494 lines
52 KiB
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494 lines
52 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/vz.73.e101329" ID-Pensoft-Pub="2625-8498-73-325" ID-Pensoft-UUID="360149275DE451CEA82EE1075A7C07B2" ID-ZooBank="2C884E3FC7BE4868A9294604C38A1757" ModsDocID="2625-8498-73-325" checkinTime="1680852563740" checkinUser="pensoft" docAuthor="Marques, Mariana P., Parrinha, Diogo, Ceriaco, Luis M. P., Brennan, Ian G., Heinicke, Matthew P. & Bauer, Aaron M." docDate="2023" docId="ED7DA2A77B69534A9359050266304689" docLanguage="en" docName="VertZool 73: 325-343" docOrigin="Vertebrate Zoology 73" docPubDate="2023-04-06" docSource="http://dx.doi.org/10.3897/vz.73.e101329" docTitle="Pachydactylus maiatoi Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer 2023, sp. nov." docType="treatment" docUuid="0C35D42A-405C-4C72-89C2-5C670FFF7FCA" docUuidSource="ZooBank" docVersion="2" id="360149275DE451CEA82EE1075A7C07B2" lastPageNumber="325" masterDocId="360149275DE451CEA82EE1075A7C07B2" masterDocTitle="A new species of thick-toed gecko (Pachydactylus) from Serra da Neve and surrounding rocky areas of southwestern Angola (Squamata: Gekkonidae)" masterLastPageNumber="343" masterPageNumber="325" pageNumber="325" updateTime="1680852865249" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>A new species of thick-toed gecko (Pachydactylus) from Serra da Neve and surrounding rocky areas of southwestern Angola (Squamata: Gekkonidae)</mods:title>
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</mods:titleInfo>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Marques, Mariana P.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1712-2632</mods:nameIdentifier>
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<mods:affiliation>CIBIO, Centro de Investigacao em Biodiversidade e Recursos Geneticos, InBIO Laboratorio Associado, Campus de Vairao, Universidade do Porto, 4485 - 661 Vairao, Portugal & BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairao, 4485 - 661, Vairao, Portugal & Departamento de Biologia, Faculdade de Ciencias da Universidade do Porto, Rua do Campo Alegre 1021, 4169 - 007 Porto, Portugal & Departamento de Zoologia e Antropologia (Museu Bocage), Museu Nacional de Historia Natural e da Ciencia, Universidade de Lisboa, Rua da Escola Politecnica, 58, 1269 - 102 Lisboa, Portugal</mods:affiliation>
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<mods:nameIdentifier type="email">mptlmarques@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Parrinha, Diogo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1302-025X</mods:nameIdentifier>
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<mods:affiliation>CIBIO, Centro de Investigacao em Biodiversidade e Recursos Geneticos, InBIO Laboratorio Associado, Campus de Vairao, Universidade do Porto, 4485 - 661 Vairao, Portugal & BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairao, 4485 - 661, Vairao, Portugal & Departamento de Biologia, Faculdade de Ciencias da Universidade do Porto, Rua do Campo Alegre 1021, 4169 - 007 Porto, Portugal</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ceriaco, Luis M. P.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0591-9978</mods:nameIdentifier>
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<mods:affiliation>CIBIO, Centro de Investigacao em Biodiversidade e Recursos Geneticos, InBIO Laboratorio Associado, Campus de Vairao, Universidade do Porto, 4485 - 661 Vairao, Portugal & BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairao, 4485 - 661, Vairao, Portugal & Departamento de Zoologia e Antropologia (Museu Bocage), Museu Nacional de Historia Natural e da Ciencia, Universidade de Lisboa, Rua da Escola Politecnica, 58, 1269 - 102 Lisboa, Portugal & Universidade Federal do Rio de Janeiro, Museu Nacional, Departamento de Vertebrados, Av. Bartolomeu de Gusmao 875, Sao Cristovao, 20941 - 160 Rio de Janeiro, Brasil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Brennan, Ian G.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4416-5224</mods:nameIdentifier>
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<mods:affiliation>Life Sciences Department, Natural History Museum, London, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Heinicke, Matthew P.</mods:namePart>
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<mods:affiliation>Department of Natural Sciences, University of Michigan-Dearborn, 4901 Evergreen Road, Dearborn, Michigan 48128, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bauer, Aaron M.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6839-8025</mods:nameIdentifier>
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<mods:affiliation>Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085 - 1699, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:title>Vertebrate Zoology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>73</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>325</mods:start>
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<mods:end>343</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/vz.73.e101329</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/vz.73.e101329</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2625-8498-73-325</mods:identifier>
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<mods:identifier type="ZooBank">2C884E3FC7BE4868A9294604C38A1757</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:0C35D42A-405C-4C72-89C2-5C670FFF7FCA" httpUri="http://treatment.plazi.org/id/ED7DA2A77B69534A9359050266304689" lastPageNumber="325" pageId="0" pageNumber="325">
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<subSubSection pageId="0" pageNumber="325" type="nomenclature">
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<paragraph pageId="0" pageNumber="325">
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<taxonomicName LSID="https://zoobank.org/0C35D42A-405C-4C72-89C2-5C670FFF7FCA" authority="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer, 2023" authorityName="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus maiatoi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maiatoi" status="sp. nov.">Pachydactylus maiatoi</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="325">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="325" type="description">
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<paragraph pageId="0" pageNumber="325">
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photos of the holotype (CAS 266485) of P. maiatoi sp. nov., (A) life and (B) preserved, from N'Dolondolo, Namibe Province, Angola. Photo by Luis M. P. Ceriaco." figureDoi="10.3897/vz.73.e101329.figure5" httpUri="https://binary.pensoft.net/fig/834449" pageId="0" pageNumber="325">Figs 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Life photo of one the paratypes (CAS 264267) of P. maiatoi sp. nov., from Maungo, Namibe Province, Angola. Photo by Luis M. P. Ceriaco." figureDoi="10.3897/vz.73.e101329.figure6" httpUri="https://binary.pensoft.net/fig/834450" pageId="0" pageNumber="325">, 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Juvenile paratype (CAS 254887) of Pachydactylus maiatoi sp. nov.. Photo by Luis M. P. Ceriaco." figureDoi="10.3897/vz.73.e101329.figure7" httpUri="https://binary.pensoft.net/fig/834451" pageId="0" pageNumber="325">, 7</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution of Pachydactylus angolensis and Pachydactylus maiatoi sp. nov. in Angola. Stars represent the respective type localities; dotted symbols denote localities represented in the phylogenetic analysis." figureDoi="10.3897/vz.73.e101329.figure8" httpUri="https://binary.pensoft.net/fig/834452" pageId="0" pageNumber="325">, 8</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Type locality of Pachydactylus maiatoi sp. nov., N'Dolondolo, Serra da Neve inselberg, northern Namibe Province. Photo by Ishan Agarwal. B Collecting locality of the paratype (CAS 264267) in Maungo region, central Namibe Province. Photo by Luis M. P. Ceriaco." figureDoi="10.3897/vz.73.e101329.figure9" httpUri="https://binary.pensoft.net/fig/834453" pageId="0" pageNumber="325">, 9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="325" type="reference_group">
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<paragraph pageId="0" pageNumber="325">
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<taxonomicName authority="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer, 2023" authorityName="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus angolensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="angolensis" status="sp. nov.">Pachydactylus angolensis</taxonomicName>
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-
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<bibRefCitation author="Ceriaco, LMP" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="325" refId="B15" refString="Ceriaco, LMP, de Sa, SAC, Bandeira, S, Valerio, H, Stanley, EL, Khun, AL, Marques, MP, Vindum, JV, Blackburn, DC, Bauer, AM, 2016. Herpetological survey of Iona National Park and Namibe Regional Natural Park, with a synoptic list of the amphibians and reptiles of Namibe Province, southwestern Angola. Proceedings of the California Academy of Sciences (Series 4) 63: 15-61." title="Herpetological survey of Iona National Park and Namibe Regional Natural Park, with a synoptic list of the amphibians and reptiles of Namibe Province, southwestern Angola. Proceedings of the California Academy of Sciences (Series 4) 63: 15 - 61." year="2016">
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<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
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et al. (2016
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</bibRefCitation>
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: 25)
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</paragraph>
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<paragraph pageId="0" pageNumber="325">
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<taxonomicName authority="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer, 2023" authorityName="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus angolensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="angolensis" status="sp. nov.">Pachydactylus angolensis</taxonomicName>
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"inland form" -
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<bibRefCitation DOI="https://doi.org/10.3897/zse.93.10915" author="Branch, WR" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="325" pagination="157 - 166" refId="B11" refString="Branch, WR, Haacke, W, Pinto, PV, Conradie, W, Baptista, N, Verburgt, L, Verissimo, L, 2017. Loveridge's Angolan geckos, Afroedura karroica bogerti and Pachydactylus scutatus angolensis (Sauria: Gekkonidae): New distribution records, comments on type localities and taxonomic status. Zoosystematics and Evolution 93: 157 - 166, DOI: https://doi.org/10.3897/zse.93.10915" title="Loveridge's Angolan geckos, Afroedura karroica bogerti and Pachydactylus scutatus angolensis (Sauria: Gekkonidae): New distribution records, comments on type localities and taxonomic status." url="https://doi.org/10.3897/zse.93.10915" volume="93" year="2017">Branch et al. (2017</bibRefCitation>
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: 164)
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</paragraph>
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<paragraph pageId="0" pageNumber="325">
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<taxonomicName authority="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer, 2023" authorityName="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus angolensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="angolensis" status="sp. nov.">Pachydactylus angolensis</taxonomicName>
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[partim] -
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<bibRefCitation author="Marques, MP" journalOrPublisher="Systematic Biology" pageId="0" pageNumber="325" refId="B38" refString="Marques, MP, Ceriaco, LMP, Blackburn, DC, Bauer, AM, 2018. Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840-2017). Proceedings of the California Academy of Sciences, Series 4, 65 Supplement 2: 1-501." title="Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 Supplement 2: 1 - 501." year="2018">Marques et al. (2018</bibRefCitation>
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: 192)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="325" type="holotype">
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<paragraph pageId="0" pageNumber="325">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="325">
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An adult male (CAS 266485, field no. AMB 10345), collected in
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<normalizedToken originalValue="N’Dolondolo">N'Dolondolo</normalizedToken>
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[-13.8133°, 13.1362°, 681 m a.s.l.], Namibe Province, Angola, by Luis M.P.
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<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
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, Suzana A. Bandeira, and Ishan Agarwal on 22 November 2016.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="325" type="paratypes">
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<paragraph pageId="0" pageNumber="325">Paratypes.</paragraph>
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<paragraph pageId="0" pageNumber="325">
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Six specimens: one adult male (CAS 254887, field no. JVV 8564), collected in Namibe-Lubango rd, 2 km E of Mangueiras [-15.0436°, 13.1600°, 625 m a.s.l.], Namibe Province, Angola, by Luis M.P.
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<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
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,
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<normalizedToken originalValue="Hilária">Hilaria</normalizedToken>
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<normalizedToken originalValue="Valério">Valerio</normalizedToken>
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, Suzana A. Bandeira, Sango de
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<normalizedToken originalValue="Sá">Sa</normalizedToken>
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, Arianna L. Kuhn, Edward L. Stanley, and Jens V. Vindum on 5 December 2013; one adult female (CAS 266484, field no. AMB 10284) with the same data as the holotype; one juvenile (CAS 266486, field no. AMB 10216) collected in Serra da Neve, base camp [-13.7770°, 13.2591°, 1488 m a.s.l.], Namibe Province, Angola, by Luis M.P.
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<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
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, Suzana A. Bandeira, and Ishan Agarwal on 18 November 2016; one adult male (CAS 264256, field no. AMB 11349) collected in Maungo [-14.5383°, 12.7474°, 363 m a.s.l.], Namibe Province, Angola, by Mariana P. Marques, Luis M.P.
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<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
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and Joyce M. Janota on 7 August 2018; one adult male (CAS 264267, field no. AMB 11402) collected in Maungo, abandoned health post [-14.5397°, 12.7447°, 368 m a.s.l.], Namibe Province, Angola, by Mariana P. Marques, Luis M.P.
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<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
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and Joyce M. Janota on 9 August 2018; one adult male (MUHNAC/MB03-001246, field no. LMPC 1182) collected in Serra da Neve, Maylowe village [-13.8355°, 13.2755°, 798 m a.s.l.], Namibe Province, Angola, by Mariana P. Marques, Luis M.P.
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<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
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, and Joyce M. Janota on 26 February 2019.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="325" type="materials_examined">
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<paragraph pageId="0" pageNumber="325">Additional material.</paragraph>
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<paragraph pageId="0" pageNumber="325">
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Namibe Province: Serra da Neve, rocky area near Catchi village [-13.7642°, 13.2573°,
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.614" unit="m" value="1614.0">1614 m</quantity>
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a.s.l.] (MUHNAC/MB03-001247); Serra da Neve base,
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<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
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N of Maylowe [-13.8280°, 13.2625°,
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<quantity metricMagnitude="2" metricUnit="m" metricValue="8.2" unit="m" value="820.0">820 m</quantity>
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a.sl.] (MUHNAC/MB03-001248); Lungo [-14.3167°, 13.2000°,
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<quantity metricMagnitude="2" metricUnit="m" metricValue="7.49" unit="m" value="749.0">749 m</quantity>
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a.s.l.] (imprecise locality) (TM 24406).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="325" type="diagnosis">
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<paragraph pageId="0" pageNumber="325">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="325">
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A small, rupiculous
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<taxonomicName class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus</emphasis>
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</taxonomicName>
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with a depressed body form. SVL at least 40.5 mm (Table
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<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphological and meristic comparisons between Pachydactylus angolensis and P. maiatoi sp. nov.. Data presented as " mean [minimum-maximum] ", measurements are presented in millimeters (mm). Abbreviations are those described in Materials and Methods section." httpUri="http://table.plazi.org/id/67DA7C3B987C70523EC823F684F658E1" pageId="0" pageNumber="325" tableUuid="67DA7C3B987C70523EC823F684F658E1">3</tableCitation>
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,
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<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Mensural (in mm) counts of the holotype and paratypes of Pachydactylus maiatoi sp. nov.. Abbreviations are the same as those described in the Materials and Methods section." httpUri="http://table.plazi.org/id/73C86F9E8B2C38E2AD4527E7D7934FE5" pageId="0" pageNumber="325" tableUuid="73C86F9E8B2C38E2AD4527E7D7934FE5">4</tableCitation>
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). Dorsum with enlarged, keeled dorsal scales. OrbD slightly greater than EyeEar. Rostral and supralabials excluded from the nostril rim. Lack of contact between the nostril and both the first supralabial and rostral. Tail indistinctly segmented with juxtaposed to subimbricate scalation. Dorsal pattern almost uniform beige to dark brown, sometimes with small and very diffuse whitish spots, with a dark mask in the lateral side of the head terminating above or just anterior to the ear. Labials white. Venter cream with brown spots on limbs and tail. Tail lighter than the dorsum. Forelimbs and hindlimbs moderately short and stout, interdigital webbing absent, free digits with four (4) subdigital, undivided lamellae.
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</paragraph>
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<paragraph pageId="0" pageNumber="325">
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Comparison with other
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<taxonomicName class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus</emphasis>
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</taxonomicName>
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species. From other members of the "northwestern clade" of
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<taxonomicName class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus</emphasis>
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</taxonomicName>
|
||
(fide
|
||
<bibRefCitation DOI="https://doi.org/10.1186/s12862-016-0846-2" author="Heinicke, MP" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="325" pagination="1 - 17" refId="B25" refString="Heinicke, MP, Jackman, TR, Bauer, AM, 2017. The measure of success: Geographic isolation promotes diversification in Pachydactylus geckos. BMC Evolutionary Biology 17: 1 - 17, DOI: https://doi.org/10.1186/s12862-016-0846-2" title="The measure of success: Geographic isolation promotes diversification in Pachydactylus geckos." url="https://doi.org/10.1186/s12862-016-0846-2" volume="17" year="2017">Heinicke et al. 2017</bibRefCitation>
|
||
) it may be easily differentiated from
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. bicolor" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="bicolor">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. bicolor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. maraisi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maraisi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. maraisi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. punctatus" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="punctatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. punctatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. scherzi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="scherzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. scherzi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. caraculicus" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="caraculicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. caraculicus</emphasis>
|
||
</taxonomicName>
|
||
by its enlarged, keeled dorsal scales (versus atuberculate or smooth scales in the latter). It can be differentiated from most other member of the northwestern group (
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. scutatus" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="scutatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. scutatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. parascutatus" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="parascutatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. parascutatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. boehmei" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="boehmei">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. boehmei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. gaiasensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="gaiasensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. gaiasensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. oreophilus" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="oreophilus">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. oreophilus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. sansteynae" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="sansteynae">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. sansteynae</emphasis>
|
||
</taxonomicName>
|
||
) by the lack of contact between the nostril and both the first supralabial and the rostral (in contact in all the latter). It can be distinguished from
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. otaviensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="otaviensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. otaviensis</emphasis>
|
||
</taxonomicName>
|
||
, endemic to the Otavi highlands in Namibia, by not having the first supralabial in contact with the nostril (versus in contact in
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. otaviensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="otaviensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. otaviensis</emphasis>
|
||
</taxonomicName>
|
||
) and by having a homogeneous cream to brown dorsal coloration pattern (versus consisting of three pale bands on the nape, just posterior to adpressed elbow, and posterior trunk, anterior to lumbar region in
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. otaviensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="otaviensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. otaviensis</emphasis>
|
||
</taxonomicName>
|
||
). Regarding its sister species,
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. angolensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. angolensis</emphasis>
|
||
</taxonomicName>
|
||
, the newly described species consistently differs in terms of its coloration, with
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. angolensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. angolensis</emphasis>
|
||
</taxonomicName>
|
||
presenting scattered large white spots and orange blotches in its light brown dorsum, while in
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. maiatoi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maiatoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. maiatoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="325">sp. nov.</emphasis>
|
||
the dorsum is generally homogenously cream to dark brown, although some specimens may present small and very diffuse whitish spots.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="325">
|
||
Regarding the other species of
|
||
<taxonomicName class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus</emphasis>
|
||
</taxonomicName>
|
||
occurring in Angola but not belonging to the "northwestern group",
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. maiatoi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maiatoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. maiatoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="325">sp. nov.</emphasis>
|
||
differs from
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. vanzyli" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="vanzyli">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. vanzyli</emphasis>
|
||
</taxonomicName>
|
||
by the presence of four (4) subdigital undivided lamellae on fourth toe (versus two (2) in the latter), from
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. rangei" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="rangei">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. rangei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. wahlbergii" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="wahlbergii">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. wahlbergii</emphasis>
|
||
</taxonomicName>
|
||
by having enlarged, keeled dorsal scales (versus atuberculate and very smooth skin). It can be further distinguished from
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. vanzyli" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="vanzyli">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. vanzyli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. rangei" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="rangei">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. rangei</emphasis>
|
||
</taxonomicName>
|
||
by its free digits (versus webbed digits of pes or both manus and pes in the latter), and its general color (pinkish in
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. rangei" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="rangei">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. rangei</emphasis>
|
||
</taxonomicName>
|
||
versus cream to brown in
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. maiatoi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maiatoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. maiatoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="325">sp. nov.</emphasis>
|
||
).
|
||
<taxonomicName baseAuthorityName="Andersson" baseAuthorityYear="1908" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus rangei" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="rangei">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus rangei</emphasis>
|
||
</taxonomicName>
|
||
only occurs in dunes and in very sandy regions including dry river beds, whereas the new species is rupiculous. It differs from
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. rugosus" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="rugosus">P. cf. rugosus</taxonomicName>
|
||
by its much less spiny appearance and by its dorsal coloration pattern (mostly homogeneous cream to brown in
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. maiatoi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maiatoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. maiatoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="325">sp. nov.</emphasis>
|
||
versus cream bands on brown dorsum in
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. rugosus" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="rugosus">P. cf. rugosus</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="325" type="description">
|
||
<paragraph pageId="0" pageNumber="325">Description of Holotype.</paragraph>
|
||
<paragraph pageId="0" pageNumber="325">
|
||
Adult male. SVL 36.1 mm (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photos of the holotype (CAS 266485) of P. maiatoi sp. nov., (A) life and (B) preserved, from N'Dolondolo, Namibe Province, Angola. Photo by Luis M. P. Ceriaco." figureDoi="10.3897/vz.73.e101329.figure5" httpUri="https://binary.pensoft.net/fig/834449" pageId="0" pageNumber="325">5</figureCitation>
|
||
; Table
|
||
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphological and meristic comparisons between Pachydactylus angolensis and P. maiatoi sp. nov.. Data presented as " mean [minimum-maximum] ", measurements are presented in millimeters (mm). Abbreviations are those described in Materials and Methods section." httpUri="http://table.plazi.org/id/67DA7C3B987C70523EC823F684F658E1" pageId="0" pageNumber="325" tableUuid="67DA7C3B987C70523EC823F684F658E1">3</tableCitation>
|
||
). Body moderately depressed, elongate (TrunkL/SVL 0.45). Head short (HeadL/SVL 0.28), moderately wide (HeadW/HeadL 0.66), depressed (HeadH/HeadL 0.38), slightly distinct from neck. Lores inflated; interorbital region flat. Snout slightly acuminate and convex, and moderately long (SnEye/HeadL 0.39), longer than eye diameter (OrbD/SnEye 0.57); scales on snout and canthus large, smooth, flattened, heterogeneous in size and shape; scales of interorbital and parietal regions heterogeneous, regularly scattered with tiny granules interspersed with larger conical to rounded tubercles. Eye small (OrbD/HeadL 0.22), with 10-12 superciliaries and posterodorsal corner of orbit bearing very small spines; pupil vertical, with crenellated margins; Ear-opening small, horizontally ovoid (EarL/HeadL 0.06). Eye to ear distance higher than the diameter of eye (EyeEar/OrbD 1.39). Rostral approximately 50% as deep as wide, no rostral groove, contacted by two enlarged supranasals and first supralabials; nostrils oval, each surrounded by two postnasals, supranasal, without contact with the first supralabial or rostral; supranasals in contact; one row of scales separates orbit from supralabials; mental elongate, deeper than wide; no enlarged postmentals or chin shields. Enlarged supralabials (left/right) counted to rictus 7/8 (6 to mid-orbit); infralabials 6/7; interorbital scale rows between superciliary rows at midpoint of orbit 19 (8 across narrowest point of frontal bone).Dorsal tubercles large (about ten times the size of adjacent scales), largest dorsolaterally and smallest along dorsal midline rounded, with a pronounced median keel, forming 16 regular longitudinal rows on trunk, grading into posteriorly-directed, conical scales on lower flanks; each tubercle surrounded by an irregular rosette of about 3 to 7 small domed to conical scales; ventral scales flattened, rounded to oval, subimbricate to imbricate, mostly homogeneous, largest on precloacal region, approximately 30 between lowest conical granular rows on flank at midbody; chin granules approximately one fifth the size of ventral scales, increasing in size rather abruptly between gular region and chest. No precloacal or femoral pores. Scales on palm, sole, and ventral surface of forelimb smooth, granular; scales on dorsal aspect of forelimb small, smooth, heterogeneous, subimbricate; scales on preaxial and ventral aspects of thighs somewhat enlarged, smooth, imbricate; scales on dorsum of thigh non-tuberculate, heterogeneous, small to large, flattened to conical; scales on dorsum of crus small, conical, with scattered large (about eight times the size of adjacent scales), keeled round tubercles. Forelimbs short, stout (ForeaL/SVL 0.13); hindlimbs short (CrusL/SVL 0.15); digits short; subdigital scansors, except for distal-most, entire, present only on distal portion of toes, approximately 1.5 times wider than more basal (non-scansorial) subdigital scales; interdigital webbing absent. Relative length of digits (manus): III ~ IV> V> II> I; (pes): IV> III> V> II> I. Subdigital scansors (excluding small distal divided scansor) I (3), II (3), III (3), IV (3), V (3) - manus; I (4), II (4), III (4), IV (4), V (3) - pes. Tail sub-cylindrical, fully regenerated; weakly depressed; postcloacal spurs present.
|
||
</paragraph>
|
||
<caption doi="10.3897/vz.73.e101329.figure5" httpUri="https://binary.pensoft.net/fig/834449" pageId="0" pageNumber="325" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="325">
|
||
<emphasis bold="true" pageId="0" pageNumber="325">Figure 5.</emphasis>
|
||
Photos of the holotype (CAS 266485) of
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. maiatoi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maiatoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. maiatoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="325">sp. nov.</emphasis>
|
||
, (
|
||
<emphasis bold="true" pageId="0" pageNumber="325">A</emphasis>
|
||
) life and (
|
||
<emphasis bold="true" pageId="0" pageNumber="325">B</emphasis>
|
||
) preserved, from
|
||
<normalizedToken originalValue="N’Dolondolo">N'Dolondolo</normalizedToken>
|
||
, Namibe Province, Angola. Photo by Luis M.P.
|
||
<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="325" type="coloration of the holotype">
|
||
<paragraph pageId="0" pageNumber="325">Coloration of the Holotype.</paragraph>
|
||
<paragraph pageId="0" pageNumber="325">
|
||
Dorsum almost homogeneous brown with small and diffuse whitish spots (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photos of the holotype (CAS 266485) of P. maiatoi sp. nov., (A) life and (B) preserved, from N'Dolondolo, Namibe Province, Angola. Photo by Luis M. P. Ceriaco." figureDoi="10.3897/vz.73.e101329.figure5" httpUri="https://binary.pensoft.net/fig/834449" pageId="0" pageNumber="325">5</figureCitation>
|
||
). Dark facial band extending from snout, through the eye and becoming more evident from the posterior part of the eye to right above the ear opening, stopping before the ear opening. Top of head beige slightly lighter than the dorsum. Supralabials and infralabials completely white. Limbs of the same color of the dorsum but digits paler. Venter cream with brown spots on limbs and tail. Tail lighter than the dorsum. Life coloration (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photos of the holotype (CAS 266485) of P. maiatoi sp. nov., (A) life and (B) preserved, from N'Dolondolo, Namibe Province, Angola. Photo by Luis M. P. Ceriaco." figureDoi="10.3897/vz.73.e101329.figure5" httpUri="https://binary.pensoft.net/fig/834449" pageId="0" pageNumber="325">5A</figureCitation>
|
||
) is similar to that in preservative (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photos of the holotype (CAS 266485) of P. maiatoi sp. nov., (A) life and (B) preserved, from N'Dolondolo, Namibe Province, Angola. Photo by Luis M. P. Ceriaco." figureDoi="10.3897/vz.73.e101329.figure5" httpUri="https://binary.pensoft.net/fig/834449" pageId="0" pageNumber="325">5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/vz.73.e101329.figure6" httpUri="https://binary.pensoft.net/fig/834450" pageId="0" pageNumber="325" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="325">
|
||
<emphasis bold="true" pageId="0" pageNumber="325">Figure 6.</emphasis>
|
||
Life photo of one the paratypes (CAS 264267) of
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. maiatoi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maiatoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. maiatoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="325">sp. nov.</emphasis>
|
||
, from Maungo, Namibe Province, Angola. Photo by Luis M.P.
|
||
<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="325" type="variation">
|
||
<paragraph pageId="0" pageNumber="325">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="325">
|
||
Variation in mensural and meristic characters of the holotype and adult paratypes are presented in Table
|
||
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphological and meristic comparisons between Pachydactylus angolensis and P. maiatoi sp. nov.. Data presented as " mean [minimum-maximum] ", measurements are presented in millimeters (mm). Abbreviations are those described in Materials and Methods section." httpUri="http://table.plazi.org/id/67DA7C3B987C70523EC823F684F658E1" pageId="0" pageNumber="325" tableUuid="67DA7C3B987C70523EC823F684F658E1">3</tableCitation>
|
||
. All paratypes agree entirely with the holotype and have a similar coloration pattern, only varying the tone of the coloration - from beige (e.g., CAS 254887, 264267; 266484, 266486) to darker brown (e.g., CAS 264256; MB03-001246). Some specimens (CAS 254887, 266484) present very faded brown spots on venter region. The juvenile paratype (CAS 254887, SVL 23.4 mm, Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Juvenile paratype (CAS 254887) of Pachydactylus maiatoi sp. nov.. Photo by Luis M. P. Ceriaco." figureDoi="10.3897/vz.73.e101329.figure7" httpUri="https://binary.pensoft.net/fig/834451" pageId="0" pageNumber="325">7</figureCitation>
|
||
) has similar coloration to that of adults, but the tail presents very vague darker bands.
|
||
</paragraph>
|
||
<caption doi="10.3897/vz.73.e101329.figure7" httpUri="https://binary.pensoft.net/fig/834451" pageId="0" pageNumber="325" start="Figure 7" startId="F7">
|
||
<paragraph pageId="0" pageNumber="325">
|
||
<emphasis bold="true" pageId="0" pageNumber="325">Figure 7.</emphasis>
|
||
Juvenile paratype (CAS 254887) of
|
||
<taxonomicName authorityName="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus maiatoi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maiatoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus maiatoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="325">sp. nov.</emphasis>
|
||
. Photo by Luis M.P.
|
||
<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="325" type="distribution">
|
||
<paragraph pageId="0" pageNumber="325">Distribution and habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="325">
|
||
<taxonomicName authorityName="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus maiatoi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maiatoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus maiatoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="325">sp. nov.</emphasis>
|
||
appears to be restricted to southwestern Angola, namely in the inland regions of northern Namibe Province and along the highlands associated with the Angolan escarpment (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution of Pachydactylus angolensis and Pachydactylus maiatoi sp. nov. in Angola. Stars represent the respective type localities; dotted symbols denote localities represented in the phylogenetic analysis." figureDoi="10.3897/vz.73.e101329.figure8" httpUri="https://binary.pensoft.net/fig/834452" pageId="0" pageNumber="325">8</figureCitation>
|
||
). Individuals were collected under rocks in areas with sparse vegetation, but always associated to inselbergs (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Type locality of Pachydactylus maiatoi sp. nov., N'Dolondolo, Serra da Neve inselberg, northern Namibe Province. Photo by Ishan Agarwal. B Collecting locality of the paratype (CAS 264267) in Maungo region, central Namibe Province. Photo by Luis M. P. Ceriaco." figureDoi="10.3897/vz.73.e101329.figure9" httpUri="https://binary.pensoft.net/fig/834453" pageId="0" pageNumber="325">9</figureCitation>
|
||
). It is expected that the species also occurs in neighboring areas of Benguela Province, but so far, no records of the species have been found in that province. The new species has a rupiculous habitat, while
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. angolensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. angolensis</emphasis>
|
||
</taxonomicName>
|
||
, as observed by the authors, is terrestrial with a habitat preference similar to that of
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. punctatus" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="punctatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. punctatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption doi="10.3897/vz.73.e101329.figure8" httpUri="https://binary.pensoft.net/fig/834452" pageId="0" pageNumber="325" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="325">
|
||
<emphasis bold="true" pageId="0" pageNumber="325">Figure 8.</emphasis>
|
||
Distribution of
|
||
<taxonomicName authorityName="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus angolensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus angolensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus maiatoi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maiatoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus maiatoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="325">sp. nov.</emphasis>
|
||
in Angola. Stars represent the respective type localities; dotted symbols denote localities represented in the phylogenetic analysis.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/vz.73.e101329.figure9" httpUri="https://binary.pensoft.net/fig/834453" pageId="0" pageNumber="325" start="Figure 9" startId="F9">
|
||
<paragraph pageId="0" pageNumber="325">
|
||
<emphasis bold="true" pageId="0" pageNumber="325">Figure 9.</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="325">A</emphasis>
|
||
Type locality of
|
||
<taxonomicName authorityName="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus maiatoi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maiatoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus maiatoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="325">sp. nov.</emphasis>
|
||
,
|
||
<normalizedToken originalValue="N’Dolondolo">N'Dolondolo</normalizedToken>
|
||
, Serra da Neve inselberg, northern Namibe Province. Photo by Ishan Agarwal.
|
||
<emphasis bold="true" pageId="0" pageNumber="325">B</emphasis>
|
||
Collecting locality of the paratype (CAS 264267) in Maungo region, central Namibe Province. Photo by Luis M. P.
|
||
<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="325" type="conservation">
|
||
<paragraph pageId="0" pageNumber="325">Conservation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="325">
|
||
<taxonomicName authorityName="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus angolensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus angolensis</emphasis>
|
||
</taxonomicName>
|
||
as previously construed was evaluated as a stable species of Least Concern (
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4747.1.3" author="Ceriaco, LMP" journalOrPublisher="Zootaxa" pageId="0" pageNumber="325" pagination="77 - 112" refId="B16" refString="Ceriaco, LMP, Heinicke, MP, Parker, KL, Marques, MP, Bauer, AM, 2020. A review of the African snake-eyed skinks (Scincidae: Panaspis) from Angola, with the description of a new species. Zootaxa 4747: 77 - 112, DOI: https://doi.org/10.11646/zootaxa.4747.1.3" title="A review of the African snake-eyed skinks (Scincidae: Panaspis) from Angola, with the description of a new species." url="https://doi.org/10.11646/zootaxa.4747.1.3" volume="4747" year="2020">
|
||
<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
|
||
et al. 2020
|
||
</bibRefCitation>
|
||
d). Although only
|
||
<taxonomicName authorityName="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus angolensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus angolensis</emphasis>
|
||
</taxonomicName>
|
||
sensu stricto is represented in a formal conservation area (i.e. Chimalavera Regional Natural Park), both species are relatively widespread in regions that are subject to minimal habitat transformation. Following the IUCN Red List criteria and guidelines (
|
||
<bibRefCitation author="Huntley, B" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="325" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B28" refString="2019. . http://www.iucnredlist.org/documents/RedListGuidelines.pdf" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2019">IUCN Standards and Petitions Committee 2019</bibRefCitation>
|
||
), and using the GeoCAT: geospatial conservation assessment tool (
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.150.2109" author="Bachman, S" journalOrPublisher="ZooKeys" pageId="0" pageNumber="325" pagination="117 - 126" refId="B1" refString="Bachman, S, Moat, J, Hill, AW, Torre, J, Scott, B, 2011. Supporting Red List threat assessment with GeoCAT: Geospatial conservation assessment tool. ZooKeys 150: 117 - 126, DOI: https://doi.org/10.3897/zookeys.150.2109" title="Supporting Red List threat assessment with GeoCAT: Geospatial conservation assessment tool." url="https://doi.org/10.3897/zookeys.150.2109" volume="150" year="2011">Bachman et al. 2011</bibRefCitation>
|
||
), we estimated an extent of occurrence of around 13.700 km2 for
|
||
<taxonomicName authorityName="Marques & Parrinha & Ceríaco & Brennan & Heinicke & Bauer" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Pachydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachydactylus maiatoi" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="maiatoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">Pachydactylus maiatoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="325">sp. nov.</emphasis>
|
||
and 18.500 km2 for
|
||
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="P. angolensis" order="Squamata" pageId="0" pageNumber="325" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="325">P. angolensis</emphasis>
|
||
</taxonomicName>
|
||
, which classify both species as Vulnerable (VU) based on their extent of occurrence. However, further research on the extent of occurrence, population trends and ecology of these species is fundamental for a formal assessment of their conservation status.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="325" type="etymology">
|
||
<paragraph pageId="0" pageNumber="325">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="325">
|
||
The species is named after the Angolan biologist Francisco Maiato
|
||
<normalizedToken originalValue="Gonçalves">Goncalves</normalizedToken>
|
||
(Huambo, 1982-), a researcher and curator of the Herbarium of Lubango (ISCED), and vice-dean for Research and International Affairs at the University Mandume Ya Ndemufayo,
|
||
<normalizedToken originalValue="Huíla">Huila</normalizedToken>
|
||
Province, Angola. Francisco Maiato
|
||
<normalizedToken originalValue="Gonçalves">Goncalves</normalizedToken>
|
||
represents a new generation of Angolan researchers and a driving force in the study and conservation of southern Angolan biodiversity. The specific epithet is a patronym in the masculine genitive singular. We propose the English common name of
|
||
<normalizedToken originalValue="Maiato’s">Maiato's</normalizedToken>
|
||
thick-toed gecko and the Portuguese common name of osga-de-dedos-grossos-de-Maiato.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |