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<document id="0FB1DFB3465F156209EBCE0FB9B1BBF1" ID-CLB-Dataset="3098" ID-DOI="10.5852/ejt.2021.782.1589" ID-GBIF-Dataset="246061ce-4a39-4b2e-95e5-e192821782e8" ID-ISSN="2118-9773" ID-Zenodo-Dep="5761502" ID-ZooBank="B4BE1ABB-A27F-4985-A330-C3352A1147A6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638795567160" checkinUser="felipe" docAuthor="Ćurčić, Srećko, Pavićević, Dragan, Vesović, Nikola, Vrbica, Maja, Kuraica, Miloš, Marković, Đorđe, Petković, Matija, Lazović, Vladimir, Pantelić, Dejan &amp; Bosco, Fabrizio" docDate="2021" docId="F2440A25FF9A4B2FDB82BCD9FAB2FCFA" docLanguage="en" docName="ejt-782_55-81.pdf.imf" docOrigin="European Journal of Taxonomy 782 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.782.1589" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Proleonhardella Jeannel 1910" docType="key" docVersion="7" lastPageNumber="76" masterDocId="0E7D725DFF8E4B3ADB3FBD0EFFC5FFF1" masterDocTitle="On the diversity of subterranean beetles of the Dinarides: new leiodid taxa (Coleoptera: Leiodidae) from Serbia" masterLastPageNumber="81" masterPageNumber="55" pageNumber="75" updateTime="1699472014327" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DFBA10CAACDF03B68C5FC8D294F5FF5E">On the diversity of subterranean beetles of the Dinarides: new leiodid taxa (Coleoptera: Leiodidae) from Serbia</mods:title>
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<mods:affiliation id="ECD57EC202095A51124B01C387493FCE">Institute of Zoology, University of Belgrade - Faculty of Biology, Studentski Trg 16, 11000 Belgrade, Serbia.</mods:affiliation>
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<mods:affiliation id="B4C769D3AA293DCA472D53581A623370">Institute of Physics Belgrade, University of Belgrade, Pregrevica 118, 11080 Belgrade, Serbia.</mods:affiliation>
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<mods:affiliation id="A91D92CF57117DBF54B77D53E519857A">Institute of Physics Belgrade, University of Belgrade, Pregrevica 118, 11080 Belgrade, Serbia. A. F. Linder Association for the Study of Karst, Via Fratelli Cervi 9 / G, 34077 Ronchi dei Legionari, Italy.</mods:affiliation>
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<mods:affiliation id="882E03D91429A89F553038890DB1355D">A. F. Linder Association for the Study of Karst, Via Fratelli Cervi 9 / G, 34077 Ronchi dei Legionari, Italy.</mods:affiliation>
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<mods:date id="E05FF7690DF124E8E4FB3A6A739BD2B8">2021</mods:date>
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<treatment id="F2440A25FF9A4B2FDB82BCD9FAB2FCFA" ID-DOI="http://doi.org/10.5281/zenodo.5762320" ID-GBIF-Taxon="191144394" ID-Zenodo-Dep="5762320" LSID="urn:lsid:plazi:treatment:F2440A25FF9A4B2FDB82BCD9FAB2FCFA" httpUri="http://treatment.plazi.org/id/F2440A25FF9A4B2FDB82BCD9FAB2FCFA" lastPageId="21" lastPageNumber="76" pageId="20" pageNumber="75">
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Key to the taxa of the genus
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<emphasis id="48996721FF9A4B2ED920BCD9FD21FE03" bold="true" box="[543,740,471,498]" italics="true" pageId="20" pageNumber="75">Proleonhardella</emphasis>
<bibRefCitation id="1E7CC6C2FF9A4B2ED9D3BCD9FC53FE00" author="Jeannel R." box="[748,918,471,497]" pageId="20" pageNumber="75" pagination="1 - 48" refId="ref13309" refString="Jeannel R. 1910. Essai d'une nouvelle classification des Silphides cavernicoles. Archives de Zoologie experimentale et generale 45 (1): 1 - 48." type="journal article" year="1910">Jeannel, 1910</bibRefCitation>
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(modified after
<bibRefCitation id="1E7CC6C2FF9A4B2EDA4CBCF4FDCAFDE4" author="Jeannel R." box="[371,527,506,533]" pageId="20" pageNumber="75" pagination="1 - 436" refId="ref13380" refString="Jeannel R. 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie experimentale et generale 63: 1 - 436." type="journal article" year="1924">Jeannel 1924</bibRefCitation>
) (
<figureCitation id="E2D6A7B6FF9A4B2ED918BCF4FD5BFDE4" box="[551,670,506,533]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="19.[191,234,1746,1772]" captionStartId-1="21.[189,232,1850,1876]" captionTargetBox-0="[190,1397,874,1707]" captionTargetBox-1="[414,1161,842,1782]" captionTargetId-0="figure-274@19.[189,1398,873,1708]" captionTargetId-1="figure-187@21.[376,1212,824,1812]" captionTargetPageId-0="19" captionTargetPageId-1="21" captionText-0="Fig. 9. Map of the distribution of taxa of the genera Bozidaria Ćurčić &amp; Pavićević gen. nov. and Proleonhardella Jeannel, 1910. White circles: B. serbooccidentalis Ćurčić &amp; Pavićević gen. et sp. nov. Turquoise star: P. (Pholeuonillus) adolfi (Reitter, 1911). Light blue sun: P. (Proleonhardella) matzenaueri matzenaueri (Apfelbeck, 1907). Dark blue sun: P. (P.) matzenaueri ottonis Müller, 1917. Purple flower: P. (P.) leonhardi (Breit, 1913). Yellow cross: P. (P.) apfelbecki Jeannel, 1924. Brown circles: P. (P.) remyi Jeannel, 1934. Green squares: P. (P.) hirtella Jeannel, 1934. Pink pentagon: P. (P.) neumanni (Apfelbeck, 1901). Red triangle: P. (P.) weiratheri (Reitter, 1913). Orange rhombuses: P. (P.) tarensis Ćurčić &amp; Pavićević sp. nov. Scale bar = 50 km." captionText-1="Fig. 10. Illustrations of morphological characters presented in the Key to the taxa of the genus Proleonhardella Jeannel, 1910 (after Jeannel 1924; Ćurčić et al. 2008a). A. Short elytra, less than twice as long as pronotum. B. Long elytra, more than twice as long as pronotum. C. Globular antennomere VIII in males. D. Slightly elongate antennomere VIII in males. E. Elytra parallel in basal half. F. Elytra narrowed in basal half." figureDoi-0="http://doi.org/10.5281/zenodo.5761520" figureDoi-1="http://doi.org/10.5281/zenodo.5761522" httpUri-0="https://zenodo.org/record/5761520/files/figure.png" httpUri-1="https://zenodo.org/record/5761522/files/figure.png" pageId="20" pageNumber="75">Figs 9–10</figureCitation>
)
</paragraph>
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1. Body more elongate, elliptical. Pronotum as wide as elytra, well constricted basally. Pronotal lateral margins well-rounded backwards. Mesosternal carina very low. Elytral punctuation rough and deep (subgenus
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<emphasis id="48996721FF9A4B2EDADABF61FD4DFD78" box="[485,648,623,649]" italics="true" pageId="20" pageNumber="75">Pholeuonillus</emphasis>
<bibRefCitation id="1E7CC6C2FF9A4B2ED9ACBF61FCE5FD78" author="Breit J." box="[659,800,623,649]" pageId="20" pageNumber="75" pagination="351 - 358" refId="ref12401" refString="Breit J. 1913. Wissenschaftliche Ergebnisse der Bearbeitung von O. Leonhard's Sammlungen. 5. Beitrage zur Blindkaferfauna von Bosnien und der Herzegowina. Entomologische Mitteilungen 2 (11): 351 - 358." type="journal article" year="1913">Breit, 1913</bibRefCitation>
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). Endogean, Mt Treskavica, southern
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.......................................................................
<taxonomicName id="BDEDC0B0FF9A4B2ED8A0BF9DFAB3FD5C" authority="(Reitter, 1911)" baseAuthorityName="Reitter" baseAuthorityYear="1911" box="[927,1398,658,685]" class="Insecta" family="Leiodidae" genus="Proleonhardella" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="adolfi" subGenus="Pholeuonillus">
<emphasis id="48996721FF9A4B2ED8A0BF9DFB01FD5C" bold="true" box="[927,1220,658,685]" pageId="20" pageNumber="75">
<emphasis id="48996721FF9A4B2ED8A0BF9DFC7CFD5C" bold="true" box="[927,953,659,685]" italics="true" pageId="20" pageNumber="75">P.</emphasis>
(
<emphasis id="48996721FF9A4B2ED8F9BF9CFBB5FD5C" bold="true" box="[966,1136,658,685]" italics="true" pageId="20" pageNumber="75">Pholeuonillus</emphasis>
)
<emphasis id="48996721FF9A4B2EDFBFBF9CFB01FD5C" bold="true" box="[1152,1220,658,685]" italics="true" pageId="20" pageNumber="75">adolfi</emphasis>
</emphasis>
(Reitter, 1911)
</taxonomicName>
</keyLead>
</key>
</paragraph>
<paragraph id="7A52BB33FF9A4B2EDBFEBFB8FAB2FCE7" blockId="20.[189,1400,553,1993]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDBFEBFB8FAB2FCE7" pageId="20" pageNumber="75">
– Body less elongate, oval. Pronotum narrower than elytra, weakly constricted basally. Pronotal lateral margins weakly rounded backwards. Elytral punctuation fine. Mesosternal carina elevated, angled (subgenus
<taxonomicName id="BDEDC0B0FF9A4B2EDA56BFF2FD17FCE7" ID-CoL="8X7RJ" authority="Jeannel, 1910" authorityName="Jeannel" authorityYear="1910" box="[361,722,764,790]" class="Insecta" family="Leiodidae" genus="Proleonhardella" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="48996721FF9A4B2EDA56BFF2FDECFCE7" box="[361,553,764,790]" italics="true" pageId="20" pageNumber="75">Proleonhardella</emphasis>
<bibRefCitation id="1E7CC6C2FF9A4B2ED90FBFF2FD17FCE7" author="Jeannel R." box="[560,722,764,790]" pageId="20" pageNumber="75" pagination="1 - 48" refId="ref13309" refString="Jeannel R. 1910. Essai d'une nouvelle classification des Silphides cavernicoles. Archives de Zoologie experimentale et generale 45 (1): 1 - 48." type="journal article" year="1910">Jeannel, 1910</bibRefCitation>
</taxonomicName>
) ....................................................................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C119A396FF9A4B2EDB82BE30FAB2FC8A" pageId="20" pageNumber="75">
<paragraph id="7A52BB33FF9A4B2EDB82BE30FAB2FCA9" blockId="20.[189,1400,553,1993]" box="[189,1399,829,856]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDB82BE30FAB2FCA9" box="[189,1399,829,856]" pageId="20" pageNumber="75">
2. Elytra shorter, more convex, less than twice as long as pronotum (
<figureCitation id="E2D6A7B6FF9A4B2ED8CFBE33FB9EFCA9" box="[1008,1115,829,856]" captionStart="Fig" captionStartId="21.[189,232,1850,1876]" captionTargetBox="[414,1161,842,1782]" captionTargetId="figure-187@21.[376,1212,824,1812]" captionTargetPageId="21" captionText="Fig. 10. Illustrations of morphological characters presented in the Key to the taxa of the genus Proleonhardella Jeannel, 1910 (after Jeannel 1924; Ćurčić et al. 2008a). A. Short elytra, less than twice as long as pronotum. B. Long elytra, more than twice as long as pronotum. C. Globular antennomere VIII in males. D. Slightly elongate antennomere VIII in males. E. Elytra parallel in basal half. F. Elytra narrowed in basal half." figureDoi="http://doi.org/10.5281/zenodo.5761522" httpUri="https://zenodo.org/record/5761522/files/figure.png" pageId="20" pageNumber="75">Fig. 10A</figureCitation>
) ................................. 3
</keyLead>
</paragraph>
<paragraph id="7A52BB33FF9A4B2EDBFEBE6FFAB2FC8A" blockId="20.[189,1400,553,1993]" box="[193,1399,865,891]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDBFEBE6FFAB2FC8A" box="[193,1399,865,891]" pageId="20" pageNumber="75">
– Elytra longer, less convex, more than twice as long as pronotum (
<figureCitation id="E2D6A7B6FF9A4B2ED8D5BE6FFB96FC8A" box="[1002,1107,865,891]" captionStart="Fig" captionStartId="21.[189,232,1850,1876]" captionTargetBox="[414,1161,842,1782]" captionTargetId="figure-187@21.[376,1212,824,1812]" captionTargetPageId="21" captionText="Fig. 10. Illustrations of morphological characters presented in the Key to the taxa of the genus Proleonhardella Jeannel, 1910 (after Jeannel 1924; Ćurčić et al. 2008a). A. Short elytra, less than twice as long as pronotum. B. Long elytra, more than twice as long as pronotum. C. Globular antennomere VIII in males. D. Slightly elongate antennomere VIII in males. E. Elytra parallel in basal half. F. Elytra narrowed in basal half." figureDoi="http://doi.org/10.5281/zenodo.5761522" httpUri="https://zenodo.org/record/5761522/files/figure.png" pageId="20" pageNumber="75">Fig. 10B</figureCitation>
) .................................. 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C119A396FF9A4B2EDB82BEACFAB2FB61" pageId="20" pageNumber="75">
<paragraph id="7A52BB33FF9A4B2EDB82BEACFAB2FBD7" blockId="20.[189,1400,553,1993]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDB82BEACFAB2FBD7" pageId="20" pageNumber="75">
3. Body of oval shape, longer (TL
<quantity id="BD1516D6FF9A4B2ED9B5BEACFCE6FC4C" box="[650,803,930,957]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="20" pageNumber="75" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
). Antennae short, not reaching middle of body. Antennomere III not longer than antennomere V. Antennomere VIII globular in males (
<figureCitation id="E2D6A7B6FF9A4B2EDFC3BECBFAAFFC11" box="[1276,1386,965,992]" captionStart="Fig" captionStartId="21.[189,232,1850,1876]" captionTargetBox="[414,1161,842,1782]" captionTargetId="figure-187@21.[376,1212,824,1812]" captionTargetPageId="21" captionText="Fig. 10. Illustrations of morphological characters presented in the Key to the taxa of the genus Proleonhardella Jeannel, 1910 (after Jeannel 1924; Ćurčić et al. 2008a). A. Short elytra, less than twice as long as pronotum. B. Long elytra, more than twice as long as pronotum. C. Globular antennomere VIII in males. D. Slightly elongate antennomere VIII in males. E. Elytra parallel in basal half. F. Elytra narrowed in basal half." figureDoi="http://doi.org/10.5281/zenodo.5761522" httpUri="https://zenodo.org/record/5761522/files/figure.png" pageId="20" pageNumber="75">Fig. 10C</figureCitation>
). Cavernicolous, Mts Bjelašnica and Igman, central
<collectingCountry id="02FAFBA3FF9A4B2ED873BEE7FBB7FBF2" box="[844,1138,1001,1027]" name="Bosnia and Herzegovina" pageId="20" pageNumber="75">Bosnia and Herzegovina</collectingCountry>
[
<taxonomicName id="BDEDC0B0FF9A4B2EDFBBBEE7FD98FBD7" ID-CoL="4MRRQ" authority="(Apfelbeck, 1907)" baseAuthorityName="Apfelbeck" baseAuthorityYear="1907" class="Insecta" family="Leiodidae" genus="Proleonhardella" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="matzenaueri" subGenus="Proleonhardella">
<emphasis id="48996721FF9A4B2EDFBBBEE7FB58FBF3" box="[1156,1181,1001,1026]" italics="true" pageId="20" pageNumber="75">P.</emphasis>
(
<emphasis id="48996721FF9A4B2EDF92BEE7FAA8FBF2" box="[1197,1389,1001,1027]" italics="true" pageId="20" pageNumber="75">Proleonhardella</emphasis>
)
<emphasis id="48996721FF9A4B2EDBD5B903FEBEFBD7" box="[234,379,1037,1062]" italics="true" pageId="20" pageNumber="75">matzenaueri</emphasis>
(Apfelbeck, 1907)
</taxonomicName>
] ....................................................................................................... 4
</keyLead>
</paragraph>
<paragraph id="7A52BB33FF9A4B2EDBFEB93EFAB2FB61" blockId="20.[189,1400,553,1993]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDBFEB93EFAB2FB61" pageId="20" pageNumber="75">
– Body of subglobular shape, shorter (TL less than
<quantity id="BD1516D6FF9A4B2ED86DB921FC73FBBB" box="[850,950,1071,1098]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="20" pageNumber="75" unit="mm" value="1.8">1.8 mm</quantity>
). Antennae long, reaching middle of body. Antennomere III longer than antennomere V. Antennomere VIII slightly elongate in males (
<figureCitation id="E2D6A7B6FF9A4B2EDBCCB97BFE9BFB61" box="[243,350,1141,1168]" captionStart="Fig" captionStartId="21.[189,232,1850,1876]" captionTargetBox="[414,1161,842,1782]" captionTargetId="figure-187@21.[376,1212,824,1812]" captionTargetPageId="21" captionText="Fig. 10. Illustrations of morphological characters presented in the Key to the taxa of the genus Proleonhardella Jeannel, 1910 (after Jeannel 1924; Ćurčić et al. 2008a). A. Short elytra, less than twice as long as pronotum. B. Long elytra, more than twice as long as pronotum. C. Globular antennomere VIII in males. D. Slightly elongate antennomere VIII in males. E. Elytra parallel in basal half. F. Elytra narrowed in basal half." figureDoi="http://doi.org/10.5281/zenodo.5761522" httpUri="https://zenodo.org/record/5761522/files/figure.png" pageId="20" pageNumber="75">Fig. 10D</figureCitation>
) ......................................................................................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C119A396FF9A4B2EDB82B9B9FAB2FA70" pageId="20" pageNumber="75">
<paragraph id="7A52BB33FF9A4B2EDB82B9B9FAB3FAE6" blockId="20.[189,1400,553,1993]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDB82B9B9FAB3FAE6" pageId="20" pageNumber="75">
4. Body of almost regular oval shape, longer (TL 2.0–
<quantity id="BD1516D6FF9A4B2ED869B9B9FC73FB20" box="[854,950,1207,1233]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="20" pageNumber="75" unit="mm" value="2.2">2.2 mm</quantity>
). Elytra not widened towards middle. Cavernicolous, Mt Bjelašnica, central
<collectingCountry id="02FAFBA3FF9A4B2ED991B9D4FC0BFB05" box="[686,974,1242,1268]" name="Bosnia and Herzegovina" pageId="20" pageNumber="75">Bosnia and Herzegovina</collectingCountry>
........................................................ .................................................
<taxonomicName id="BDEDC0B0FF9A4B2ED95BB9F0FAB3FAE6" ID-CoL="8YFC2" authority="matzenaueri (Apfelbeck, 1907)" authorityName="matzenaueri (Apfelbeck" authorityYear="1907" baseAuthorityName="Apfelbeck" baseAuthorityYear="1907" box="[612,1398,1277,1304]" class="Insecta" family="Leiodidae" genus="Proleonhardella" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="75" phylum="Arthropoda" rank="subSpecies" species="matzenaueri" subGenus="Proleonhardella" subSpecies="matzenaueri">
<emphasis id="48996721FF9A4B2ED95BB9F0FB5DFAE6" bold="true" box="[612,1176,1277,1304]" pageId="20" pageNumber="75">
<emphasis id="48996721FF9A4B2ED95BB9F0FDB3FAE9" bold="true" box="[612,630,1278,1304]" italics="true" pageId="20" pageNumber="75">P</emphasis>
. (
<emphasis id="48996721FF9A4B2ED9B0B9F3FC91FAE9" bold="true" box="[655,852,1277,1304]" italics="true" pageId="20" pageNumber="75">Proleonhardella</emphasis>
)
<emphasis id="48996721FF9A4B2ED85AB9F3FB5DFAE6" bold="true" box="[869,1176,1277,1303]" italics="true" pageId="20" pageNumber="75">matzenaueri matzenaueri</emphasis>
</emphasis>
(Apfelbeck, 1907)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A52BB33FF9A4B2EDBFEB82FFAB2FA70" blockId="20.[189,1400,553,1993]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDBFEB82FFAB2FA70" pageId="20" pageNumber="75">
– Body narrower anteriorly, shorter (TL 1.8–2.0 mm). Elytra clearly widened medially. Cavernicolous, Mt Igman, central
<collectingCountry id="02FAFBA3FF9A4B2EDAFCB84AFD21FAAF" box="[451,740,1348,1374]" name="Bosnia and Herzegovina" pageId="20" pageNumber="75">Bosnia and Herzegovina</collectingCountry>
....................................................................................... ...................................................................
<taxonomicName id="BDEDC0B0FF9A4B2ED9D7B869FAB2FA70" ID-CoL="7HW7J" authority="Muller, 1917" authorityName="Muller" authorityYear="1917" box="[744,1399,1382,1409]" class="Insecta" family="Leiodidae" genus="Proleonhardella" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="75" phylum="Arthropoda" rank="subSpecies" species="matzenaueri" subGenus="Proleonhardella" subSpecies="ottonis">
<emphasis id="48996721FF9A4B2ED9D7B869FB1DFA70" bold="true" box="[744,1240,1382,1409]" pageId="20" pageNumber="75">
<emphasis id="48996721FF9A4B2ED9D7B869FD3FFA70" bold="true" box="[744,762,1383,1409]" italics="true" pageId="20" pageNumber="75">P</emphasis>
. (
<emphasis id="48996721FF9A4B2ED82DB868FC12FA70" bold="true" box="[786,983,1382,1409]" italics="true" pageId="20" pageNumber="75">Proleonhardella</emphasis>
)
<emphasis id="48996721FF9A4B2ED8D7B869FB1DFA70" bold="true" box="[1000,1240,1383,1409]" italics="true" pageId="20" pageNumber="75">matzenaueri ottonis</emphasis>
</emphasis>
Müller, 1917
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C119A396FF9A4B2EDB82B8A6FAB2F948" pageId="20" pageNumber="75">
<paragraph id="7A52BB33FF9A4B2EDB82B8A6FAB3F9DD" blockId="20.[189,1400,553,1993]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDB82B8A6FAB3F9DD" pageId="20" pageNumber="75">
5. Body more elongate and convex, longer (TL
<quantity id="BD1516D6FF9A4B2ED9D0B8A6FC88FA32" box="[751,845,1448,1475]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="75" unit="mm" value="1.5">1.5 mm</quantity>
). Elytral punctuation finer and denser.Antennae shorter, with apical antennomeres thicker. Antennomere VIII barely longer than wide, antennomere IX as long as wide and antennomere X transverse in females. Cavernicolous, Mt Treskavica, southern
<collectingCountry id="02FAFBA3FF9A4B2EDBD5BB1CFDCEF9DD" box="[234,523,1554,1580]" name="Bosnia and Herzegovina" pageId="20" pageNumber="75">Bosnia and Herzegovina</collectingCountry>
............................................
<taxonomicName id="BDEDC0B0FF9A4B2ED85DBB1CFAB3F9DD" ID-CoL="4MRRP" authority="(Breit, 1913)" baseAuthorityName="Breit" baseAuthorityYear="1913" box="[866,1398,1553,1580]" class="Insecta" family="Leiodidae" genus="Proleonhardella" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="leonhardi" subGenus="Proleonhardella">
<emphasis id="48996721FF9A4B2ED85DBB1CFB12F9DD" bold="true" box="[866,1239,1553,1580]" pageId="20" pageNumber="75">
<emphasis id="48996721FF9A4B2ED85DBB1CFCB1F9DD" bold="true" box="[866,884,1554,1580]" italics="true" pageId="20" pageNumber="75">P</emphasis>
. (
<emphasis id="48996721FF9A4B2ED8B3BB1FFB94F9DD" bold="true" box="[908,1105,1553,1580]" italics="true" pageId="20" pageNumber="75">Proleonhardella</emphasis>
)
<emphasis id="48996721FF9A4B2EDF5DBB1FFB12F9DD" bold="true" box="[1122,1239,1553,1580]" italics="true" pageId="20" pageNumber="75">leonhardi</emphasis>
</emphasis>
(
<bibRefCitation id="1E7CC6C2FF9A4B2EDFD8BB1CFAABF9DD" author="Breit J." box="[1255,1390,1554,1580]" pageId="20" pageNumber="75" pagination="351 - 358" refId="ref12401" refString="Breit J. 1913. Wissenschaftliche Ergebnisse der Bearbeitung von O. Leonhard's Sammlungen. 5. Beitrage zur Blindkaferfauna von Bosnien und der Herzegowina. Entomologische Mitteilungen 2 (11): 351 - 358." type="journal article" year="1913">Breit, 1913</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A52BB33FF9A4B2EDBFEBB38FAB2F948" blockId="20.[189,1400,553,1993]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDBFEBB38FAB2F948" pageId="20" pageNumber="75">
– Body less elongate and convex, shorter (TL
<quantity id="BD1516D6FF9A4B2ED9C4BB3BFC9EF9A1" box="[763,859,1589,1616]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="20" pageNumber="75" unit="mm" value="1.3">1.3 mm</quantity>
). Elytral punctuation stronger and less dense. Antennae longer, with apical antennomeres less thick. Antennomeres VIII, IX and X longer than wide in females. Cavernicolous, village of Trnovo, vicinity of the city of Sarajevo, central
<collectingCountry id="02FAFBA3FF9A4B2EDE1ABB75FE76F948" name="Bosnia and Herzegovina" pageId="20" pageNumber="75">Bosnia and Herzegovina</collectingCountry>
......................................................
<taxonomicName id="BDEDC0B0FF9A4B2ED86EBB91FAB2F948" ID-CoL="4MRRM" authority="Jeannel, 1924" authorityName="Jeannel" authorityYear="1924" box="[849,1399,1694,1721]" class="Insecta" family="Leiodidae" genus="Proleonhardella" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="apfelbecki" subGenus="Proleonhardella">
<emphasis id="48996721FF9A4B2ED86EBB91FB0BF948" bold="true" box="[849,1230,1694,1721]" pageId="20" pageNumber="75">
<emphasis id="48996721FF9A4B2ED86EBB91FCA6F948" bold="true" box="[849,867,1695,1721]" italics="true" pageId="20" pageNumber="75">P</emphasis>
. (
<emphasis id="48996721FF9A4B2ED843BB90FB84F948" bold="true" box="[892,1089,1694,1721]" italics="true" pageId="20" pageNumber="75">Proleonhardella</emphasis>
)
<emphasis id="48996721FF9A4B2EDF6DBB90FB0BF948" bold="true" box="[1106,1230,1694,1721]" italics="true" pageId="20" pageNumber="75">apfelbecki</emphasis>
</emphasis>
<bibRefCitation id="1E7CC6C2FF9A4B2EDFEABB91FAB2F948" author="Jeannel R." box="[1237,1399,1695,1721]" pageId="20" pageNumber="75" pagination="1 - 436" refId="ref13380" refString="Jeannel R. 1924. Monographie des Bathysciinae. Biospeologica L. Archives de Zoologie experimentale et generale 63: 1 - 436." type="journal article" year="1924">Jeannel, 1924</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C119A396FF9A4B2EDB82BBEEFAB2F895" pageId="20" pageNumber="75">
<paragraph id="7A52BB33FF9A4B2EDB82BBEEFAB2F8B0" blockId="20.[189,1400,553,1993]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDB82BBEEFAB2F8B0" pageId="20" pageNumber="75">
6. Body longer (TL 3.0–
<quantity id="BD1516D6FF9A4B2EDAD6BBEEFD83F90A" box="[489,582,1760,1787]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="20" pageNumber="75" unit="mm" value="3.5">3.5 mm</quantity>
). Cavernicolous, Kamena Gora and vicinity of the towns of
<collectingRegion id="B82975D1FF9A4B2EDFC2BBEEFAB7F90B" box="[1277,1394,1760,1786]" country="Serbia" name="Prijepolje" pageId="20" pageNumber="75">Prijepolje</collectingRegion>
,
<collectingRegion id="B82975D1FF9A4B2EDBD6BA0DFEF4F8EC" box="[233,305,1795,1821]" country="Serbia" name="Priboj" pageId="20" pageNumber="75">Priboj</collectingRegion>
and Pljevlja, southwestern
<collectingCountry id="02FAFBA3FF9A4B2ED94ABA0DFD05F8EC" box="[629,704,1795,1821]" name="Serbia" pageId="20" pageNumber="75">Serbia</collectingCountry>
and northern
<collectingCountry id="02FAFBA3FF9A4B2ED85CBA0AFC37F8EF" box="[867,1010,1796,1822]" name="Serbia" pageId="20" pageNumber="75">Montenegro</collectingCountry>
.................................................. ..........................................................................................
<taxonomicName id="BDEDC0B0FF9A4B2ED8B2BA29FAB2F8B0" ID-CoL="4MRRL" authority="Jeannel, 1934" authorityName="Jeannel" authorityYear="1934" box="[909,1399,1830,1857]" class="Insecta" family="Leiodidae" genus="Proleonhardella" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="remyi" subGenus="Proleonhardella">
<emphasis id="48996721FF9A4B2ED8B2BA29FB08F8B1" bold="true" box="[909,1229,1830,1857]" pageId="20" pageNumber="75">
<emphasis id="48996721FF9A4B2ED8B2BA29FC63F8B0" bold="true" box="[909,934,1831,1857]" italics="true" pageId="20" pageNumber="75">P.</emphasis>
(
<emphasis id="48996721FF9A4B2ED88CBA28FBBDF8B0" bold="true" box="[947,1144,1830,1857]" italics="true" pageId="20" pageNumber="75">Proleonhardella</emphasis>
)
<emphasis id="48996721FF9A4B2EDFB6BA28FB08F8B1" bold="true" box="[1161,1229,1830,1856]" italics="true" pageId="20" pageNumber="75">remyi</emphasis>
</emphasis>
<bibRefCitation id="1E7CC6C2FF9A4B2EDFEABA28FAB2F8B0" author="Jeannel R." box="[1237,1399,1830,1857]" pageId="20" pageNumber="75" pagination="89 - 103" refId="ref13457" refString="Jeannel R. 1934. Bathysciinae recueillis par M. M. Remy et R. Husson dans le Sandjak de Novi-Pazar et la Macedoine grecque. Revue francaise d'entomologie 1: 89 - 103." type="journal article" year="1934">Jeannel, 1934</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A52BB33FF9A4B2EDBFEBA44FAB2F895" blockId="20.[189,1400,553,1993]" box="[193,1399,1865,1892]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDBFEBA44FAB2F895" box="[193,1399,1865,1892]" pageId="20" pageNumber="75">
– Body shorter (TL less than
<quantity id="BD1516D6FF9A4B2ED914BA47FD61F895" box="[555,676,1865,1892]" metricMagnitude="-3" metricUnit="m" metricValue="2.435" pageId="20" pageNumber="75" unit="mm" value="2.435">2.435 mm</quantity>
) ............................................................................................. 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C119A396FF9A4B2EDB82BA82FAB2F839" pageId="20" pageNumber="75">
<paragraph id="7A52BB33FF9A4B2EDB82BA82FAB2F854" blockId="20.[189,1400,553,1993]" box="[189,1399,1931,1958]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDB82BA82FAB2F854" box="[189,1399,1931,1958]" pageId="20" pageNumber="75">
7. Elytra parallel in basal half (
<figureCitation id="E2D6A7B6FF9A4B2ED907BA85FD65F854" box="[568,672,1931,1957]" captionStart="Fig" captionStartId="21.[189,232,1850,1876]" captionTargetBox="[414,1161,842,1782]" captionTargetId="figure-187@21.[376,1212,824,1812]" captionTargetPageId="21" captionText="Fig. 10. Illustrations of morphological characters presented in the Key to the taxa of the genus Proleonhardella Jeannel, 1910 (after Jeannel 1924; Ćurčić et al. 2008a). A. Short elytra, less than twice as long as pronotum. B. Long elytra, more than twice as long as pronotum. C. Globular antennomere VIII in males. D. Slightly elongate antennomere VIII in males. E. Elytra parallel in basal half. F. Elytra narrowed in basal half." figureDoi="http://doi.org/10.5281/zenodo.5761522" httpUri="https://zenodo.org/record/5761522/files/figure.png" pageId="20" pageNumber="75">Fig. 10E</figureCitation>
) .............................................................................................. 8
</keyLead>
</paragraph>
<paragraph id="7A52BB33FF9A4B2EDBFEBAA1FAB2F839" blockId="20.[189,1400,553,1993]" box="[193,1399,1966,1993]" pageId="20" pageNumber="75">
<keyLead id="C11C1806FF9A4B2EDBFEBAA1FAB2F839" box="[193,1399,1966,1993]" pageId="20" pageNumber="75">
– Elytra narrowed in basal half (
<figureCitation id="E2D6A7B6FF9A4B2ED970BAA0FD70F839" box="[591,693,1966,1992]" captionStart="Fig" captionStartId="21.[189,232,1850,1876]" captionTargetBox="[414,1161,842,1782]" captionTargetId="figure-187@21.[376,1212,824,1812]" captionTargetPageId="21" captionText="Fig. 10. Illustrations of morphological characters presented in the Key to the taxa of the genus Proleonhardella Jeannel, 1910 (after Jeannel 1924; Ćurčić et al. 2008a). A. Short elytra, less than twice as long as pronotum. B. Long elytra, more than twice as long as pronotum. C. Globular antennomere VIII in males. D. Slightly elongate antennomere VIII in males. E. Elytra parallel in basal half. F. Elytra narrowed in basal half." figureDoi="http://doi.org/10.5281/zenodo.5761522" httpUri="https://zenodo.org/record/5761522/files/figure.png" pageId="20" pageNumber="75">Fig. 10F</figureCitation>
) ........................................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C119A396FF9B4B2FDB82BC07FAB3FE07" pageId="21" pageNumber="76">
<paragraph id="7A52BB33FF9B4B2FDB82BC07FAB2FE7C" blockId="21.[189,1399,265,780]" pageId="21" pageNumber="76">
<keyLead id="C11C1806FF9B4B2FDB82BC07FAB2FE7C" pageId="21" pageNumber="76">
8. Pubescence long. Punctuation less fine. Apical antennomeres more widened. Pronotum less rounded. Apex of median lobe of aedeagus less bent ventrally. Elytral apex wide, obtuse and oblique. Cavernicolous, vicinity of the towns of
<collectingRegion id="B82975D1FF9B4B2FD9EDBC41FC81FE98" box="[722,836,335,361]" country="Serbia" name="Prijepolje" pageId="21" pageNumber="76">Prijepolje</collectingRegion>
and
<collectingRegion id="B82975D1FF9B4B2FD8BABC41FC08FE98" box="[901,973,335,361]" country="Serbia" name="Priboj" pageId="21" pageNumber="76">Priboj</collectingRegion>
(southwestern
<collectingCountry id="02FAFBA3FF9B4B2FDFB8BC41FB11FE9B" box="[1159,1236,335,362]" name="Serbia" pageId="21" pageNumber="76">Serbia</collectingCountry>
) and Pljevlja (northern
<collectingCountry id="02FAFBA3FF9B4B2FDA60BC7DFE35FE7C" box="[351,496,371,397]" name="Serbia" pageId="21" pageNumber="76">Montenegro</collectingCountry>
) ..................................................
<taxonomicName id="BDEDC0B0FF9B4B2FD846BC7DFAB2FE7C" ID-CoL="4MRRN" authority="Jeannel, 1934" authorityName="Jeannel" authorityYear="1934" box="[889,1399,370,397]" class="Insecta" family="Leiodidae" genus="Proleonhardella" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="hirtella" subGenus="Proleonhardella">
<emphasis id="48996721FF9B4B2FD846BC7DFB0BFE7C" bold="true" box="[889,1230,370,397]" pageId="21" pageNumber="76">
<emphasis id="48996721FF9B4B2FD846BC7DFC57FE7C" bold="true" box="[889,914,371,397]" italics="true" pageId="21" pageNumber="76">P.</emphasis>
(
<emphasis id="48996721FF9B4B2FD8A0BC7CFBA1FE7C" bold="true" box="[927,1124,370,397]" italics="true" pageId="21" pageNumber="76">Proleonhardella</emphasis>
)
<emphasis id="48996721FF9B4B2FDF49BC7CFB0BFE7C" bold="true" box="[1142,1230,370,397]" italics="true" pageId="21" pageNumber="76">hirtella</emphasis>
</emphasis>
<bibRefCitation id="1E7CC6C2FF9B4B2FDFEABC7CFAB2FE7C" author="Jeannel R." box="[1237,1399,370,397]" pageId="21" pageNumber="76" pagination="89 - 103" refId="ref13457" refString="Jeannel R. 1934. Bathysciinae recueillis par M. M. Remy et R. Husson dans le Sandjak de Novi-Pazar et la Macedoine grecque. Revue francaise d'entomologie 1: 89 - 103." type="journal article" year="1934">Jeannel, 1934</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A52BB33FF9B4B2FDBFEBC98FAB3FE07" blockId="21.[189,1399,265,780]" pageId="21" pageNumber="76">
<keyLead id="C11C1806FF9B4B2FDBFEBC98FAB3FE07" pageId="21" pageNumber="76">
– Pubescence short. Punctuation finer. Apical antennomeres less widened. Pronotum more rounded. Apex of median lobe of aedeagus more bent ventrally. Elytral apex attenuated. Cavernicolous, Mt Sjemeć, eastern
<collectingCountry id="02FAFBA3FF9B4B2FDAECBCD2FD31FE07" box="[467,756,476,502]" name="Bosnia and Herzegovina" pageId="21" pageNumber="76">Bosnia and Herzegovina</collectingCountry>
.........
<taxonomicName id="BDEDC0B0FF9B4B2FD874BCD2FAB3FE07" ID-CoL="4MRRT" authority="(Reitter, 1913)" baseAuthorityName="Reitter" baseAuthorityYear="1913" box="[843,1398,476,503]" class="Insecta" family="Leiodidae" genus="Proleonhardella" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="Proleonhardella">
<emphasis id="48996721FF9B4B2FD874BCD2FB01FE06" bold="true" box="[843,1220,476,503]" pageId="21" pageNumber="76">
<emphasis id="48996721FF9B4B2FD874BCD2FCA1FE07" bold="true" box="[843,868,476,502]" italics="true" pageId="21" pageNumber="76">P.</emphasis>
(
<emphasis id="48996721FF9B4B2FD84EBCD2FBF3FE06" bold="true" box="[881,1078,476,503]" italics="true" pageId="21" pageNumber="76">Proleonhardella</emphasis>
)
<emphasis id="48996721FF9B4B2FDF77BCD2FB01FE06" bold="true" box="[1096,1220,476,503]" italics="true" pageId="21" pageNumber="76">weiratheri</emphasis>
</emphasis>
(
<bibRefCitation id="1E7CC6C2FF9B4B2FDFECBCD2FAABFE07" author="Reitter E." box="[1235,1390,476,502]" pageId="21" pageNumber="76" pagination="153 - 157" refId="ref14256" refString="Reitter E. 1913. Sechs neue von Herrn Weirather in Bosnien entdeckte Hohlensilphiden. Koleopterologische Rundschau 10: 153 - 157." type="journal article" year="1913">Reitter, 1913</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C119A396FF9B4B2FDB82BF10FAB2FCFA" pageId="21" pageNumber="76">
<paragraph id="7A52BB33FF9B4B2FDB82BF10FAB2FD50" blockId="21.[189,1399,265,780]" pageId="21" pageNumber="76">
<keyLead id="C11C1806FF9B4B2FDB82BF10FAB2FD50" pageId="21" pageNumber="76">
9. A1/A2 M 0.80. Mesosternal carina almost right-angled. Elytral lateral margins below humeral angles clearly sinuate. Maximum width of elytra prior to middle. Cavernicolous, village of Podromanija, near the town of Sokolac, eastern
<collectingCountry id="02FAFBA3FF9B4B2FD945BF6AFC5EFD8F" box="[634,923,612,638]" name="Bosnia and Herzegovina" pageId="21" pageNumber="76">Bosnia and Herzegovina</collectingCountry>
............................................................... ..........................................................................
<taxonomicName id="BDEDC0B0FF9B4B2FD823BF86FAB2FD50" ID-CoL="4MRRR" authority="(Apfelbeck, 1901)" baseAuthorityName="Apfelbeck" baseAuthorityYear="1901" box="[796,1399,647,674]" class="Insecta" family="Leiodidae" genus="Proleonhardella" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="neumanni" subGenus="Proleonhardella">
<emphasis id="48996721FF9B4B2FD823BF86FB5DFD50" bold="true" box="[796,1176,647,674]" pageId="21" pageNumber="76">
<emphasis id="48996721FF9B4B2FD823BF86FCEBFD53" bold="true" box="[796,814,648,674]" italics="true" pageId="21" pageNumber="76">P</emphasis>
. (
<emphasis id="48996721FF9B4B2FD879BF89FBCEFD53" bold="true" box="[838,1035,647,674]" italics="true" pageId="21" pageNumber="76">Proleonhardella</emphasis>
)
<emphasis id="48996721FF9B4B2FDF23BF89FB5DFD50" bold="true" box="[1052,1176,647,673]" italics="true" pageId="21" pageNumber="76">neumanni</emphasis>
</emphasis>
(
<bibRefCitation id="1E7CC6C2FF9B4B2FDF98BF89FAABFD50" author="Apfelbeck V." box="[1191,1390,647,673]" pageId="21" pageNumber="76" pagination="14 - 16" refId="ref12344" refString="Apfelbeck V. 1901. Drei neue Hohlenkafer aus Bosnien. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 51: 14 - 16." type="journal article" year="1901">Apfelbeck, 1901</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7A52BB33FF9B4B2FDBFEBFA5FAB2FCFA" blockId="21.[189,1399,265,780]" pageId="21" pageNumber="76">
<keyLead id="C11C1806FF9B4B2FDBFEBFA5FAB2FCFA" pageId="21" pageNumber="76">
– A1/A2 M 0.76. Mesosternal carina obtuse-angled. Elytral lateral margins below humeral angles sub-parallel. Maximum width of elytra slightly after basal third. Cavernicolous, Mt Tara, western
<collectingCountry id="02FAFBA3FF9B4B2FDBD3BFFFFEF2FCFA" box="[236,311,753,779]" name="Serbia" pageId="21" pageNumber="76">Serbia</collectingCountry>
....................................................
<taxonomicName id="BDEDC0B0FF9B4B2FD9F6BFFFFAD0FCFA" ID-CoL="9GCPC" authority="Curcic &amp; Pavicevic" authorityName="Curcic &amp; Pavicevic" authorityYear="2021" box="[713,1301,748,780]" class="Insecta" family="Leiodidae" genus="Proleonhardella" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="tarensis" status="sp. nov." subGenus="Proleonhardella">
<emphasis id="48996721FF9B4B2FD9F6BFFFFBEDFCFA" bold="true" box="[713,1064,752,779]" pageId="21" pageNumber="76">
<emphasis id="48996721FF9B4B2FD9F6BFFFFD1EFCFA" bold="true" box="[713,731,753,779]" italics="true" pageId="21" pageNumber="76">P</emphasis>
. (
<emphasis id="48996721FF9B4B2FD9CCBFFEFC7DFCFA" bold="true" box="[755,952,752,779]" italics="true" pageId="21" pageNumber="76">Proleonhardella</emphasis>
)
<emphasis id="48996721FF9B4B2FD8F6BFFFFBEDFCFA" bold="true" box="[969,1064,753,779]" italics="true" pageId="21" pageNumber="76">tarensis</emphasis>
</emphasis>
Ćurčić &amp; Pavićević
</taxonomicName>
<taxonomicNameLabel id="53AADA5AFF9B4B2FDE23BFFFFAB2FCFA" box="[1308,1399,753,779]" pageId="21" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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