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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.114.28799" ID-GBIF-Dataset="175ade4f-e320-4f41-ab85-7811f4147be4" ID-PMC="PMC6323114" ID-Pensoft-Pub="1314-2003-114-123" ID-Pensoft-UUID="CB70FFE82A46DE6DFFEBFFF8F43A6230" ID-PubMed="30627045" ID-Zenodo-Dep="2533527" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-2003-114-123" ModsDocOrigin="PhytoKeys " ModsDocTitle="Revision of Muhlenbergia (Poaceae, Chloridoideae, Cynodonteae, Muhlenbergiinae) in Peru: classification, phylogeny, and a new species, M.romaschenkoi" checkinTime="1553125316505" checkinUser="pensoft" docAuthor="Peterson, Paul M., Vega, Isidoro Sanchez, Romaschenko, Konstantin, Giraldo-Canas, Diego &amp; Rodriguez, Nancy F. Refulio" docDate="2018" docId="FBB293145D8DF5226EA36FAD6347FB3E" docLanguage="en" docName="PhytoKeys 114: 123-206" docOrigin="PhytoKeys 114" docSource="http://dx.doi.org/10.3897/phytokeys.114.28799" docTitle="Muhlenbergia maxima Laegaard &amp; Sanchez Vega, Nordic J. Bot. 10: 439. 1990" docType="treatment" docVersion="6" id="CB70FFE82A46DE6DFFEBFFF8F43A6230" lastPageNumber="160" masterDocId="CB70FFE82A46DE6DFFEBFFF8F43A6230" masterDocTitle="Revision of Muhlenbergia (Poaceae, Chloridoideae, Cynodonteae, Muhlenbergiinae) in Peru: classification, phylogeny, and a new species, M. romaschenkoi" masterLastPageNumber="206" masterPageNumber="123" pageNumber="158" updateTime="1668139139961" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of Muhlenbergia (Poaceae, Chloridoideae, Cynodonteae, Muhlenbergiinae) in Peru: classification, phylogeny, and a new species, M. romaschenkoi</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Peterson, Paul M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vega, Isidoro Sanchez</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Romaschenko, Konstantin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Giraldo-Canas, Diego</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rodriguez, Nancy F. Refulio</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>114</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>123</mods:start>
<mods:end>206</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.114.28799</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.114.28799</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-114-123</mods:identifier>
<mods:identifier type="Pensoft-UUID">CB70FFE82A46DE6DFFEBFFF8F43A6230</mods:identifier>
<mods:identifier type="Zenodo-Dep">2533527</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="154472445" LSID="urn:lsid:plazi:treatment:FBB293145D8DF5226EA36FAD6347FB3E" httpUri="http://treatment.plazi.org/id/FBB293145D8DF5226EA36FAD6347FB3E" lastPageId="37" lastPageNumber="160" pageId="35" pageNumber="158">
<subSubSection pageId="35" pageNumber="158" type="nomenclature">
<paragraph pageId="35" pageNumber="158">
12.
<taxonomicName LSID="23109C2B-5545-51B3-A3A9-CFA20B8FE9A4" authority="Laegaard &amp; Sanchez Vega, Nordic J. Bot. 10: 439. 1990." class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia maxima" order="Poales" pageId="35" pageNumber="158" phylum="Tracheophyta" rank="species" species="maxima">
Muhlenbergia maxima
<normalizedToken originalValue="Lægaard">Laegaard</normalizedToken>
&amp;
<normalizedToken originalValue="Sánchez">Sanchez</normalizedToken>
Vega, Nordic J. Bot. 10: 439. 1990.
</taxonomicName>
<figureCitation captionStart="Figure 2" captionStartId="F3" captionText="Figure 2. A-E Muhlenbergia beyrichiana Kunth A habit B ligule with auricles C glumes D floret E stamens and pistil F-H Muhlenbergia maxima Laegaard &amp; Sanchez Vega F habit G end of panicle branch H floret. A-E drawn from A. S. Hitchcock 20635 (US) F-H drawings from Laegaard &amp; Sanchez Vega (1990) drawn from the holotype collection Sanchez Vega &amp; Ruiz Vigo 3561." figureDoi="10.3897/phytokeys.114.28799.figure2" httpUri="https://binary.pensoft.net/fig/254043" pageId="35" pageNumber="158">Fig. 2F-H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="159" type="materials_examined">
<paragraph pageId="36" pageNumber="159">
<pageBreakToken pageId="36" pageNumber="159" start="start">Type</pageBreakToken>
.
</paragraph>
<paragraph pageId="36" pageNumber="159">
Peru, Prov. Cajamarca, Choten between Paso El
<normalizedToken originalValue="Gavilán">Gavilan</normalizedToken>
and San Juan at km 153 on the road to the coast, 2900 m, 29 May 1984,
<emphasis italics="true" pageId="36" pageNumber="159">
I.
<normalizedToken originalValue="Sánchez">Sanchez</normalizedToken>
Vega &amp; Ruiz Vigo 3561
</emphasis>
(holotype: CPUN!; isotypes: AAU!, K!, US!, USM!).
</paragraph>
</subSubSection>
<subSubSection lastPageId="37" lastPageNumber="160" pageId="36" pageNumber="159" type="description">
<paragraph pageId="36" pageNumber="159">Description.</paragraph>
<paragraph lastPageId="37" lastPageNumber="160" pageId="36" pageNumber="159">
Loosely caespitose
<emphasis italics="true" pageId="36" pageNumber="159">perennials</emphasis>
.
<emphasis italics="true" pageId="36" pageNumber="159">Culms</emphasis>
100-140 cm tall, erect, rigid, terete, ca. 2-3.2 mm diameter near base with 3 or 4 glabrous to pubescent nodes; in
<pageBreakToken pageId="37" pageNumber="160" start="start">ternodes</pageBreakToken>
terete above.
<emphasis italics="true" pageId="37" pageNumber="160">Leaf sheaths</emphasis>
longer than the internodes, mostly basally inserted, compressed-keeled near base, becoming fibrous with age, scaberulous, finely striate;
<emphasis italics="true" pageId="37" pageNumber="160">ligules</emphasis>
3-5 mm long, apex irregularly toothed to lacinate, margins entire and extended above to form auricles;
<emphasis italics="true" pageId="37" pageNumber="160">blades</emphasis>
25-45 cm long, 2.8-4 mm wide, flat to folded, striate, scabrous, apex attenuate, midvein prominent.
<emphasis italics="true" pageId="37" pageNumber="160">Panicles</emphasis>
30-46 cm long, 3-6 cm wide, narrow;
<emphasis italics="true" pageId="37" pageNumber="160">branches</emphasis>
6-11 cm long, scabrous, ascending, appressed to loosely spreading, mostly floriferous to base, except on lowest branches;
<emphasis italics="true" pageId="37" pageNumber="160">pedicels</emphasis>
mostly shorter than the spikelets.
<emphasis italics="true" pageId="37" pageNumber="160">Spikelets</emphasis>
2.5-2.8 mm long, 1-flowered, greenish with reddish-purple tinting;
<emphasis italics="true" pageId="37" pageNumber="160">glumes</emphasis>
2.2-2.6 mm long, almost equal or shorter than the floret, hyaline to membranous, faintly 1-veined, subequal, the lower usually slightly shorter, glabrous to scattered pubescent, scaberulous;
<emphasis italics="true" pageId="37" pageNumber="160">lemmas</emphasis>
2.5-2.8 mm long, ovate, 3-veined, awned, with scattered sericeous hairs more numerous along the veins, scaberulous, the callus pilose, the flexuous awn inserted just below the obtuse to acute apex, the awn 4-9 mm long;
<emphasis italics="true" pageId="37" pageNumber="160">paleas</emphasis>
about as long as the lemma, faintly 2-veined, sericeous between the veins below;
<emphasis italics="true" pageId="37" pageNumber="160">lodicules</emphasis>
about 0.1 mm long, truncate, glabrous;
<emphasis italics="true" pageId="37" pageNumber="160">stamens</emphasis>
3; anthers 1.3-1.8 mm long, purple.
<emphasis italics="true" pageId="37" pageNumber="160">Caryopses</emphasis>
1.4-1.7 mm long, fusiform, brownish.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="160" type="distribution">
<paragraph pageId="37" pageNumber="160">Distribution.</paragraph>
<paragraph pageId="37" pageNumber="160">A Peruvian endemic known only from Amazonas and Cajamarca departments.</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="160" type="ecology">
<paragraph pageId="37" pageNumber="160">Ecology.</paragraph>
<paragraph pageId="37" pageNumber="160">
Rocky slopes near rivers and grassy slopes in disturbed ground along roads associated with
<taxonomicName class="Dicotyledoneae" family="Fabaceae" genus="Acacia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Acacia" order="Rosales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="37" pageNumber="160">Acacia</emphasis>
</taxonomicName>
,
<taxonomicName class="Dicotyledoneae" family="Begoniaceae" genus="Begonia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Begonia" order="Violales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="37" pageNumber="160">Begonia</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Pilg" authorityYear="1906" class="Liliopsida" family="Poaceae" genus="Cortaderia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cortaderia bifida" order="Poales" pageId="37" pageNumber="160" phylum="Magnoliophyta" rank="species" species="bifida">
<emphasis italics="true" pageId="37" pageNumber="160">Cortaderia bifida</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="C. jubata" pageId="37" pageNumber="160" rank="species" species="jubata">
<emphasis italics="true" pageId="37" pageNumber="160">C. jubata</emphasis>
</taxonomicName>
,
<taxonomicName class="Dicotyledoneae" family="Fabaceae" genus="Desmodium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Desmodium" order="Rosales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="37" pageNumber="160">Desmodium</emphasis>
</taxonomicName>
,
<taxonomicName class="Dicotyledoneae" family="Sapindaceae" genus="Dodonaea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Dodonaea viscosa" order="Sapindales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="species" species="viscosa">
<emphasis italics="true" pageId="37" pageNumber="160">Dodonaea viscosa</emphasis>
</taxonomicName>
Jacq.,
<taxonomicName class="Dicotyledoneae" family="Lamiaceae" genus="Hyptis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hyptis" order="Tubiflorae" pageId="37" pageNumber="160" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="37" pageNumber="160">Hyptis</emphasis>
</taxonomicName>
,
<taxonomicName class="Equisetopsida" family="Lamiaceae" genus="Lepechinia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Lepechinia" order="Lamiales" pageId="37" pageNumber="160" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="37" pageNumber="160">Lepechinia</emphasis>
</taxonomicName>
,
<taxonomicName class="Dicotyledoneae" family="Oxalidaceae" genus="Oxalis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Oxalis" order="Geraniales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="37" pageNumber="160">Oxalis</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Molina" authorityYear="1782" class="Liliopsida" family="Bromeliaceae" genus="Puya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Puya" order="Poales" pageId="37" pageNumber="160" phylum="Liliopsida" rank="genus">
<emphasis italics="true" pageId="37" pageNumber="160">Puya</emphasis>
</taxonomicName>
,
<taxonomicName class="Dicotyledoneae" family="Lamiaceae" genus="Salvia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Salvia" order="Tubiflorae" pageId="37" pageNumber="160" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="37" pageNumber="160">Salvia</emphasis>
</taxonomicName>
and
<taxonomicName class="Equisetopsida" family="Poaceae" genus="Schizachyrium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Schizachyrium" order="Poales" pageId="37" pageNumber="160" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="37" pageNumber="160">Schizachyrium</emphasis>
</taxonomicName>
; 2100-2900 m.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="160" type="comments">
<paragraph pageId="37" pageNumber="160">Comments.</paragraph>
<paragraph pageId="37" pageNumber="160">
<taxonomicName class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia maxima" order="Poales" pageId="37" pageNumber="160" phylum="Tracheophyta" rank="species" species="maxima">
<emphasis italics="true" pageId="37" pageNumber="160">Muhlenbergia maxima</emphasis>
</taxonomicName>
is a member of
<taxonomicName authority="subg. Trichochloa" class="Magnoliopsida" family="Poaceae" genus="M." higherTaxonomySource="GBIF" infraspecific-rank="subg." kingdom="Plantae" lsidName="M." order="Poales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="genus">M. subg. Trichochloa</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.3732/ajb.0900359" author="Peterson, PM" journalOrPublisher="American Journal of Botany" pageId="59" pageNumber="182" pagination="1532 - 1554" refId="B38" refString="Peterson, PM, Romaschenko, K, Johnson, G, 2010b. A phylogeny and classification of the Muhlenbergiinae (Poaceae: Chloridoideae: Cynodonteae) based on plastid and nuclear DNA sequences. American Journal of Botany 97 (9): 1532 - 1554, DOI: https://doi.org/10.3732/ajb.0900359" title="A phylogeny and classification of the Muhlenbergiinae (Poaceae: Chloridoideae: Cynodonteae) based on plastid and nuclear DNA sequences." url="https://doi.org/10.3732/ajb.0900359" volume="97" year="2010 b">Peterson et al. 2010b</bibRefCitation>
).
<taxonomicName authorityName="Soderstr" authorityYear="1967" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia inaequalis" order="Poales" pageId="37" pageNumber="160" phylum="Tracheophyta" rank="species" species="inaequalis">
<emphasis italics="true" pageId="37" pageNumber="160">Muhlenbergia inaequalis</emphasis>
</taxonomicName>
Soderstr. endemic to Colombia and Venezuela and
<taxonomicName class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. lehmanniana" order="Poales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="species" species="lehmanniana">
<emphasis italics="true" pageId="37" pageNumber="160">M. lehmanniana</emphasis>
</taxonomicName>
Henr. ranging from Costa Rica to Colombia (
<bibRefCitation DOI="https://doi.org/10.1111/j.1756-1051.1990.tb01786.x" author="Laegaard, S" journalOrPublisher="Nordic Journal of Botany" pageId="58" pageNumber="181" pagination="437 - 441" refId="B18" refString="Laegaard, S, Sanchez Vega, I, 1990. Three new species of Muhlenbergia and Uniola (Poaceae) from northern Peru. Nordic Journal of Botany 10 (4): 437 - 441, DOI: https://doi.org/10.1111/j.1756-1051.1990.tb01786.x" title="Three new species of Muhlenbergia and Uniola (Poaceae) from northern Peru." url="https://doi.org/10.1111/j.1756-1051.1990.tb01786.x" volume="10" year="1990">
<normalizedToken originalValue="Lægaard">Laegaard</normalizedToken>
and
<normalizedToken originalValue="Sánchez">Sanchez</normalizedToken>
Vega 1990
</bibRefCitation>
;
<bibRefCitation author="Giraldo-Canas, D" journalOrPublisher="Caldasia" pageId="58" pageNumber="181" pagination="269 - 302" refId="B7" refString="Giraldo-Canas, D, Peterson, P, 2009. El genero Muhlenbergia (Poaceae: Chloridoideae: Cynodonteae: Muhlenbergiinae) en Colombia. Caldasia 31: 269 - 302" title="El genero Muhlenbergia (Poaceae: Chloridoideae: Cynodonteae: Muhlenbergiinae) en Colombia." volume="31" year="2009">
<normalizedToken originalValue="Giraldo-Cañas">Giraldo-Canas</normalizedToken>
and Peterson 2009
</bibRefCitation>
) are morphologically similar to
<taxonomicName class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. maxima" order="Poales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="species" species="maxima">
<emphasis italics="true" pageId="37" pageNumber="160">M. maxima</emphasis>
</taxonomicName>
.
<taxonomicName class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia maxima" order="Poales" pageId="37" pageNumber="160" phylum="Tracheophyta" rank="species" species="maxima">
<emphasis italics="true" pageId="37" pageNumber="160">Muhlenbergia maxima</emphasis>
</taxonomicName>
differs from
<taxonomicName class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. inaequalis" order="Poales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="species" species="inaequalis">
<emphasis italics="true" pageId="37" pageNumber="160">M. inaequalis</emphasis>
</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. lehmanniana" order="Poales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="species" species="lehmanniana">
<emphasis italics="true" pageId="37" pageNumber="160">M. lehmanniana</emphasis>
</taxonomicName>
in having shorter lemmatal awns and a longer ligule than
<taxonomicName class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. inaequalis" order="Poales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="species" species="inaequalis">
<emphasis italics="true" pageId="37" pageNumber="160">M. inaequalis</emphasis>
</taxonomicName>
(0.5-1.5 mm long) and a shorter ligule than
<taxonomicName class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="M. lehmanniana" order="Poales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="species" species="lehmanniana">
<emphasis italics="true" pageId="37" pageNumber="160">M. lehmanniana</emphasis>
</taxonomicName>
(1-2.5 cm long). There is not much genetic variation among all members of
<taxonomicName authority="subg. Trichochloa" class="Magnoliopsida" family="Poaceae" genus="M." higherTaxonomySource="GBIF" infraspecific-rank="subg." kingdom="Plantae" lsidName="M." order="Poales" pageId="37" pageNumber="160" phylum="Angiospermae" rank="genus">M. subg. Trichochloa</taxonomicName>
in
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2010.01.018" author="Peterson, PM" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="58" pageNumber="181" pagination="580 - 598" refId="B37" refString="Peterson, PM, Romaschenko, K, Johnson, G, 2010a. A classification of the Chloridoideae (Poaceae) based on multi-gene phylogenetic trees. Molecular Phylogenetics and Evolution 55 (2): 580 - 598, DOI: https://doi.org/10.1016/j.ympev.2010.01.018" title="A classification of the Chloridoideae (Poaceae) based on multi-gene phylogenetic trees." url="https://doi.org/10.1016/j.ympev.2010.01.018" volume="55" year="2010 a">Peterson et al. (2010a)</bibRefCitation>
and a detailed population genetic study of these three South American species is needed to interpret their evolutionary history.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="160" type="materials_examined">
<paragraph pageId="37" pageNumber="160">Specimens examined.</paragraph>
<paragraph pageId="37" pageNumber="160">
Peru.
<emphasis bold="true" pageId="37" pageNumber="160">Amazonas</emphasis>
: 27 km from Balsas towards Leymabamba, 2250 m, 23 Mar 1988, S.A. Renvoize &amp; S.
<normalizedToken originalValue="Lægaard">Laegaard</normalizedToken>
4908 (CPUN, K); 66 km from Leymebamba towards Balsas, 2100 m, 25 Mar 1988, S.A. Renvoize &amp; S.
<normalizedToken originalValue="Lægaard">Laegaard</normalizedToken>
4935 (CPUN, K, MO, US).
<emphasis bold="true" pageId="37" pageNumber="160">Cajamarca</emphasis>
: Prov. Cajabamba, 14 air km WSW of Cajabamba and 2.5 km on road W of Araqueda,
<geoCoordinate degrees="7" direction="south" minutes="39" orientation="latitude" precision="1" seconds="9.1" value="-7.652528">7°39'9.1&quot;S</geoCoordinate>
,
<geoCoordinate degrees="78" direction="west" minutes="9" orientation="longitude" precision="1" seconds="58.9" value="-78.16636">78°9'58.9&quot;W</geoCoordinate>
, 2242 m, 23 Mar 2008, P.M. Peterson &amp; R.J. Soreng 21853 (MO, US, USM); Prov. Cajamarca, S of Paso El
<normalizedToken originalValue="Gavelán">Gavelan</normalizedToken>
, ca. 10 air km S of Cajamarca, just above km post 150 on hwy 8 above San Juan,
<geoCoordinate degrees="7" direction="south" minutes="15" orientation="latitude" precision="1" seconds="37.2" value="-7.2603335">7°15'37.2&quot;S</geoCoordinate>
,
<geoCoordinate degrees="78" direction="west" minutes="30" orientation="longitude" precision="1" seconds="34.5" value="-78.50958">78°30'34.5&quot;W</geoCoordinate>
, 2544 m, 26 Mar 2008, P.M. Peterson, R.J. Soreng &amp; I
<normalizedToken originalValue="Sánchez">Sanchez</normalizedToken>
Vega 21884 (US, USM); Dist. San Juan, entre San Juan y Paso el
<normalizedToken originalValue="Gavilán">Gavilan</normalizedToken>
, 2650 m, 30 May 2003, I.
<normalizedToken originalValue="Sánchez">Sanchez</normalizedToken>
Vega 11965, M.
<normalizedToken originalValue="Sánchez">Sanchez</normalizedToken>
M. y R. Cueva R. (CPUN, HAO); Celendin,
<normalizedToken originalValue="Marañón">Maranon</normalizedToken>
River Valley, Chachapoyas-Cajamarca road, 2100 m, 28 May 1984, D.N. Smith &amp; J.M. Cabanillas S. 7271 (MO, US).
</paragraph>
</subSubSection>
</treatment>
</document>