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<document id="8BD787583CA93905717DAF360BBE35C6" ID-CLB-Dataset="39033" ID-DOI="10.11646/zootaxa.3658.1.1" ID-GBIF-Dataset="0adf09ba-5301-4828-abf6-8dbe89f041ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="246551" ID-ZooBank="C02D1C74-25C0-41DD-B098-62098EB7B62A" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1463647879570" checkinUser="plazi" docAuthor="Dragan P. Chobanov, Beata Grzywacz, Ionuţ Ş. Iorgu, Battal Cιplak, Maya B. Ilieva &amp; Elżbieta Warchałowska-Śliwa" docDate="2013" docId="F26F31283923FFBCB1B00CAAFEA29D02" docLanguage="en" docName="zt03658p081.pdf.imf" docOrigin="Zootaxa 3658 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Isophya obtusa Brunner" docType="treatment" docVersion="12" lastPageNumber="47" masterDocId="0E5649503908FF92B1270A03FFD39875" masterDocTitle="Review of the Balkan Isophya (Orthoptera: Phaneropteridae) with particular emphasis on the Isophya modesta group and remarks on the systematics of the genus based on morphological and acoustic data" masterLastPageNumber="81" masterPageNumber="1" pageNumber="44" updateTime="1698662571276" updateUser="ExternalLinkService">
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<mods:title id="4D16017C4EFF48454FE9ECC840311C75">Review of the Balkan Isophya (Orthoptera: Phaneropteridae) with particular emphasis on the Isophya modesta group and remarks on the systematics of the genus based on morphological and acoustic data</mods:title>
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<mods:namePart id="EFE0ABD80BA0E696F4AB434674CBED29">Dragan P. Chobanov</mods:namePart>
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<mods:namePart id="C4FB64150DF8C9AEAF90DC455F35E2E3">Beata Grzywacz</mods:namePart>
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<mods:namePart id="8C28BB38BEA294F3B07F05FECCE38DBB">Ionuţ Ş. Iorgu</mods:namePart>
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<mods:namePart id="89497C446F0B41AD71BF851C3C91EB65">Battal Cιplak</mods:namePart>
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<mods:namePart id="280817A31E30FE37AC3D1439F498C77E">Maya B. Ilieva</mods:namePart>
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<mods:namePart id="BC83678022568E2C38DBCC644E4BACDF">Elżbieta Warchałowska-Śliwa</mods:namePart>
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<treatment id="F26F31283923FFBCB1B00CAAFEA29D02" ID-DOI="http://doi.org/10.5281/zenodo.5617396" ID-GBIF-Taxon="120682315" ID-Zenodo-Dep="5617396" LSID="urn:lsid:plazi:treatment:F26F31283923FFBCB1B00CAAFEA29D02" httpUri="http://treatment.plazi.org/id/F26F31283923FFBCB1B00CAAFEA29D02" lastPageId="46" lastPageNumber="47" pageId="43" pageNumber="44">
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<paragraph id="7A79803E3923FFB9B1B00CAAFD1F9EB6" blockId="43.[151,770,1705,1764]" box="[151,716,1705,1731]" pageId="43" pageNumber="44">
<heading id="213137523923FFB9B1B00CAAFD1F9EB6" bold="true" box="[151,716,1705,1731]" fontSize="11" level="1" pageId="43" pageNumber="44" reason="1">
<emphasis id="48B25C2C3923FFB9B1B00CAAFD1F9EB6" bold="true" box="[151,716,1705,1731]" pageId="43" pageNumber="44">
<taxonomicName id="BDC6FBBD3923FFB9B1B00CAAFE129EB6" authority="Brunner" authorityName="Brunner" box="[151,449,1705,1731]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="obtusa">
<emphasis id="48B25C2C3923FFB9B1B00CAAFE9D9EB6" bold="true" box="[151,334,1705,1731]" italics="true" pageId="43" pageNumber="44">Isophya obtusa</emphasis>
Brunner
</taxonomicName>
von Wattenwyl, 1882
</emphasis>
</heading>
</paragraph>
<paragraph id="7A79803E3923FFB9B1B00CCFFCD19E91" blockId="43.[151,770,1705,1764]" box="[151,770,1740,1764]" pageId="43" pageNumber="44">
(
<figureCitation id="E2FD9CBB3923FFB9B1B80CCFFEF39E91" box="[159,288,1740,1764]" captionStart="FIGURES 19 30" captionStartId="6.[151,264,1632,1655]" captionTargetBox="[154,1433,187,1606]" captionTargetId="figure@6.[151,1436,183,1611]" captionTargetPageId="6" captionText="FIGURES 19 30. General appearance of some Balkan Isophya; 19 — I. modestior (male); 20, 21 — I. dobrogensis (20 — male, 21 — female); 22, 23 — I. zubowskii (22 — male, 23 — female); 24, 25 — I. camptoxypha (24 — male, 25 — female); 26, 27 — I. gulae (26 — male, 27 — female); 28, 29 — I. obtusa (dark— 28, and green— 29 colour forms; males); 30 — I. speciosa (malebelow, and female—above)." httpUri="https://zenodo.org/record/246553/files/figure.png" pageId="43" pageNumber="44">Figs 28, 29</figureCitation>
,
<figureCitation id="E2FD9CBB3923FFB9B00C0CCFFE9A9E91" box="[299,329,1740,1764]" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="43" pageNumber="44">53</figureCitation>
,
<figureCitation id="E2FD9CBB3923FFB9B0720CCEFEA09E91" box="[341,371,1741,1764]" captionStart="FIGURES 56 79" captionStartId="13.[151,264,1769,1792]" captionTargetBox="[178,1421,662,1756]" captionTargetId="figure@13.[178,1422,662,1756]" captionTargetPageId="13" captionText="FIGURES 56 79. Morphology of female head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55 unless specifically given): 56 — I. hospodar; 57 — I. rectipennis; 58 — I. pavelii (paratype of I. rammei Peshev); 59 — I. thracica; 60 — I. bureschi; 61 — I. yaraligozi; 62 — I. tosevski; 63 — I. andreevae (paratype); 64 — I. clara; 65 — I. miksici (paratype); 66 — I. plevnensis (paratype); 67 — I. longicaudata adamovici (paratype, BG: Sliven, 1000 m, NMNHS); 68 — I. l. longicaudata; 69 — I. rhodopensis leonorae (BG: “ Alibotush, 1500 m ”, 13.07. 1959, NMNHS); 70 — I. rh. rhodopensis (as in Fig. 46 but 17.07.1956); 71 — I. rh. petkovi; 72 — I. modestior; 73 — I. dobrogensis; 74 — I. zubowskii; 75 — I. aff. camptoxypha (MK: Bistra Mt., 2.08. 2004, CC); 76 — I. gulae (paratype, BG: Dolna Topchiya, 10.07. 1974, NMNHS); 77 — I. obtusa (BG: Vezhen Peak, 15.07. 1969, NMNHS); 78 — I. amplipennis (TR: “ Turkey | 1960 | Sureya Bey ”, NHM); 79 — I. speciosa (BG: Sakar Mt., 14.06. 1952, NMNHS). Scale (if present; black line right of the specimen) = 10 mm." httpUri="https://zenodo.org/record/246554/files/figure.png" pageId="43" pageNumber="44">77</figureCitation>
,
<figureCitation id="E2FD9CBB3923FFB9B0580CCFFE789E91" box="[383,427,1740,1764]" captionStart="FIGURES 80 104" captionStartId="14.[151,264,1835,1858]" captionTargetBox="[226,1360,681,1815]" captionTargetId="figure@14.[226,1362,681,1816]" captionTargetPageId="14" captionText="FIGURES 80 104. Male abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55): 80 — I. hospodar; 81 — I. rectipennis; 82 — I. pavelii; 83 — I. thracica; 84 — I. bureschi; 85 — I. yaraligozi; 86 — I. tosevski; 87 — I. andreevae; 88 — I. clara; 89 — I. miksici; 90 — I. plevnensis; 91 — I. longicaudata adamovici; 92 — I. l. longicaudata; 93 — I. m. modesta (= intermedia); 94 — I. rhodopensis leonorae; 95 — I. rh. rhodopensis; 96 — I. rh. petkovi; 97 — I. modestior; 98 — I. dobrogensis; 99 — I. zubowskii; 100 — I. aff. camptoxypha; 101 — I. gulae; 102 — I. obtusa; 103 — I. amplipennis; 104 — I. speciosa. Scale (if present) = 2 mm." httpUri="https://zenodo.org/record/246555/files/figure.png" pageId="43" pageNumber="44">102</figureCitation>
,
<figureCitation id="E2FD9CBB3923FFB9B0900CCFFE309E91" box="[439,483,1740,1764]" captionStart="FIGURES 105 129" captionStartId="17.[151,264,1173,1196]" captionTargetBox="[151,1436,195,1152]" captionTargetId="figure@17.[151,1436,195,1152]" captionTargetPageId="17" captionText="FIGURES 105 129. Female abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55 unless specifically given); figures of the whole ovipositor are followed by enlarged figure of the ovipositor base: 105 — I. hospodar (BG: Dolni Glavanak, 25.04. 2005, CC); 106 — I. rectipennis (BG: Byala Vill., 27.06. 2002, CC); 107 — I. pavelii (BG: Chernogorovo, 29.05. 2006, CC); 108 — I. thracica; 109 — I. bureschi (BG: Bansko, 8.08. 2006, CC); 110 — I. yaraligozi; 111 — I. tosevski; 112 — I. andreevae (BG: Kresnensko Hanche, 19.04. 2006, CC); 113 — I. clara; 114 — I. miksici (BG: Gorski Dom Lodge, 24.06. 2006, CC); 115 — I. plevnensis (BG: Levishte, 14.07. 2009, CC); 116 — I. longicaudata adamovici (BG: Karandila, 24.06. 2008, CC); 117 — I. l. longicaudata (BG: Sofia University Botanical garden, 26.06. 1.07.2004, CC); 118 — I. m. modesta (= intermedia); 119 — I. rhodopensis leonorae (BG: Livade, 9.08. 2006, CC); 120 — I. rh. rhodopensis (BG: Bachkovo, 600 m, 23.05. 2004, CC); 121 — I. rh. petkovi (BG: Gluhite Kamani, 23.06. 2008, CC); 122 — I. modestior (BG: Bankya, 24.05. 2006, CC); 123 — I. dobrogensis; 124 — I. zubowskii; 125 — I. aff. camptoxypha (data as in Fig. 75); 126 — I. gulae; 127 — I. obtusa (BG: Ravnets Ridge, 14.07. 2009, CC); 128 — I. amplipennis (as in Fig. 78); 129 — I. speciosa (BG: Bankya, 24.05. 2006, CC). Indications on Fig. 127 show, respectively: G—gonangulum; L—lamella; P—lateral pit between lamella and gonangulum. Scale (if present) = 5 mm." httpUri="https://zenodo.org/record/246556/files/figure.png" pageId="43" pageNumber="44">127</figureCitation>
,
<figureCitation id="E2FD9CBB3923FFB9B0C80CCFFD809E91" box="[495,595,1740,1764]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="43" pageNumber="44">178, 179</figureCitation>
,
<figureCitation id="E2FD9CBB3923FFB9B3780CCFFD109E91" box="[607,707,1740,1764]" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="43" pageNumber="44">188, 189</figureCitation>
,
<figureCitation id="E2FD9CBB3923FFB9B3E80CCFFD299E91" box="[719,762,1740,1764]" captionStart="FIGURE 194" captionStartId="49.[151,250,1945,1968]" captionTargetBox="[151,1434,1141,1924]" captionTargetId="figure@49.[151,1436,1141,1924]" captionTargetPageId="49" captionText="FIGURE 194. Distribution of I. modestior, I. dobrogensis, I. zubowskii, I. aff. camptoxypha, I. gulae, and I. obtusa on the Balkan Peninsula." httpUri="https://zenodo.org/record/246571/files/figure.png" pageId="43" pageNumber="44">194</figureCitation>
)
</paragraph>
<paragraph id="7A79803E3923FFB9B1B00D17FCC59F3C" blockId="43.[151,944,1812,1865]" pageId="43" pageNumber="44">
<taxonomicName id="BDC6FBBD3923FFB9B1B00D17FE599F5F" authority="Brunner" authorityName="Brunner" box="[151,394,1812,1834]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="obtusa">
<emphasis id="48B25C2C3923FFB9B1B00D17FEE29F5F" box="[151,305,1812,1834]" italics="true" pageId="43" pageNumber="44">Isophya obtusa</emphasis>
Brunner
</taxonomicName>
von Wattenwyl: Brunner von Wattenwyl 1882 (sp.n.).
<taxonomicName id="BDC6FBBD3923FFB9B1B00D30FE239F3D" authority="Peshev" authorityName="Peshev" box="[151,496,1842,1865]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subSpecies" species="pravdini" subSpecies="bazyluki">
<emphasis id="48B25C2C3923FFB9B1B00D30FE4D9F3C" box="[151,414,1843,1865]" italics="true" pageId="43" pageNumber="44">Isophya pravdini bazyluki</emphasis>
Peshev
</taxonomicName>
: Peshev 1985 (ssp.n.),
<emphasis id="48B25C2C3923FFB9B3F00D30FCC39F3D" bold="true" box="[727,784,1843,1864]" pageId="43" pageNumber="44">syn.n</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="32DCD3B53923FFBEB1B00D76FBC59D91" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="description">
<paragraph id="7A79803E3923FFB9B1B00D76FE8F9FC4" blockId="43.[151,1436,1909,2006]" pageId="43" pageNumber="44">
<emphasis id="48B25C2C3923FFB9B1B00D76FE159FF8" box="[151,454,1909,1933]" italics="true" pageId="43" pageNumber="44">Morphological description</emphasis>
: see the references above; Bey-Bienko 1954; Harz 1969.
<emphasis id="48B25C2C3923FFB9B54F0D74FB0E9FFB" box="[1128,1245,1911,1934]" italics="true" pageId="43" pageNumber="44">Karyotype</emphasis>
: Warchałowska- Śliwa
<emphasis id="48B25C2C3923FFB9B1FA0D99FEC59FC4" box="[221,278,1945,1969]" italics="true" pageId="43" pageNumber="44">et al.</emphasis>
2008.
</paragraph>
<paragraph id="7A79803E3923FFBEB1E00DBDFCF999FD" blockId="43.[151,1436,1909,2006]" lastBlockId="44.[151,1437,151,1976]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
<emphasis id="48B25C2C3923FFB9B1E00DBDFEEA9FA0" box="[199,313,1982,2005]" italics="true" pageId="43" pageNumber="44">Synonymy</emphasis>
: Peshev (1985) described
<taxonomicName id="BDC6FBBD3923FFB9B3440DBCFCED9FA0" box="[611,830,1981,2006]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subSpecies" species="pravdini" subSpecies="bazyluki">
<emphasis id="48B25C2C3923FFB9B3440DBCFCED9FA0" box="[611,830,1981,2006]" italics="true" pageId="43" pageNumber="44">I. pravdini bazyluki</emphasis>
</taxonomicName>
, comparing it with the other subspecies of
<taxonomicName id="BDC6FBBD3923FFB9B4020DBCFA4F9FA0" box="[1317,1436,1981,2006]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pravdini">
<emphasis id="48B25C2C3923FFB9B4020DBCFA4F9FA0" box="[1317,1436,1981,2006]" italics="true" pageId="43" pageNumber="44">I. pravdini</emphasis>
</taxonomicName>
and thus placing it within the presently regarded
<taxonomicName id="BDC6FBBD3924FFBEB3E60A9AFCE498C5" box="[705,823,152,176]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="48B25C2C3924FFBEB3E60A9AFCE498C5" box="[705,823,152,176]" italics="true" pageId="44" pageNumber="45">I. modesta</emphasis>
</taxonomicName>
group. After the revision has been done the other two subspecies of
<taxonomicName id="BDC6FBBD3924FFBEB0120ABEFE7E98A1" box="[309,429,188,212]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pravdini">
<emphasis id="48B25C2C3924FFBEB0120ABEFE7E98A1" box="[309,429,188,212]" italics="true" pageId="44" pageNumber="45">I. pravdini</emphasis>
</taxonomicName>
became synonyms of related taxa (
<taxonomicName id="BDC6FBBD3924FFBEB2190ABEFC1998A1" box="[830,970,188,212]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="plevnensis">
<emphasis id="48B25C2C3924FFBEB2190ABEFC1998A1" box="[830,970,188,212]" italics="true" pageId="44" pageNumber="45">I. plevnensis</emphasis>
</taxonomicName>
and
<taxonomicName id="BDC6FBBD3924FFBEB5240ABEFB7C98A1" box="[1027,1199,188,212]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="48B25C2C3924FFBEB5240ABEFB7C98A1" box="[1027,1199,188,212]" italics="true" pageId="44" pageNumber="45">I. longicaudata</emphasis>
</taxonomicName>
; see above). Yet, the
<typeStatus id="A57D3E9C3924FFBEB1B00AE3FF14988D" box="[151,199,224,248]" pageId="44" pageNumber="45">type</typeStatus>
series of
<taxonomicName id="BDC6FBBD3924FFBEB0130AE2FE15988D" box="[308,454,224,248]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="pravdini" subSpecies="bazyluki">
<emphasis id="48B25C2C3924FFBEB0130AE2FE15988D" box="[308,454,224,248]" italics="true" pageId="44" pageNumber="45">I. p. bazyluki</emphasis>
</taxonomicName>
has not been found. We visited its
<typeStatus id="A57D3E9C3924FFBEB2750AE3FC51988D" box="[850,898,224,248]" pageId="44" pageNumber="45">type</typeStatus>
locality where, in an isolated spot, an abundant population fitting the description of
<taxonomicName id="BDC6FBBD3924FFBEB3100B06FD1E9969" box="[567,717,260,284]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="pravdini" subSpecies="bazyluki">
<emphasis id="48B25C2C3924FFBEB3100B06FD1E9969" box="[567,717,260,284]" italics="true" pageId="44" pageNumber="45">I. p. bazyluki</emphasis>
</taxonomicName>
was discovered. The animals had the same measurements and tegminal structure as given by Peshev (1985) (though the figures in this paper are quite schematic). After comparison of the morphology and song it appeared obvious that we had collected animals of
<taxonomicName id="BDC6FBBD3924FFBEB59B0B4EFACC9911" box="[1212,1311,332,356]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="obtusa">
<emphasis id="48B25C2C3924FFBEB59B0B4EFACC9911" box="[1212,1311,332,356]" italics="true" pageId="44" pageNumber="45">I. obtusa</emphasis>
</taxonomicName>
and thus
<taxonomicName id="BDC6FBBD3924FFBEB4AB0B4EFE8999FD" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="pravdini" subSpecies="bazyluki">
<emphasis id="48B25C2C3924FFBEB4AB0B4EFE8999FD" italics="true" pageId="44" pageNumber="45">I. pravdini bazyluki</emphasis>
</taxonomicName>
is here considered as its junior synonym.
</paragraph>
<paragraph id="7A79803E3924FFBEB1E00B97FAFF9BBD" blockId="44.[151,1437,151,1976]" pageId="44" pageNumber="45">
<emphasis id="48B25C2C3924FFBEB1E00B97FD6699D9" box="[199,693,404,428]" italics="true" pageId="44" pageNumber="45">Supplement to the description and diagnosis</emphasis>
: The body is moderately stout. Male pronotum (
<figureCitation id="E2FD9CBB3924FFBEB5F60B97FA8299D9" box="[1233,1361,404,428]" captionStart="FIGURES 19 30" captionStartId="6.[151,264,1632,1655]" captionTargetBox="[154,1433,187,1606]" captionTargetId="figure@6.[151,1436,183,1611]" captionTargetPageId="6" captionText="FIGURES 19 30. General appearance of some Balkan Isophya; 19 — I. modestior (male); 20, 21 — I. dobrogensis (20 — male, 21 — female); 22, 23 — I. zubowskii (22 — male, 23 — female); 24, 25 — I. camptoxypha (24 — male, 25 — female); 26, 27 — I. gulae (26 — male, 27 — female); 28, 29 — I. obtusa (dark— 28, and green— 29 colour forms; males); 30 — I. speciosa (malebelow, and female—above)." httpUri="https://zenodo.org/record/246553/files/figure.png" pageId="44" pageNumber="45">Figs 28, 29</figureCitation>
,
<figureCitation id="E2FD9CBB3924FFBEB47A0B97FAA999D9" box="[1373,1402,404,428]" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="44" pageNumber="45">53</figureCitation>
) is saddle-shaped, uplifted in prozone and metazone and constricted in its middle; the lateral keels in metazone are sinuate. Male tegmina (
<figureCitation id="E2FD9CBB3924FFBEB0840BDFFE259981" box="[419,502,476,500]" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="44" pageNumber="45">Fig. 53</figureCitation>
) are equal or slightly longer than pronotum and usually wider than the metazone. The anal edge of left tegmen (at the apical end of CuP) is almost right, frequently protruding over the lateral side of pronotum. CuP is moderately widened and bulged, considerably approximated to CuA, the latter being situated higher than CuP. The stridulatory file (
<figureCitation id="E2FD9CBB3924FFBEB36D0844FD7C9A2A" box="[586,687,583,607]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="44" pageNumber="45">Figs 178</figureCitation>
A, 179A) has dense teeth and a length from
<quantity id="BD3E2DDB3924FFBEB5840844FB2A9A15" box="[1187,1273,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="44" pageNumber="45" unit="mm" value="3.1">3.1 mm</quantity>
with 120125 teeth (specimen from Central Stara Planina Mts) to
<quantity id="BD3E2DDB3924FFBEB225086FFC8E9AF1" box="[770,861,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="44" pageNumber="45" unit="mm" value="3.8">3.8 mm</quantity>
with 161 teeth (specimen from Lyulin Mt.); the
<typeStatus id="A57D3E9C3924FFBEB1B0088CFF2A9AD2" box="[151,249,655,679]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
(“Ilatibor”, ex. coll. Br.v.W., NMW), investigated by Sigfrid Ingrisch, has 137 teeth (according to a label attached to the specimen). Female tegmina (
<figureCitation id="E2FD9CBB3924FFBEB3A208B7FD0E9AB9" box="[645,733,692,716]" captionStart="FIGURES 56 79" captionStartId="13.[151,264,1769,1792]" captionTargetBox="[178,1421,662,1756]" captionTargetId="figure@13.[178,1422,662,1756]" captionTargetPageId="13" captionText="FIGURES 56 79. Morphology of female head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55 unless specifically given): 56 — I. hospodar; 57 — I. rectipennis; 58 — I. pavelii (paratype of I. rammei Peshev); 59 — I. thracica; 60 — I. bureschi; 61 — I. yaraligozi; 62 — I. tosevski; 63 — I. andreevae (paratype); 64 — I. clara; 65 — I. miksici (paratype); 66 — I. plevnensis (paratype); 67 — I. longicaudata adamovici (paratype, BG: Sliven, 1000 m, NMNHS); 68 — I. l. longicaudata; 69 — I. rhodopensis leonorae (BG: “ Alibotush, 1500 m ”, 13.07. 1959, NMNHS); 70 — I. rh. rhodopensis (as in Fig. 46 but 17.07.1956); 71 — I. rh. petkovi; 72 — I. modestior; 73 — I. dobrogensis; 74 — I. zubowskii; 75 — I. aff. camptoxypha (MK: Bistra Mt., 2.08. 2004, CC); 76 — I. gulae (paratype, BG: Dolna Topchiya, 10.07. 1974, NMNHS); 77 — I. obtusa (BG: Vezhen Peak, 15.07. 1969, NMNHS); 78 — I. amplipennis (TR: “ Turkey | 1960 | Sureya Bey ”, NHM); 79 — I. speciosa (BG: Sakar Mt., 14.06. 1952, NMNHS). Scale (if present; black line right of the specimen) = 10 mm." httpUri="https://zenodo.org/record/246554/files/figure.png" pageId="44" pageNumber="45">Figs 77</figureCitation>
,
<figureCitation id="E2FD9CBB3924FFBEB3CF08B7FCC09AB9" box="[744,787,692,716]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="44" pageNumber="45">179</figureCitation>
C) have reticulate venation and are moderately truncate in their apical part.
<collectingCountry id="02D1C0AE3924FFBEB07208D4FE5E9A9A" box="[341,397,727,751]" name="India" pageId="44" pageNumber="45">Hind</collectingCountry>
femora have at least one spine on their internal ventral keel. Male cerci (
<figureCitation id="E2FD9CBB3924FFBEB5E308D4FAF79A9A" box="[1220,1316,727,751]" captionStart="FIGURES 80 104" captionStartId="14.[151,264,1835,1858]" captionTargetBox="[226,1360,681,1815]" captionTargetId="figure@14.[226,1362,681,1816]" captionTargetPageId="14" captionText="FIGURES 80 104. Male abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55): 80 — I. hospodar; 81 — I. rectipennis; 82 — I. pavelii; 83 — I. thracica; 84 — I. bureschi; 85 — I. yaraligozi; 86 — I. tosevski; 87 — I. andreevae; 88 — I. clara; 89 — I. miksici; 90 — I. plevnensis; 91 — I. longicaudata adamovici; 92 — I. l. longicaudata; 93 — I. m. modesta (= intermedia); 94 — I. rhodopensis leonorae; 95 — I. rh. rhodopensis; 96 — I. rh. petkovi; 97 — I. modestior; 98 — I. dobrogensis; 99 — I. zubowskii; 100 — I. aff. camptoxypha; 101 — I. gulae; 102 — I. obtusa; 103 — I. amplipennis; 104 — I. speciosa. Scale (if present) = 2 mm." httpUri="https://zenodo.org/record/246555/files/figure.png" pageId="44" pageNumber="45">Fig. 102</figureCitation>
) are thick, slightly angularly incurved in their apical third (similarly to these of the
<taxonomicName id="BDC6FBBD3924FFBEB2FC08FEFB819B61" box="[987,1106,764,788]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="48B25C2C3924FFBEB2FC08FEFB819B61" box="[987,1106,764,788]" italics="true" pageId="44" pageNumber="45">I. modesta</emphasis>
</taxonomicName>
group and
<taxonomicName id="BDC6FBBD3924FFBEB5F008FEFAB29B61" box="[1239,1377,764,788]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="modestior">
<emphasis id="48B25C2C3924FFBEB5F008FEFAB29B61" box="[1239,1377,764,788]" italics="true" pageId="44" pageNumber="45">I. modestior</emphasis>
</taxonomicName>
) and apically widely obtuse; the apical tooth (
<figureCitation id="E2FD9CBB3924FFBEB35C091CFD0C9B42" box="[635,735,799,823]" captionStart="FIGURES 174 181" captionStartId="36.[151,264,1078,1101]" captionTargetBox="[151,1436,193,1056]" captionTargetId="figure@36.[151,1436,193,1057]" captionTargetPageId="36" captionText="FIGURES 174 181. SEM of taxonomically important structures of Isophya: 174 — I. dobrogensis (RO: Popina Island); 175 — I. zubowskii (RO: Babadag, NMB); 176 — I. camptoxypha (RO: “ Bucec b. Sinaia, 13 1800 m, 25.8. 41, Ramme ”, NMB); 177 — I. gulae (BG: Elhovo, Dolna Topchiya Resrve); 178, 179 — I. obtusa (178 — BG: Lyulin Mt.; 179 — Pleven Lodge); 180 — I. amplipennis (TR: near Sogut); 181 — I. speciosa (BG: Kovach Locality). A—male stridulatory file (174 179 — scale 200 μm; 180, 181 — scale 100 μm); B—ventral / ventro-apical view of apex of male cerci (scale 100 μm); C—female stridulatory apparatus (scale 100 μm; for 175 C— 200 μm)." httpUri="https://zenodo.org/record/246565/files/figure.png" pageId="44" pageNumber="45">Fig. 178</figureCitation>
B) is wide, short and surrounded by dense hairs. The main colouration is dark green to violet-green with brownish-violet tibiae, tegmina and cerci. The disc of tegmina is brownish to brownish-violet with darker CuP and only the apical part of tegmina may be green. The song (
<figureCitation id="E2FD9CBB3924FFBEB44D0964FF289BD1" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="44" pageNumber="45">Figs 188, 189</figureCitation>
) consists of groups of 24 to many (rarely single) gradually decrescending syllables consisting of dense impulses and lasting 100500 ms. Within the groups the syllables are separated by intervals of 1.5
<geoCoordinate id="1FF2E6F93924FFBEB5C809ACFAC09BB2" box="[1263,1299,943,967]" direction="south" orientation="latitude" pageId="44" pageNumber="45" precision="5555" value="-4.5">4.5</geoCoordinate>
s.
</paragraph>
<paragraph id="7A79803E3924FFBEB1E009D6FCCE9C65" blockId="44.[151,1437,151,1976]" pageId="44" pageNumber="45">
<emphasis id="48B25C2C3924FFBEB1E009D6FE859B99" box="[199,342,981,1004]" italics="true" pageId="44" pageNumber="45">Bioacoustics</emphasis>
: The specimens from two separated populations showed some intrapopulation variation and slight interpopulation differentiation of their calling song.
</paragraph>
<paragraph id="7A79803E3924FFBEB1E00E1FFED99D79" blockId="44.[151,1437,151,1976]" pageId="44" pageNumber="45">
The syllables in the song of specimens from Middle Stara Planina Mountains (
<figureCitation id="E2FD9CBB3924FFBEB5490E1FFB009C41" box="[1134,1235,1052,1076]" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="44" pageNumber="45">Fig. 188</figureCitation>
) were frequently separated into a main and an additional part, the latter present or absent within the same recording of an individual. At 2527°С the main part consisted of 4070 dense impulses and lasted 100250 ms (3 33). The impulse period lasted 2.13.2 ms at 25°С to 3.5 ms at 27°С. The additional part (when present) followed the main part at 50170 ms and had 26 impulses (after-clicks) and thus the total syllable length became 250400 ms (3 33). At 28°С the temporal song characteristics of the population from the
<typeStatus id="A57D3E9C3924FFBEB2070ED3FC839C9D" box="[800,848,1232,1256]" pageId="44" pageNumber="45">type</typeStatus>
locality of
<taxonomicName id="BDC6FBBD3924FFBEB2F20ED2FB609C9D" box="[981,1203,1232,1256]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="pravdini" subSpecies="bazyluki">
<emphasis id="48B25C2C3924FFBEB2F20ED2FB609C9D" box="[981,1203,1232,1256]" italics="true" pageId="44" pageNumber="45">I. pravdini bazyluki</emphasis>
</taxonomicName>
fit within the above described.
</paragraph>
<paragraph id="7A79803E3924FFBEB1E00F14FBC59D91" blockId="44.[151,1437,151,1976]" pageId="44" pageNumber="45">
The specimens from Vitosha Mountain showed a song (
<figureCitation id="E2FD9CBB3924FFBEB27E0F14FC6F9D5A" box="[857,956,1303,1327]" captionStart="FIGURES 182 189" captionStartId="38.[151,264,1858,1881]" captionTargetBox="[151,1436,555,1837]" captionTargetId="figure@38.[151,1436,555,1837]" captionTargetPageId="38" captionText="FIGURES 182 189. Male calling song at two different speeds (air temperature at recording shown; time scale below): 182, 183 — I. modestior (182 — BG: Vitosha Mt.; 183 — Serbia: Novi Sad); 184 — I. dobrogensis (RO: Popina Island); 185 — I. zubowskii (RO: Dobrogea, Niculiţel); 186 — I. camptoxypha (RO: Ciucaş Mt.); 187 — I. gulae (BG: Elhovo, Dolna Topchiya Reserve); 188, 189 — I. obtusa (188 — Middle Stara Planina Mts, Pleven Lodge; 189 — BG: Vitosha Mt.). Arrow and a “ Ƥ ” symbol on fig. 187 indicate the response of a female I. gulae." httpUri="https://zenodo.org/record/246566/files/figure.png" pageId="44" pageNumber="45">Fig. 189</figureCitation>
) consisting of compact syllables without clearly detached after-clicks, though in some specimens there was a tendence for separating the last few impulses by intervals of 1080 ms. The syllables lasted 140420 ms (few 33 at 22 and 2526°С), whereas the dependence of the length of the syllable of temperature was masked by the high individual variability in the number of impulses (from 30 to 72 counted). The influence of temperature was more clearly recognised in the impulse period, which in the main part at 22°С had an average length of 5.9 ms and at 2526°С—3.7 ms.
</paragraph>
</subSubSection>
<subSubSection id="32DCD3B53924FFBEB1E00FF3FEF69FCD" pageId="44" pageNumber="45" type="distribution">
<paragraph id="7A79803E3924FFBEB1E00FF3FAE89EE2" blockId="44.[151,1437,151,1976]" pageId="44" pageNumber="45">
<emphasis id="48B25C2C3924FFBEB1E00FF3FDA49E7D" box="[199,631,1520,1544]" italics="true" pageId="44" pageNumber="45">
Distribution (
<figureCitation id="E2FD9CBB3924FFBEB0460FF3FE129E7D" box="[353,449,1520,1544]" captionStart="FIGURE 194" captionStartId="49.[151,250,1945,1968]" captionTargetBox="[151,1434,1141,1924]" captionTargetId="figure@49.[151,1436,1141,1924]" captionTargetPageId="49" captionText="FIGURE 194. Distribution of I. modestior, I. dobrogensis, I. zubowskii, I. aff. camptoxypha, I. gulae, and I. obtusa on the Balkan Peninsula." httpUri="https://zenodo.org/record/246571/files/figure.png" pageId="44" pageNumber="45">Fig. 194</figureCitation>
) and phenology
</emphasis>
: The species is fragmentary distributed in the high parts of Middle and Western Stara Planina Mountains and the low- and middle mointain belt of Lyulin, Vitosha and Plana Mountains between 750 and
<quantity id="BD3E2DDB3924FFBEB0470C34FE659E3A" box="[352,438,1591,1615]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="44" pageNumber="45" unit="m" value="2150.0">2150 m</quantity>
alt. (Central and Western
<collectingCountry id="02D1C0AE3924FFBEB3C50C34FC919E3A" box="[738,834,1591,1615]" name="Bulgaria" pageId="44" pageNumber="45">Bulgaria</collectingCountry>
and extreme Eastern
<collectingCountry id="02D1C0AE3924FFBEB51C0C34FB5B9E3A" box="[1083,1160,1591,1615]" name="Serbia" pageId="44" pageNumber="45">Serbia</collectingCountry>
). In the lower part of its range it inhabits lush humid plant associations in forests (e.g. observed on
<taxonomicName id="BDC6FBBD3924FFBEB2F80C5EFB979E01" box="[991,1092,1629,1652]" class="Magnoliopsida" family="Asteraceae" genus="Petasites" kingdom="Plantae" order="Asterales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="genus">
<emphasis id="48B25C2C3924FFBEB2F80C5EFB979E01" box="[991,1092,1629,1652]" italics="true" pageId="44" pageNumber="45">Petasites</emphasis>
</taxonomicName>
) while at the upper limit of its distribution may be common in pseudosubalpine mesophyte meadows. Nymphs—IVVI, imago—VIXI.
</paragraph>
<paragraph id="7A79803E3924FFBEB1E00CA6FEF69FCD" blockId="44.[151,1437,151,1976]" pageId="44" pageNumber="45">
<emphasis id="48B25C2C3924FFBEB1E00CA6FD529EC9" box="[199,641,1700,1724]" italics="true" pageId="44" pageNumber="45">Notes to the literature distribution data</emphasis>
: Early records for this species refer to a highly diverse material due to incorrect identification. Data for Sarajevo (
<collectingCountry id="02D1C0AE3924FFBEB3A20CC4FC729EAA" box="[645,929,1735,1759]" name="Bosnia and Herzegovina" pageId="44" pageNumber="45">Bosnia and Herzegovina</collectingCountry>
) by Burr (1898) (cited by Us and Matvejev 1967) concern
<taxonomicName id="BDC6FBBD3924FFBEB0640CEEFE4B9F71" box="[323,408,1772,1796]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="clara">
<emphasis id="48B25C2C3924FFBEB0640CEEFE4B9F71" box="[323,408,1772,1796]" italics="true" pageId="44" pageNumber="45">I. clara</emphasis>
</taxonomicName>
. The records for
<collectingCountry id="02D1C0AE3924FFBEB3440CEFFD1B9F71" box="[611,712,1772,1796]" name="Romania" pageId="44" pageNumber="45">Romania</collectingCountry>
(Frey-Gessner 1897; Burr 1899) are also very probably wrong though it is unclear which species is/are concerned in these cases. Data by Ramme (1951) citing Retowskii (1889) for
<collectingCountry id="02D1C0AE3924FFBEB1980D37FECC9F39" box="[191,287,1844,1868]" name="Turkey" pageId="44" pageNumber="45">Anatolia</collectingCountry>
(Sinope; Samsun) have been referred to
<taxonomicName id="BDC6FBBD3924FFBEB3D70D36FC429F39" box="[752,913,1844,1868]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="amplipennis">
<emphasis id="48B25C2C3924FFBEB3D70D36FC429F39" box="[752,913,1844,1868]" italics="true" pageId="44" pageNumber="45">I. amplipennis</emphasis>
</taxonomicName>
by Bey-Bienko (1954). The record “Zlatibor” (Brunner von Wattenwyl 1882; Adamoviċ 1975) referred to Zlatibor Mt. in Western
<collectingCountry id="02D1C0AE3924FFBEB5650D54FB5A9F1A" box="[1090,1161,1879,1903]" name="Serbia" pageId="44" pageNumber="45">Serbia</collectingCountry>
concerns wrong reading the label by Pančiċ where the actual typing is “Ilatibor”. The latter should be referred to a place in Western Stara Planina Mts.
</paragraph>
</subSubSection>
<caption id="2EB9D0B63925FFBFB1B00E28FF269C0A" httpUri="https://zenodo.org/record/246567/files/figure.png" pageId="45" pageNumber="46" targetBox="[151,1436,195,1046]" targetPageId="45">
<paragraph id="7A79803E3925FFBFB1B00E28FF269C0A" blockId="45.[151,1436,1067,1151]" pageId="45" pageNumber="46">
<emphasis id="48B25C2C3925FFBFB1B00E28FEFE9C34" bold="true" box="[151,301,1067,1090]" pageId="45" pageNumber="46">FIGURE 190.</emphasis>
Male calling song of
<taxonomicName id="BDC6FBBD3925FFBFB32B0E2EFDAA9C37" box="[524,633,1069,1090]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="48B25C2C3925FFBFB32B0E2EFDAA9C37" box="[524,633,1069,1090]" italics="true" pageId="45" pageNumber="46">I. speciosa</emphasis>
</taxonomicName>
(BG: Vitosha Mt., Zheleznitsa) at two different speeds (air temperature during recording 24ºC; time scale below). Break-signs in Fig. 190B indicate the place of interruption of the syllable (no part of it is missing).
</paragraph>
</caption>
<subSubSection id="32DCD3B53925FFBCB1B00EA4FEA29D02" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
<paragraph id="7A79803E3925FFBFB1B00EA4FDB99CCA" blockId="45.[151,1437,1190,2007]" box="[151,618,1190,1215]" pageId="45" pageNumber="46">
<emphasis id="48B25C2C3925FFBFB1B00EA4FDB99CCA" bold="true" box="[151,618,1190,1215]" pageId="45" pageNumber="46">
III.
<taxonomicName id="BDC6FBBD3925FFBFB1E20EA5FE5B9CCA" box="[197,392,1190,1215]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="48B25C2C3925FFBFB1E20EA5FE5B9CCA" bold="true" box="[197,392,1190,1215]" italics="true" pageId="45" pageNumber="46">Isophya speciosa</emphasis>
</taxonomicName>
species group (s.l.)
</emphasis>
</paragraph>
<paragraph id="7A79803E3925FFBFB1B00EC6FC569EE7" blockId="45.[151,1437,1190,2007]" pageId="45" pageNumber="46">
Warchałowska-Śliwa
<emphasis id="48B25C2C3925FFBFB0AB0EC8FE159C97" box="[396,454,1226,1250]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
(2008) outlined the
<taxonomicName id="BDC6FBBD3925FFBFB3970EC8FCEF9C97" box="[688,828,1226,1250]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="48B25C2C3925FFBFB3970EC8FCEF9C97" box="[688,828,1226,1250]" italics="true" pageId="45" pageNumber="46">I. schneideri</emphasis>
</taxonomicName>
group for a few morphologically similar taxa though they do not give the exact content of the group. Ünal (2010) split this grouping into
<taxonomicName id="BDC6FBBD3925FFBFB5420EF3FB209D73" box="[1125,1267,1262,1287]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="48B25C2C3925FFBFB5420EF3FB209D73" box="[1125,1267,1262,1287]" italics="true" pageId="45" pageNumber="46">I. schneideri</emphasis>
</taxonomicName>
group (with
<emphasis id="48B25C2C3925FFBFB4AB0EF3FF089D5F" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="BDC6FBBD3925FFBFB4AB0EF3FF0B9D5F" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="cania">I. cania</taxonomicName>
,
</emphasis>
<taxonomicName id="BDC6FBBD3925FFBFB1C30F10FDC99D5F" authority="Karabag, 1962" authorityName="Karabag" authorityYear="1962" box="[228,538,1298,1322]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="hakkarica">
<emphasis id="48B25C2C3925FFBFB1C30F10FEBF9D5F" box="[228,364,1298,1322]" italics="true" pageId="45" pageNumber="46">I. hakkarica</emphasis>
Karabag, 1962
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3925FFBFB30D0F10FC859D5F" authority="Uvarov, 1940" authorityName="Uvarov" authorityYear="1940" box="[554,854,1298,1322]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="karabaghi">
<emphasis id="48B25C2C3925FFBFB30D0F10FD669D5F" box="[554,693,1298,1322]" italics="true" pageId="45" pageNumber="46">I. karabaghi</emphasis>
Uvarov, 1940
</taxonomicName>
,
<emphasis id="48B25C2C3925FFBFB2420F10FC2A9D5F" box="[869,1017,1298,1322]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="BDC6FBBD3925FFBFB2420F10FC269D5F" box="[869,1013,1298,1322]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="schneideri">I. schneideri</taxonomicName>
,
</emphasis>
<taxonomicName id="BDC6FBBD3925FFBFB5260F10FADD9D5F" authority="Ramme, 1951" authorityName="Ramme" authorityYear="1951" box="[1025,1294,1298,1322]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="sikorai">
<emphasis id="48B25C2C3925FFBFB5260F10FBBB9D5F" box="[1025,1128,1298,1322]" italics="true" pageId="45" pageNumber="46">I. sikorai</emphasis>
Ramme, 1951
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3925FFBFB43A0F10FA419D5F" box="[1309,1426,1298,1322]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="thracica">
<emphasis id="48B25C2C3925FFBFB43A0F10FA419D5F" box="[1309,1426,1298,1322]" italics="true" pageId="45" pageNumber="46">I. thracica</emphasis>
</taxonomicName>
) and
<taxonomicName id="BDC6FBBD3925FFBFB1E00F3BFEBB9D3B" box="[199,360,1334,1359]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="amplipennis">
<emphasis id="48B25C2C3925FFBFB1E00F3BFEBB9D3B" box="[199,360,1334,1359]" italics="true" pageId="45" pageNumber="46">I. amplipennis</emphasis>
</taxonomicName>
group (with
<emphasis id="48B25C2C3925FFBFB0DB0F3BFD709D3B" box="[508,675,1334,1359]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="BDC6FBBD3925FFBFB0DB0F3BFD739D3B" box="[508,672,1334,1359]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="amplipennis">I. amplipennis</taxonomicName>
,
</emphasis>
<taxonomicName id="BDC6FBBD3925FFBFB38C0F3BFC5F9D3B" authority="Unal, 2010" authorityName="Unal" authorityYear="2010" box="[683,908,1334,1359]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="artvin">
<emphasis id="48B25C2C3925FFBFB38C0F3BFCD49D3B" box="[683,775,1335,1359]" italics="true" pageId="45" pageNumber="46">I. artvin</emphasis>
Ünal, 2010
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3925FFBFB2BD0F3BFB709D3B" authority="Miram, 1938" authorityName="Miram" authorityYear="1938" box="[922,1187,1334,1359]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="48B25C2C3925FFBFB2BD0F3BFBDA9D3B" box="[922,1033,1334,1359]" italics="true" pageId="45" pageNumber="46">I. gracilis</emphasis>
Miram, 1938
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3925FFBFB5950F3BFAAE9D3B" authority="Unal, 2010" authorityName="Unal" authorityYear="2010" box="[1202,1405,1334,1359]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="hitit">
<emphasis id="48B25C2C3925FFBFB5950F3BFB249D3B" box="[1202,1271,1334,1359]" italics="true" pageId="45" pageNumber="46">I. hitit</emphasis>
Ünal, 2010
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3925FFBFB4AB0F3BFE319D07" authority="Adelung, 1907" authorityName="Adelung" authorityYear="1907" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="redtenbacheri">
<emphasis id="48B25C2C3925FFBFB4AB0F3BFEE09D07" italics="true" pageId="45" pageNumber="46">I. redtenbacheri</emphasis>
Adelung, 1907
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3925FFBFB0D60F58FCF29D07" authority="Bolivar, 1899" authorityName="Bolivar" authorityYear="1899" box="[497,801,1370,1394]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rodsjankoi">
<emphasis id="48B25C2C3925FFBFB0D60F58FD539D07" box="[497,640,1370,1394]" italics="true" pageId="45" pageNumber="46">I. rodsjankoi</emphasis>
Bolívar, 1899
</taxonomicName>
,
<emphasis id="48B25C2C3925FFBFB2170F58FB129D07" box="[816,1217,1370,1394]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="BDC6FBBD3925FFBFB2170F58FC7B9D07" box="[816,936,1371,1394]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="savignyi">I. savignyi</taxonomicName>
,
<taxonomicName id="BDC6FBBD3925FFBFB2920F58FBE39D07" box="[949,1072,1371,1394]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="speciosa">I. speciosa</taxonomicName>
,
<taxonomicName id="BDC6FBBD3925FFBFB51B0F58FB129D07" authority="Naskrecki et Unal, 1995" authorityName="Naskrecki et Unal" authorityYear="1995" box="[1084,1217,1370,1394]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="splendida">I. splendida</taxonomicName>
</emphasis>
Naskrecki et Ünal, 1995,
<taxonomicName id="BDC6FBBD3925FFBFB1FB0F83FE3F9DE3" authority="Ramme, 1951" authorityName="Ramme" authorityYear="1951" box="[220,492,1406,1431]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="sureyai">
<emphasis id="48B25C2C3925FFBFB1FB0F83FE979DE3" box="[220,324,1407,1431]" italics="true" pageId="45" pageNumber="46">I. sureyai</emphasis>
Ramme, 1951
</taxonomicName>
). However, we reconsidered the relationships of few taxa (see above) and related the species
<taxonomicName id="BDC6FBBD3925FFBFB1D60FA0FE9A9DCF" box="[241,329,1443,1466]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="cania">
<emphasis id="48B25C2C3925FFBFB1D60FA0FE9A9DCF" box="[241,329,1443,1466]" italics="true" pageId="45" pageNumber="46">I. cania</emphasis>
</taxonomicName>
and
<taxonomicName id="BDC6FBBD3925FFBFB0A30FA0FE289DCF" box="[388,507,1442,1466]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="thracica">
<emphasis id="48B25C2C3925FFBFB0A30FA0FE289DCF" box="[388,507,1442,1466]" italics="true" pageId="45" pageNumber="46">I. thracica</emphasis>
</taxonomicName>
to the
<taxonomicName id="BDC6FBBD3925FFBFB3680FA0FD379DCF" box="[591,740,1443,1466]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rectipennis">
<emphasis id="48B25C2C3925FFBFB3680FA0FD379DCF" box="[591,740,1443,1466]" italics="true" pageId="45" pageNumber="46">I. rectipennis</emphasis>
</taxonomicName>
group.
<taxonomicName id="BDC6FBBD3925FFBFB2670FA0FC7B9DCF" box="[832,936,1442,1466]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="sikorai">
<emphasis id="48B25C2C3925FFBFB2670FA0FC7B9DCF" box="[832,936,1442,1466]" italics="true" pageId="45" pageNumber="46">I. sikorai</emphasis>
</taxonomicName>
should also be excluded from
<taxonomicName id="BDC6FBBD3925FFBFB42A0FA0FA489DCF" box="[1293,1435,1442,1466]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="48B25C2C3925FFBFB42A0FA0FA489DCF" box="[1293,1435,1442,1466]" italics="true" pageId="45" pageNumber="46">I. schneideri</emphasis>
</taxonomicName>
group on account of its morphological position between
<taxonomicName id="BDC6FBBD3925FFBFB2360FCBFC579DAB" box="[785,900,1478,1503]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="straubei">
<emphasis id="48B25C2C3925FFBFB2360FCBFC579DAB" box="[785,900,1478,1503]" italics="true" pageId="45" pageNumber="46">I. straubei</emphasis>
</taxonomicName>
and
<taxonomicName id="BDC6FBBD3925FFBFB29C0FCBFBC69DAB" box="[955,1045,1479,1503]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="major">
<emphasis id="48B25C2C3925FFBFB29C0FCBFBC69DAB" box="[955,1045,1479,1503]" italics="true" pageId="45" pageNumber="46">I. major</emphasis>
</taxonomicName>
groups (especially considering the wide fastigium verticis, long tegmina, widely separated CuA and CuP, parallel venation of female tegmina, shape of male subgenital plate and short hind femora with high number of ventral spines); a study of the song is necessary for more precise evaluation of its relationships. Finally, the position of
<taxonomicName id="BDC6FBBD3925FFBFB2ED0C30FB859E3F" box="[970,1110,1586,1610]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="karabaghi">
<emphasis id="48B25C2C3925FFBFB2ED0C30FB859E3F" box="[970,1110,1586,1610]" italics="true" pageId="45" pageNumber="46">I. karabaghi</emphasis>
</taxonomicName>
is also under question since according to the male and female tegminal morphology and male cerci the latter is related to
<taxonomicName id="BDC6FBBD3925FFBFB5830C5BFF019EE7" authority="Karabag, 1957" authorityName="Karabag" authorityYear="1957" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="bicarinata">
<emphasis id="48B25C2C3925FFBFB5830C5BFAFC9E1B" box="[1188,1327,1622,1647]" italics="true" pageId="45" pageNumber="46">I. bicarinata</emphasis>
Karabag, 1957
</taxonomicName>
, a species placed by Ünal in another group (
<taxonomicName id="BDC6FBBD3925FFBFB3E50C78FCF89EE7" box="[706,811,1659,1682]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="zernovi">
<emphasis id="48B25C2C3925FFBFB3E50C78FCF89EE7" box="[706,811,1659,1682]" italics="true" pageId="45" pageNumber="46">I. zernovi</emphasis>
</taxonomicName>
group).
</paragraph>
<paragraph id="7A79803E3925FFBFB1E00C9CFD6B9E8B" blockId="45.[151,1437,1190,2007]" pageId="45" pageNumber="46">
In
<taxonomicName id="BDC6FBBD3925FFBFB1C00CA3FE5A9EC3" box="[231,393,1694,1719]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="amplipennis">
<emphasis id="48B25C2C3925FFBFB1C00CA3FE5A9EC3" box="[231,393,1694,1719]" italics="true" pageId="45" pageNumber="46">I. amplipennis</emphasis>
</taxonomicName>
group Ünal (2010) included only the taxa occurring in
<collectingCountry id="02D1C0AE3925FFBFB5200C9DFB849EC3" box="[1031,1111,1694,1718]" name="Turkey" pageId="45" pageNumber="46">Turkey</collectingCountry>
while significant number of related species remains unconsidered. Thus, including all morphologically similar species we propose to use the name
<taxonomicName id="BDC6FBBD3925FFBFB1FD0CEBFE819E8B" box="[218,338,1767,1791]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="48B25C2C3925FFBFB1FD0CEBFE819E8B" box="[218,338,1767,1791]" italics="true" pageId="45" pageNumber="46">I. speciosa</emphasis>
</taxonomicName>
, the latter being published first.
</paragraph>
<paragraph id="7A79803E3925FFBCB1E00D09FCD19C2D" blockId="45.[151,1437,1190,2007]" lastBlockId="46.[151,1436,151,1399]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<taxonomicName id="BDC6FBBD3925FFBFB1E00D09FE5A9F57" box="[199,393,1802,1826]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="48B25C2C3925FFBFB1E00D09FE5A9F57" box="[199,393,1802,1826]" italics="true" pageId="45" pageNumber="46">Isophya speciosa</emphasis>
</taxonomicName>
group includes high number of taxa expressing the most specialised characters within this genus. The species have small to moderate body size for the genus and slender appearance.
<collectingCountry id="02D1C0AE3925FFBFB5FB0D2DFAC79F33" box="[1244,1300,1838,1862]" name="India" pageId="45" pageNumber="46">Hind</collectingCountry>
femora are usually shorter than
<quantity id="BD3E2DDB3925FFBFB0AF0D51FE0C9F1F" box="[392,479,1874,1898]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="45" pageNumber="46" unit="mm" value="18.0">18 mm</quantity>
. The fastigium is very narrow, less than 1/2 to 1/4 of the width of scapus. Male pronotum is constricted in the middle, widened and raised in pro- and metazone, sometimes clearly saddle-shaped. Male tegmina show smooth transition from a primitive state for the group (mostly in the primitive forms, e.g.
<emphasis id="48B25C2C3925FFBFB4AB0D98FE479FA3" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="BDC6FBBD3925FFBFB4AB0D98FF249FA3" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="savignyi">I. savignyi</taxonomicName>
,
<taxonomicName id="BDC6FBBD3925FFBFB0230DC3FE479FA3" box="[260,404,1982,2007]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rodsjankoi">I. rodsjankoi</taxonomicName>
</emphasis>
and
<taxonomicName id="BDC6FBBD3925FFBFB0EB0DC3FDE59FA3" box="[460,566,1983,2007]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="sureyai">
<emphasis id="48B25C2C3925FFBFB0EB0DC3FDE59FA3" box="[460,566,1983,2007]" italics="true" pageId="45" pageNumber="46">I. sureyai</emphasis>
</taxonomicName>
)—equal or slightly longer and hardly wider than pronotum, slightly bulged, with flat disc and well separated CuA and CuP (or even almost invisible smooth CuA), to a progressive state (in some northeastern and the northwestern forms, e.g.
<emphasis id="48B25C2C3926FFBCB3CF0ABEFBC098A1" box="[744,1043,188,212]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="BDC6FBBD3926FFBCB3CF0ABEFCB798A1" box="[744,868,189,212]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="speciosa">I. speciosa</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB2560ABEFBC098A1" box="[881,1043,188,212]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="amplipennis">I. amplipennis</taxonomicName>
</emphasis>
)—strongly bulged tegmina, wider and clearly longer than pronotum (up to 2 times longer in
<taxonomicName id="BDC6FBBD3926FFBCB2140AE2FC1C988D" box="[819,975,224,248]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="kalishevskii">
<emphasis id="48B25C2C3926FFBCB2140AE2FC1C988D" box="[819,975,224,248]" italics="true" pageId="46" pageNumber="47">I. kalishevskii</emphasis>
</taxonomicName>
), with sharp venation and approximated CuP and CuA (due to a fold in the area between these veins and uplifting the CuA vein). CuP is thin (about as wide as the third antennal limb) and long (usually over 3/4 of the hind margin of metazone). The stridulatory file has 75 180 teeth (usually about 100); the teeth gradually increase in size towards the apex of file being fine at its base and middle. Female (as well as male) tegmina have clearly developed coarse reticulate venation; their apical part is widely rounded or, rarely, slightly blunt. The lower keels of hind femora do not possess spines. The width and curvature of male cerci vary in different species; the apical spine may be apically or subapically situated, thin or thick, pointed. The ovipositor is very short, shorter than
<quantity id="BD3E2DDB3926FFBCB20E0BDFFCA89981" box="[809,891,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="mm" value="10.0">10 mm</quantity>
(usually less than
<quantity id="BD3E2DDB3926FFBCB57F0BDFFB739981" box="[1112,1184,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="46" pageNumber="47" unit="mm" value="9.0">9 mm</quantity>
). The pit between the lamella and gonangulum is usually moderately closed by the lamella; the latter may have or have not an excision apically. The body colouration may be green or variable; very typical for the group is developing of melanistic forms with different share of black. The green forms have reddish stripes at the lateral margins of metazone situated above the light band, while in the melanistic forms this stripe is masked by black and red spots. The colouration of male tegmina usually differs from the main body colour and may be variable with yellow, white, red, etc. The calling song consists of highly elaborated single syllables. The song producing movement is generally separated to two consecutive incomplete open-and-closing movements (cf. Heller 1988, 1990) resulting in a syllable consisting of two or more parts. The syllables of the eastern representatives (
<emphasis id="48B25C2C3926FFBCB5A908FEFA589B61" box="[1166,1419,764,788]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="BDC6FBBD3926FFBCB5A908FEFAD99B61" box="[1166,1290,765,788]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="savignyi">I. savignyi</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB43D08FEFA589B61" box="[1306,1419,764,788]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="gracilis">I. gracilis</taxonomicName>
</emphasis>
), corresponding to a more primitive habitus, have short first part and crescending/crescending-decrescending second part, altogether lasting about 500 ms (27.5°С in
<taxonomicName id="BDC6FBBD3926FFBCB3F50946FC999B29" box="[722,842,837,860]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis id="48B25C2C3926FFBCB3F50946FC999B29" box="[722,842,837,860]" italics="true" pageId="46" pageNumber="47">I. savignyi</emphasis>
</taxonomicName>
; possibly up to one second at lower temperatures) (Heller 1990). In the northwestern taxa (
<emphasis id="48B25C2C3926FFBCB34F096AFC469BF5" box="[616,917,872,896]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="BDC6FBBD3926FFBCB34F096AFD369BF5" box="[616,741,873,896]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="speciosa">I. speciosa</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB3D5096AFC469BF5" box="[754,917,872,896]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="amplipennis">I. amplipennis</taxonomicName>
</emphasis>
), showing more progressive morphology, the syllables have three (or even four in
<taxonomicName id="BDC6FBBD3926FFBCB373098EFD029BD1" box="[596,721,909,932]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="48B25C2C3926FFBCB373098EFD029BD1" box="[596,721,909,932]" italics="true" pageId="46" pageNumber="47">I. speciosa</emphasis>
</taxonomicName>
) recognisable parts, produced by distinct slowing down and pausing of each of the two closing movements of the syllable (compare Heller 1988: Abb. 33); altogether the syllable may last three to over five seconds. The X-chromosome is acro- or subacrocentric (
<typeStatus id="A57D3E9C3926FFBCB5CD09D7FACF9B99" box="[1258,1308,980,1004]" pageId="46" pageNumber="47">type</typeStatus>
1B or 2A according to Warchałowska-Śliwa
<emphasis id="48B25C2C3926FFBCB33909FAFD849C65" box="[542,599,1016,1040]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
2008). At least two complexes may be separated within the group, though their exact composition is still ambiguous and needs further investigation (especially of the song structures). The group is most closely related to the
<taxonomicName id="BDC6FBBD3926FFBCB3010E42FD619C2D" box="[550,690,1088,1112]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="48B25C2C3926FFBCB3010E42FD619C2D" box="[550,690,1088,1112]" italics="true" pageId="46" pageNumber="47">I. schneideri</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="7A79803E3926FFBCB1E00E66FEA29D02" blockId="46.[151,1436,151,1399]" pageId="46" pageNumber="47">
<taxonomicName id="BDC6FBBD3926FFBCB1E00E66FE939C09" box="[199,320,1125,1148]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="48B25C2C3926FFBCB1E00E66FE939C09" box="[199,320,1125,1148]" italics="true" pageId="46" pageNumber="47">I. speciosa</emphasis>
</taxonomicName>
group as here considered in a broad sense envelops the highest number of taxa within the genus, distributed from the Caucasus throughout
<collectingCountry id="02D1C0AE3926FFBCB3490E84FD1C9CEA" box="[622,719,1159,1183]" name="Turkey" pageId="46" pageNumber="47">Anatolia</collectingCountry>
to the Balkan Peninsula. About 20 taxa and several groups (e.g. Ünal 2010: partim) are presently considered within the group, most of them occurring in
<collectingCountry id="02D1C0AE3926FFBCB5520EAFFB0B9CB1" box="[1141,1240,1196,1220]" name="Turkey" pageId="46" pageNumber="47">Anatolia</collectingCountry>
, with two species reaching the Balkan Peninsula:
<taxonomicName id="BDC6FBBD3926FFBCB30C0ED2FD179C9D" box="[555,708,1233,1256]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="acuminata">
<emphasis id="48B25C2C3926FFBCB30C0ED2FD179C9D" box="[555,708,1233,1256]" italics="true" pageId="46" pageNumber="47">I. acuminata</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB3F80ED2FC589C9D" box="[735,907,1232,1256]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="amplipennis">
<emphasis id="48B25C2C3926FFBCB3F80ED2FC589C9D" box="[735,907,1232,1256]" italics="true" pageId="46" pageNumber="47">I. amplipennis</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB2820ED2FBDF9C9D" box="[933,1036,1233,1256]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="artvin">
<emphasis id="48B25C2C3926FFBCB2820ED2FBDF9C9D" box="[933,1036,1233,1256]" italics="true" pageId="46" pageNumber="47">I. artvin</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB5010ED2FADB9C9D" box="[1062,1288,1233,1256]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="subSpecies" species="caspica" subSpecies="caspica">
<emphasis id="48B25C2C3926FFBCB5010ED2FADB9C9D" box="[1062,1288,1233,1256]" italics="true" pageId="46" pageNumber="47">I. caspica caspica</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB4040ED2FE9E9D79" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="subSpecies" species="caspica" subSpecies="stshelkanovtzevi">
<emphasis id="48B25C2C3926FFBCB4040ED2FE9E9D79" italics="true" pageId="46" pageNumber="47">I. caspica stshelkanovtzevi</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB0780EF6FE1C9D79" box="[351,463,1268,1292]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="48B25C2C3926FFBCB0780EF6FE1C9D79" box="[351,463,1268,1292]" italics="true" pageId="46" pageNumber="47">I. gracilis</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB0C50EF6FDF99D79" box="[482,554,1268,1292]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="hitit">
<emphasis id="48B25C2C3926FFBCB0C50EF6FDF99D79" box="[482,554,1268,1292]" italics="true" pageId="46" pageNumber="47">I. hitit</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB31B0EF6FD089D79" box="[572,731,1268,1292]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="kalishevskii">
<emphasis id="48B25C2C3926FFBCB31B0EF6FD089D79" box="[572,731,1268,1292]" italics="true" pageId="46" pageNumber="47">I. kalishevskii</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB3CA0EF6FC459D79" box="[749,918,1269,1292]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="nigrosignata">
<emphasis id="48B25C2C3926FFBCB3CA0EF6FC459D79" box="[749,918,1269,1292]" italics="true" pageId="46" pageNumber="47">I. nigrosignata</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB28E0EF6FBC49D79" box="[937,1047,1268,1292]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pylnovi">
<emphasis id="48B25C2C3926FFBCB28E0EF6FBC49D79" box="[937,1047,1268,1292]" italics="true" pageId="46" pageNumber="47">I. pylnovi</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB50E0EF6FB339D79" box="[1065,1248,1268,1292]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="redtenbacheri">
<emphasis id="48B25C2C3926FFBCB50E0EF6FB339D79" box="[1065,1248,1268,1292]" italics="true" pageId="46" pageNumber="47">I. redtenbacheri</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB5D50EF6FAAA9D79" box="[1266,1401,1268,1292]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="48B25C2C3926FFBCB5D50EF6FAAA9D79" box="[1266,1401,1268,1292]" italics="true" pageId="46" pageNumber="47">I. reticulata</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB4AB0EF6FF2A9D45" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="rizeensis">
<emphasis id="48B25C2C3926FFBCB4AB0EF6FF2A9D45" italics="true" pageId="46" pageNumber="47">I. rizeensis</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB0370F1AFE7B9D45" box="[272,424,1304,1328]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="rodsjankoi">
<emphasis id="48B25C2C3926FFBCB0370F1AFE7B9D45" box="[272,424,1304,1328]" italics="true" pageId="46" pageNumber="47">I. rodsjankoi</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB0E70F1AFDEE9D45" box="[448,573,1305,1328]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis id="48B25C2C3926FFBCB0E70F1AFDEE9D45" box="[448,573,1305,1328]" italics="true" pageId="46" pageNumber="47">I. savignyi</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB3720F1AFD059D45" box="[597,726,1305,1328]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="48B25C2C3926FFBCB3720F1AFD059D45" box="[597,726,1305,1328]" italics="true" pageId="46" pageNumber="47">I. speciosa</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB3C90F1AFCAF9D45" box="[750,892,1304,1328]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="splendida">
<emphasis id="48B25C2C3926FFBCB3C90F1AFCAF9D45" box="[750,892,1304,1328]" italics="true" pageId="46" pageNumber="47">I. splendida</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB2B30F1AFB5B9D45" box="[916,1160,1305,1328]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="subSpecies" species="sureyai" subSpecies="amazonae">
<emphasis id="48B25C2C3926FFBCB2B30F1AFB5B9D45" box="[916,1160,1305,1328]" italics="true" pageId="46" pageNumber="47">I. sureyai amazonae</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB5870F1AFAA79D45" box="[1184,1396,1305,1328]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="subSpecies" species="sureyai" subSpecies="sureyai">
<emphasis id="48B25C2C3926FFBCB5870F1AFAA79D45" box="[1184,1396,1305,1328]" italics="true" pageId="46" pageNumber="47">I. sureyai sureyai</emphasis>
</taxonomicName>
,
<taxonomicName id="BDC6FBBD3926FFBCB4AB0F1AFEF89D21" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="uludaghensis">
<emphasis id="48B25C2C3926FFBCB4AB0F1AFEF89D21" italics="true" pageId="46" pageNumber="47">I. uludaghensis</emphasis>
</taxonomicName>
. Refining the relatioships within the group is at the moment difficult and requires broader phylogenetic study.
</paragraph>
</subSubSection>
</treatment>
</document>