178 lines
22 KiB
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178 lines
22 KiB
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<document id="5EB323C9427EB36FF51275D7CE597919" ID-CLB-Dataset="7301" ID-DOI="10.1051/acarologia/20174164" ID-GBIF-Dataset="64acea26-7e13-44f4-906a-c723ec908b22" ID-ISSN="2107-7207" ID-Zenodo-Dep="4640309" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616776713369" checkinUser="carolina" docAuthor="Gwiazdowicz, D. J. & Teodorowicz, E." docDate="2017" docId="27302249F3352A046FC0F9093E8B2A65" docLanguage="en" docName="Acarologia.57.2.393-406.pdf" docOrigin="Acarologia 57 (2)" docSource="http://dx.doi.org/10.1051/acarologia/20174164" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Halolaelaps Berlese et Trouessart 1889" docType="treatment" docVersion="8" lastPageNumber="394" masterDocId="DB095A31F3372A076F77FFD93C1F2B17" masterDocTitle="Description of Halolaelaps coulsoni n. sp. (Acari, Halolaelapidae) from the High Arctic" masterLastPageNumber="406" masterPageNumber="393" pageNumber="394" updateTime="1698935196593" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:titleInfo id="634D4FD9AB2492307176833F233D0D19">
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<mods:title id="0EB50E4702A3C5E597E70C7E95695DD4">Description of Halolaelaps coulsoni n. sp. (Acari, Halolaelapidae) from the High Arctic</mods:title>
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<mods:namePart id="9ADCCAA17E85392DC2045736A23E1904">Gwiazdowicz, D. J.</mods:namePart>
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<mods:namePart id="1E76A9D72868EB829925E56C765AC16F">Teodorowicz, E.</mods:namePart>
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<mods:title id="8C3EAC5967201738BEF45D9A8BB44D33">Acarologia</mods:title>
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<mods:part id="3CCA3F365B2642AB1D447CDF6854AFC2">
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<mods:date id="7EBB7CE3C405145E62995E84F309D65D">2017</mods:date>
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<mods:number id="F8D9A47A480383A1C4D9A693AEEB02C8">2017-04-20</mods:number>
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<mods:number id="5B42309D3F1A4E71362053DAFBF20A10">57</mods:number>
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<treatment id="27302249F3352A046FC0F9093E8B2A65" ID-DOI="http://doi.org/10.5281/zenodo.4697164" ID-GBIF-Taxon="181020967" ID-Zenodo-Dep="4697164" LSID="urn:lsid:plazi:treatment:27302249F3352A046FC0F9093E8B2A65" httpUri="http://treatment.plazi.org/id/27302249F3352A046FC0F9093E8B2A65" lastPageId="3" lastPageNumber="394" pageId="2" pageNumber="394">
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<subSubSection id="E783C0D4F3352A056FC0F9093EFE2DFF" box="[183,737,1744,1769]" pageId="2" pageNumber="394" type="nomenclature">
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<paragraph id="AF26935FF3352A056FC0F9093EFE2DFF" blockId="2.[183,737,1744,1769]" box="[183,737,1744,1769]" pageId="2" pageNumber="394">
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<heading id="F46E2433F3352A056FC0F9093EFE2DFF" bold="true" box="[183,737,1744,1769]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="394" reason="2">
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<emphasis id="9DED4F4DF3352A056FC0F9093EFE2DFF" bold="true" box="[183,737,1744,1769]" pageId="2" pageNumber="394">
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Genus
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<taxonomicName id="6899E8DCF3352A056E79F9093EFE2DFF" authority="Berlese et Trouessart, 1889" authorityName="Berlese et Trouessart" authorityYear="1889" box="[270,737,1744,1769]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
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<emphasis id="9DED4F4DF3352A056E79F9093D802DFF" bold="true" box="[270,415,1744,1768]" italics="true" pageId="2" pageNumber="394">Halolaelaps</emphasis>
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<bibRefCitation id="CB08EEAEF3352A056ED2F9083EFE2DFF" author="Berlese A. & Trouessart E." box="[421,737,1744,1769]" pageId="2" pageNumber="394" pagination="121 - 143" refId="ref4881" refString="Berlese A., Trouessart E. 1889 - Diagnoses d'Acariens nouveaux oupeuconnus - Bull. Bibl. Scient. Ouest, 2: 121 - 143." type="journal article" year="1889">Berlese et Trouessart, 1889</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E783C0D4F3352A056FE8F8D73D7F2C5E" pageId="2" pageNumber="394" type="reference_group">
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<paragraph id="AF26935FF3352A056FE8F8D73D7F2C5E" blockId="2.[159,762,1806,1949]" pageId="2" pageNumber="394">
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Synonyms:
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<taxonomicName id="6899E8DCF3352A056E5AF8D73E962C31" authority="Oudemans, 1939" authorityName="Oudemans" authorityYear="1939" box="[301,649,1806,1830]" class="Arachnida" family="Halolaelapidae" genus="Saintdidieria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
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<emphasis id="9DED4F4DF3352A056E5AF8D73DA72C31" box="[301,440,1806,1830]" italics="true" pageId="2" pageNumber="394">Saintdidieria</emphasis>
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Oudemans, 1939
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</taxonomicName>
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;
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<taxonomicName id="6899E8DCF3352A056DE1F8D73D7F2C5E" authority="Leitner 1946" authorityName="Leitner" authorityYear="1946" class="Arachnida" family="Halolaelapidae" genus="Saprolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
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<emphasis id="9DED4F4DF3352A056DE1F8D73CD52C5E" italics="true" pageId="2" pageNumber="394">Saprolaelaps</emphasis>
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Leitner 1946
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="E783C0D4F3352A056FB2F8BB3EE92C8A" pageId="2" pageNumber="394" type="type_taxon">
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<paragraph id="AF26935FF3352A056FB2F8BB3EE92C8A" blockId="2.[159,762,1806,1949]" pageId="2" pageNumber="394">
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<typeStatus id="70222DFDF3352A056FB2F8BB3CE12C6D" box="[197,254,1890,1914]" pageId="2" pageNumber="394">Type</typeStatus>
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species:
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<taxonomicName id="6899E8DCF3352A056E02F8BB3ECA2C6E" authority="Brady, 1875" authorityName="Brady" authorityYear="1875" box="[373,725,1889,1913]" class="Arachnida" family="Ascidae" genus="Gamasus" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="marinus">
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<emphasis id="9DED4F4DF3352A056E02F8BB3E222C6E" box="[373,573,1890,1913]" italics="true" pageId="2" pageNumber="394">Gamasus marinus</emphasis>
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Brady, 1875
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</taxonomicName>
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(=
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<taxonomicName id="6899E8DCF3352A056FE8F85D3EF12C8A" authority="Berlese et Trouessart, 1889" authorityName="Berlese et Trouessart" authorityYear="1889" box="[159,750,1924,1949]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="glabriusculus">
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<emphasis id="9DED4F4DF3352A056FE8F85D3DAC2C8B" box="[159,435,1924,1948]" italics="true" pageId="2" pageNumber="394">Halolaelaps glabriusculus</emphasis>
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<bibRefCitation id="CB08EEAEF3352A056ECDF85D3EF12C8A" author="Berlese A. & Trouessart E." box="[442,750,1924,1949]" pageId="2" pageNumber="394" pagination="121 - 143" refId="ref4881" refString="Berlese A., Trouessart E. 1889 - Diagnoses d'Acariens nouveaux oupeuconnus - Bull. Bibl. Scient. Ouest, 2: 121 - 143." type="journal article" year="1889">Berlese et Trouessart, 1889</bibRefCitation>
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</taxonomicName>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="E783C0D4F3352A056C17FEEE3F942E4D" pageId="2" pageNumber="394" type="description">
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<paragraph id="AF26935FF3352A056C17FEEE395C2FAC" blockId="2.[826,1429,311,1212]" pageId="2" pageNumber="394">
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The genus
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<taxonomicName id="6899E8DCF3352A056CAAFEEE38462A58" authorityName="Berlese et Trouessart" authorityYear="1889" box="[989,1113,311,335]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
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<emphasis id="9DED4F4DF3352A056CAAFEEE38462A58" box="[989,1113,311,335]" italics="true" pageId="2" pageNumber="394">Halolaelaps</emphasis>
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</taxonomicName>
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was created by
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<bibRefCitation id="CB08EEAEF3352A056A78FEEE38182A65" author="Berlese A. & Trouessart E." pageId="2" pageNumber="394" pagination="121 - 143" refId="ref4881" refString="Berlese A., Trouessart E. 1889 - Diagnoses d'Acariens nouveaux oupeuconnus - Bull. Bibl. Scient. Ouest, 2: 121 - 143." type="journal article" year="1889">Berlese and Trouessart (1889)</bibRefCitation>
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, and later complimented by succeeding authors (
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<bibRefCitation id="CB08EEAEF3352A056B7FFEA438C32A82" author="Hirschmann W." box="[1032,1244,381,405]" pageId="2" pageNumber="394" pagination="21 - 24" refId="ref5824" refString="Hirschmann W. 1966 - Die Gattung Halolaelaps Berlese & Trouessart 1889 nov. comb - Acarologie, 9: 21 - 24." type="journal article" year="1966">Hirschmann 1966</bibRefCitation>
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,
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<bibRefCitation id="CB08EEAEF3352A056B9EFEA43F6D2AAF" author="Evans G. O. & Till W. M." pageId="2" pageNumber="394" pagination="139 - 270" refId="ref5608" refString="Evans G. O., Till W. M. 1979 - Mesostigmatic mites of Britain and Ireland (Chelicerata: Acari- Parasitiformes). An introduction to their external morphology and classification - Trans. Zool. Soc. Lond., 35: 139 - 270. doi: 10.1111 / j. 1096 - 3642.1979. tb 00059. x" type="journal article" year="1979">Evans and Till 1979</bibRefCitation>
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,
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<bibRefCitation id="CB08EEAEF3352A056CF3FE79391F2AAF" author="Bregetova N. G. & Shcherbak G. I." box="[900,1280,416,440]" pageId="2" pageNumber="394" pagination="256 - 307" refId="ref5127" refString="Bregetova N. G., Shcherbak G. I. 1977 - Fam. Rhodacaridae - In: Gilyarov M. S., Bregetova N. G. (Eds) A Key to the Soil-Inhabiting Mites of the Mesostigmata - Nauka, Leningrad, p. 256 - 307. [In Russian]" type="book chapter" year="1977">
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<taxonomicName id="6899E8DCF3352A056CF3FE793FE52AAF" authority="(Eds)" authorityName="Gwiazdowicz & Teodorowicz" authorityYear="2017" baseAuthorityName="Eds" box="[900,1018,416,440]" genus="Bregetova" pageId="2" pageNumber="406" rank="genus" status="gen. nov.">Bregetova</taxonomicName>
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and Shcherbak 1977
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</bibRefCitation>
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,
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<bibRefCitation id="CB08EEAEF3352A056A64FE79398E2AAF" author="Karg W." box="[1299,1425,416,440]" pageId="2" pageNumber="394" pagination="523" refId="ref5870" refString="Karg W. 1993 - Acari (Acarina), Milben Parasitiformes (Anactinochaeta), Cohors Gamasina Leach. Raubmilben - Die Tierwelt Deutschlands, 59 Teil. Gustav Fischer Verlag, Jena, pp. 523." type="journal article" year="1993">Karg 1993</bibRefCitation>
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,
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<bibRefCitation id="CB08EEAEF3352A056C4DFE1A3FF92ACC" author="Halliday R. B." box="[826,998,451,475]" pageId="2" pageNumber="394" pagination="214 - 230" refId="ref5783" refString="Halliday R. B. 2008 - Two new species of littoral mites from Australia (Acari: Halolaelapidae) - Syst. Appl. Acarol., 13: 214 - 230. doi: 10.11158 / saa. 13.3.8" type="journal article" year="2008">Halliday 2008</bibRefCitation>
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). A typical character of
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<taxonomicName id="6899E8DCF3352A056A6EFE1A398A2ACC" authorityName="Berlese et Trouessart" authorityYear="1889" box="[1305,1429,451,475]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
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<emphasis id="9DED4F4DF3352A056A6EFE1A398A2ACC" box="[1305,1429,451,475]" italics="true" pageId="2" pageNumber="394">Halolaelaps</emphasis>
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</taxonomicName>
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are two subequal dorsal shields, the idiosoma lacking horn-like caudal processes. The opisthonotal shield of many species bears a median incision running posteriorly from an anterior boarder towards the inner area of the shield. Dorsal setae commonly are simple or pilose. Tritosternum biflagellate with columnar base. Sternal shield subrectangular, fused with a presternal plate but not with endopodal plates, bearing three pairs of simple setae (st1-3) and two pairs of lyrifissures (iv1- 2). Pair of st4 setae situated in a soft membrane or on the metasternal plates and pair st5 on a trapezoidal genital shield. Peritremes either short or long and peritrematal shields not expanded posteriorly; ventrianal shield broad, bearing 3-4 pairs of ventral setae or reduced to anal shield with three circumanal setae; metapodal plates varied in shape; palp genu with 6 setae, palp apotele 3-tined, corniculi horn-like. Chelicerae chelate-dentate; movable digit without excrescences. Tectum with a median prong either evenly trifurcate or denticulate.
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</paragraph>
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<paragraph id="AF26935FF3352A056C17FB003F942E4D" blockId="2.[826,1429,1240,1370]" pageId="2" pageNumber="394">Tarsus I with claws; trochanter I with 6 setae; femur I with 13 setae; genu I with 12 setae; tibia I with 11 setae; genu III with 9 setae; coxae sometimes with spines.</paragraph>
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</subSubSection>
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<subSubSection id="E783C0D4F3352A046C17FAAF3E8B2A65" lastPageId="3" lastPageNumber="395" pageId="2" pageNumber="394" type="discussion">
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<paragraph id="AF26935FF3352A046C17FAAF3E8B2A65" blockId="2.[826,1429,1398,1949]" lastBlockId="3.[159,762,311,370]" lastPageId="3" lastPageNumber="395" pageId="2" pageNumber="394">
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Remarks — A division of the genus
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<taxonomicName id="6899E8DCF3352A056A6EFAAF398A2E99" authorityName="Berlese et Trouessart" authorityYear="1889" box="[1305,1429,1398,1422]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
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<emphasis id="9DED4F4DF3352A056A6EFAAF398A2E99" box="[1305,1429,1398,1422]" italics="true" pageId="2" pageNumber="394">Halolaelaps</emphasis>
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</taxonomicName>
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was made by
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<bibRefCitation id="CB08EEAEF3352A056CA8FA4339532EA5" author="Blaszak C. & Ehrnsberger R." box="[991,1356,1434,1458]" pageId="2" pageNumber="394" pagination="143 - 267" refId="ref4909" refString="Blaszak C., Ehrnsberger R. 1993 - Beitrage zur Kenntnis von Halolaelaps (Saprogamasellus) Gotz, 1952 (Acari: Gamasida: Halolaelapidae) - Genus, 4: 143 - 267." type="journal article" year="1993">Błaszak and Ehrnsberger (1993</bibRefCitation>
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,
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<bibRefCitation id="CB08EEAEF3352A056A2EFA40398E2EA6" author="Blaszak C. & Ehrnsberger R." box="[1369,1425,1433,1457]" pageId="2" pageNumber="394" pagination="25 - 94" refId="ref4945" refString="Blaszak C., Ehrnsberger R. 1995 - Beitrage zur Kenntnis von Halolaelaps (Halogamasellus) subgen. nov., (Acari: Gamasida: Halolaelapidae) - Osnab. Nat. Mit., 20 / 21: 25 - 94." type="journal article" year="1995">1995</bibRefCitation>
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,
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<bibRefCitation id="CB08EEAEF3352A056C4DFA643F9F2EC2" author="Blaszak C. & Ehrnsberger R." box="[826,896,1469,1493]" pageId="2" pageNumber="394" pagination="159 - 181" refId="ref4990" refString="Blaszak C., Ehrnsberger R. 1998 a - Beitrage zur Kenntnis von Halolaelaps (Halolaelaps s. str.), (Acari: Gamasida: Halolaelapidae) - Osnab. Nat. Mit., 24: 159 - 181." type="journal article" year="1998">1998a</bibRefCitation>
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,b) into 4 subgenera (
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<taxonomicName id="6899E8DCF3352A056B0FFA6438EB2EC2" authorityName="Berlese et Trouessart" authorityYear="1889" box="[1144,1268,1469,1493]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="9DED4F4DF3352A056B0FFA6438EB2EC2" box="[1144,1268,1469,1493]" italics="true" pageId="2" pageNumber="394">Halolaelaps</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="86DEF236F3352A056B8CFA64392D2EC3" box="[1275,1330,1469,1492]" pageId="2" pageNumber="394" sensu="stricto">s.str.</taxonomicNameLabel>
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,
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<emphasis id="9DED4F4DF3352A056A48FA643F882EEF" italics="true" pageId="2" pageNumber="394">Halogamasellus</emphasis>
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,
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<taxonomicName id="6899E8DCF3352A056CDBFA39380A2EEF" authority="Gwiazdowicz & Teodorowicz, 2017" authorityName="Gwiazdowicz & Teodorowicz" authorityYear="2017" box="[940,1045,1504,1528]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="405" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Haloseius">
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<emphasis id="9DED4F4DF3352A056CDBFA39380A2EEF" box="[940,1045,1504,1528]" italics="true" pageId="2" pageNumber="394">Haloseius</emphasis>
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</taxonomicName>
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,
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<emphasis id="9DED4F4DF3352A056B5DFA3938C12EEF" box="[1066,1246,1504,1528]" italics="true" pageId="2" pageNumber="394">Saprogamasellus</emphasis>
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) based on the shape of epistome, the number of setae on femur I, and a spine on coxa II. Those designations and the characters used raised some multiple questions and controversies. For instance, why a type-species for
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<taxonomicName id="6899E8DCF3352A056C11F9563FFD2DB0" authorityName="Berlese et Trouessart" authorityYear="1889" box="[870,994,1679,1703]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus">
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<emphasis id="9DED4F4DF3352A056C11F9563FFD2DB0" box="[870,994,1679,1703]" italics="true" pageId="2" pageNumber="394">Halolaelaps</emphasis>
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</taxonomicName>
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, that is
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<taxonomicName id="6899E8DCF3352A056B3DF95639352DB0" baseAuthorityName="Brady" baseAuthorityYear="1875" box="[1098,1322,1679,1703]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="394" phylum="Arthropoda" rank="species" species="marinus">
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<emphasis id="9DED4F4DF3352A056B3DF95639352DB0" box="[1098,1322,1679,1703]" italics="true" pageId="2" pageNumber="394">Halolaelaps marinus</emphasis>
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</taxonomicName>
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, has not been placed into the subgenus
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<taxonomicName id="6899E8DCF3352A056BDEF96B393A2DDD" authorityName="Berlese et Trouessart" authorityYear="1889" box="[1193,1317,1714,1738]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="394" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="9DED4F4DF3352A056BDEF96B393A2DDD" box="[1193,1317,1714,1738]" italics="true" pageId="2" pageNumber="394">Halolaelaps</emphasis>
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</taxonomicName>
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|
<taxonomicNameLabel id="86DEF236F3352A056A5AF96A397D2DDD" box="[1325,1378,1715,1738]" pageId="2" pageNumber="394" sensu="stricto">s.str.</taxonomicNameLabel>
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but into subgenus
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<emphasis id="9DED4F4DF3352A056C99F90C38872DFA" box="[1006,1176,1749,1773]" italics="true" pageId="2" pageNumber="394">Hologamasellus</emphasis>
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(
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<bibRefCitation id="CB08EEAEF3352A056BDDF90C3FDA2C07" author="Blaszak C. & Ehrnsberger R." pageId="2" pageNumber="394" pagination="25 - 94" refId="ref4945" refString="Blaszak C., Ehrnsberger R. 1995 - Beitrage zur Kenntnis von Halolaelaps (Halogamasellus) subgen. nov., (Acari: Gamasida: Halolaelapidae) - Osnab. Nat. Mit., 20 / 21: 25 - 94." type="journal article" year="1995">Błaszak and Ehrnsberger 1995</bibRefCitation>
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) or why a
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<typeStatus id="70222DFDF3352A056B38F920389C2C06" box="[1103,1155,1785,1809]" pageId="2" pageNumber="394">type</typeStatus>
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species have not been designated for the four subgenera that had been erected. All the controversial issues on this subject have been presented by
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<bibRefCitation id="CB08EEAEF3352A056B09F8B8392B2C6D" author="Halliday R. B." box="[1150,1332,1889,1914]" pageId="2" pageNumber="394" pagination="214 - 230" refId="ref5783" refString="Halliday R. B. 2008 - Two new species of littoral mites from Australia (Acari: Halolaelapidae) - Syst. Appl. Acarol., 13: 214 - 230. doi: 10.11158 / saa. 13.3.8" type="journal article" year="2008">Halliday (2008)</bibRefCitation>
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, and the authors of this paper agree with them completely. Therefore the discussion here does not follow the division of
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<taxonomicName id="6899E8DCF3342A046E54FE833D802A65" authorityName="Berlese et Trouessart" authorityYear="1889" box="[291,415,346,370]" class="Arachnida" family="Halolaelapidae" genus="Halolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="395" phylum="Arthropoda" rank="genus">
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<emphasis id="9DED4F4DF3342A046E54FE833D802A65" box="[291,415,346,370]" italics="true" pageId="3" pageNumber="395">Halolaelaps</emphasis>
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</taxonomicName>
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into four subgenera.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |