="[667,894,431,449]" italics="true" pageId="19" pageNumber="358">....................................</emphasis>
<taxonomicName id="4C34B2ADFFCBFFDC3CABDE91FA56FE18" authority="Daugeron &amp; Grootaert" authorityName=", Daugeron &amp; Grootaert" authorityYear="2003" box="[899,1426,430,449]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="358" phylum="Arthropoda" rank="species" species="miranda" subGenus="Coptophlebia">
<emphasis id="B940153CFFCBFFDC3CABDE91FC5EFE18" box="[899,922,431,449]" italics="true" pageId="19" pageNumber="358">E.</emphasis>
(
<emphasis id="B940153CFFCBFFDC3C80DE90FC7AFE18" box="[936,958,430,449]" italics="true" pageId="19" pageNumber="358">C.</emphasis>
)
<emphasis id="B940153CFFCBFFDC3CE3DE90FBE2FE18" box="[971,1062,430,449]" italics="true" pageId="19" pageNumber="358">miranda</emphasis>
Daugeron &amp; Grootaert (
<collectingCountry id="F32389BEFFCBFFDC3A03DE90FA4FFE18" box="[1323,1419,430,449]" name="Thailand" pageId="19" pageNumber="358">Thailand</collectingCountry>
)
</taxonomicName>
</paragraph>
<paragraph id="8B8BC92EFFCBFFDC3F9BDEF5FA50FE25" blockId="19.[179,1428,254,713]" pageId="19" pageNumber="358">
3. Wing clear to feebly dark....................................................................
<taxonomicName id="4C34B2ADFFCBFFDC3CB9DEF2FA56FE07" authority="Daugeron &amp; Grootaert" authorityName="Daugeron &amp; Grootaert" authorityYear="2003" box="[913,1426,459,479]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="358" phylum="Arthropoda" rank="species" species="nganga" subGenus="Coptophlebia">
<emphasis id="B940153CFFCBFFDC3CB9DEF2FC6CFE07" box="[913,936,460,478]" italics="true" pageId="19" pageNumber="358">E.</emphasis>
(
<emphasis id="B940153CFFCBFFDC3C9DDEF2FC0FFE06" box="[949,971,460,479]" italics="true" pageId="19" pageNumber="358">C.</emphasis>
)
<emphasis id="B940153CFFCBFFDC3CF1DEF2FBE3FE07" box="[985,1063,460,478]" italics="true" pageId="19" pageNumber="358">nganga</emphasis>
Daugeron &amp; Grootaert (
<collectingCountry id="F32389BEFFCBFFDC3A03DEF5FA4FFE07" box="[1323,1419,459,478]" name="Thailand" pageId="19" pageNumber="358">Thailand</collectingCountry>
)
</taxonomicName>
Wing brown.......................................................................................................................................................................... 4
</paragraph>
<paragraph id="8B8BC92EFFCBFFDC3F9BDD38FA50FDEF" blockId="19.[179,1428,254,713]" pageId="19" pageNumber="358">
4. Epandrium with a very long dorsal projection (
<figureCitation id="130FD5ABFFCBFFDC3DECDD38FCC9FDC0" box="[708,781,518,537]" captionStart="Figure 14" captionStartId="21.[162,240,1127,1146]" captionTargetBox="[178,760,197,1093]" captionTargetId="figure-593@21.[164,772,181,1122]" captionTargetPageId="21" captionText="Figure 14. The Empis (Coptophlebia) aspina clade, E. (C.) urumae sp. nov. male hypopygium in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435292" httpUri="https://zenodo.org/record/5435292/files/figure.png" pageId="19" pageNumber="358">Fig. 14</figureCitation>
) .............................
<emphasis id="B940153CFFCBFFDC3CFDDD38FB19FDC0" bold="true" box="[981,1245,518,537]" pageId="19" pageNumber="358">
<taxonomicName id="4C34B2ADFFCBFFDC3CFDDD38FBB9FDC0" authority="Daugeron &amp; Grootaert, 2005" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[981,1149,518,537]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="358" phylum="Arthropoda" rank="species" species="urumae" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B940153CFFCBFFDC3CFDDD38FC28FDC0" bold="true" box="[981,1004,518,537]" italics="true" pageId="19" pageNumber="358">E.</emphasis>
(
<emphasis id="B940153CFFCBFFDC3CD3DD38FBD6FDC0" bold="true" box="[1019,1042,518,537]" italics="true" pageId="19" pageNumber="358">C.</emphasis>
)
<emphasis id="B940153CFFCBFFDC3B0ADD38FBB9FDC0" bold="true" box="[1058,1149,518,537]" italics="true" pageId="19" pageNumber="358">urumae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273A847FFCBFFDC3BADDD38FB19FDC0" box="[1157,1245,518,537]" pageId="19" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F32389BEFFCBFFDC3BC4DD38FAEAFDC0" box="[1260,1326,518,537]" name="Japan" pageId="19" pageNumber="358">Japan</collectingCountry>
: Ryukyu) Epandrium with a shorter dorsal projection..................................................................................................................... 5
</paragraph>
<paragraph id="8B8BC92EFFCBFFDC3F9BDD7EFA50FD57" blockId="19.[179,1428,254,713]" pageId="19" pageNumber="358">
5. Fore tibia with 3 short anteroventral spines at middle, phallus with lateral projection (
<figureCitation id="130FD5ABFFCBFFDC3B54DD7FFB05FD8D" box="[1148,1217,577,596]" captionStart="Figure 13" captionStartId="20.[142,220,1827,1846]" captionTargetBox="[203,697,1307,1793]" captionTargetId="figure-642@20.[189,701,1290,1811]" captionTargetPageId="20" captionText="Figure 13. The Empis (Coptophlebia) aspina clade, E. (C.) projecta sp. nov. phallus in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435290" httpUri="https://zenodo.org/record/5435290/files/figure.png" pageId="19" pageNumber="358">Fig. 13</figureCitation>
)............................... .........................................................................................................................
<emphasis id="B940153CFFCBFFDC3CFADD60FB1BFDA8" bold="true" box="[978,1247,606,625]" pageId="19" pageNumber="358">
<taxonomicName id="4C34B2ADFFCBFFDC3CFADD60FBBBFDA8" authority="Daugeron &amp; Grootaert, 2005" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[978,1151,606,625]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="358" phylum="Arthropoda" rank="species" species="projecta" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B940153CFFCBFFDC3CFADD60FC2DFDA8" bold="true" box="[978,1001,606,625]" italics="true" pageId="19" pageNumber="358">E.</emphasis>
(
<emphasis id="B940153CFFCBFFDC3CD1DD60FBD4FDA8" bold="true" box="[1017,1040,606,625]" italics="true" pageId="19" pageNumber="358">C.</emphasis>
)
<emphasis id="B940153CFFCBFFDC3B37DD60FBBBFDA8" bold="true" box="[1055,1151,606,625]" italics="true" pageId="19" pageNumber="358">projecta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273A847FFCBFFDC3BA0DD60FB1BFDA8" box="[1160,1247,606,625]" pageId="19" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F32389BEFFCBFFDC3BC6DD60FAE5FDA8" box="[1262,1313,606,625]" name="Laos" pageId="19" pageNumber="358">Laos</collectingCountry>
,
<collectingCountry id="F32389BEFFCBFFDC3A04DD60FA48FDA8" box="[1324,1420,606,625]" name="Thailand" pageId="19" pageNumber="358">Thailand</collectingCountry>
) Fore tibia only ventrally covered with short bristly hairs, phallus without lateral projection .................................... 6
</paragraph>
<paragraph id="8B8BC92EFFCBFFDC3F9BDDA7FA56FD10" blockId="19.[179,1428,254,713]" pageId="19" pageNumber="358">
6. Fore tibia with a group of strong dorsal bristles apically ..............................................
<emphasis id="B940153CFFCBFFDC3B66DDA7FA8FFD75" bold="true" box="[1102,1355,664,684]" pageId="19" pageNumber="358">
<taxonomicName id="4C34B2ADFFCBFFDC3B66DDA7FB2FFD72" authority="Daugeron &amp; Grootaert, 2005" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[1102,1259,664,684]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="358" phylum="Arthropoda" rank="species" species="aspina" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B940153CFFCBFFDC3B66DDA7FBA1FD75" bold="true" box="[1102,1125,665,684]" italics="true" pageId="19" pageNumber="358">E.</emphasis>
(
<emphasis id="B940153CFFCBFFDC3B5CDDA6FB48FD72" bold="true" box="[1140,1164,664,683]" italics="true" pageId="19" pageNumber="358">C.</emphasis>
)
<emphasis id="B940153CFFCBFFDC3BB3DDA6FB2FFD72" bold="true" box="[1179,1259,664,683]" italics="true" pageId="19" pageNumber="358">aspina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273A847FFCBFFDC3BDBDDA7FA8FFD75" box="[1267,1355,665,684]" pageId="19" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F32389BEFFCBFFDC3A71DDA7FA48FD75" box="[1369,1420,665,684]" name="Laos" pageId="19" pageNumber="358">Laos</collectingCountry>
) Fore tibia without a group of strong dorsal bristles apical
<emphasis id="B940153CFFCBFFDC3C11DD89FCA5FD10" box="[825,865,695,713]" italics="true" pageId="19" pageNumber="358">......</emphasis>
<taxonomicName id="4C34B2ADFFCBFFDC3C49DD89FA56FD10" authority="Daugeron &amp; Grootaert" authorityName="Daugeron &amp; Grootaert" authorityYear="2003" box="[865,1426,694,713]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="358" phylum="Arthropoda" rank="species" species="ratburiensis" subGenus="Coptophlebia">
<emphasis id="B940153CFFCBFFDC3C49DD89FCB3FD10" box="[865,887,695,713]" italics="true" pageId="19" pageNumber="358">E.</emphasis>
(
<emphasis id="B940153CFFCBFFDC3CACDD88FC5FFD10" box="[900,923,694,713]" italics="true" pageId="19" pageNumber="358">C.</emphasis>
)
<emphasis id="B940153CFFCBFFDC3C80DD88FBE3FD10" box="[936,1063,694,713]" italics="true" pageId="19" pageNumber="358">ratburiensis</emphasis>
Daugeron &amp; Grootaert (
<collectingCountry id="F32389BEFFCBFFDC3A03DD88FA4FFD10" box="[1323,1419,694,713]" name="Thailand" pageId="19" pageNumber="358">Thailand</collectingCountry>
)
</taxonomicName>
</paragraph>
</table>
<caption id="DF4B99A6FFCBFFDC3F8ADAEFFC4EF9DB" ID-DOI="http://doi.org/10.5281/zenodo.5435288" ID-Zenodo-Dep="5435288" httpUri="https://zenodo.org/record/5435288/files/figure.png" pageId="19" pageNumber="358" startId="19.[162,240,1489,1508]" targetBox="[338,1266,798,1455]" targetPageId="19">
<paragraph id="8B8BC92EFFCBFFDC3F8ADAEFFC4EF9DB" blockId="19.[162,1442,1489,1538]" pageId="19" pageNumber="358">
<emphasis id="B940153CFFCBFFDC3F8ADAEFFED3FA3D" bold="true" box="[162,279,1489,1508]" pageId="19" pageNumber="358">Figure 12.</emphasis>
The
<taxonomicName id="4C34B2ADFFCBFFDC3E7BDAECFD40FA3D" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[339,644,1489,1509]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="358" phylum="Arthropoda" rank="species" species="aspina" subGenus="Coptophlebia">
<emphasis id="B940153CFFCBFFDC3E7BDAECFE52FA3D" box="[339,406,1490,1508]" italics="true" pageId="19" pageNumber="358">Empis</emphasis>
(
<emphasis id="B940153CFFCBFFDC3E8FDAEFFDF5FA3D" box="[423,561,1489,1508]" italics="true" pageId="19" pageNumber="358">Coptophlebia</emphasis>
)
<emphasis id="B940153CFFCBFFDC3D16DAECFD40FA3D" box="[574,644,1490,1508]" italics="true" pageId="19" pageNumber="358">aspina</emphasis>
</taxonomicName>
clade, male of
<taxonomicName id="4C34B2ADFFCBFFDC3C00DAECFC07FA3D" authority="Daugeron &amp; Grootaert, 2005" authorityName="Daugeron &amp; Grootaert" authorityYear="2005" box="[808,963,1489,1509]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="358" phylum="Arthropoda" rank="species" species="aspina" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B940153CFFCBFFDC3C00DAECFCFBFA3C" bold="true" box="[808,831,1490,1509]" italics="true" pageId="19" pageNumber="358">E.</emphasis>
(
<emphasis id="B940153CFFCBFFDC3C65DAEFFCA0FA3D" bold="true" box="[845,868,1489,1508]" italics="true" pageId="19" pageNumber="358">C.</emphasis>
) aspina
</taxonomicName>
<emphasis id="B940153CFFCBFFDC3CE3DAECFBE0FA3C" bold="true" box="[971,1060,1490,1509]" pageId="19" pageNumber="358">
<taxonomicNameLabel id="A273A847FFCBFFDC3CE3DAECFBE0FA3C" box="[971,1060,1490,1509]" pageId="19" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, epandrium and cercus in lateral view; B, hypandrium and phallus in lateral view. Scale bars = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C32E9AA5FFCCFFDB3F8FDE42FE51FE48" box="[167,405,379,401]" pageId="20" pageNumber="359" type="description">
<paragraph id="8B8BC92EFFCCFFDB3F8FDE42FE51FE48" blockId="20.[142,758,195,401]" box="[167,405,379,401]" pageId="20" pageNumber="359">FEMALE: unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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