578 lines
73 KiB
XML
578 lines
73 KiB
XML
<document id="C44D7717D9CA3A1AA05AC165C915EAB7" ID-DOI="10.25674/so95iss1id311" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes & Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFD3FFEDBF14FAB50FAB07D0" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docTitle="Scopaeus diversilobatus Frisch 2022, spec. nov." docType="treatment" docVersion="2" lastPageNumber="48" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="42" updateTime="1709147743548" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C5A55077964155C28C848A37BB69EF61">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:name id="CD8FC3B6888D467D9EBA80B9069E24AC" type="personal">
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<mods:namePart id="170B88B12650C319642168501BD3D07A">Frisch, Johannes</mods:namePart>
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<mods:name id="BCE53EF7659A1BAE5C7E20DB2FBDCEF8" type="personal">
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<mods:roleTerm id="E2B574D3918EE4D1E2CC4C9BB14FE650">Author</mods:roleTerm>
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<mods:namePart id="DCA24E55D18A857DCE7F35421B0166B4">Narakusumo, Raden Pramesa</mods:namePart>
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<mods:affiliation id="37E35B35CDBE553547A43D4D21947E39">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:typeOfResource id="2FE8098786AE2951A11E5A220F857F33">text</mods:typeOfResource>
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<mods:title id="757153FD8FB2988754EF61EE4D9053C6">Soil Organisms</mods:title>
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<mods:part id="0443393B75ADF28D21E47DAB834974D9">
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<mods:date id="55733EC067639BBEF5881AD0BA738A70">2022</mods:date>
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<mods:number id="2850D3692F70EAF6D08E6BEC9FDA8834">2022-04-01</mods:number>
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<mods:number id="D90C767611202A0D5FE539773051E2C1">95</mods:number>
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<mods:url id="75E5E42E5B266DD2C40915CA0E31C89F">http://dx.doi.org/10.25674/so95iss1id311</mods:url>
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<mods:classification id="1634C1C10B5D0FD9889F9E0A7D436040">journal article</mods:classification>
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<mods:identifier id="64A4D1666F7BF0915E79966DA1E00F4B" type="DOI">10.25674/so95iss1id311</mods:identifier>
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<treatment id="03B7C77AFFD3FFEDBF14FAB50FAB07D0" ID-DOI="http://doi.org/10.5281/zenodo.10724307" ID-Zenodo-Dep="10724307" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFEDBF14FAB50FAB07D0" httpUri="http://treatment.plazi.org/id/03B7C77AFFD3FFEDBF14FAB50FAB07D0" lastPageId="25" lastPageNumber="48" pageId="19" pageNumber="42">
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<subSubSection id="C30425E7FFD3FFE7BF14FAB508530165" box="[847,1358,1357,1384]" pageId="19" pageNumber="42" type="nomenclature">
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<paragraph id="8BA1766CFFD3FFE7BF14FAB508530165" blockId="19.[847,1358,1357,1417]" box="[847,1358,1357,1384]" pageId="19" pageNumber="42">
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<heading id="D0E9C100FFD3FFE7BF14FAB508530165" box="[847,1358,1357,1384]" fontSize="11" level="2" pageId="19" pageNumber="42" reason="5">
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<taxonomicName id="4C1E0DEFFFD3FFE7BF14FAB509D1016A" authority="Frisch" authorityName="Frisch" authorityYear="2022" box="[847,1228,1357,1384]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="diversilobatus" status="sp. nov.">
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<emphasis id="B96AAA7EFFD3FFE7BF14FAB509D1016A" bold="true" box="[847,1228,1357,1384]" pageId="19" pageNumber="42">
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<emphasis id="B96AAA7EFFD3FFE7BF14FAB509690165" bold="true" box="[847,1140,1357,1384]" italics="true" pageId="19" pageNumber="42">Scopaeus diversilobatus</emphasis>
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Frisch
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</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2591705FFD3FFE7B881FAB608530165" box="[1242,1358,1358,1384]" pageId="19" pageNumber="42" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30425E7FFD3FFE7BF14FA9609FF0184" box="[847,1250,1390,1417]" pageId="19" pageNumber="42" type="description">
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<paragraph id="8BA1766CFFD3FFE7BF14FA9609FF0184" blockId="19.[847,1358,1357,1417]" box="[847,1250,1390,1417]" pageId="19" pageNumber="42">
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(
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<figureCitation id="13256AE9FFD3FFE7BF0CFA960EB30184" box="[855,942,1390,1417]" captionStart="Figures 14–16" captionStartId="9.[189,262,1930,1952]" captionTargetBox="[262,1384,910,1880]" captionTargetId="figure-57@9.[189,1436,894,1911]" captionTargetPageId="9" captionText="Figures 14–16. Habitus of Scopaeus heronifer, paratype, Jawa Barat: Mt Salak (14); S. diversilobatus, paratype, Jawa Barat, Sukabumi: Djampang Kulon (15); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (16)." figureDoi="http://doi.org/10.5281/zenodo.10724246" httpUri="https://zenodo.org/record/10724246/files/figure.png" pageId="19" pageNumber="42">Figs 15</figureCitation>
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,
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<figureCitation id="13256AE9FFD3FFE7BFE0FA970EC40184" box="[955,985,1391,1417]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="19" pageNumber="42">39</figureCitation>
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,
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<figureCitation id="13256AE9FFD3FFE7BFBEFA9709290184" box="[997,1076,1391,1417]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="19" pageNumber="42">89–91</figureCitation>
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,
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<figureCitation id="13256AE9FFD3FFE7B81BFA9709710184" box="[1088,1132,1391,1417]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="19" pageNumber="42">133</figureCitation>
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,
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<figureCitation id="13256AE9FFD3FFE7B823FA9609C60184" box="[1144,1243,1390,1417]" captionStart="Figures 163–172" captionStartId="42.[151,224,1839,1861]" captionTargetBox="[225,1322,230,1817]" captionTargetId="figure-168@42.[157,1398,210,1820]" captionTargetPageId="42" captionText="Figures 163–172. Sperm pump (163–172) and bursa (165, 167, 170, 172) of ♀ of Scopaeus grandis, paratype, Jawa Barat, Sukabumi: Cidahu (163); S.heronifer, paratype, Jawa Barat, Sukabumi:Sirnarasa (164); S. diversilobatus, paratype, Jawa Barat, Sukabumi, Djampang Kulon (165, with lateral gonocoxal plates IX folded laterally; 166); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (167, with lateral gonocoxal plates IX, 168); S. velifer, paratype, Banten, Lebak: Citorek Kidul (169); S. elegantulus, Jawa Barat, Sukabumi: Djampang Kulon (170, with lateral gonocoxal plates IX, 171), S. uncinatus, paratype, Banten, Lebak: Citorek Kidul (172). Long scale bar: 163, 164, 166, 168, 169, 171, 172; short scale bar: 165, 167, 170." figureDoi="http://doi.org/10.5281/zenodo.10724315" httpUri="https://zenodo.org/record/10724315/files/figure.png" pageId="19" pageNumber="42">165, 166</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30425E7FFD3FFE7BF14FA4A091502FF" pageId="19" pageNumber="42" type="materials_examined">
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<paragraph id="8BA1766CFFD3FFE7BF14FA4A091502FF" blockId="19.[824,1437,1458,1975]" pageId="19" pageNumber="42">
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||
<emphasis id="B96AAA7EFFD3FFE7BF14FA4A090101C7" bold="true" box="[847,1052,1458,1482]" pageId="19" pageNumber="42">Type specimens:</emphasis>
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||
<materialsCitation id="3B767C31FFD3FFE7B86FFA4A0E9A0220" collectingDate="2015-09-25" collectionCode="MZB" collectorName="Sukabumi, N & Frisch" country="Indonesia" elevation="250" latitude="-7.2341666" location="Cikarang River" longLatPrecision="21" longitude="106.61361" pageId="19" pageNumber="42" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Barat" typeStatus="holotype">
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||
<typeStatus id="54A5C8CEFFD3FFE7B86FFA4A098501C7" box="[1076,1176,1458,1482]" pageId="19" pageNumber="42" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D18BDE5FFD3FFE7B8EBFA4B09D901C7" box="[1200,1220,1459,1482]" count="1" pageId="19" pageNumber="42" type="male">♂</specimenCount>
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,
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||
<collectingCountry id="F30936FCFFD3FFE7B8BBFA4A085101C7" box="[1248,1356,1458,1482]" name="Indonesia" pageId="19" pageNumber="42">Indonesia</collectingCountry>
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,
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||
<collectingRegion id="49DAB88EFFD3FFE7B933FA4B0E6B01E7" country="Indonesia" name="Jawa Barat" pageId="19" pageNumber="42">Jawa Barat</collectingRegion>
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,
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||
<collectorName id="26EB13BAFFD3FFE7BFDFFA2B090A01E7" box="[900,1047,1491,1515]" pageId="19" pageNumber="42">Sukabumi, N</collectorName>
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Djampang Kulon:
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<location id="8EC120B7FFD3FFE7B8A9FA2B088101E6" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFEDBF14FAB50FAB07D0:8EC120B7FFD3FFE7B8A9FA2B088101E6" box="[1266,1436,1491,1515]" country="Indonesia" latitude="-7.2341666" longLatPrecision="21" longitude="106.61361" name="Cikarang River" pageId="19" pageNumber="42" stateProvince="Jawa Barat">Cikarang River</location>
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(
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<geoCoordinate id="EE2A10ABFFD3FFE7BF1BFA0C0EA20201" box="[832,959,1524,1548]" degrees="07" direction="south" minutes="14" orientation="latitude" pageId="19" pageNumber="42" precision="15" seconds="03" value="-7.2341666">07°14’03’’S</geoCoordinate>
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,
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<geoCoordinate id="EE2A10ABFFD3FFE7BF91FA0C094A0201" box="[970,1111,1524,1548]" degrees="106" direction="east" minutes="36" orientation="longitude" pageId="19" pageNumber="42" precision="15" seconds="49" value="106.61361">106°36’49’’E</geoCoordinate>
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),
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<quantity id="4CE6DB89FFD3FFE7B831FA0C09B20206" box="[1130,1199,1524,1548]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="42" unit="m" value="250.0">
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<elevation id="0033915FFFD3FFE7B831FA0C09B20206" box="[1130,1199,1524,1548]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="42" unit="m" value="250.0">250 m</elevation>
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</quantity>
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,
|
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<date id="FFA050ACFFD3FFE7B8E1FA0C083C0201" box="[1210,1313,1524,1548]" pageId="19" pageNumber="42" value="2015-09-25">
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<collectingDate id="EFE4A944FFD3FFE7B8E1FA0C083C0201" box="[1210,1313,1524,1548]" pageId="19" pageNumber="42" value="2015-09-25">25.9.2015</collectingDate>
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</date>
|
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, leg.
|
||
<collectorName id="26EB13BAFFD3FFE7B902FA0C08810201" box="[1369,1436,1524,1548]" pageId="19" pageNumber="42">Frisch</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD3FFE7BF1BF9ED0E620220" box="[832,895,1557,1581]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="19" pageNumber="42" type="Museum">MZB</collectionCode>
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)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="54A5C8CEFFD3FFE7BFCBF9ED0EE60220" box="[912,1019,1557,1581]" pageId="19" pageNumber="42" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<specimenCount id="9D18BDE5FFD3FFE7B852F9ED09800220" box="[1033,1181,1557,1581]" count="10" pageId="19" pageNumber="42" type="generic">10 specimens</specimenCount>
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||
):
|
||
<materialsCitation id="3B767C31FFD3FFE7B8F5F9ED09E90243" collectingDate="2015-09-25" collectionCode="MFNB, MZB" collectorName="Sukabumi, N & Frisch" country="Indonesia" elevation="250" latitude="-7.2341666" location="Cikarang River" longLatPrecision="21" longitude="106.61361" pageId="19" pageNumber="42" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Bali" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFD3FFE7B8F5F9ED09CB0221" box="[1198,1238,1557,1581]" count="3" pageId="19" pageNumber="42" type="male">3 ♂</specimenCount>
|
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,
|
||
<specimenCount id="9D18BDE5FFD3FFE7B8BAF9ED081A0221" box="[1249,1287,1557,1581]" count="4" pageId="19" pageNumber="42" type="female">4 ♀</specimenCount>
|
||
, same data as holotype (
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<collectionCode id="ED0FEEA9FFD3FFE7BFFFF9CE0EEB0243" box="[932,1014,1590,1614]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="19" pageNumber="42">MFNB</collectionCode>
|
||
,
|
||
<collectionCode id="ED0FEEA9FFD3FFE7BFA4F9CE09200243" box="[1023,1085,1590,1614]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="19" pageNumber="42" type="Museum">MZB</collectionCode>
|
||
).
|
||
<collectingCountry id="F30936FCFFD3FFE7B815F9CE09A70243" box="[1102,1210,1590,1614]" name="Indonesia" pageId="19" pageNumber="42">Indonesia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49DAB88EFFD3FFE7B898F9CE09E90243" box="[1219,1268,1590,1614]" country="Indonesia" name="Bali" pageId="19" pageNumber="42">Bali</collectingRegion>
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="3B767C31FFD3FFE7B8A7F9CE09170282" collectionCode="PWCM" collectorName="Jembrana & Cekik & Wunderle" country="Malaysia" elevation="1994300" location="Malaysia" pageId="19" pageNumber="42" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFD3FFE7B8A7F9CE083E0240" box="[1276,1315,1590,1614]" count="1" pageId="19" pageNumber="42" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectorName id="26EB13BAFFD3FFE7B975F9CE08870243" box="[1326,1434,1590,1614]" pageId="19" pageNumber="42">Jembrana</collectorName>
|
||
:
|
||
<collectorName id="26EB13BAFFD3FFE7BF63F9AE0E610263" box="[824,892,1622,1646]" pageId="19" pageNumber="42">Cekik</collectorName>
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||
,
|
||
<date id="FFA050ACFFD3FFE7BFC8F9AE0EE00263" box="[915,1021,1622,1646]" pageId="19" pageNumber="42" value="1994-06-20">
|
||
20.6.
|
||
<quantity id="4CE6DB89FFD3FFE7BF9CF9AE097B0263" box="[967,1126,1622,1647]" metricMagnitude="6" metricUnit="m" metricValue="1.9943" pageId="19" pageNumber="42" unit="m" value="1994300.0">
|
||
<elevation id="0033915FFFD3FFE7BF9CF9AE097B0263" box="[967,1126,1622,1647]" metricMagnitude="6" metricUnit="m" metricValue="1.9943" pageId="19" pageNumber="42" unit="m" value="1994300.0">1994, 300 m</elevation>
|
||
</quantity>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFD3FFE7B8ECF9AE083D0263" box="[1207,1312,1622,1646]" pageId="19" pageNumber="42">Wunderle</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD3FFE7B960F9AF088F0262" box="[1339,1426,1623,1647]" pageId="19" pageNumber="42">PWCM</collectionCode>
|
||
).
|
||
<collectingCountry id="F30936FCFFD3FFE7BF63F98F0E800282" box="[824,925,1655,1679]" name="Malaysia" pageId="19" pageNumber="42">Malaysia</collectingCountry>
|
||
, Sarawak
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="3B767C31FFD3FFE7B84EF98F09A202BD" collectingDate="1981-04-17" collectionCode="HECO" collectorName="Rougemont" country="Malaysia" location="San Uncap Undong" municipality="Undong" pageId="19" pageNumber="42" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFD3FFE7B84EF98F09230282" box="[1045,1086,1655,1679]" count="1" pageId="19" pageNumber="42" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC5EC16FFD3FFE7B811F98F09BD0282" box="[1098,1184,1655,1679]" pageId="19" pageNumber="42">Undong</collectingMunicipality>
|
||
(
|
||
<location id="8EC120B7FFD3FFE7B8F4F98F089A0282" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFEDBF14FAB50FAB07D0:8EC120B7FFD3FFE7B8F4F98F089A0282" box="[1199,1415,1655,1679]" country="Malaysia" municipality="Undong" name="San Uncap Undong" pageId="19" pageNumber="42">San Uncap Undong</location>
|
||
?),
|
||
<date id="FFA050ACFFD3FFE7BF63F9600E8102BD" box="[824,924,1688,1712]" pageId="19" pageNumber="42" value="1981-04-17">
|
||
<collectingDate id="EFE4A944FFD3FFE7BF63F9600E8102BD" box="[824,924,1688,1712]" pageId="19" pageNumber="42" value="1981-04-17">17.4.1981</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFD3FFE7BF80F961094A02BC" box="[987,1111,1689,1713]" pageId="19" pageNumber="42">Rougemont</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD3FFE7B832F96009AA02BD" box="[1129,1207,1688,1712]" pageId="19" pageNumber="42">HECO</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B767C31FFD3FFE7B896F960091902FF" collectingDate="1978-08-05" collectionCode="NHML" collectorName="Hammond & Marshall" country="Malaysia" elevation="10" location="Mt Mulu" pageId="19" pageNumber="42" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFD3FFE7B896F96009E402BD" box="[1229,1273,1688,1712]" count="1" pageId="19" pageNumber="42" type="male">1 ♂</specimenCount>
|
||
,
|
||
<location id="8EC120B7FFD3FFE7B953F961087602BD" LSID="urn:lsid:plazi:treatment:03B7C77AFFD3FFEDBF14FAB50FAB07D0:8EC120B7FFD3FFE7B953F961087602BD" box="[1288,1387,1688,1712]" country="Malaysia" name="Mt Mulu" pageId="19" pageNumber="42">Mt Mulu</location>
|
||
NP, near base camp, 50 –
|
||
<quantity id="4CE6DB89FFD3FFE7B873F941097F02DC" box="[1064,1122,1721,1745]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="42" unit="m" value="10.0">
|
||
<elevation id="0033915FFFD3FFE7B873F941097F02DC" box="[1064,1122,1721,1745]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="42" unit="m" value="10.0">10 m</elevation>
|
||
</quantity>
|
||
<date id="FFA050ACFFD3FFE7B836F94109D502DC" box="[1133,1224,1721,1745]" pageId="19" pageNumber="42" value="1978-08-05">
|
||
<collectingDate id="EFE4A944FFD3FFE7B836F94109D502DC" box="[1133,1224,1721,1745]" pageId="19" pageNumber="42" value="1978-08-05">5.8.1978</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFD3FFE7B951F941086102DC" box="[1290,1404,1721,1745]" pageId="19" pageNumber="42">Hammond</collectorName>
|
||
&
|
||
<collectorName id="26EB13BAFFD3FFE7BF63F9220E8402FF" box="[824,921,1754,1778]" pageId="19" pageNumber="42">Marshall</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD3FFE7BFFCF9220EE102FF" box="[935,1020,1754,1778]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="19" pageNumber="42">NHML</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFD3FFEDBF14F9030E1406B8" lastPageId="25" lastPageNumber="48" pageId="19" pageNumber="42" type="description">
|
||
<paragraph id="8BA1766CFFD3FFE0BF14F9030CCA05C3" blockId="19.[824,1437,1458,1975]" lastBlockId="20.[151,764,241,528]" lastPageId="20" lastPageNumber="43" pageId="19" pageNumber="42">
|
||
<emphasis id="B96AAA7EFFD3FFE7BF14F9030EC0031E" bold="true" box="[847,989,1787,1811]" pageId="19" pageNumber="42">Description:</emphasis>
|
||
Habitus and coloring as in
|
||
<figureCitation id="13256AE9FFD3FFE7B957F9030844031E" box="[1292,1369,1787,1811]" captionStart="Figures 14–16" captionStartId="9.[189,262,1930,1952]" captionTargetBox="[262,1384,910,1880]" captionTargetId="figure-57@9.[189,1436,894,1911]" captionTargetPageId="9" captionText="Figures 14–16. Habitus of Scopaeus heronifer, paratype, Jawa Barat: Mt Salak (14); S. diversilobatus, paratype, Jawa Barat, Sukabumi: Djampang Kulon (15); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (16)." figureDoi="http://doi.org/10.5281/zenodo.10724246" httpUri="https://zenodo.org/record/10724246/files/figure.png" pageId="19" pageNumber="42">Fig. 15</figureCitation>
|
||
. Head subquadrate. Penultimate antennal segment elongate. Mesotibia moderately thickened. Body surface with fine, dense, setose punctation, without microreticulation, shiny. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body dark brown except for slightly lighter pronotum, yellow-brown posterior fourth of elytra, medium brown abdomen with darkened disk of tergites and light brown tip, yellow brown maxillary palpi with darker penultimate segment, medium brown antennae with darker median segments, and yellow-brown legs. Total body length
|
||
<quantity id="4CE6DB89FFD4FFE0BE34FE6E0FE505A0" box="[623,760,406,430]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.6" pageId="20" pageNumber="43" unit="mm" value="3.75" valueMax="3.9" valueMin="3.6">3.6 – 3.9 mm</quantity>
|
||
; forebody length
|
||
<quantity id="4CE6DB89FFD4FFE0BD12FE4F0CCE05C3" box="[329,467,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="20" pageNumber="43" unit="mm" value="2.2" valueMax="2.3" valueMin="2.1">2.1 – 2.3 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFD4FFEDBCF5FE200F9F05FD" blockId="20.[151,764,241,528]" lastBlockId="25.[189,801,241,989]" lastPageId="25" lastPageNumber="48" pageId="20" pageNumber="43">
|
||
Male: Abdominal sternite VII with unmodified posterior margin. Emargination in posterior fifth of abdominal sternite VIII strongly convex in median third (
|
||
<figureCitation id="13256AE9FFD4FFE0BF47FEEA0E610527" box="[796,892,274,298]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="20" pageNumber="43">Fig. 133</figureCitation>
|
||
). Aedeagus (
|
||
<figureCitation id="13256AE9FFD4FFE0B84DFEEA09730527" box="[1046,1134,274,298]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="20" pageNumber="43">Figs 39</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD4FFE0B8DBFEEA09DA0527" box="[1152,1223,274,298]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="20" pageNumber="43">89 – 91</figureCitation>
|
||
) about 1.0 mm long with distal lobes asymmetrical; dextral apical lobe strongly reduced, as long as dorsal lobe, in lateral view with undulate, convex apical and ventral margins (
|
||
<figureCitation id="13256AE9FFD4FFE0B917FE8D0E3205A3" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="20" pageNumber="43">Fig. 39</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD4FFE0BF60FE6E0E4A05A3" box="[827,855,406,430]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="20" pageNumber="43">89</figureCitation>
|
||
), in ventral and dorsal view almost vestigial with truncate apex (
|
||
<figureCitation id="13256AE9FFD4FFE0BF9AFE4F095805C2" box="[961,1093,439,463]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="20" pageNumber="43">Figs 90, 91</figureCitation>
|
||
); sinistral apical lobe well developed, testaceous, twice as long as dextral apical lobe, in lateral view triangular (
|
||
<figureCitation id="13256AE9FFD4FFE0B838FE0009B3061C" box="[1123,1198,504,529]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="20" pageNumber="43">Fig. 39</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD4FFE0B8ECFE0109CE061C" box="[1207,1235,505,529]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="20" pageNumber="43">89</figureCitation>
|
||
), in ventral and dorsal view convex laterally with somewhat retracted, acute apex (
|
||
<figureCitation id="13256AE9FFD9FFEDBD60FEEA0CAC0527" box="[315,433,274,298]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="25" pageNumber="48">Figs 90, 91</figureCitation>
|
||
); flagellum curved ventrosinistrad with thin end projecting from apical lobes ventrally (
|
||
<figureCitation id="13256AE9FFD9FFEDBEABFECB0DC50561" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="25" pageNumber="48">Figs 39</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD9FFEDBCB1FEAC0C1B0561" box="[234,262,340,364]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="25" pageNumber="48">89</figureCitation>
|
||
); lateral lobes reduced, indicated by group of setae (
|
||
<figureCitation id="13256AE9FFD9FFEDBD5EFE8D0C980580" box="[261,389,373,397]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="25" pageNumber="48">Figs 89, 90</figureCitation>
|
||
); ventral lobe long, slender, adjacent to aedeagus (
|
||
<figureCitation id="13256AE9FFD9FFEDBD08FE6E0CB505A3" box="[339,424,406,430]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="25" pageNumber="48">Figs 39</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD9FFEDBDE3FE6E0CC905A3" box="[440,468,406,430]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="25" pageNumber="48">89</figureCitation>
|
||
), in ventral view narrow and shifted sinistrad (
|
||
<figureCitation id="13256AE9FFD9FFEDBDDFFE4F0CC505C2" box="[388,472,439,463]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="25" pageNumber="48">Fig. 90</figureCitation>
|
||
); transverse ridge of median foramen not projecting ventrally (
|
||
<figureCitation id="13256AE9FFD9FFEDBE73FE200F6A05FD" box="[552,631,472,496]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="25" pageNumber="48">Fig. 89</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6126E4FFD4FFE0BCCDF8A60EE803B5" ID-DOI="http://doi.org/10.5281/zenodo.10724264" ID-Zenodo-Dep="10724264" httpUri="https://zenodo.org/record/10724264/files/figure.png" pageId="20" pageNumber="43" startId="20.[150,223,1886,1908]" targetBox="[222,1327,582,1831]" targetPageId="20" targetType="figure">
|
||
<paragraph id="8BA1766CFFD4FFE0BCCDF8A60EE803B5" blockId="20.[150,1399,1886,1976]" pageId="20" pageNumber="43">
|
||
<emphasis id="B96AAA7EFFD4FFE0BCCDF8A60C39037E" bold="true" box="[150,292,1886,1908]" pageId="20" pageNumber="43">Figures 50–55.</emphasis>
|
||
Apex of aedeagus in lateral (
|
||
<emphasis id="B96AAA7EFFD4FFE0BE6DF8A60F50037E" bold="true" box="[566,589,1886,1907]" pageId="20" pageNumber="43">50</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD4FFE0BE00F8A60F6F037E" bold="true" box="[603,626,1886,1907]" pageId="20" pageNumber="43">53</emphasis>
|
||
), ventral (
|
||
<emphasis id="B96AAA7EFFD4FFE0BE88F8A60FF7037E" bold="true" box="[723,746,1886,1907]" pageId="20" pageNumber="43">51</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD4FFE0BEADF8A60E10037E" bold="true" box="[758,781,1886,1907]" pageId="20" pageNumber="43">54</emphasis>
|
||
), dorsal view (
|
||
<emphasis id="B96AAA7EFFD4FFE0BFC1F8A60EAC037E" bold="true" box="[922,945,1886,1907]" pageId="20" pageNumber="43">52</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD4FFE0BFE6F8A60EC9037E" bold="true" box="[957,980,1886,1907]" pageId="20" pageNumber="43">55</emphasis>
|
||
) of
|
||
<taxonomicName id="4C1E0DEFFFD4FFE0BFA7F8A609BD0379" authorityName="Cameron" authorityYear="1936" box="[1020,1184,1886,1908]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="43" phylum="Arthropoda" rank="species" species="javanus">
|
||
<emphasis id="B96AAA7EFFD4FFE0BFA7F8A609BD0379" box="[1020,1184,1886,1908]" italics="true" pageId="20" pageNumber="43">Scopaeus javanus</emphasis>
|
||
</taxonomicName>
|
||
, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (
|
||
<emphasis id="B96AAA7EFFD4FFE0BDB0F88D0F370387" bold="true" box="[491,554,1909,1930]" pageId="20" pageNumber="43">50 – 52</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFD4FFE0BE64F88D0FCA0387" authorityName="Frisch" authorityYear="2022" box="[575,727,1909,1930]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="43" phylum="Arthropoda" rank="species" species="batukaruensis">
|
||
<emphasis id="B96AAA7EFFD4FFE0BE64F88D0F570387" box="[575,586,1909,1930]" italics="true" pageId="20" pageNumber="43">S</emphasis>
|
||
.
|
||
<emphasis id="B96AAA7EFFD4FFE0BE03F88D0FCA0387" box="[600,727,1909,1930]" italics="true" pageId="20" pageNumber="43">batukaruensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Bali, Tabanan: Bedugul (
|
||
<emphasis id="B96AAA7EFFD4FFE0B871F88D09740387" bold="true" box="[1066,1129,1909,1930]" pageId="20" pageNumber="43">53 – 55</emphasis>
|
||
).
|
||
<emphasis id="B96AAA7EFFD4FFE0B826F88D081B0386" bold="true" box="[1149,1286,1909,1931]" pageId="20" pageNumber="43">Abbreviations</emphasis>
|
||
:
|
||
<emphasis id="B96AAA7EFFD4FFE0B94FF88D083B0387" bold="true" box="[1300,1318,1909,1930]" pageId="20" pageNumber="43">al</emphasis>
|
||
– apical lobes,
|
||
<emphasis id="B96AAA7EFFD4FFE0BC88F8740DFB03AC" bold="true" box="[211,230,1932,1953]" pageId="20" pageNumber="43">dl</emphasis>
|
||
– dorsal lobe,
|
||
<emphasis id="B96AAA7EFFD4FFE0BD35F8740C6603AC" bold="true" box="[366,379,1932,1953]" pageId="20" pageNumber="43">ll</emphasis>
|
||
– lateral lobes,
|
||
<emphasis id="B96AAA7EFFD4FFE0BE57F8730F3B03AC" bold="true" box="[524,550,1931,1953]" pageId="20" pageNumber="43">mf</emphasis>
|
||
– median foramen,
|
||
<emphasis id="B96AAA7EFFD4FFE0BE85F8740FED03AC" bold="true" box="[734,752,1932,1953]" pageId="20" pageNumber="43">tr</emphasis>
|
||
– transverse ridge of median foramen.
|
||
<emphasis id="B96AAA7EFFD4FFE0B80CF87309BD03AD" bold="true" box="[1111,1184,1931,1952]" pageId="20" pageNumber="43">Arrows</emphasis>
|
||
: apex of dorsal lobe (
|
||
<emphasis id="B96AAA7EFFD4FFE0B933F8740DAD03BA" bold="true" pageId="20" pageNumber="43">I, IV</emphasis>
|
||
), apicoventral emargination of apical lobes (
|
||
<emphasis id="B96AAA7EFFD4FFE0BE1CF85A0F7C03BA" bold="true" box="[583,609,1954,1975]" pageId="20" pageNumber="43">III</emphasis>
|
||
), apicolateral margin of apical lobes (
|
||
<emphasis id="B96AAA7EFFD4FFE0BFE0F85A0EF503BA" bold="true" box="[955,1000,1954,1975]" pageId="20" pageNumber="43">II, V</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6126E4FFD5FFE1BCE6F8C90C8F03B5" ID-DOI="http://doi.org/10.5281/zenodo.10724267" ID-Zenodo-Dep="10724267" httpUri="https://zenodo.org/record/10724267/files/figure.png" pageId="21" pageNumber="44" startId="21.[189,262,1841,1863]" targetBox="[261,1359,384,1727]" targetPageId="21" targetType="figure">
|
||
<paragraph id="8BA1766CFFD5FFE1BCE6F8C90C8F03B5" blockId="21.[189,1437,1841,1976]" pageId="21" pageNumber="44">
|
||
<emphasis id="B96AAA7EFFD5FFE1BCE6F8C90C57034B" bold="true" box="[189,330,1841,1863]" pageId="21" pageNumber="44">Figures 56–61.</emphasis>
|
||
Apex of aedeagus in lateral (56, 59), ventral (
|
||
<emphasis id="B96AAA7EFFD5FFE1BEB4F8C90E1B034B" bold="true" box="[751,774,1841,1862]" pageId="21" pageNumber="44">57</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD5FFE1BF4AF8C90E35034B" bold="true" box="[785,808,1841,1862]" pageId="21" pageNumber="44">60</emphasis>
|
||
), dorsal view (
|
||
<emphasis id="B96AAA7EFFD5FFE1BFE8F8C90ED7034B" bold="true" box="[947,970,1841,1862]" pageId="21" pageNumber="44">58</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD5FFE1BF8EF8C90EF1034B" bold="true" box="[981,1004,1841,1862]" pageId="21" pageNumber="44">61</emphasis>
|
||
) of
|
||
<taxonomicName id="4C1E0DEFFFD5FFE1B849F8C909B4034B" authorityName="Frisch" authorityYear="2022" box="[1042,1193,1841,1862]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="44" phylum="Arthropoda" rank="species" species="riedeli">
|
||
<emphasis id="B96AAA7EFFD5FFE1B849F8C909B4034B" box="[1042,1193,1841,1862]" italics="true" pageId="21" pageNumber="44">Scopaeus riedeli</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Sumatra, South Lampung: Rajabasa Mts (
|
||
<emphasis id="B96AAA7EFFD5FFE1BDF1F8B00CF40350" bold="true" box="[426,489,1864,1885]" pageId="21" pageNumber="44">56 – 58</emphasis>
|
||
<emphasis id="B96AAA7EFFD5FFE1BDB3F8B00CED0350" box="[488,496,1864,1885]" italics="true" pageId="21" pageNumber="44">)</emphasis>
|
||
;
|
||
<taxonomicName id="4C1E0DEFFFD5FFE1BDA6F8B00F710350" authorityName="Cameron" authorityYear="1930" box="[509,620,1864,1885]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="44" phylum="Arthropoda" rank="species" species="jacobsoni">
|
||
<emphasis id="B96AAA7EFFD5FFE1BDA6F8B00F710350" box="[509,620,1864,1885]" italics="true" pageId="21" pageNumber="44">S. jacobsoni</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Sumatra: Padang (
|
||
<emphasis id="B96AAA7EFFD5FFE1BF26F8B00EA60350" bold="true" box="[893,955,1864,1885]" pageId="21" pageNumber="44">59– 61</emphasis>
|
||
).
|
||
<emphasis id="B96AAA7EFFD5FFE1BF94F8B009450353" bold="true" box="[975,1112,1864,1886]" pageId="21" pageNumber="44">Abbreviations</emphasis>
|
||
:
|
||
<emphasis id="B96AAA7EFFD5FFE1B83FF8B0096B0350" bold="true" box="[1124,1142,1864,1885]" pageId="21" pageNumber="44">al</emphasis>
|
||
– apical lobes,
|
||
<emphasis id="B96AAA7EFFD5FFE1B95CF8B008070350" bold="true" box="[1287,1306,1864,1885]" pageId="21" pageNumber="44">dl</emphasis>
|
||
– dorsal lobe,
|
||
<emphasis id="B96AAA7EFFD5FFE1BCE6F8A70DD70379" bold="true" box="[189,202,1887,1908]" pageId="21" pageNumber="44">ll</emphasis>
|
||
– lateral lobes,
|
||
<emphasis id="B96AAA7EFFD5FFE1BD04F8A60C640379" bold="true" box="[351,377,1886,1908]" pageId="21" pageNumber="44">mf</emphasis>
|
||
– median foramen,
|
||
<emphasis id="B96AAA7EFFD5FFE1BE6EF8A70F230379" bold="true" box="[565,574,1887,1908]" pageId="21" pageNumber="44">s</emphasis>
|
||
– groups of setae of lateral lobes,
|
||
<emphasis id="B96AAA7EFFD5FFE1BFDBF8A70E8F0379" bold="true" box="[896,914,1887,1908]" pageId="21" pageNumber="44">tr</emphasis>
|
||
– transverse ridge of median foramen,
|
||
<emphasis id="B96AAA7EFFD5FFE1B8A4F8A7080C0379" bold="true" box="[1279,1297,1887,1908]" pageId="21" pageNumber="44">vl</emphasis>
|
||
– ventral lobe.
|
||
<emphasis id="B96AAA7EFFD5FFE1BCE6F88D0C1B0387" bold="true" box="[189,262,1909,1930]" pageId="21" pageNumber="44">Arrows</emphasis>
|
||
: ventral margin of apical lobes (
|
||
<emphasis id="B96AAA7EFFD5FFE1BE76F88D0F2B0387" bold="true" box="[557,566,1909,1930]" pageId="21" pageNumber="44">I</emphasis>
|
||
); apex of dorsal lobe (
|
||
<emphasis id="B96AAA7EFFD5FFE1BF58F88D0E080387" bold="true" box="[771,789,1909,1930]" pageId="21" pageNumber="44">II</emphasis>
|
||
); apicolateral margins of apical lobes (
|
||
<emphasis id="B96AAA7EFFD5FFE1B823F88D098F0387" bold="true" box="[1144,1170,1909,1930]" pageId="21" pageNumber="44">III</emphasis>
|
||
); ventral lobes of dorsal lobe (
|
||
<emphasis id="B96AAA7EFFD5FFE1BC9EF8740DC303AC" bold="true" box="[197,222,1932,1953]" pageId="21" pageNumber="44">IV</emphasis>
|
||
); apical extension of dextral, ventral spine of dorsal lobe (
|
||
<emphasis id="B96AAA7EFFD5FFE1BEA9F8740E1F03AC" bold="true" box="[754,770,1932,1953]" pageId="21" pageNumber="44">V</emphasis>
|
||
); apical extension of sinistral, ventral spine of dorsal lobe (
|
||
<emphasis id="B96AAA7EFFD5FFE1B97BF874082403AC" bold="true" box="[1312,1337,1932,1953]" pageId="21" pageNumber="44">VI</emphasis>
|
||
); unpaired lobe of aedeagus (
|
||
<emphasis id="B96AAA7EFFD5FFE1BD38F85B0C9803B5" bold="true" box="[355,389,1955,1976]" pageId="21" pageNumber="44">VII</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6126E4FFD6FFE2BCCCF8A50E6203BA" ID-DOI="http://doi.org/10.5281/zenodo.10724270" ID-Zenodo-Dep="10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="22" pageNumber="45" startId="22.[151,224,1885,1907]" targetBox="[220,1321,396,1832]" targetPageId="22" targetType="figure">
|
||
<paragraph id="8BA1766CFFD6FFE2BCCCF8A50E6203BA" blockId="22.[151,1400,1885,1976]" pageId="22" pageNumber="45">
|
||
<emphasis id="B96AAA7EFFD6FFE2BCCCF8A50C3E037F" bold="true" box="[151,291,1885,1907]" pageId="22" pageNumber="45">Figures 62–67.</emphasis>
|
||
Apex of aedeagus in lateral (
|
||
<emphasis id="B96AAA7EFFD6FFE2BE77F8A50F5E037F" bold="true" box="[556,579,1885,1906]" pageId="22" pageNumber="45">62</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD6FFE2BE16F8A50F79037F" bold="true" box="[589,612,1885,1906]" pageId="22" pageNumber="45">65</emphasis>
|
||
), ventral (
|
||
<emphasis id="B96AAA7EFFD6FFE2BE99F8A50FC4037F" bold="true" box="[706,729,1885,1906]" pageId="22" pageNumber="45">63</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD6FFE2BEBFF8A50FE6037F" bold="true" box="[740,763,1885,1906]" pageId="22" pageNumber="45">66</emphasis>
|
||
), dorsal view (
|
||
<emphasis id="B96AAA7EFFD6FFE2BFD9F8A50E84037F" bold="true" box="[898,921,1885,1906]" pageId="22" pageNumber="45">64</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD6FFE2BFFFF8A50EA6037F" bold="true" box="[932,955,1885,1906]" pageId="22" pageNumber="45">67</emphasis>
|
||
) of
|
||
<taxonomicName id="4C1E0DEFFFD6FFE2BFBBF8A60968037E" authorityName="Cameron" authorityYear="1941" box="[992,1141,1886,1907]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="45" phylum="Arthropoda" rank="species" species="mixtus">
|
||
<emphasis id="B96AAA7EFFD6FFE2BFBBF8A60968037E" box="[992,1141,1886,1907]" italics="true" pageId="22" pageNumber="45">Scopaeus mixtus</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Malaysia, Sabah: Sandakan (
|
||
<emphasis id="B96AAA7EFFD6FFE2BCA4F88C0C200384" bold="true" box="[255,317,1908,1929]" pageId="22" pageNumber="45">62– 64</emphasis>
|
||
<emphasis id="B96AAA7EFFD6FFE2BD66F88D0C580387" box="[317,325,1909,1930]" italics="true" pageId="22" pageNumber="45">)</emphasis>
|
||
;
|
||
<taxonomicName id="4C1E0DEFFFD6FFE2BD08F88D0F0F0384" authorityName="Frisch" authorityYear="2022" box="[339,530,1908,1930]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="45" phylum="Arthropoda" rank="species" species="halimunsalakensis">
|
||
<emphasis id="B96AAA7EFFD6FFE2BD08F88D0F0F0384" box="[339,530,1908,1930]" italics="true" pageId="22" pageNumber="45">S. halimunsalakensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Jawa Barat, Sukabumi: Sirnarasa (
|
||
<emphasis id="B96AAA7EFFD6FFE2BFE5F88C0EE00384" bold="true" box="[958,1021,1908,1929]" pageId="22" pageNumber="45">65 – 67</emphasis>
|
||
).
|
||
<emphasis id="B96AAA7EFFD6FFE2B849F88C09860387" bold="true" box="[1042,1179,1908,1930]" pageId="22" pageNumber="45">Abbreviations</emphasis>
|
||
:
|
||
<emphasis id="B96AAA7EFFD6FFE2B8F2F88C09A60384" bold="true" box="[1193,1211,1908,1929]" pageId="22" pageNumber="45">al</emphasis>
|
||
– apical lobes,
|
||
<emphasis id="B96AAA7EFFD6FFE2B90BF88C087E0384" bold="true" box="[1360,1379,1908,1929]" pageId="22" pageNumber="45">dl</emphasis>
|
||
– dorsal lobe,
|
||
<emphasis id="B96AAA7EFFD6FFE2BD53F8730C0D03AC" bold="true" box="[264,272,1931,1953]" pageId="22" pageNumber="45">f</emphasis>
|
||
– flagellum,
|
||
<emphasis id="B96AAA7EFFD6FFE2BDD2F8730C8B03AD" bold="true" box="[393,406,1931,1952]" pageId="22" pageNumber="45">ll</emphasis>
|
||
– lateral lobes,
|
||
<emphasis id="B96AAA7EFFD6FFE2BE73F8730F5F03AC" bold="true" box="[552,578,1931,1953]" pageId="22" pageNumber="45">mf</emphasis>
|
||
– median foramen,
|
||
<emphasis id="B96AAA7EFFD6FFE2BEA0F8730E1903AD" bold="true" box="[763,772,1931,1952]" pageId="22" pageNumber="45">s</emphasis>
|
||
– groups of setae,
|
||
<emphasis id="B96AAA7EFFD6FFE2BFE9F8730ED903AD" bold="true" box="[946,964,1931,1952]" pageId="22" pageNumber="45">vl</emphasis>
|
||
– ventral lobe.
|
||
<emphasis id="B96AAA7EFFD6FFE2B808F873098103AD" bold="true" box="[1107,1180,1931,1952]" pageId="22" pageNumber="45">Arrows</emphasis>
|
||
: dorsal emargination of apical lobes, dorsal lobe and flagellum (
|
||
<emphasis id="B96AAA7EFFD6FFE2BE5AF85A0F1703BA" bold="true" box="[513,522,1954,1975]" pageId="22" pageNumber="45">I</emphasis>
|
||
); apical emargination of dorsal lobe (
|
||
<emphasis id="B96AAA7EFFD6FFE2BF3BF85A0E6F03BA" bold="true" box="[864,882,1954,1975]" pageId="22" pageNumber="45">II</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6126E4FFD7FFE3BCE6F8B10E1403B4" ID-DOI="http://doi.org/10.5281/zenodo.10724272" ID-Zenodo-Dep="10724272" httpUri="https://zenodo.org/record/10724272/files/figure.png" pageId="23" pageNumber="46" startId="23.[189,262,1865,1887]" targetBox="[261,1375,305,1707]" targetPageId="23" targetType="figure">
|
||
<paragraph id="8BA1766CFFD7FFE3BCE6F8B10E1403B4" blockId="23.[189,1437,1865,1978]" pageId="23" pageNumber="46">
|
||
<emphasis id="B96AAA7EFFD7FFE3BCE6F8B10C540353" bold="true" box="[189,329,1865,1887]" pageId="23" pageNumber="46">Figures 68–73.</emphasis>
|
||
Apex of aedeagus in lateral (
|
||
<emphasis id="B96AAA7EFFD7FFE3BE08F8B10F770353" bold="true" box="[595,618,1865,1886]" pageId="23" pageNumber="46">68</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD7FFE3BE2EF8B10F910353" bold="true" box="[629,652,1865,1886]" pageId="23" pageNumber="46">71</emphasis>
|
||
), ventral (
|
||
<emphasis id="B96AAA7EFFD7FFE3BEB1F8B10E1C0353" bold="true" box="[746,769,1865,1886]" pageId="23" pageNumber="46">69</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD7FFE3BF57F8B10E3E0353" bold="true" box="[780,803,1865,1886]" pageId="23" pageNumber="46">72</emphasis>
|
||
), dorsal view (
|
||
<emphasis id="B96AAA7EFFD7FFE3BFF0F8B10EDF0353" bold="true" box="[939,962,1865,1886]" pageId="23" pageNumber="46">70</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD7FFE3BF96F8B10EF90353" bold="true" box="[973,996,1865,1886]" pageId="23" pageNumber="46">72</emphasis>
|
||
) of
|
||
<taxonomicName id="4C1E0DEFFFD7FFE3B852F8B108140353" authorityName="Frisch" authorityYear="2022" box="[1033,1289,1865,1886]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="46" phylum="Arthropoda" rank="species" species="tortuosiflagellatus">
|
||
<emphasis id="B96AAA7EFFD7FFE3B852F8B108140353" box="[1033,1289,1865,1886]" italics="true" pageId="23" pageNumber="46">Scopaeus tortuosiflagellatus</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Jawa Barat, Sukabumi: Cisalimas (
|
||
<emphasis id="B96AAA7EFFD7FFE3BD9CF8A70F1B0379" bold="true" box="[455,518,1887,1909]" pageId="23" pageNumber="46">68 – 70</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFD7FFE3BE42F8980F980378" authorityName="Frisch" authorityYear="2022" box="[537,645,1888,1909]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="46" phylum="Arthropoda" rank="species" species="posoanus">
|
||
<emphasis id="B96AAA7EFFD7FFE3BE42F8980F390378" box="[537,548,1888,1909]" italics="true" pageId="23" pageNumber="46">S</emphasis>
|
||
.
|
||
<emphasis id="B96AAA7EFFD7FFE3BE74F8980F980378" box="[559,645,1888,1909]" italics="true" pageId="23" pageNumber="46">posoanus</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Sulawesi Tengah: Lake Poso (
|
||
<emphasis id="B96AAA7EFFD7FFE3BFA0F8A709290379" bold="true" box="[1019,1076,1887,1909]" pageId="23" pageNumber="46">71–73</emphasis>
|
||
).
|
||
<emphasis id="B96AAA7EFFD7FFE3B81DF8A709D20378" bold="true" box="[1094,1231,1887,1909]" pageId="23" pageNumber="46">Abbreviations</emphasis>
|
||
:
|
||
<emphasis id="B96AAA7EFFD7FFE3B881F89809F10378" bold="true" box="[1242,1260,1888,1909]" pageId="23" pageNumber="46">al</emphasis>
|
||
– apical lobes,
|
||
<emphasis id="B96AAA7EFFD7FFE3B922F89808910378" bold="true" box="[1401,1420,1888,1909]" pageId="23" pageNumber="46">dl</emphasis>
|
||
– dorsal lobe,
|
||
<emphasis id="B96AAA7EFFD7FFE3BD77F88E0C290381" bold="true" box="[300,308,1910,1932]" pageId="23" pageNumber="46">f</emphasis>
|
||
– flagellum,
|
||
<emphasis id="B96AAA7EFFD7FFE3BDF0F88E0CA50386" bold="true" box="[427,440,1910,1931]" pageId="23" pageNumber="46">ll</emphasis>
|
||
– lateral lobes,
|
||
<emphasis id="B96AAA7EFFD7FFE3BE13F88E0F7F0381" bold="true" box="[584,610,1910,1932]" pageId="23" pageNumber="46">mf</emphasis>
|
||
– median foramen,
|
||
<emphasis id="B96AAA7EFFD7FFE3BF43F88E0E3C0386" bold="true" box="[792,801,1910,1931]" pageId="23" pageNumber="46">s</emphasis>
|
||
– groups of setae,
|
||
<emphasis id="B96AAA7EFFD7FFE3BF97F88E0EC30386" bold="true" box="[972,990,1910,1931]" pageId="23" pageNumber="46">tr</emphasis>
|
||
– transverse ridge,
|
||
<emphasis id="B96AAA7EFFD7FFE3B8D2F88E09860386" bold="true" box="[1161,1179,1910,1931]" pageId="23" pageNumber="46">vl</emphasis>
|
||
– ventral lobe,
|
||
<emphasis id="B96AAA7EFFD7FFE3B973F88E085B0386" bold="true" box="[1320,1350,1910,1931]" pageId="23" pageNumber="46">vld</emphasis>
|
||
– dextral portion of ventral lobe,
|
||
<emphasis id="B96AAA7EFFD7FFE3BDD4F8750CB403AF" bold="true" box="[399,425,1933,1954]" pageId="23" pageNumber="46">vls</emphasis>
|
||
– sinistral portion of ventral lobe.
|
||
<emphasis id="B96AAA7EFFD7FFE3BE86F8750E3B03AF" bold="true" box="[733,806,1933,1954]" pageId="23" pageNumber="46">Arrows</emphasis>
|
||
: ventral spine of apical lobe (
|
||
<emphasis id="B96AAA7EFFD7FFE3B875F875092A03AF" bold="true" box="[1070,1079,1933,1954]" pageId="23" pageNumber="46">I</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD7FFE3B81AF875094E03AF" bold="true" box="[1089,1107,1933,1954]" pageId="23" pageNumber="46">II</emphasis>
|
||
), proximal enlargement of flagellum (
|
||
<emphasis id="B96AAA7EFFD7FFE3BC9EF85C0DC203B4" bold="true" box="[197,223,1956,1977]" pageId="23" pageNumber="46">III</emphasis>
|
||
), ventral tooth of apical lobe (
|
||
<emphasis id="B96AAA7EFFD7FFE3BDA8F85C0F1103B4" bold="true" box="[499,524,1956,1977]" pageId="23" pageNumber="46">IV</emphasis>
|
||
), tip of dorsal lobe (
|
||
<emphasis id="B96AAA7EFFD7FFE3BE9CF85C0FCA03B4" bold="true" box="[711,727,1956,1977]" pageId="23" pageNumber="46">V</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD7FFE3BEB9F85C0FE603B4" bold="true" box="[738,763,1956,1977]" pageId="23" pageNumber="46">VI</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6126E4FFD8FFECBCCCF8B20F0703B6" pageId="24" pageNumber="47" startId="24.[151,224,1866,1888]" targetType="figure">
|
||
<paragraph id="8BA1766CFFD8FFECBCCCF8B20F0703B6" blockId="24.[151,1399,1866,1979]" pageId="24" pageNumber="47">
|
||
<emphasis id="B96AAA7EFFD8FFECBCCCF8B20C3A0352" bold="true" box="[151,295,1866,1888]" pageId="24" pageNumber="47">Figures 74–79.</emphasis>
|
||
Apex of aedeagus in lateral (
|
||
<emphasis id="B96AAA7EFFD8FFECBE1FF8B20F460352" bold="true" box="[580,603,1866,1887]" pageId="24" pageNumber="47">74</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD8FFECBE32F8B20F9D0352" bold="true" box="[617,640,1866,1887]" pageId="24" pageNumber="47">77</emphasis>
|
||
), ventral (
|
||
<emphasis id="B96AAA7EFFD8FFECBEBEF8B20FE10352" bold="true" box="[741,764,1866,1887]" pageId="24" pageNumber="47">75</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD8FFECBF51F8B20E3C0352" bold="true" box="[778,801,1866,1887]" pageId="24" pageNumber="47">78</emphasis>
|
||
), dorsal view (
|
||
<emphasis id="B96AAA7EFFD8FFECBFE8F8B20ED70352" bold="true" box="[947,970,1866,1887]" pageId="24" pageNumber="47">76</emphasis>
|
||
,
|
||
<emphasis id="B96AAA7EFFD8FFECBF83F8B20EF20352" bold="true" box="[984,1007,1866,1887]" pageId="24" pageNumber="47">79</emphasis>
|
||
) of
|
||
<taxonomicName id="4C1E0DEFFFD8FFECB840F8B2080E0352" authorityName="Frisch" authorityYear="2022" box="[1051,1299,1866,1887]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="47" phylum="Arthropoda" rank="species" species="spiraliflagellatus">
|
||
<emphasis id="B96AAA7EFFD8FFECB840F8B2080E0352" box="[1051,1299,1866,1887]" italics="true" pageId="24" pageNumber="47">Scopaeus spiraliflagellatus</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Sulawesi Tengah: Lake Poso (
|
||
<emphasis id="B96AAA7EFFD8FFECBDF0F8990CF4037B" bold="true" box="[427,489,1889,1910]" pageId="24" pageNumber="47">74– 76</emphasis>
|
||
<emphasis id="B96AAA7EFFD8FFECBDB2F8990CEC037B" box="[489,497,1889,1910]" italics="true" pageId="24" pageNumber="47">)</emphasis>
|
||
;
|
||
<taxonomicName id="4C1E0DEFFFD8FFECBDA6F8990FBC037B" authorityName="Frisch" authorityYear="2022" box="[509,673,1889,1910]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="47" phylum="Arthropoda" rank="species" species="anuliflagellatus">
|
||
<emphasis id="B96AAA7EFFD8FFECBDA6F8990FBC037B" box="[509,673,1889,1910]" italics="true" pageId="24" pageNumber="47">S. anuliflagellatus</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Jawa Barat, Sukabumi: Cidahu (
|
||
<emphasis id="B96AAA7EFFD8FFECB86BF8990972037B" bold="true" box="[1072,1135,1889,1910]" pageId="24" pageNumber="47">77 – 79</emphasis>
|
||
).
|
||
<emphasis id="B96AAA7EFFD8FFECB8D9F8990816037A" bold="true" box="[1154,1291,1889,1911]" pageId="24" pageNumber="47">Abbreviations</emphasis>
|
||
:
|
||
<emphasis id="B96AAA7EFFD8FFECB94CF8990834037B" bold="true" box="[1303,1321,1889,1910]" pageId="24" pageNumber="47">al</emphasis>
|
||
– apical lobes,
|
||
<emphasis id="B96AAA7EFFD8FFECBC88F88F0C1C0380" bold="true" box="[211,257,1911,1933]" pageId="24" pageNumber="47">al(d)</emphasis>
|
||
– dextral apical lobe,
|
||
<emphasis id="B96AAA7EFFD8FFECBD94F88F0CE50380" bold="true" box="[463,504,1911,1933]" pageId="24" pageNumber="47">al(s)</emphasis>
|
||
– sinistral apical lobe,
|
||
<emphasis id="B96AAA7EFFD8FFECBE94F8800FFF0380" bold="true" box="[719,738,1912,1933]" pageId="24" pageNumber="47">dl</emphasis>
|
||
– dorsal lobe,
|
||
<emphasis id="B96AAA7EFFD8FFECBF37F88F0E860380" bold="true" box="[876,923,1911,1933]" pageId="24" pageNumber="47">dl(d)</emphasis>
|
||
– dextral end of dorsal lobe,
|
||
<emphasis id="B96AAA7EFFD8FFECB8F0F88F09CB0380" bold="true" box="[1195,1238,1911,1933]" pageId="24" pageNumber="47">dl(s)</emphasis>
|
||
– sinistral end of dorsal lobe,
|
||
<emphasis id="B96AAA7EFFD8FFECBD5DF8760C1303A9" bold="true" box="[262,270,1934,1956]" pageId="24" pageNumber="47">f</emphasis>
|
||
– flagellum,
|
||
<emphasis id="B96AAA7EFFD8FFECBDDFF8760C8C03AE" bold="true" box="[388,401,1934,1955]" pageId="24" pageNumber="47">ll</emphasis>
|
||
– lateral lobes,
|
||
<emphasis id="B96AAA7EFFD8FFECBE44F8760F2403A9" bold="true" box="[543,569,1934,1956]" pageId="24" pageNumber="47">mf</emphasis>
|
||
– median foramen,
|
||
<emphasis id="B96AAA7EFFD8FFECBEB6F8760FE203AE" bold="true" box="[749,767,1934,1955]" pageId="24" pageNumber="47">tr</emphasis>
|
||
– transverse ridge of median foramen,
|
||
<emphasis id="B96AAA7EFFD8FFECB83BF876096303AE" bold="true" box="[1120,1150,1934,1955]" pageId="24" pageNumber="47">vld</emphasis>
|
||
– dextral portion of ventral lobe,
|
||
<emphasis id="B96AAA7EFFD8FFECBC91F85D0DF903B7" bold="true" box="[202,228,1957,1978]" pageId="24" pageNumber="47">vls</emphasis>
|
||
– sinistral portion of ventral lobe.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA1766CFFD9FFEDBC8FFE000F57067E" blockId="25.[189,801,241,989]" pageId="25" pageNumber="48">
|
||
Female: Sperm pump with process segment and apophysis of chamber long, slender and curved at ends (
|
||
<figureCitation id="13256AE9FFD9FFEDBC9EFDC20C3B065F" box="[197,294,570,594]" captionStart="Figures 163–172" captionStartId="42.[151,224,1839,1861]" captionTargetBox="[225,1322,230,1817]" captionTargetId="figure-168@42.[157,1398,210,1820]" captionTargetPageId="42" captionText="Figures 163–172. Sperm pump (163–172) and bursa (165, 167, 170, 172) of ♀ of Scopaeus grandis, paratype, Jawa Barat, Sukabumi: Cidahu (163); S.heronifer, paratype, Jawa Barat, Sukabumi:Sirnarasa (164); S. diversilobatus, paratype, Jawa Barat, Sukabumi, Djampang Kulon (165, with lateral gonocoxal plates IX folded laterally; 166); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (167, with lateral gonocoxal plates IX, 168); S. velifer, paratype, Banten, Lebak: Citorek Kidul (169); S. elegantulus, Jawa Barat, Sukabumi: Djampang Kulon (170, with lateral gonocoxal plates IX, 171), S. uncinatus, paratype, Banten, Lebak: Citorek Kidul (172). Long scale bar: 163, 164, 166, 168, 169, 171, 172; short scale bar: 165, 167, 170." figureDoi="http://doi.org/10.5281/zenodo.10724315" httpUri="https://zenodo.org/record/10724315/files/figure.png" pageId="25" pageNumber="48">Fig. 166</figureCitation>
|
||
); bursal duct about twice as long as sperm pump; bursa membranous (
|
||
<figureCitation id="13256AE9FFD9FFEDBDB8FDA30F23067E" box="[483,574,603,627]" captionStart="Figures 163–172" captionStartId="42.[151,224,1839,1861]" captionTargetBox="[225,1322,230,1817]" captionTargetId="figure-168@42.[157,1398,210,1820]" captionTargetPageId="42" captionText="Figures 163–172. Sperm pump (163–172) and bursa (165, 167, 170, 172) of ♀ of Scopaeus grandis, paratype, Jawa Barat, Sukabumi: Cidahu (163); S.heronifer, paratype, Jawa Barat, Sukabumi:Sirnarasa (164); S. diversilobatus, paratype, Jawa Barat, Sukabumi, Djampang Kulon (165, with lateral gonocoxal plates IX folded laterally; 166); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (167, with lateral gonocoxal plates IX, 168); S. velifer, paratype, Banten, Lebak: Citorek Kidul (169); S. elegantulus, Jawa Barat, Sukabumi: Djampang Kulon (170, with lateral gonocoxal plates IX, 171), S. uncinatus, paratype, Banten, Lebak: Citorek Kidul (172). Long scale bar: 163, 164, 166, 168, 169, 171, 172; short scale bar: 165, 167, 170." figureDoi="http://doi.org/10.5281/zenodo.10724315" httpUri="https://zenodo.org/record/10724315/files/figure.png" pageId="25" pageNumber="48">Fig. 165</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFD9FFEDBC8FFD840E1406B8" blockId="25.[189,801,241,989]" pageId="25" pageNumber="48">
|
||
<emphasis id="B96AAA7EFFD9FFEDBC8FFD840C4F0699" bold="true" box="[212,338,636,660]" pageId="25" pageNumber="48">Phylogeny:</emphasis>
|
||
<taxonomicName id="4C1E0DEFFFD9FFEDBD02FD840F7D0699" authorityName="Frisch" authorityYear="2022" box="[345,608,636,660]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="diversilobatus">
|
||
<emphasis id="B96AAA7EFFD9FFEDBD02FD840F7D0699" box="[345,608,636,660]" italics="true" pageId="25" pageNumber="48">Scopaeus diversilobatus</emphasis>
|
||
</taxonomicName>
|
||
is to be classified in the
|
||
<taxonomicName id="4C1E0DEFFFD9FFEDBD5AFD650C7206B8" authorityName="Oke" authorityYear="1933" box="[257,367,669,693]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B96AAA7EFFD9FFEDBD5AFD650C7206B8" box="[257,367,669,693]" italics="true" pageId="25" pageNumber="48">S. gracilis</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<bibRefCitation id="EF8F0B9DFFD9FFEDBE4CFD650FE006B8" author="Frisch, J. & D. Burckhardt & V. Wolters" box="[535,765,669,693]" pageId="25" pageNumber="48" pagination="27 - 53" refId="ref31920" refString="Frisch, J., D. Burckhardt & V. Wolters (2002): Rove beetles of the subtribe Scopaeina Mulsant & Rey (Coleoptera: Staphylinidae) in the West Palaearctic: Phylogeny, biogeography and species catalogue. - Organisms, Diversity & Evolution 2: 27 - 53." type="journal article" year="2002">Frisch et al. 2002: 39</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFD9FFEDBC8FFD460F3C0715" pageId="25" pageNumber="48" type="distribution">
|
||
<paragraph id="8BA1766CFFD9FFEDBC8FFD460F3C0715" blockId="25.[189,801,241,989]" pageId="25" pageNumber="48">
|
||
<emphasis id="B96AAA7EFFD9FFEDBC8FFD460C7706DB" bold="true" box="[212,362,702,726]" pageId="25" pageNumber="48">Distribution:</emphasis>
|
||
Hitherto recorded from western
|
||
<collectingRegion id="49DAB88EFFD9FFEDBEB2FD460E0606D8" box="[745,795,702,725]" country="Indonesia" name="Jawa Timur" pageId="25" pageNumber="48">Java</collectingRegion>
|
||
,
|
||
<collectingRegion id="49DAB88EFFD9FFEDBCE6FD270DF706FA" box="[189,234,735,759]" country="Indonesia" name="Bali" pageId="25" pageNumber="48">Bali</collectingRegion>
|
||
and northern Borneo,
|
||
<taxonomicName id="4C1E0DEFFFD9FFEDBDAEFD270E1C06FA" authorityName="Frisch" authorityYear="2022" box="[501,769,735,759]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="diversilobatus">
|
||
<emphasis id="B96AAA7EFFD9FFEDBDAEFD270E1C06FA" box="[501,769,735,759]" italics="true" pageId="25" pageNumber="48">Scopaeus diversilobatus</emphasis>
|
||
</taxonomicName>
|
||
is widespread in the Sunda Islands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFD9FFEDBC8FFCD90FAB07D0" pageId="25" pageNumber="48" type="etymology">
|
||
<paragraph id="8BA1766CFFD9FFEDBC8FFCD90FAB07D0" blockId="25.[189,801,241,989]" pageId="25" pageNumber="48">
|
||
<emphasis id="B96AAA7EFFD9FFEDBC8FFCD90C470734" bold="true" box="[212,346,801,825]" pageId="25" pageNumber="48">Etymology:</emphasis>
|
||
The epithet
|
||
<taxonomicName id="4C1E0DEFFFD9FFEDBE5EFCD80F830735" authorityName="Frisch" authorityYear="2022" box="[517,670,800,824]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="diversilobatus">
|
||
<emphasis id="B96AAA7EFFD9FFEDBE5EFCD80F830735" box="[517,670,800,824]" italics="true" pageId="25" pageNumber="48">diversilobatus</emphasis>
|
||
</taxonomicName>
|
||
[adjective, Latin, composed of the adjectives
|
||
<emphasis id="B96AAA7EFFD9FFEDBE15FCB90FB50754" box="[590,680,833,857]" italics="true" pageId="25" pageNumber="48">diversus</emphasis>
|
||
(different) and
|
||
<emphasis id="B96AAA7EFFD9FFEDBCAAFC9A0C5C0777" box="[241,321,866,890]" italics="true" pageId="25" pageNumber="48">lobatus</emphasis>
|
||
(indicating a characteristic/feature of the lobe)] describes the different size and shape of the apical lobes of the aedeagus of this new species, one of which is strongly reduced unlike the other (
|
||
<figureCitation id="13256AE9FFD9FFEDBE6AFC3D0FB707D0" box="[561,682,965,989]" captionStart="Figures 86–91" captionStartId="27.[189,262,1863,1885]" captionTargetBox="[261,1365,582,1806]" captionTargetId="figure-268@27.[189,1436,563,1844]" captionTargetPageId="27" captionText="Figures 86–91. Apex of aedeagus in lateral (86, 89), ventral (87, 90), dorsal view (88, 91) of Scopaeus heronifer, holotype, Banten, Lebak: Majasari (86– 88); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (89–91). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: distoventral incision of dextral apical lobe (I); sinistral, apical tooth of flagellum (II); distal emargination of dorsal lobe (III)." figureDoi="http://doi.org/10.5281/zenodo.10724278" httpUri="https://zenodo.org/record/10724278/files/figure.png" pageId="25" pageNumber="48">Figs 87, 88</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |