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<document id="0BC3BB16572EAEB513A89398E6B6A8D8" ID-DOI="10.5281/zenodo.5393630" ID-ISSN="1638-9387" ID-Zenodo-Dep="5393630" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630319175263" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; Spiegel, Didier Van Den" docDate="2007" docId="B36487D8FFE4D51ED10CFF28FF53F8B4" docLanguage="en" docName="Zoosystema.2007.29.3.417-456.pdf" docOrigin="Zoosystema 29 (3)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Glyphiulus Gervais 1847" docType="treatment" docVersion="4" lastPageNumber="455" masterDocId="4F5DFFA0FFC1D538D20EFFF0FFA5FC3D" masterDocTitle="Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group" masterLastPageNumber="456" masterPageNumber="417" pageNumber="454" updateTime="1699225420998" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1BD6B2191C9069C022765AD2B40D7AF3">Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group</mods:title>
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<mods:namePart id="DDB6D2CE75E82CA89E46286AD7231ECC">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="205F2B8C1E4AAB9D66588CAF42444208">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
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<mods:namePart id="C9B56B437C1CE93C702A9B6B5EBE424F">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="D91AD61F365F1A5BC467168C5D216500">Muséum national dHistoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
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<mods:namePart id="E73508D7A93D2F0A9E7AC8446EA731D4">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="07046F4B6D832989D603F624D1777E87">Muséum national dHistoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
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<mods:namePart id="3BA6D1C39644F71CB93BC3B9DC23C820">Spiegel, Didier Van Den</mods:namePart>
<mods:affiliation id="348139E873D3C916A425C151B117AFB7">Musée royal de lAfrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be</mods:affiliation>
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<paragraph id="3B7236CEFFE4D51DD10CFF28FBF9FCCF" blockId="37.[770,1116,216,242]" box="[770,1116,216,242]" pageId="37" pageNumber="454">
<heading id="603A81A2FFE4D51DD10CFF28FBF9FCCF" box="[770,1116,216,242]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="454" reason="2">
Genus
<taxonomicName id="FCCD4D4DFFE4D51DD140FF28FBF9FCCF" ID-CoL="7P3VR" authority="Gervais, 1847" authorityName="Gervais" authorityYear="1847" box="[846,1116,216,242]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="09B9EADCFFE4D51DD140FF28FC1BFCCF" box="[846,958,216,242]" italics="true" pageId="37" pageNumber="454">Glyphiulus</emphasis>
Gervais, 1847
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="73D76545FFE4D51DD0ABFEE1FBBEFD4E" pageId="37" pageNumber="454" type="materials_examined">
<paragraph id="3B7236CEFFE4D51DD0ABFEE1FD7BFD7E" blockId="37.[677,1210,273,323]" pageId="37" pageNumber="454">
<collectionCode id="5DDCAE0BFFE4D51DD0ABFEE1FC86FD1A" box="[677,803,273,296]" pageId="37" pageNumber="454">
<typeStatus id="E476886CFFE4D51DD0ABFEE1FD70FD1A" box="[677,725,273,296]" pageId="37" pageNumber="454">TYPE</typeStatus>
SPECIES
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. —
<taxonomicName id="FCCD4D4DFFE4D51DD141FEE1FCD8FD14" box="[847,893,273,297]" class="Diplopoda" family="Julidae" genus="Iulus" kingdom="Animalia" order="Julida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="09B9EADCFFE4D51DD141FEE1FCD8FD14" box="[847,893,273,297]" italics="true" pageId="37" pageNumber="454">Iulus</emphasis>
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(recte:
<taxonomicName id="FCCD4D4DFFE4D51DD1CBFEE1FC56FD14" authorityName="Linnaeus" authorityYear="1758" box="[965,1011,273,297]" class="Diplopoda" family="Julidae" genus="Julus" kingdom="Animalia" order="Julida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="09B9EADCFFE4D51DD1CBFEE1FC56FD14" box="[965,1011,273,297]" italics="true" pageId="37" pageNumber="454">Julus</emphasis>
</taxonomicName>
)
<taxonomicName id="FCCD4D4DFFE4D51DD60CFEE1FD7EFD7E" authority="Gervais, 1847" authorityName="Gervais" authorityYear="1847" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="09B9EADCFFE4D51DD60CFEE1FBC1FD14" box="[1026,1124,273,297]" italics="true" pageId="37" pageNumber="454">granulatus</emphasis>
Gervais, 1847
</taxonomicName>
.
</paragraph>
<paragraph id="3B7236CEFFE4D51DD0ABFEABFBBEFD4E" blockId="37.[677,1051,347,371]" box="[677,1051,347,371]" pageId="37" pageNumber="454">
<collectionCode id="5DDCAE0BFFE4D51DD0ABFEABFC92FD4C" box="[677,823,347,370]" pageId="37" pageNumber="454">
<typeStatus id="E476886CFFE4D51DD0ABFEABFD70FD4C" box="[677,725,347,370]" pageId="37" pageNumber="454">TYPE</typeStatus>
LOCALITY
</collectionCode>
. — Mascarene Islands.
</paragraph>
</subSubSection>
<subSubSection id="73D76545FFE4D51DD0ABFE7BFB13F89F" pageId="37" pageNumber="454" type="diagnosis">
<paragraph id="3B7236CEFFE4D51DD0ABFE7BFBE2FF12" blockId="37.[677,1210,394,1186]" pageId="37" pageNumber="454">
DIAGNOSIS. —
<taxonomicName id="FCCD4D4DFFE4D51DD139FE7AFC6EFD9F" authorityName="Cook" authorityYear="1895" box="[823,971,394,418]" class="Diplopoda" family="Cambalopsidae" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="family">Cambalopsidae</taxonomicName>
with tergal crests evident at least on postcollar segments; similar crests often present also on collum, these not fragmented into more than two parts. Carinotaxy formulae of collum highly variable, often complex, but median crest, even if abbreviated anteriorly, usually clear. Carinotaxy formulae of postcollar segments usually 2(1)/2(1)+I/i+4/3+I/i+2(1)/2(1) (
<taxonomicName id="FCCD4D4DFFE4D51DD0A3FDB4FCAAFE61" authorityName="Gervais" authorityYear="1847" box="[685,783,580,604]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="09B9EADCFFE4D51DD0A3FDB4FCAAFE61" box="[685,783,580,604]" italics="true" pageId="37" pageNumber="454">granulatus</emphasis>
</taxonomicName>
-group) or 2(1)/2(1)+I/i+3/3+I/i+2(1)/2(1) (
<taxonomicName id="FCCD4D4DFFE4D51DD0A3FDAFFCA6FE4B" authorityName="Carl" authorityYear="1911" box="[685,771,607,630]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="09B9EADCFFE4D51DD0A3FDAFFCA6FE4B" box="[685,771,607,630]" italics="true" pageId="37" pageNumber="454">javanicus</emphasis>
</taxonomicName>
-group), crests undivided or divided into three tubercles occurring only occasionally, same as three crests below I/i. Basalmost crests on ozoporiferous segments never hypertrophied, instead always being smallest, sometimes even abbreviated. Ozopores starting from segment 5, always lying on particularly prominent tubercles. Eyes usually present, though often reduced and sometimes unpigmented, especially so in cavernicoles.
</paragraph>
<paragraph id="3B7236CEFFE4D51DD0B5FCC2FBAFF80A" blockId="37.[677,1210,394,1186]" pageId="37" pageNumber="454">
<collectingRegion id="F909F82CFFE4D51DD0B5FCC2FCB7FF74" box="[699,786,818,841]" country="Myanmar" name="Chin" pageId="37" pageNumber="454">Mentum</collectingRegion>
of gnathochilarium usually divided. Male legs 1 from little modified to strongly reduced, either with coxal outgrowths contiguous but virtually never completely fused medially (
<taxonomicName id="FCCD4D4DFFE4D51DD1A3FC72FBA1FFA4" authorityName="Carl" authorityYear="1911" box="[941,1028,898,921]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="09B9EADCFFE4D51DD1A3FC72FBA1FFA4" box="[941,1028,898,921]" italics="true" pageId="37" pageNumber="454">javanicus</emphasis>
</taxonomicName>
-group) or with leg remains (more rarely, retaining a few segments) widely separated due to a sternum devoid of median strictures (
<taxonomicName id="FCCD4D4DFFE4D51DD0A3FC20FCB0FFD5" authorityName="Gervais" authorityYear="1847" box="[685,789,976,1000]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="09B9EADCFFE4D51DD0A3FC20FCB0FFD5" box="[685,789,976,1000]" italics="true" pageId="37" pageNumber="454">granulatus</emphasis>
</taxonomicName>
-group). Male legs 2 nearly normal, not incrassate, coxae somewhat modified, and prefemora abbreviated frontally, for accommodation of penes. Male legs 3 with strongly elongated coxae.
</paragraph>
<paragraph id="3B7236CEFFE4D51DD0B5FBCAFB13F89F" blockId="37.[677,1210,394,1186]" pageId="37" pageNumber="454">Anterior gonopods usually with a shield-like coxosternum and a moveable, lateral, 1-segmented, often curved telopodite; flagella absent. Posterior gonopods compact, not elongated, often with apical flagelloid outgrowths.</paragraph>
</subSubSection>
<subSubSection id="73D76545FFE4D51ED0ABFB3EFF53F8B4" lastPageId="38" lastPageNumber="455" pageId="37" pageNumber="454" type="discussion">
<paragraph id="3B7236CEFFE4D51DD0ABFB3EFCACF8DB" blockId="37.[677,1210,1230,1700]" box="[677,777,1230,1256]" pageId="37" pageNumber="454">REMARKS</paragraph>
<paragraph id="3B7236CEFFE4D51ED0ABFB1DFEABFD2C" blockId="37.[677,1210,1230,1700]" lastBlockId="38.[113,647,216,1161]" lastPageId="38" lastPageNumber="455" pageId="37" pageNumber="454">
The general distribution of the
<taxonomicName id="FCCD4D4DFFE4D51DD60CFB1DFBD5F93A" authorityName="Gervais" authorityYear="1847" box="[1026,1136,1261,1287]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="09B9EADCFFE4D51DD60CFB1DFBD5F93A" box="[1026,1136,1261,1287]" italics="true" pageId="37" pageNumber="454">granulatus</emphasis>
</taxonomicName>
-group shows a quite coherent pattern, occupying a region between Sichuan, Hubei and Shanghai in the North, and central
<collectingCountry id="43DA765EFFE4D51DD126FABCFC23F95B" box="[808,902,1356,1382]" name="Vietnam" pageId="37" pageNumber="454">Vietnam</collectingCountry>
in the South. This matches very closely the distribution of the
<taxonomicName id="FCCD4D4DFFE4D51DD61FFA9DFBD4F9BB" authorityName="Carl" authorityYear="1911" box="[1041,1137,1389,1414]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="09B9EADCFFE4D51DD61FFA9DFBD4F9BB" box="[1041,1137,1389,1414]" italics="true" pageId="37" pageNumber="454">javanicus</emphasis>
</taxonomicName>
-group (
<figureCitation id="A3F62A4BFFE4D51DD0A2FA7CFD5EF99B" box="[684,763,1420,1446]" captionStart="FIG" captionStartId="36.[677,688,1159,1176]" captionText="FIG. 27. — Distribution of species of the javanicus-group, and of Glyphiulus Gervais,1847 in general (that of G. granulatus (Gervais, 1847) being pantropical, it is omitted here), shown more or less from north to south: 1, G. intermedius n. sp.; 2, G. parobliteratus n. sp.; 3, G. recticullus Zhang &amp; Li, 1982; 4, G. zorzini Mauriès &amp; Nguyen Duy-Jacquemin, 1997; 5, G. paracostulifer n. sp.; 6, G. sinensis (Meng &amp; Zhang, 1993); 7, G. obliteratoides n. sp.; 8, G. obliteratus n. sp.; 9, G. subobliteratus n. sp.; 10, G. pulcher (Loksa, 1960); 11, G. formosus (Pocock, 1895); 12, G. costulifer n. sp.; 13, G. mediobliteratus n. sp.; 14, G. subcostulifer n. sp.; 15, G. siamensis Mauriès, 1983; 16, G. percostulifer n. sp.; 17, G. vietnamicus Mauriès,1977; 18, G. mediator Attems, 1938; 19, G. javanicus Carl, 1911." pageId="37" pageNumber="454">Fig. 27</figureCitation>
), which ranges from between Sichuan and Zhejiang in the North to Java in the South. In other words, the distribution of the
<taxonomicName id="FCCD4D4DFFE4D51DD1E3FA3BFBFCF9D8" authorityName="Gervais" authorityYear="1847" box="[1005,1113,1483,1509]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="09B9EADCFFE4D51DD1E3FA3BFBFCF9D8" box="[1005,1113,1483,1509]" italics="true" pageId="37" pageNumber="454">granulatus</emphasis>
</taxonomicName>
-group is somewhat more restricted and falls entirely within that of the
<taxonomicName id="FCCD4D4DFFE4D51DD12BF9FBFC22FA19" authorityName="Carl" authorityYear="1911" box="[805,903,1547,1572]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="09B9EADCFFE4D51DD12BF9FBFC22FA19" box="[805,903,1547,1572]" italics="true" pageId="37" pageNumber="454">javanicus</emphasis>
</taxonomicName>
-group, the latter thus fully reflecting the general range of
<taxonomicName id="FCCD4D4DFFE4D51DD1D7F9DAFBE3FA79" authorityName="Gervais" authorityYear="1847" box="[985,1094,1578,1604]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="37" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="09B9EADCFFE4D51DD1D7F9DAFBE3FA79" box="[985,1094,1578,1604]" italics="true" pageId="37" pageNumber="454">Glyphiulus</emphasis>
</taxonomicName>
. Even after this review, there is no doubt that these generally subtropical to tropical, highly montane and often karstic areas contain many new species of this widespread and large genus waiting to be found and described.
</paragraph>
<paragraph id="3B7236CEFFE7D51ED286FEE7FE6DFE32" blockId="38.[113,647,216,1161]" pageId="38" pageNumber="455">
The only outlying member of the
<taxonomicName id="FCCD4D4DFFE7D51ED3ECFEE8FDE5FD0C" authorityName="Carl" authorityYear="1911" box="[482,576,280,305]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="38" pageNumber="455" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="09B9EADCFFE7D51ED3ECFEE8FDE5FD0C" box="[482,576,280,305]" italics="true" pageId="38" pageNumber="455">javanicus</emphasis>
</taxonomicName>
-group appears to be
<taxonomicName id="FCCD4D4DFFE7D51ED2F2FEC7FEDCFD6C" authorityName="Carl" authorityYear="1911" box="[252,377,311,337]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="38" pageNumber="455" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="09B9EADCFFE7D51ED2F2FEC7FEDCFD6C" box="[252,377,311,337]" italics="true" pageId="38" pageNumber="455">G. javanicus</emphasis>
</taxonomicName>
itself.The strong isolation of this species from the remaining bulk, which is restricted to continental Southeast Asia (
<figureCitation id="A3F62A4BFFE7D51ED027FE86FDDCFDAD" box="[553,633,374,400]" captionStart="FIG" captionStartId="36.[677,688,1159,1176]" captionText="FIG. 27. — Distribution of species of the javanicus-group, and of Glyphiulus Gervais,1847 in general (that of G. granulatus (Gervais, 1847) being pantropical, it is omitted here), shown more or less from north to south: 1, G. intermedius n. sp.; 2, G. parobliteratus n. sp.; 3, G. recticullus Zhang &amp; Li, 1982; 4, G. zorzini Mauriès &amp; Nguyen Duy-Jacquemin, 1997; 5, G. paracostulifer n. sp.; 6, G. sinensis (Meng &amp; Zhang, 1993); 7, G. obliteratoides n. sp.; 8, G. obliteratus n. sp.; 9, G. subobliteratus n. sp.; 10, G. pulcher (Loksa, 1960); 11, G. formosus (Pocock, 1895); 12, G. costulifer n. sp.; 13, G. mediobliteratus n. sp.; 14, G. subcostulifer n. sp.; 15, G. siamensis Mauriès, 1983; 16, G. percostulifer n. sp.; 17, G. vietnamicus Mauriès,1977; 18, G. mediator Attems, 1938; 19, G. javanicus Carl, 1911." pageId="38" pageNumber="455">Fig. 27</figureCitation>
), might well represent an accidental introduction from the mainland.
<bibRefCitation id="5F5C4B3FFFE7D51ED34EFE45FE1EFDED" author="CARL J." box="[320,443,437,464]" pageId="38" pageNumber="455" pagination="397 - 407" refId="ref17242" refString="CARL J. 1911. - Drei neue Diplopoden des Genfer Museums. Revue suisse de Zoologie 19: 397 - 407." type="journal article" year="1911">Carl (1911)</bibRefCitation>
explicitly indicated a sugar cane plantation as the place whence the
<typeStatus id="E476886CFFE7D51ED057FE26FD23FDCD" box="[601,646,470,496]" pageId="38" pageNumber="455">type</typeStatus>
series of
<taxonomicName id="FCCD4D4DFFE7D51ED2C0FE05FEF5FE32" authorityName="Carl" authorityYear="1911" box="[206,336,501,527]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="38" pageNumber="455" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="09B9EADCFFE7D51ED2C0FE05FEF5FE32" box="[206,336,501,527]" italics="true" pageId="38" pageNumber="455">G. javanicus</emphasis>
</taxonomicName>
had come.
</paragraph>
<paragraph id="3B7236CEFFE7D51ED286FDE5FF53F8B4" blockId="38.[113,647,216,1161]" pageId="38" pageNumber="455">
The fact that the species of
<taxonomicName id="FCCD4D4DFFE7D51ED3AAFDE5FDB1FE12" authorityName="Gervais" authorityYear="1847" box="[420,532,533,559]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="38" pageNumber="455" phylum="Arthropoda" rank="genus">
<emphasis id="09B9EADCFFE7D51ED3AAFDE5FDB1FE12" box="[420,532,533,559]" italics="true" pageId="38" pageNumber="455">Glyphiulus</emphasis>
</taxonomicName>
are mostly highly local in distribution deserves attention. This can often be accounted for by their being troglobites, naturally restricted to one single cave or a limited group of caves. However, the impression that the majority of the species are scarcely more than troglophiles suggests that a wealthy, barely sampled fauna also exists epigeically. This undersampling is due to the fact that most of the expeditions to these often remote and poorly accessible places have been speleological, concentrating on the “mystique” of the underground world, thus giving the biased impression that cambalopsids in general, and
<taxonomicName id="FCCD4D4DFFE7D51ED30FFC40FE63FFF7" box="[257,454,944,970]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="38" pageNumber="455" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="09B9EADCFFE7D51ED30FFC40FED6FFF7" box="[257,371,944,970]" italics="true" pageId="38" pageNumber="455">Glyphiulus</emphasis>
species
</taxonomicName>
in particular, are especially common in caves. Above-ground explorations will certainly reveal a far richer fauna of forest-dwelling species in the region. Summarising the poor state of current knowledge, we have still only touched the tip of the iceberg (
<bibRefCitation id="5F5C4B3FFFE7D51ED01FFBBFFF4FF8B4" author="GOLOVATCH S. I. &amp; GEOFFROY J. - J. &amp; MAURIES J. - P. &amp; VAN DEN SPIEGEL D." pageId="38" pageNumber="455" pagination="7 - 49" refId="ref17405" refString="GOLOVATCH S. I., GEOFFROY J. - J., MAURIES J. - P. &amp; VAN DEN SPIEGEL D. 2007. - Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatusgroup. Zoosystema 29 (1): 7 - 49." type="journal article" year="2007">
Golovatch
<emphasis id="09B9EADCFFE7D51ED27FFB9FFF03F8B5" box="[113,166,1134,1160]" italics="true" pageId="38" pageNumber="455">et al.</emphasis>
2007
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>