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<document id="75530374074EBC88A407A948CEA0B6FF" ID-DOI="10.11646/zootaxa.3999.1.10" ID-GBIF-Dataset="6a8fd7a3-057e-4ea0-940c-c0dcce976917" ID-ISSN="1175-5326" ID-Zenodo-Dep="231712" ID-ZooBank="2D81FF91-D26B-434C-87BC-B993D50702CC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461280431214" checkinUser="plazi" docAuthor="Heppner, John B. &amp; Bae, Yang-Seop" docDate="2015" docId="FF6B87ADFF8EFFCFFF385F57B5147F3A" docLanguage="en" docName="zt03999p150.pdf" docOrigin="Zootaxa 3999 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Compsocommosis vietnamensis Heppner &amp; Bae, new species" docType="treatment" docVersion="8" lastPageNumber="149" masterDocId="0352FFD5FF8DFFCAFFAF5C72B7227E5F" masterDocTitle="Compsocommosis new genus, with a new species in Vietnam, and Transfer of Mictocommosis to Archipini (Lepidoptera: Tortricidae: Tortricinae: Archipini)" masterLastPageNumber="150" masterPageNumber="144" pageNumber="147" updateTime="1698613881048" updateUser="plazi">
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<mods:title id="2F8D579FAEDCC52448E48F1ECC6B584C">Compsocommosis new genus, with a new species in Vietnam, and Transfer of Mictocommosis to Archipini (Lepidoptera: Tortricidae: Tortricinae: Archipini)</mods:title>
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<mods:name id="410F1D50C3D5CC2EA5D8326A50DD8D5E" type="personal">
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<mods:namePart id="C192D125AC77A6511596C81F782FE542">Heppner, John B.</mods:namePart>
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<mods:namePart id="A81F34170E124356750A02796D04B0A2">Bae, Yang-Seop</mods:namePart>
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<mods:title id="BE0A584D850D78038E5CA80A45B7EF80">Zootaxa</mods:title>
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<mods:date id="E9DD014BE456D021E62B6995D6C6BF51">2015</mods:date>
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<treatment id="FF6B87ADFF8EFFCFFF385F57B5147F3A" ID-DOI="http://doi.org/10.5281/zenodo.6117289" ID-GBIF-Taxon="119640381" ID-Zenodo-Dep="6117289" LSID="urn:lsid:plazi:treatment:FF6B87ADFF8EFFCFFF385F57B5147F3A" httpUri="http://treatment.plazi.org/id/FF6B87ADFF8EFFCFFF385F57B5147F3A" lastPageId="5" lastPageNumber="149" pageId="3" pageNumber="147">
<subSubSection id="3FD86530FF8EFFC9FF385F57B6307D3E" pageId="3" pageNumber="147" type="nomenclature">
<paragraph id="777D36BBFF8EFFC9FF385F57B45C7D60" blockId="3.[151,894,805,865]" box="[151,894,805,831]" pageId="3" pageNumber="147">
<heading id="2C3581D7FF8EFFC9FF385F57B45C7D60" bold="true" box="[151,894,805,831]" fontSize="11" level="1" pageId="3" pageNumber="147" reason="1">
<emphasis id="45B6EAA9FF8EFFC9FF385F57B45C7D60" bold="true" box="[151,894,805,831]" pageId="3" pageNumber="147">
<taxonomicName id="B0C24D38FF8EFFC9FF385F57B5C37D60" ID-CoL="B4QV5" authority="Heppner &amp; Bae" authorityName="Heppner &amp; Bae" box="[151,737,805,831]" class="Insecta" family="Tortricidae" genus="Compsocommosis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="147" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="45B6EAA9FF8EFFC9FF385F57B5307D60" bold="true" box="[151,530,805,831]" italics="true" pageId="3" pageNumber="147">Compsocommosis vietnamensis</emphasis>
Heppner &amp; Bae
</taxonomicName>
,
<taxonomicNameLabel id="5E8557D2FF8EFFC9FD415F57B45C7D60" box="[750,894,805,831]" pageId="3" pageNumber="147" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="777D36BBFF8EFFC9FF385F3AB6307D3E" blockId="3.[151,894,805,865]" box="[151,274,840,865]" pageId="3" pageNumber="147">
(
<figureCitation id="EFF92A3EFF8EFFC9FF305F3AB62A7D3E" box="[159,264,840,865]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 3" captionStart-2="FIGURES 4 5" captionStartId-0="1.[151,250,1674,1696]" captionStartId-1="2.[151,264,1054,1077]" captionStartId-2="2.[151,264,1857,1880]" captionTargetBox-0="[170,1412,906,1635]" captionTargetBox-1="[150,1436,445,1033]" captionTargetBox-2="[176,1406,1168,1813]" captionTargetId-0="figure@1.[152,1435,873,1650]" captionTargetId-1="figure@2.[137,1450,445,1034]" captionTargetId-2="figure@2.[171,1431,1127,1835]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionText-0="FIGURE 1. COmpSOcOmmOSiS vietnamenSiS n. sp., female holotype (14.5 mm wingspan)." captionText-1="FIGURES 2 3. COmpSOcOmmOSiS vietnamenSiS n. sp., head close-ups (lateral and dorsal views), female." captionText-2="FIGURES 4 5. Wing venations: 4) COmpSOcOmmOSiS n. gen. (C. vietnamenSiS n. sp., female); 5) MictOcOmmOSiS Diakonoff (M. nigrOmaculata (Issiki), male, redrawn from Diakonoff, 1977)." httpUri-0="https://zenodo.org/record/231713/files/figure.png" httpUri-1="https://zenodo.org/record/231714/files/figure.png" httpUri-2="https://zenodo.org/record/231715/files/figure.png" pageId="3" pageNumber="147">Figs. 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3FD86530FF8EFFC9FF385FE2B38B7DF6" box="[151,1193,912,937]" pageId="3" pageNumber="147" type="materials_examined">
<paragraph id="777D36BBFF8EFFC9FF385FE2B38B7DF6" blockId="3.[151,1437,912,1981]" box="[151,1193,912,937]" pageId="3" pageNumber="147">
<emphasis id="45B6EAA9FF8EFFC9FF385FE2B6177DF6" bold="true" box="[151,309,912,937]" pageId="3" pageNumber="147">
<typeStatus id="A8798819FF8EFFC9FF385FE2B7F07DF6" box="[151,210,912,937]" pageId="3" pageNumber="147">Type</typeStatus>
locality.
</emphasis>
Cuc Phuong National Park, Mac Lake (
<quantity id="B03A9B5EFF8EFFC9FD595FE2B4627DF7" box="[758,832,912,937]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="147" unit="m" value="155.0">155 m</quantity>
), Ninh Binh Province,
<collectingCountry id="0FD5762BFF8EFFC9FBEC5FE2B3847DF6" box="[1091,1190,912,937]" name="Vietnam" pageId="3" pageNumber="147">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3FD86530FF8EFFC9FF685FC6B41E7A4A" pageId="3" pageNumber="147" type="diagnosis">
<paragraph id="777D36BBFF8EFFC9FF685FC6B41E7A4A" blockId="3.[151,1437,912,1981]" pageId="3" pageNumber="147">
<emphasis id="45B6EAA9FF8EFFC9FF685FC6B6187D92" bold="true" box="[199,314,948,973]" pageId="3" pageNumber="147">Diagnosis</emphasis>
. Diagnostic characters for this species have already been noted under the generic discussion, but in the female the size of the lamellae postvaginalis, the ductus bursae-bursal border plates, the form of the bursa, and the size of the signum will likely be specific for this species.
</paragraph>
</subSubSection>
<subSubSection id="3FD86530FF8EFFC9FF685852B33A79C7" pageId="3" pageNumber="147" type="description">
<paragraph id="777D36BBFF8EFFC9FF685852B3C278BB" blockId="3.[151,1437,912,1981]" pageId="3" pageNumber="147">
<emphasis id="45B6EAA9FF8EFFC9FF685852B6737A66" bold="true" box="[199,337,1056,1081]" pageId="3" pageNumber="147">Description</emphasis>
. Wing expanse: 14.514.6 mm (n = 3). Male unknown. Female (
<figureCitation id="EFF92A3EFF8EFFC9FB845852B34F7A67" box="[1067,1133,1056,1081]" captionStart="FIGURE 1" captionStartId="1.[151,250,1674,1696]" captionTargetBox="[170,1412,906,1635]" captionTargetId="figure@1.[152,1435,873,1650]" captionTargetPageId="1" captionText="FIGURE 1. COmpSOcOmmOSiS vietnamenSiS n. sp., female holotype (14.5 mm wingspan)." httpUri="https://zenodo.org/record/231713/files/figure.png" pageId="3" pageNumber="147">Fig. 1</figureCitation>
).
<emphasis id="45B6EAA9FF8EFFC9FB2A5852B3E07A66" box="[1157,1218,1056,1081]" italics="true" pageId="3" pageNumber="147">Head</emphasis>
(
<figureCitation id="EFF92A3EFF8EFFC9FB7E5852B21C7A67" box="[1233,1342,1056,1081]" captionStart="FIGURES 2 3" captionStartId="2.[151,264,1054,1077]" captionTargetBox="[150,1436,445,1033]" captionTargetId="figure@2.[137,1450,445,1034]" captionTargetPageId="2" captionText="FIGURES 2 3. COmpSOcOmmOSiS vietnamenSiS n. sp., head close-ups (lateral and dorsal views), female." httpUri="https://zenodo.org/record/231714/files/figure.png" pageId="3" pageNumber="147">Figs. 23</figureCitation>
): Vertex tan-fuscous; frons tan, white at base; antenna brown-fuscous alternating with tan, ventrally setous but unscaled, scape tan-fuscous; labial palpus cream white on basal segment, dark brown ventrolaterally on middle segment, with apical segment dark tan fuscous at middle, and mesally the same but less fuscous on second segment.
<emphasis id="45B6EAA9FF8EFFC9FAEC58FEB2B67AFA" box="[1347,1428,1164,1189]" italics="true" pageId="3" pageNumber="147">Thorax</emphasis>
: Brown-fuscous; tegulae same; venter white, with light tan at neck; legs white, with brown-fuscous dorsally on femur and tibia, with brown-fuscous on tarsal segments, and tarsi tan ventrally. Forewing dorsally brown, with irrorated white area on dorsal middle, narrowing to tornus and half of basal area; costal margin with brown alternating with 67 pale pairs of strigulae, more whitish nearer apical quarter, with yellow line at apical 1/4 curved to tornus, margined by iridescent silver-fuscous; silver subterminal line; black spot near tornus with small light yellow spots; fringe dark fuscous, with white near apex and near tornus; venter of wing like dorsum but duller and more yellow marks on costa and apex. Hindwing white basal half brown-fuscous at base and some brown-fuscous irrorations near anal margin and angled beyond base; apical 1/3 brown; a large black patch along termen and tornus, with small yellow and silver spots towards tornus and subterminal silver line; fringe white except fuscous near apex; venter of wing like dorsum but duller.
<emphasis id="45B6EAA9FF8EFFC9FD6C5986B40F7852" box="[707,813,1524,1549]" italics="true" pageId="3" pageNumber="147">Abdomen</emphasis>
: Brown-fuscous with pale white scale row on posterior margin of each tergite; venter cream-white; anal tuft fuscous. Female genitalia (
<figureCitation id="EFF92A3EFF8EFFC9FBE55A6AB3B0786F" box="[1098,1170,1560,1585]" captionStart="FIGURES 4 5" captionStartId="2.[151,264,1857,1880]" captionTargetBox="[176,1406,1168,1813]" captionTargetId="figure@2.[171,1431,1127,1835]" captionTargetPageId="2" captionText="FIGURES 4 5. Wing venations: 4) COmpSOcOmmOSiS n. gen. (C. vietnamenSiS n. sp., female); 5) MictOcOmmOSiS Diakonoff (M. nigrOmaculata (Issiki), male, redrawn from Diakonoff, 1977)." httpUri="https://zenodo.org/record/231715/files/figure.png" pageId="3" pageNumber="147">Fig. 4</figureCitation>
) elongated, with short ovipositor, papilla anales setous and flattened; sterigma split with two rectangular spinose plates, sharply truncated posteriorly; ostium bursae ovate, simple, wider than deep, with bulbous anterior end as deep as wide; ductus bursae simple, narrowed at junction with anterior of ostium, then widening anteriorly to width subequal to bursa opening; ductus seminalis large, emergent dorsally from ductus bursae near subostium; corpus bursa simple, elongated, with two spined, pyramidal plates at ductus border; signum a small sharply pointed, thorn-like process.
</paragraph>
<paragraph id="777D36BBFF8EFFC9FF685A82B6C0790E" blockId="3.[151,1437,912,1981]" pageId="3" pageNumber="147">
<emphasis id="45B6EAA9FF8EFFC9FF685A82B6807956" bold="true" box="[199,418,1776,1801]" pageId="3" pageNumber="147">Specimens studied</emphasis>
.
<emphasis id="45B6EAA9FF8EFFC9FE1F5A82B54A7956" box="[432,616,1776,1801]" italics="true" pageId="3" pageNumber="147">
<typeStatus id="A8798819FF8EFFC9FE1F5A82B5357956" box="[432,535,1776,1801]" pageId="3" pageNumber="147" type="holotype">Holotype</typeStatus>
female
</emphasis>
: Cuc Phuong Natl. Pk., Mac Lake (
<quantity id="B03A9B5EFF8EFFC9FC5C5A82B31C7957" box="[1011,1086,1776,1801]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="147" unit="m" value="155.0">155 m</quantity>
), Ninh Binh Prov.,
<collectingCountry id="0FD5762BFF8EFFC9FABB5A82B2547956" box="[1300,1398,1776,1801]" name="Vietnam" pageId="3" pageNumber="147">Vietnam</collectingCountry>
,
<date id="037C107BFF8EFFC9FA2F5A82B6017972" pageId="3" pageNumber="147" value="2009-05-04" valueMax="2009-05-07" valueMin="2009-05-04">4 7 May 2009</date>
, J. B. Heppner (genitalia slide JBH 2859). Deposited with FSCA/McGuire Center, on indefinite loan from PPRI (Hanoi,
<collectingCountry id="0FD5762BFF8EFFC9FEDD5B4AB6F5790E" box="[370,471,1848,1873]" name="Vietnam" pageId="3" pageNumber="147">Vietnam</collectingCountry>
).
</paragraph>
<paragraph id="777D36BBFF8EFFC9FF385B2FB33A79C7" blockId="3.[151,1437,912,1981]" pageId="3" pageNumber="147">
<typeStatus id="A8798819FF8EFFC9FF385B2FB62B792B" box="[151,265,1885,1908]" pageId="3" pageNumber="147" type="paratype">
<emphasis id="45B6EAA9FF8EFFC9FF385B2FB62B792B" box="[151,265,1885,1908]" italics="true" pageId="3" pageNumber="147">Paratypes</emphasis>
</typeStatus>
(2♀):
<collectingCountry id="0FD5762BFF8EFFC9FEFC5B2FB6FB792A" box="[339,473,1885,1909]" name="Vietnam" pageId="3" pageNumber="147">VIETNAM</collectingCountry>
:
<emphasis id="45B6EAA9FF8EFFC9FE4C5B2EB5B4792B" box="[483,662,1884,1909]" italics="true" pageId="3" pageNumber="147">Ninh Binh Prov.</emphasis>
: same data as
<typeStatus id="A8798819FF8EFFC9FC985B2EB4BB792A" box="[823,921,1884,1909]" pageId="3" pageNumber="147" type="holotype">holotype</typeStatus>
(1♀) (FSCA/MGLB).
<emphasis id="45B6EAA9FF8EFFC9FB345B2FB24D792B" box="[1179,1391,1885,1908]" italics="true" pageId="3" pageNumber="147">
Hanoi/
<collectingCountry id="0FD5762BFF8EFFC9FB445B2FB20C792B" box="[1259,1326,1885,1908]" name="Turkey" pageId="3" pageNumber="147">Hatay</collectingCountry>
Prov.
</emphasis>
: Ba Vi Natl. Park [
<quantity id="B03A9B5EFF8EFFC9FE985BF2B6B179C7" box="[311,403,1920,1944]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="3" pageNumber="147" unit="m" value="1080.0">1080 m</quantity>
],
<date id="037C107BFF8EFFC9FE095BF2B55E79C7" box="[422,636,1920,1945]" pageId="3" pageNumber="147" value="2010-07-19" valueMax="2010-08-05" valueMin="2010-07-19">19 Jul5 Aug 2010</date>
(1♀), Y.-S. Bae &amp; L.-X. Vi (INUC).
</paragraph>
</subSubSection>
<subSubSection id="3FD86530FF8EFFC9FF685BD6B5C979E2" box="[199,747,1956,1981]" pageId="3" pageNumber="147" type="etymology">
<paragraph id="777D36BBFF8EFFC9FF685BD6B5C979E2" blockId="3.[151,1437,912,1981]" box="[199,747,1956,1981]" pageId="3" pageNumber="147">
<emphasis id="45B6EAA9FF8EFFC9FF685BD6B66479E2" bold="true" box="[199,326,1956,1981]" pageId="3" pageNumber="147">Etymology</emphasis>
. The species is named after
<collectingCountry id="0FD5762BFF8EFFC9FD2B5BD6B5C579E2" box="[644,743,1956,1981]" name="Vietnam" pageId="3" pageNumber="147">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="23BD6633FF89FFCEFF385B2DB3EF79CC" httpUri="https://zenodo.org/record/231716/files/figure.png" pageId="4" pageNumber="148" targetBox="[150,1436,193,1864]" targetPageId="4">
<paragraph id="777D36BBFF89FFCEFF385B2DB3EF79CC" blockId="4.[151,1436,1887,1939]" pageId="4" pageNumber="148">
<emphasis id="45B6EAA9FF89FFCEFF385B2DB637792B" bold="true" box="[151,277,1887,1909]" pageId="4" pageNumber="148">FIGURE 6.</emphasis>
<taxonomicName id="B0C24D38FF89FFCEFEB25B12B542792A" box="[285,608,1888,1909]" class="Insecta" family="Tortricidae" genus="Compsocommosis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="45B6EAA9FF89FFCEFEB25B12B542792A" box="[285,608,1888,1909]" italics="true" pageId="4" pageNumber="148">COmpSOcOmmOSiS vietnamenSiS</emphasis>
</taxonomicName>
<emphasis id="45B6EAA9FF89FFCEFDC75B2DB581792B" bold="true" box="[616,675,1887,1908]" pageId="4" pageNumber="148">n. sp.</emphasis>
, female genitalia, with details a) sterigma and ostium, with anterior split colliculum below subostium; b) ductus bursae cestum-like dentate plates; and c) signum (slide JBH 2859).
</paragraph>
</caption>
<subSubSection id="3FD86530FF88FFCFFF685CE5B4837EA7" pageId="5" pageNumber="149" type="biology_ecology">
<paragraph id="777D36BBFF88FFCFFF685CE5B4837EA7" blockId="5.[151,1436,151,357]" pageId="5" pageNumber="149">
<emphasis id="45B6EAA9FF88FFCFFF685CE5B63D7EEF" bold="true" box="[199,287,151,176]" pageId="5" pageNumber="149">Biology</emphasis>
. These moths are likely diurnal, as far as is known, like the
<taxonomicName id="B0C24D38FF88FFCFFC645CE5B20F7EEF" authority="(Heppner 1982)" baseAuthorityName="Heppner" baseAuthorityYear="1982" box="[971,1325,151,176]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="149" phylum="Arthropoda" rank="tribe" tribe="Hilarographini">
Hilarographini (
<bibRefCitation id="13534B4AFF88FFCFFB2F5CEAB2077EEF" author="Heppner" box="[1152,1317,152,176]" pageId="5" pageNumber="149" refString="Heppner, J. B. (1982) Synopsis of the Hilarographini (Lepidoptera: Tortricidae) of the world. PrOceedingS Of the EntOmOlOgical SOciety Of WaShingtOn, 84, 704 - 715." type="journal article" year="1982">Heppner 1982</bibRefCitation>
)
</taxonomicName>
, although specimens have all been taken at lights. Otherwise, nothing is known of the life history and biology of the species.
<emphasis id="45B6EAA9FF88FFCFFF685CADB67B7EA7" bold="true" box="[199,345,223,248]" pageId="5" pageNumber="149">Distribution</emphasis>
. The species is known only from northern
<collectingCountry id="0FD5762BFF88FFCFFC955CADB4BF7EA7" box="[826,925,223,248]" name="Vietnam" pageId="5" pageNumber="149">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3FD86530FF88FFCFFF685D76B5147F3A" pageId="5" pageNumber="149" type="discussion">
<paragraph id="777D36BBFF88FFCFFF685D76B5147F3A" blockId="5.[151,1436,151,357]" pageId="5" pageNumber="149">
<emphasis id="45B6EAA9FF88FFCFFF685D76B6677F42" bold="true" box="[199,325,260,285]" pageId="5" pageNumber="149">Discussion</emphasis>
. The new species superficially looks much like
<taxonomicName id="B0C24D38FF88FFCFFC075D77B37A7F43" box="[936,1112,261,284]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="45B6EAA9FF88FFCFFC075D77B37A7F43" box="[936,1112,261,284]" italics="true" pageId="5" pageNumber="149">Mictocommosis</emphasis>
</taxonomicName>
species in terms of adult maculation. The female appears more like members of the genus
<taxonomicName id="B0C24D38FF88FFCFFC275D5BB4F87F1F" box="[904,986,297,320]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="45B6EAA9FF88FFCFFC275D5BB4F87F1F" box="[904,986,297,320]" italics="true" pageId="5" pageNumber="149">Nexosa</emphasis>
</taxonomicName>
, but genitalia and wing venation differ considerably (
<bibRefCitation id="13534B4AFF88FFCFFE9B5D3EB5087F3A" author="Heppner" box="[308,554,332,357]" pageId="5" pageNumber="149" refString="Heppner, J. B. &amp; Bae, Y. S. (2015) Review of NexOSa Diakonoff in Vietnam, with a new species and a new subspecies, and transfer to the tribe Archipini (Lepidoptera: Tortricidae: Tortricini: Archipini). ZOOtaxa, 3999 (1), 32 - 40. http: // dx. doi. org / 10.11646 / zootaxa. 3999.1.2" type="journal article" year="2015">Heppner &amp; Bae 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>